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<mods:title id="DD96CA89073BF4DC9EAD1BA44F0EF875">Notorhinus floresi sp. nov. gen. nov.: The first records of Siphonophorida in Chile and Siphonorhinidae in South America (Colobognatha)</mods:title>
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<mods:namePart id="E40F72EEDE0AE3F0998C77E13B910CAB">Moritz, Leif</mods:namePart>
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<mods:affiliation id="9FFD8D8D9DA291B070319AF22C298A79">Leibniz Institute for the Analysis of Biodiversity Change, Zoological Research Museum Alexander Koenig, Adenauerallee 127, Bonn, Germany &amp; Institute of Evolutionary Biology and Animal Ecology, University of Bonn, An der Immenburg, Bonn, Germany</mods:affiliation>
<mods:nameIdentifier id="7F4A35F1907832B90E3DD7A67E6A86E8" type="email">moritz.leif@gmail.com</mods:nameIdentifier>
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<mods:namePart id="DAEA813A03B8D21599DDB052E1CBA609">Parra-Gomez, Antonio</mods:namePart>
<mods:nameIdentifier id="AB0BD1047CFC5528BF3E0C238E01600D" type="ORCID">https://orcid.org/0000-0002-4144-6610</mods:nameIdentifier>
<mods:affiliation id="73AEB87F2568DAEDCE032DC01F5B1423">Facultad de Ciencias, Universidad Austral de Chile, Av. Rector Eduardo Morales Miranda 23, Valdivia, Chile</mods:affiliation>
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<mods:date id="FB17DA95283D74111A7A7960A2195E63">2023</mods:date>
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<subSubSection id="ADF6C1066FE4CDE7FD004BC43946EF10" pageId="0" pageNumber="565" type="nomenclature">
<paragraph id="48544986D8A02B4C2A4A3BD835F0230E" pageId="0" pageNumber="565">
3.1.1. Family
<taxonomicName id="598A9D837E07AE12E501D45F04445FC6" ID-CoL="7NL2T" ID-ENA="421862" LSID="41B14038-BC48-5721-ABC9-AE1AD1A0012F" authority="Cook, 1895" authorityName="Moritz &amp; Parra-Gómez" authorityYear="2023" class="Diplopoda" family="Siphonorhinidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Siphonophorida" pageId="0" pageNumber="565" rank="family">Siphonorhinidae Cook, 1895</taxonomicName>
</paragraph>
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<paragraph id="C180D20B0109513A7FC3E295AA1AD551" pageId="0" pageNumber="565">Remarks.</paragraph>
<paragraph id="0D75CEF4F6CEF93EED3CA5D9D482D0DA" pageId="0" pageNumber="565">
The specimens (
<taxonomicName id="A9FE14938777C4A280266231ED1D28A4" authorityName="Moritz &amp; Parra-Gómez" authorityYear="2023" class="Diplopoda" family="Siphonorhinidae" genus="Notorhinus" kingdom="Animalia" lsidName="Notorhinus floresi" order="Platydesmida" pageId="0" pageNumber="565" rank="species" species="floresi">
<emphasis id="CFB0ADE3BCC43809CD35F1FB9FC0CAF9" italics="true" pageId="0" pageNumber="565">Notorhinus floresi</emphasis>
</taxonomicName>
<emphasis id="AA71CDBE2BEF1C36C78DBE659C615D07" bold="true" pageId="0" pageNumber="565">sp. nov.</emphasis>
and
<taxonomicName id="7A2003071F4DE1E3ED223DB0619FD38D" authorityName="Moritz &amp; Parra-Gómez" authorityYear="2023" class="Diplopoda" family="Siphonorhinidae" genus="Notorhinus" kingdom="Animalia" lsidName="Notorhinus" order="Platydesmida" pageId="0" pageNumber="565" rank="genus">
<emphasis id="B89AE50218DD5B92EBD0754F1787457D" italics="true" pageId="0" pageNumber="565">Notorhinus</emphasis>
</taxonomicName>
sp. (MNHNC 8390)) can be placed in the family
<taxonomicName id="BC60E4E1C46A0AFD9EFB8800ECC4B030" authorityName="Cook" authorityYear="1895" class="Diplopoda" family="Siphonorhinidae" kingdom="Animalia" lsidName="" order="Siphonophorida" pageId="0" pageNumber="565" phylum="Arthropoda" rank="family">Siphonorhinidae</taxonomicName>
based on the following characters: Head pyriform, not extended into beak (rostrum) (Figs
<figureCitation id="83C41A18D615C600678773EE482EB2CB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Notorhinus floresi sp. nov., male holotype (MNHNC 8387), head, SEM. A Head, frontal view. B Head, lateral view. C Labrum, frontal view. D Labrum and gnathochilarium, ventral view. E Apical antennomeres, lateral view. F Apical antennomere, apical view. G Gnathochilarium and leg-pair 1, ventral view. Abbreviations: I-VII = antennomeres, ac = apical cones, cl = claw, co = collum, cx = coxa, ep = epicranium, fe = femur, fh = forehead, ge = genae, in = inscision of the labrum, la = labrum, LL = lamella lingualis, lp = spinous protuberences of labrum, lt = teeth lining labral incision, me = mentum, pa = palp-like structure, po = pores of the salivary glands, pof = postfemur, prf = prefemur, sb = sensilla basiconica, ssb = spiniform sensilla basiconica, sti = gnathochilarial stipes, ta = tarsus, ti = tibia." figureDoi="10.3897/asp.81.e100520.figure3" httpUri="https://binary.pensoft.net/fig/861788" pageId="0" pageNumber="565">3A</figureCitation>
,
<figureCitation id="46CA12B20FDE32DE8169648D909BC6EA" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Notorhinus sp., female (MNHNC 8390), SEM. A Head and anterior body-rings, frontal view. B Apical parts of gnathochilarium and labrum, ventral view. C Left antenna, lateral view. D Antennomere 7 with apical sensory structures. E Tarsal claw of mid-body leg. F Mid-body tergite, dorsal view. G Ozopore, lateral view, arrows indicate spines behind ozopore. H Posterior tergal margin (limbus), dorsal view, arrow indicates fluke-shaped sculpture. I Posterior body-rings and telson, ventral view. Abbreviations: I-VII = antennomeres, ac = apical cone, cl = tarsal claw, co = collum, ep = epicranium, fh = forehead, gn = gnathochilarium, hp = hypoproct, L 1 = leg-pair 1, L 2 = leg-pair 2, la = labrum, LL = lamella lingualis, md = mandible gnathal lobe, me = mentum, mz = metazonite, oz = ozopore, pp = paraproct, pr = preanal ring, pz = prozonite, sb = sensilla basiconica, ssb = spiniform sensilla basiconica, sti = gnathochilarial stipes, ta = tarsus." figureDoi="10.3897/asp.81.e100520.figure8" httpUri="https://binary.pensoft.net/fig/861793" pageId="0" pageNumber="565">8A</figureCitation>
); antennae elbowed; antennomere 2 longer than wide (Figs
<figureCitation id="708298B92A26154DA9722DA8A1CCB7AD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Notorhinus floresi sp. nov., male holotype (MNHNC 8387), head, SEM. A Head, frontal view. B Head, lateral view. C Labrum, frontal view. D Labrum and gnathochilarium, ventral view. E Apical antennomeres, lateral view. F Apical antennomere, apical view. G Gnathochilarium and leg-pair 1, ventral view. Abbreviations: I-VII = antennomeres, ac = apical cones, cl = claw, co = collum, cx = coxa, ep = epicranium, fe = femur, fh = forehead, ge = genae, in = inscision of the labrum, la = labrum, LL = lamella lingualis, lp = spinous protuberences of labrum, lt = teeth lining labral incision, me = mentum, pa = palp-like structure, po = pores of the salivary glands, pof = postfemur, prf = prefemur, sb = sensilla basiconica, ssb = spiniform sensilla basiconica, sti = gnathochilarial stipes, ta = tarsus, ti = tibia." figureDoi="10.3897/asp.81.e100520.figure3" httpUri="https://binary.pensoft.net/fig/861788" pageId="0" pageNumber="565">3B</figureCitation>
,
<figureCitation id="536920DBFEF661A88905C81C05F07227" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Notorhinus sp., female (MNHNC 8390), SEM. A Head and anterior body-rings, frontal view. B Apical parts of gnathochilarium and labrum, ventral view. C Left antenna, lateral view. D Antennomere 7 with apical sensory structures. E Tarsal claw of mid-body leg. F Mid-body tergite, dorsal view. G Ozopore, lateral view, arrows indicate spines behind ozopore. H Posterior tergal margin (limbus), dorsal view, arrow indicates fluke-shaped sculpture. I Posterior body-rings and telson, ventral view. Abbreviations: I-VII = antennomeres, ac = apical cone, cl = tarsal claw, co = collum, ep = epicranium, fh = forehead, gn = gnathochilarium, hp = hypoproct, L 1 = leg-pair 1, L 2 = leg-pair 2, la = labrum, LL = lamella lingualis, md = mandible gnathal lobe, me = mentum, mz = metazonite, oz = ozopore, pp = paraproct, pr = preanal ring, pz = prozonite, sb = sensilla basiconica, ssb = spiniform sensilla basiconica, sti = gnathochilarial stipes, ta = tarsus." figureDoi="10.3897/asp.81.e100520.figure8" httpUri="https://binary.pensoft.net/fig/861793" pageId="0" pageNumber="565">8C</figureCitation>
); gnathochilarium consisting of separate plates (Figs
<figureCitation id="4F11E4D5BE1E23FDE4433D88CB6354C1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Notorhinus floresi sp. nov., male holotype (MNHNC 8387), head, SEM. A Head, frontal view. B Head, lateral view. C Labrum, frontal view. D Labrum and gnathochilarium, ventral view. E Apical antennomeres, lateral view. F Apical antennomere, apical view. G Gnathochilarium and leg-pair 1, ventral view. Abbreviations: I-VII = antennomeres, ac = apical cones, cl = claw, co = collum, cx = coxa, ep = epicranium, fe = femur, fh = forehead, ge = genae, in = inscision of the labrum, la = labrum, LL = lamella lingualis, lp = spinous protuberences of labrum, lt = teeth lining labral incision, me = mentum, pa = palp-like structure, po = pores of the salivary glands, pof = postfemur, prf = prefemur, sb = sensilla basiconica, ssb = spiniform sensilla basiconica, sti = gnathochilarial stipes, ta = tarsus, ti = tibia." figureDoi="10.3897/asp.81.e100520.figure3" httpUri="https://binary.pensoft.net/fig/861788" pageId="0" pageNumber="565">3G</figureCitation>
,
<figureCitation id="B6C56630B37F5BE0CCB781C2F19DA488" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Notorhinus sp., female (MNHNC 8390), SEM. A Head and anterior body-rings, frontal view. B Apical parts of gnathochilarium and labrum, ventral view. C Left antenna, lateral view. D Antennomere 7 with apical sensory structures. E Tarsal claw of mid-body leg. F Mid-body tergite, dorsal view. G Ozopore, lateral view, arrows indicate spines behind ozopore. H Posterior tergal margin (limbus), dorsal view, arrow indicates fluke-shaped sculpture. I Posterior body-rings and telson, ventral view. Abbreviations: I-VII = antennomeres, ac = apical cone, cl = tarsal claw, co = collum, ep = epicranium, fh = forehead, gn = gnathochilarium, hp = hypoproct, L 1 = leg-pair 1, L 2 = leg-pair 2, la = labrum, LL = lamella lingualis, md = mandible gnathal lobe, me = mentum, mz = metazonite, oz = ozopore, pp = paraproct, pr = preanal ring, pz = prozonite, sb = sensilla basiconica, ssb = spiniform sensilla basiconica, sti = gnathochilarial stipes, ta = tarsus." figureDoi="10.3897/asp.81.e100520.figure8" httpUri="https://binary.pensoft.net/fig/861793" pageId="0" pageNumber="565">8B</figureCitation>
); anterior margin of collum straight (compare
<bibRefCitation id="D03FC47A62E1B508A6C9A31FA8436259" DOI="https://doi.org/10.1163/9789004188273_017" author="Enghoff, H" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="565" refId="B11" refString="Enghoff, H, Golovatch, SI, Short, M, Stoev, P, Wesener, T, 2015. Diplo-poda - taxonomic overview. In: Minelli A (Ed.) Treatise on Zoology - Anatomy, Taxonomy, Biology. The Myriapoda, Volume 2. Brill, Leiden, pp. 363-453. https://doi.org/10.1163/9789004188273_017" title="Diplo-poda - taxonomic overview. In: Minelli A (Ed.) Treatise on Zoology - Anatomy, Taxonomy, Biology. The Myriapoda, Volume 2. Brill, Leiden, pp. 363 - 453." url="https://doi.org/10.1163/9789004188273_017" year="2015">Enghoff et al. 2015</bibRefCitation>
;
<bibRefCitation id="B9D5A696831C1046167B3FD1844FD345" DOI="https://doi.org/10.3897/zookeys.626.9681" author="Marek, PE" journalOrPublisher="ZooKeys" pageId="0" pageNumber="565" pagination="1 - 43" refId="B30" refString="Marek, PE, Krejca, JK, Shear, WA, 2016. A new species of Illacme Cook &amp; Loomis, 1928 from Sequoia National Park, California, with a world catalog of the Siphonorhinidae (Diplopoda, Siphonophorida). ZooKeys 626: 1 - 43, DOI: https://doi.org/10.3897/zookeys.626.9681" title="A new species of Illacme Cook &amp; Loomis, 1928 from Sequoia National Park, California, with a world catalog of the Siphonorhinidae (Diplopoda, Siphonophorida)." url="https://doi.org/10.3897/zookeys.626.9681" volume="626" year="2016">Marek et al. 2016</bibRefCitation>
). According to a key provided by
<bibRefCitation id="C3DB1DDA18A0E9C0284509B2A95C7C81" author="Shelley, RM" journalOrPublisher="Myriapodologica" pageId="0" pageNumber="565" pagination="22 - 33" refId="B51" refString="Shelley, RM, 1996. The milliped order Siphonophorida in the United States and northern Mexico. Myriapodologica 4 (4): 22 - 33" title="The milliped order Siphonophorida in the United States and northern Mexico." volume="4" year="1996">Shelley (1996</bibRefCitation>
: 23) members of the family
<taxonomicName id="CD2339122D8AAA7E132A252EC79128C5" authorityName="Cook" authorityYear="1895" class="Diplopoda" family="Siphonorhinidae" kingdom="Animalia" lsidName="" order="Siphonophorida" pageId="0" pageNumber="565" phylum="Arthropoda" rank="family">Siphonorhinidae</taxonomicName>
lack sensory pits on antennomeres 5 and 6 (see also
<bibRefCitation id="D80BD49D2C980AECE4AE6959795BAF33" DOI="https://doi.org/10.3897/zookeys.241.3831" author="Marek, PE" journalOrPublisher="ZooKeys" pageId="0" pageNumber="565" pagination="77 - 112" refId="B31" refString="Marek, PE, Shear, WA, Bond, JE, 2012. A redescription of the leggiest animal, the millipede Illacme plenipes, with notes on its natural history and biogeography (Diplopoda, Siphonophorida, Siphonorhinidae). ZooKeys 241: 77 - 112, DOI: https://doi.org/10.3897/zookeys.241.3831" title="A redescription of the leggiest animal, the millipede Illacme plenipes, with notes on its natural history and biogeography (Diplopoda, Siphonophorida, Siphonorhinidae)." url="https://doi.org/10.3897/zookeys.241.3831" volume="241" year="2012">Marek et al. 2012</bibRefCitation>
), as is the case in the specimens studied here (Figs
<figureCitation id="55E7BBBD649DCD6DBD9E36A1354B9574" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Notorhinus floresi sp. nov., male holotype (MNHNC 8387), head, SEM. A Head, frontal view. B Head, lateral view. C Labrum, frontal view. D Labrum and gnathochilarium, ventral view. E Apical antennomeres, lateral view. F Apical antennomere, apical view. G Gnathochilarium and leg-pair 1, ventral view. Abbreviations: I-VII = antennomeres, ac = apical cones, cl = claw, co = collum, cx = coxa, ep = epicranium, fe = femur, fh = forehead, ge = genae, in = inscision of the labrum, la = labrum, LL = lamella lingualis, lp = spinous protuberences of labrum, lt = teeth lining labral incision, me = mentum, pa = palp-like structure, po = pores of the salivary glands, pof = postfemur, prf = prefemur, sb = sensilla basiconica, ssb = spiniform sensilla basiconica, sti = gnathochilarial stipes, ta = tarsus, ti = tibia." figureDoi="10.3897/asp.81.e100520.figure3" httpUri="https://binary.pensoft.net/fig/861788" pageId="0" pageNumber="565">3E</figureCitation>
,
<figureCitation id="290F83B436CC2661E1745677207C8C2B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Notorhinus sp., female (MNHNC 8390), SEM. A Head and anterior body-rings, frontal view. B Apical parts of gnathochilarium and labrum, ventral view. C Left antenna, lateral view. D Antennomere 7 with apical sensory structures. E Tarsal claw of mid-body leg. F Mid-body tergite, dorsal view. G Ozopore, lateral view, arrows indicate spines behind ozopore. H Posterior tergal margin (limbus), dorsal view, arrow indicates fluke-shaped sculpture. I Posterior body-rings and telson, ventral view. Abbreviations: I-VII = antennomeres, ac = apical cone, cl = tarsal claw, co = collum, ep = epicranium, fh = forehead, gn = gnathochilarium, hp = hypoproct, L 1 = leg-pair 1, L 2 = leg-pair 2, la = labrum, LL = lamella lingualis, md = mandible gnathal lobe, me = mentum, mz = metazonite, oz = ozopore, pp = paraproct, pr = preanal ring, pz = prozonite, sb = sensilla basiconica, ssb = spiniform sensilla basiconica, sti = gnathochilarial stipes, ta = tarsus." figureDoi="10.3897/asp.81.e100520.figure8" httpUri="https://binary.pensoft.net/fig/861793" pageId="0" pageNumber="565">8C</figureCitation>
). However, lack of these sensory pits seems to apply only to the North American genus
<taxonomicName id="5022D396E59BEA9A5A260A3E5AF2BA72" authorityName="Cook &amp; Loomis" authorityYear="1928" class="Diplopoda" family="Siphonorhinidae" genus="Illacme" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Illacme" order="Siphonophorida" pageId="0" pageNumber="565" phylum="Echinodermata" rank="genus">
<emphasis id="55C96FB5D8AFDBAF3F600EEBE0528730" italics="true" pageId="0" pageNumber="565">Illacme</emphasis>
</taxonomicName>
(
<bibRefCitation id="6BFC943CB05984977EA59EE5A71CEECE" DOI="https://doi.org/10.3897/zookeys.241.3831" author="Marek, PE" journalOrPublisher="ZooKeys" pageId="0" pageNumber="565" pagination="77 - 112" refId="B31" refString="Marek, PE, Shear, WA, Bond, JE, 2012. A redescription of the leggiest animal, the millipede Illacme plenipes, with notes on its natural history and biogeography (Diplopoda, Siphonophorida, Siphonorhinidae). ZooKeys 241: 77 - 112, DOI: https://doi.org/10.3897/zookeys.241.3831" title="A redescription of the leggiest animal, the millipede Illacme plenipes, with notes on its natural history and biogeography (Diplopoda, Siphonophorida, Siphonorhinidae)." url="https://doi.org/10.3897/zookeys.241.3831" volume="241" year="2012">Marek et al. 2012</bibRefCitation>
,
<bibRefCitation id="225D4B5FFAF4988E65E861B022D7AB41" DOI="https://doi.org/10.3897/zookeys.626.9681" author="Marek, PE" journalOrPublisher="ZooKeys" pageId="0" pageNumber="565" pagination="1 - 43" refId="B30" refString="Marek, PE, Krejca, JK, Shear, WA, 2016. A new species of Illacme Cook &amp; Loomis, 1928 from Sequoia National Park, California, with a world catalog of the Siphonorhinidae (Diplopoda, Siphonophorida). ZooKeys 626: 1 - 43, DOI: https://doi.org/10.3897/zookeys.626.9681" title="A new species of Illacme Cook &amp; Loomis, 1928 from Sequoia National Park, California, with a world catalog of the Siphonorhinidae (Diplopoda, Siphonophorida)." url="https://doi.org/10.3897/zookeys.626.9681" volume="626" year="2016">2016</bibRefCitation>
), the Malagasy genus
<taxonomicName id="6B2DF383BE9CBFCCABA355B0068E7FB3" authorityName="Wesener" authorityYear="2023" class="Diplopoda" genus="Madagascarhinus" kingdom="Animalia" lsidName="Madagascarhinus" order="Siphonophorida" pageId="0" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B372BDFCACBB25C3BB4EA0FD28DA94E8" italics="true" pageId="0" pageNumber="565">Madagascarhinus</emphasis>
</taxonomicName>
(
<bibRefCitation id="03FD0F68131C9EF70C07E278B567DDC4" author="Wesener, T" journalOrPublisher="Zootaxa" pageId="0" pageNumber="565" pagination="163 - 175" publicationUrl="https://doi.org/­10.11646/zootaxa.5278.1.9" refId="B61" refString="Wesener, T, 2023. Madagascarhinus, a new genus of the family Siphonorhinidae with two new species from Madagascar (Diplo-poda, Siphonophorida). Zootaxa 5278 (1): 163 - 175, DOI: https://doi.org/-10.11646/zootaxa.5278.1.9" title="Madagascarhinus, a new genus of the family Siphonorhinidae with two new species from Madagascar (Diplo-poda, Siphonophorida)." url="https://doi.org/­10.11646/zootaxa.5278.1.9" volume="5278" year="2023">Wesener 2023</bibRefCitation>
) and the South African genus
<taxonomicName id="7B1DDEC2D9EBC4D3B1B0D346C923F716" authorityName="Verhoeff" authorityYear="1939" class="Diplopoda" family="Siphonorhinidae" genus="Nematozonium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nematozonium" order="Siphonophorida" pageId="0" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="7D4249E35BF99822DE4EA17461E2B55B" italics="true" pageId="0" pageNumber="565">Nematozonium</emphasis>
</taxonomicName>
(
<bibRefCitation id="EBB665080A2683955F9BE9EE5272201F" author="Shelley, RM" journalOrPublisher="African Entomology" pageId="0" pageNumber="565" pagination="217 - 222" refId="B52" refString="Shelley, RM, Hoffman, RL, 2004. A contribution on the South African millipede genus, Nematozonium Verhoeff, 1939 (Siphonophorida: Siphonorhinidae). African Entomology 12 (2): 217 - 222" title="A contribution on the South African millipede genus, Nematozonium Verhoeff, 1939 (Siphonophorida: Siphonorhinidae)." volume="12" year="2004">Shelley and Hoffman 2004</bibRefCitation>
), as these sensory pits have been reported for
<taxonomicName id="E09FD0AC94D83BA20CFB24DFFBF5D24F" authorityName="Pocock" authorityYear="1894" class="Diplopoda" family="Siphonorhinidae" genus="Siphonorhinus" kingdom="Animalia" lsidName="Siphonorhinus" order="Siphonophorida" pageId="0" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="4E7E1FC418BCFF9A2FD62E49320899E9" italics="true" pageId="0" pageNumber="565">Siphonorhinus</emphasis>
</taxonomicName>
(
<bibRefCitation id="BCA2537C346A1CA85ACFD59E2698B5B9" author="Attems, CMT" journalOrPublisher="Mitteilungen aus dem Zoologischen Museum in Berlin" pageId="0" pageNumber="565" pagination="117 - 184" refId="B2" refString="Attems, CMT, 1930. Myriopoden der Kleinen Sunda-Inseln, gesammelt von der Expedition Dr. Rensch. Mitteilungen aus dem Zoologischen Museum in Berlin 16 (1): 117 - 184" title="Myriopoden der Kleinen Sunda-Inseln, gesammelt von der Expedition Dr. Rensch." volume="16" year="1930">Attems 1930</bibRefCitation>
,
<bibRefCitation id="EA645E4AECDDEE0988F5CF535998BD88" author="Attems, CMT" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle" pageId="0" pageNumber="565" pagination="187 - 353" refId="B4" refString="Attems, CMT, 1938. Die von Dr. C. Dawydoff in franzoesisch Indochina gesammelten Myriopoden. Memoires du Museum National d'Histoire Naturelle 6 (2): 187 - 353" title="Die von Dr. C. Dawydoff in franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">1938</bibRefCitation>
) and
<taxonomicName id="C1080B9618F0E624E668D1936DF99DB5" authorityName="Attems" authorityYear="1938" class="Diplopoda" family="Siphonorhinidae" genus="Kleruchus" kingdom="Animalia" lsidName="Kleruchus" order="Siphonophorida" pageId="0" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="1153214238399A07DC3F8CB7205FBF68" italics="true" pageId="0" pageNumber="565">Kleruchus</emphasis>
</taxonomicName>
(Attems, 1938).
</paragraph>
</subSubSection>
</treatment>
</document>