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<document id="2BE5305024667B57EE66538347EAB594" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629612136760" checkinUser="felipe" docAuthor="Ballantyne, Lesley A. &amp; Lambkin, Christine" docDate="2009" docId="0394D665BE06FF97FF3C52F42171EDCA" docLanguage="en" docName="zt01997p188.pdf" docOrigin="Zootaxa 1997" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Atyphella dalmatia Ballantyne &amp; Lambkin 2009, sp. n." docType="treatment" docVersion="3" lastPageNumber="44" masterDocId="FFADAE1DBE2CFFBCFFAB57492227EE30" masterDocTitle="Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae)" masterLastPageNumber="188" masterPageNumber="1" pageNumber="43" updateTime="1699083011144" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8141883B319C1A7B4918E4C5DC57C57A">Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae)</mods:title>
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<mods:namePart id="4B5AC38908B9AB240287A80710BCFFCB">Ballantyne, Lesley A.</mods:namePart>
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<mods:namePart id="F14CF5F62CFB7CCF0322421F55DBBFD0">Lambkin, Christine</mods:namePart>
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<mods:date id="1C41E6DAC3F39D17FBFCDBC33AC80116">2009</mods:date>
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<treatment id="0394D665BE06FF97FF3C52F42171EDCA" ID-DOI="http://doi.org/10.5281/zenodo.5324266" ID-Zenodo-Dep="5324266" LSID="urn:lsid:plazi:treatment:0394D665BE06FF97FF3C52F42171EDCA" httpUri="http://treatment.plazi.org/id/0394D665BE06FF97FF3C52F42171EDCA" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<subSubSection id="C32734F8BE06FF96FF3C52F423E2EBE7" box="[151,453,1469,1495]" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="8B826773BE06FF96FF3C52F423E2EBE7" blockId="42.[151,453,1469,1530]" box="[151,453,1469,1495]" pageId="42" pageNumber="43">
<heading id="D0CAD01FBE06FF96FF3C52F423E2EBE7" bold="true" box="[151,453,1469,1495]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
<emphasis id="B949BB61BE06FF96FF3C52F423E2EBE7" bold="true" box="[151,453,1469,1495]" pageId="42" pageNumber="43">
<taxonomicName id="4C3D1CF0BE06FF96FF3C52F4235BEBE7" ID-CoL="5W9RM" authority="Ballantyne &amp; Lambkin, 2009" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[151,380,1469,1495]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="dalmatia" status="sp. nov.">
<emphasis id="B949BB61BE06FF96FF3C52F4235BEBE7" bold="true" box="[151,380,1469,1495]" italics="true" pageId="42" pageNumber="43">Atyphella dalmatia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A061ABE06FF96FE2852F423E2EBE7" box="[387,453,1469,1495]" pageId="42" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE06FF96FF3C52A923A1EBCA" box="[151,390,1504,1530]" pageId="42" pageNumber="43" type="description">
<paragraph id="8B826773BE06FF96FF3C52A923A1EBCA" blockId="42.[151,453,1469,1530]" box="[151,390,1504,1530]" pageId="42" pageNumber="43">
(
<figureCitation id="13067BF6BE06FF96FF3452A9235AEBCA" box="[159,381,1504,1530]" captionStart="FIGURES 130149" captionStartId="157.[151,269,1828,1852]" captionTargetBox="[239,1340,178,1804]" captionTargetId="figure-118@157.[239,1348,174,1804]" captionTargetPageId="157" captionText="FIGURES 130149. Atyphella spp. 130, 131 A. flammans female, dorsal ventral (emerged from larvae taken at Mt Blackwood, QMBA). 132136 diagrammatic representation flightless Atyphella females; 132, 135 A. scintillans, 133 A. inconspicua; 134 A. similis; 136 probably A. lewisi. 137141, 147 A. dalmatia sp. n. 137139 dorsal and ventral paratype males; 140, 141 aedeagus in sheath, ventral (140) and dorsal; 147 pronotum paratype male. 142146, 148, 149 A. kirakira sp. n. 142, 144 dorsal, ventral; 143 ventral terminal abdomen ventrites 57; 145, 148 aedeagus in sheath ventral (145) and dorsal; 146, 149 aedeagus ventral (146) and dorsal." pageId="42" pageNumber="43">Figs 137141, 147</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE06FF96FF3C516621C0E8AA" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="8B826773BE06FF96FF3C51662339E842" blockId="42.[151,1436,1583,2010]" pageId="42" pageNumber="43">
<materialsCitation id="3B556D2EBE06FF96FF3C51662339E842" collectionCode="SAM" collectorName="Pr. &amp; Finschhaven &amp; Wareo, L" country="Papua New Guinea" latitude="-6.35" location="Morobe" longLatPrecision="783" longitude="147.51" pageId="42" pageNumber="43" specimenCount="1" stateProvince="Morobe" typeStatus="holotype">
<emphasis id="B949BB61BE06FF96FF3C51662337E879" bold="true" box="[151,272,1583,1609]" pageId="42" pageNumber="43">
<typeStatus id="5486D9D1BE06FF96FF3C5166232CE879" box="[151,267,1583,1609]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male.
<emphasis id="B949BB61BE06FF96FECE516620B5E879" bold="true" box="[357,658,1583,1609]" pageId="42" pageNumber="43">
<collectingCountry id="F32A27E3BE06FF96FECE516620B5E879" box="[357,658,1583,1609]" name="Papua New Guinea" pageId="42" pageNumber="43">PAPUA NEW GUINEA</collectingCountry>
</emphasis>
:
<geoCoordinate id="EE0901B4BE06FF96FD08517920CDE87A" box="[675,746,1584,1610]" degrees="6.35" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="555" value="-6.35">6.35S</geoCoordinate>
,
<geoCoordinate id="EE0901B4BE06FF96FD5C5179217BE87A" box="[759,860,1584,1610]" degrees="147.51" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="555" value="147.51">147.51E</geoCoordinate>
,
<collectingRegion id="49F9A991BE06FF96FCC1517921E1E87A" box="[874,966,1584,1610]" country="Papua New Guinea" name="Morobe" pageId="42" pageNumber="43">Morobe</collectingRegion>
<collectorName id="26C802A5BE06FF96FC64517921D5E87A" box="[975,1010,1584,1610]" pageId="42" pageNumber="43">Pr.</collectorName>
,
<collectorName id="26C802A5BE06FF96FC54517926B4E87A" box="[1023,1171,1584,1610]" pageId="42" pageNumber="43">Finschhaven</collectorName>
(
<emphasis id="B949BB61BE06FF96FB0E517926E2E879" box="[1189,1221,1584,1609]" italics="true" pageId="42" pageNumber="43">sic</emphasis>
),
<collectorName id="26C802A5BE06FF96FB7451792777E87A" box="[1247,1360,1584,1610]" pageId="42" pageNumber="43">Wareo, L</collectorName>
. Wagner (
<collectionCode id="ED2CFFB6BE06FF96FF6751112335E842" box="[204,274,1624,1650]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="42" pageNumber="43">SAM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B826773BE06FF96FF6C513621C0E8AA" blockId="42.[151,1436,1583,2010]" box="[199,999,1663,1690]" pageId="42" pageNumber="43">
<materialsCitation id="3B556D2EBE06FF96FF6C513621C4E8AA" box="[199,995,1663,1690]" collectionCode="SAM" pageId="42" pageNumber="43" specimenCount="2" typeStatus="paratype">
<emphasis id="B949BB61BE06FF96FF6C51362363E8A9" bold="true" box="[199,324,1663,1689]" pageId="42" pageNumber="43">
<typeStatus id="5486D9D1BE06FF96FF6C51362363E8A9" box="[199,324,1663,1689]" pageId="42" pageNumber="43" type="paratype">Paratypes</typeStatus>
</emphasis>
(24). Same locality, collector, as
<typeStatus id="5486D9D1BE06FF96FD6051C92112E8AA" box="[715,821,1664,1690]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
, males (
<collectionCode id="ED2CFFB6BE06FF96FC3E51C921FDE8AA" box="[917,986,1664,1690]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="42" pageNumber="43">SAM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE06FF97FF6D51EE2021ED9A" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="8B826773BE06FF96FF6D51EE2313E8DA" blockId="42.[151,1436,1583,2010]" pageId="42" pageNumber="43">
<emphasis id="B949BB61BE06FF96FF6D51EE2361E8F1" bold="true" box="[198,326,1703,1729]" pageId="42" pageNumber="43">Diagnosis.</emphasis>
Distinguished from all other
<taxonomicName id="4C3D1CF0BE06FF96FD0F51E12133E8F1" authorityName="Olliff, Madagascan" authorityYear="1890" box="[676,788,1704,1729]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B949BB61BE06FF96FD0F51E12133E8F1" box="[676,788,1704,1729]" italics="true" pageId="42" pageNumber="43">Atyphella</emphasis>
</taxonomicName>
by the scattered small pale spots on the brown to dark brown elytra.
</paragraph>
<paragraph id="8B826773BE06FF97FF6D51BE22D8EDB2" blockId="42.[151,1436,1583,2010]" lastBlockId="43.[151,1437,152,1018]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<emphasis id="B949BB61BE06FF96FF6D51BE2386E921" bold="true" box="[198,417,1783,1809]" pageId="42" pageNumber="43">Male description.</emphasis>
<quantity id="4CC5CA96BE06FF96FE0251B12011E922" box="[425,566,1784,1810]" metricMagnitude="-3" metricUnit="m" metricValue="7.95" metricValueMax="8.8" metricValueMin="7.1" pageId="42" pageNumber="43" unit="mm" value="7.95" valueMax="8.8" valueMin="7.1">7.18.8 mm</quantity>
long;
<quantity id="4CC5CA96BE06FF96FD2F51B12136E922" box="[644,785,1784,1810]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.4" metricValueMin="2.8" pageId="42" pageNumber="43" unit="mm" value="3.0999999999999996" valueMax="3.4" valueMin="2.8">2.83.4 mm</quantity>
wide; W/L 0.4. Colour: Pronotum largely yellow, usually with small paired median brown areas (20/25) (
<figureCitation id="13067BF6BE06FF96FCA95069214EE90A" box="[770,873,1824,1850]" captionStart="FIGURES 130149" captionStartId="157.[151,269,1828,1852]" captionTargetBox="[239,1340,178,1804]" captionTargetId="figure-118@157.[239,1348,174,1804]" captionTargetPageId="157" captionText="FIGURES 130149. Atyphella spp. 130, 131 A. flammans female, dorsal ventral (emerged from larvae taken at Mt Blackwood, QMBA). 132136 diagrammatic representation flightless Atyphella females; 132, 135 A. scintillans, 133 A. inconspicua; 134 A. similis; 136 probably A. lewisi. 137141, 147 A. dalmatia sp. n. 137139 dorsal and ventral paratype males; 140, 141 aedeagus in sheath, ventral (140) and dorsal; 147 pronotum paratype male. 142146, 148, 149 A. kirakira sp. n. 142, 144 dorsal, ventral; 143 ventral terminal abdomen ventrites 57; 145, 148 aedeagus in sheath ventral (145) and dorsal; 146, 149 aedeagus ventral (146) and dorsal." pageId="42" pageNumber="43">Fig. 147</figureCitation>
), median brown area occupying about half surface area and widening posteriorly (3/25) or occupying more than half of the whole area (2/25); MS, MN yellow; elytra dark brown (24/25, pale brown in
<specimenCount id="9D3BACFABE06FF96FD7950392149E9BA" box="[722,878,1904,1930]" pageId="42" pageNumber="43" type="generic" typeStatus="paratype">one paratype</specimenCount>
), with paler spots which are concentrated in the inner (sutural) half (
<figureCitation id="13067BF6BE06FF96FE2B50D12001E982" box="[384,550,1944,1970]" captionStart="FIGURES 130149" captionStartId="157.[151,269,1828,1852]" captionTargetBox="[239,1340,178,1804]" captionTargetId="figure-118@157.[239,1348,174,1804]" captionTargetPageId="157" captionText="FIGURES 130149. Atyphella spp. 130, 131 A. flammans female, dorsal ventral (emerged from larvae taken at Mt Blackwood, QMBA). 132136 diagrammatic representation flightless Atyphella females; 132, 135 A. scintillans, 133 A. inconspicua; 134 A. similis; 136 probably A. lewisi. 137141, 147 A. dalmatia sp. n. 137139 dorsal and ventral paratype males; 140, 141 aedeagus in sheath, ventral (140) and dorsal; 147 pronotum paratype male. 142146, 148, 149 A. kirakira sp. n. 142, 144 dorsal, ventral; 143 ventral terminal abdomen ventrites 57; 145, 148 aedeagus in sheath ventral (145) and dorsal; 146, 149 aedeagus ventral (146) and dorsal." pageId="42" pageNumber="43">Figs 137138</figureCitation>
), minimum of 3 pale spots on left elytron (1/25) to maximum of 18 spots on each elytron;
<specimenCount id="9D3BACFABE06FF96FE91508923B1E9EA" box="[314,406,1984,2010]" pageId="42" pageNumber="43" type="male">4 males</specimenCount>
with spots mostly concentrated around inner two interstitial lines; head, antennae and palpi mid-brown; ventral aspect of pro and mesosterna yellowish to light brown, metasternum dark brown; all legs yellow with tibiae and tarsi dark brown; V25 very dark brown, 6 and 7 white; tergites dark brown, T7 pale and slightly darker mottled, T8 pale. Pronotum: (
<figureCitation id="13067BF6BE07FF97FC8D57A121AAEF32" box="[806,909,232,258]" captionStart="FIGURES 130149" captionStartId="157.[151,269,1828,1852]" captionTargetBox="[239,1340,178,1804]" captionTargetId="figure-118@157.[239,1348,174,1804]" captionTargetPageId="157" captionText="FIGURES 130149. Atyphella spp. 130, 131 A. flammans female, dorsal ventral (emerged from larvae taken at Mt Blackwood, QMBA). 132136 diagrammatic representation flightless Atyphella females; 132, 135 A. scintillans, 133 A. inconspicua; 134 A. similis; 136 probably A. lewisi. 137141, 147 A. dalmatia sp. n. 137139 dorsal and ventral paratype males; 140, 141 aedeagus in sheath, ventral (140) and dorsal; 147 pronotum paratype male. 142146, 148, 149 A. kirakira sp. n. 142, 144 dorsal, ventral; 143 ventral terminal abdomen ventrites 57; 145, 148 aedeagus in sheath ventral (145) and dorsal; 146, 149 aedeagus ventral (146) and dorsal." pageId="43" pageNumber="44">Fig. 147</figureCitation>
)
<quantity id="4CC5CA96BE07FF97FC0B57A12608EF32" box="[928,1071,232,258]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="43" pageNumber="44" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.51.8 mm</quantity>
long,
<quantity id="4CC5CA96BE07FF97FB2B57A12736EF32" box="[1152,1297,232,258]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.2" metricValueMin="2.5" pageId="43" pageNumber="44" unit="mm" value="2.85" valueMax="3.2" valueMin="2.5">2.53.2 mm</quantity>
wide; W/L 1.61.7; lateral margins divergent posteriorly along their length (21/25) with slight rounded convergence in 4/ 25; posterolateral corners &lt;90°, rounded, projecting beyond median posterior margin; anterior half of hypomeron not flat but approaching closely, posterior area widely flat and closely adpressed. Elytron: parallel-sided; 5.67.0 mm long; 3 well-defined interstitial lines; epipleuron and suture continue around apex (visible both from above and beneath) neither further thickened in apical half. Head: cannot be retracted into prothoracic cavity; GHW 1.52.0 mm; SIW
<quantity id="4CC5CA96BE07FF97FD0D56912162EFC2" box="[678,837,472,498]" metricMagnitude="-4" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="43" pageNumber="44" unit="mm" value="0.275" valueMax="0.3" valueMin="0.25">0.250.3 mm</quantity>
; SIW/GHW 0.150.16; ASD&lt;ASW (sockets close but not contiguous); frons-vertex junction not defined, this area rounded. Antennae slightly longer than GHW up to almost twice GHW; 11 segmented. Mouthparts functional; apical segment of labial palpi ovoid, longer than wide. Abdomen, ventrites (
<figureCitation id="13067BF6BE07FF97FDBB55192050EC5A" box="[528,631,592,618]" captionStart="FIGURES 130149" captionStartId="157.[151,269,1828,1852]" captionTargetBox="[239,1340,178,1804]" captionTargetId="figure-118@157.[239,1348,174,1804]" captionTargetPageId="157" captionText="FIGURES 130149. Atyphella spp. 130, 131 A. flammans female, dorsal ventral (emerged from larvae taken at Mt Blackwood, QMBA). 132136 diagrammatic representation flightless Atyphella females; 132, 135 A. scintillans, 133 A. inconspicua; 134 A. similis; 136 probably A. lewisi. 137141, 147 A. dalmatia sp. n. 137139 dorsal and ventral paratype males; 140, 141 aedeagus in sheath, ventral (140) and dorsal; 147 pronotum paratype male. 142146, 148, 149 A. kirakira sp. n. 142, 144 dorsal, ventral; 143 ventral terminal abdomen ventrites 57; 145, 148 aedeagus in sheath ventral (145) and dorsal; 146, 149 aedeagus ventral (146) and dorsal." pageId="43" pageNumber="44">Fig. 139</figureCitation>
): LO entire in V6 and V7, in V7 LO reaching sides but not posterior margin (LO closely following posterior outline and disposition in fresh specimen may be interpreted differently). MPP short, rounded. Tergites: T8 as wide as long; anterolateral prolongations narrow, not as long as posterior exposed portion, not obviously expanded vertically. Aedeagal sheath (
<figureCitation id="13067BF6BE07FF97FBDC5581270DECD2" box="[1143,1322,712,738]" captionStart="FIGURES 130149" captionStartId="157.[151,269,1828,1852]" captionTargetBox="[239,1340,178,1804]" captionTargetId="figure-118@157.[239,1348,174,1804]" captionTargetPageId="157" captionText="FIGURES 130149. Atyphella spp. 130, 131 A. flammans female, dorsal ventral (emerged from larvae taken at Mt Blackwood, QMBA). 132136 diagrammatic representation flightless Atyphella females; 132, 135 A. scintillans, 133 A. inconspicua; 134 A. similis; 136 probably A. lewisi. 137141, 147 A. dalmatia sp. n. 137139 dorsal and ventral paratype males; 140, 141 aedeagus in sheath, ventral (140) and dorsal; 147 pronotum paratype male. 142146, 148, 149 A. kirakira sp. n. 142, 144 dorsal, ventral; 143 ventral terminal abdomen ventrites 57; 145, 148 aedeagus in sheath ventral (145) and dorsal; 146, 149 aedeagus ventral (146) and dorsal." pageId="43" pageNumber="44">Figs 140, 141</figureCitation>
): lacking rounded projection along posterior margin of sternite. Aedeagus (
<figureCitation id="13067BF6BE07FF97FC3855B9261FED3A" box="[915,1080,752,778]" captionStart="FIGURES 130149" captionStartId="157.[151,269,1828,1852]" captionTargetBox="[239,1340,178,1804]" captionTargetId="figure-118@157.[239,1348,174,1804]" captionTargetPageId="157" captionText="FIGURES 130149. Atyphella spp. 130, 131 A. flammans female, dorsal ventral (emerged from larvae taken at Mt Blackwood, QMBA). 132136 diagrammatic representation flightless Atyphella females; 132, 135 A. scintillans, 133 A. inconspicua; 134 A. similis; 136 probably A. lewisi. 137141, 147 A. dalmatia sp. n. 137139 dorsal and ventral paratype males; 140, 141 aedeagus in sheath, ventral (140) and dorsal; 147 pronotum paratype male. 142146, 148, 149 A. kirakira sp. n. 142, 144 dorsal, ventral; 143 ventral terminal abdomen ventrites 57; 145, 148 aedeagus in sheath ventral (145) and dorsal; 146, 149 aedeagus ventral (146) and dorsal." pageId="43" pageNumber="44">Figs 140, 141</figureCitation>
): L/W&lt;3/1; LL/ML moderate; LL not divergent along most of length dorsally and slightly shorter than ML; apices of LL rounded, not outturned, width subequal to that of ML; inner apical area of LL obliquely truncate; base of LL produced, rounded.
</paragraph>
<paragraph id="8B826773BE07FF97FF6D54C62021ED9A" blockId="43.[151,1437,152,1018]" box="[198,518,911,938]" pageId="43" pageNumber="44">
<emphasis id="B949BB61BE07FF97FF6D54C623A3ED99" bold="true" box="[198,388,911,937]" pageId="43" pageNumber="44">Female, Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE07FF97FF6D54FE2171EDCA" pageId="43" pageNumber="44" type="etymology">
<paragraph id="8B826773BE07FF97FF6D54FE2171EDCA" blockId="43.[151,1437,152,1018]" pageId="43" pageNumber="44">
<emphasis id="B949BB61BE07FF97FF6D54FE236CEDE1" bold="true" box="[198,331,951,977]" pageId="43" pageNumber="44">Etymology</emphasis>
. The specific name is latinised from the name of the spotted dog (dalmatian) and is a play on words to highlight the spotted nature of the dorsal surface.
</paragraph>
</subSubSection>
</treatment>
</document>