242 lines
39 KiB
XML
242 lines
39 KiB
XML
<document id="AE1950AD17802AF55F79E39D6DFF43B1" ID-DOI="10.11646/zootaxa.5372.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10146342" ID-ZooBank="1300D562-3E0B-4F72-933F-8FA7D28F6853" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700186674367" checkinUser="plazi" docAuthor="Mendonça, Maria Thayane Da Silva, Silva, Valéria Juliete Da & Fernandes, José Antônio Marin" docDate="2023" docId="414B87B5FF94980DFF63DC0177B4F83A" docLanguage="en" docName="zootaxa.5372.1.1.pdf" docOrigin="Zootaxa 5372 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5372.1.1/52272" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Edessa (Edessa) nigroacantha Mendonça & Silva & Fernandes 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="72" masterDocId="BD72FFCDFFD3984AFFF4DE477513FFC1" masterDocTitle="Description of fifty-one new species and new taxonomic arrangement for the E. sexdens group of the subgenus Edessa (Heteroptera, Pentatomidae, Edessinae, Edessa)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="72" updateTime="1700188109802" updateUser="ExternalLinkService">
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<mods:title id="C281543C8F1EE019C589BDC2C1A706B4">Description of fifty-one new species and new taxonomic arrangement for the E. sexdens group of the subgenus Edessa (Heteroptera, Pentatomidae, Edessinae, Edessa)</mods:title>
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</mods:titleInfo>
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<mods:name id="A5F0C36F8FB81FE6E611D41D9A247C21" type="personal">
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<mods:roleTerm id="77760BAF083DE157510654D98176E98C">Author</mods:roleTerm>
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<mods:namePart id="F3C1E15035A447468501F4FBA2EF72A4">Mendonça, Maria Thayane Da Silva</mods:namePart>
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<mods:affiliation id="8EDB6199B6E4598F7310F69993C55BFD">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil & Programa de Pós-Graduação em Zoologia UFPA / MPEG</mods:affiliation>
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<mods:roleTerm id="3471975859B9335DFCC15B2728036399">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7F2EEF3456D5EEEC078D32E620112A90">Silva, Valéria Juliete Da</mods:namePart>
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<mods:nameIdentifier id="028A1D5D5C125476B278F063206AEE4C" type="ORCID">0000-0002-1845-5228</mods:nameIdentifier>
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<mods:affiliation id="5993A1A88AACC4F42F899A4CF94E523F">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil & valeriajuliete @ hotmail. com; https: // orcid. org / 0000 - 0002 - 1845 - 5228</mods:affiliation>
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<mods:nameIdentifier id="0F6DDAFA05F66C06B4949BE48A0FC1E3" type="email">valeriajuliete@hotmail.com</mods:nameIdentifier>
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<mods:name id="8DADFD3150939D57C0B6FAC31AC53DE4" type="personal">
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</mods:role>
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<mods:namePart id="DCDE7F62E060BB0DA1505C88BAD9A54D">Fernandes, José Antônio Marin</mods:namePart>
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<mods:nameIdentifier id="EFD21785996AB53A5839FF8DF256C80F" type="ORCID">0000-0001-7450-5296</mods:nameIdentifier>
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<mods:affiliation id="C681D3F5989D3ED05430BA63154B5938">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil & Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil & Corresponding author: joseamf @ ufpa. br; https: // orcid. org / 0000 - 0001 - 7450 - 5296</mods:affiliation>
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<mods:nameIdentifier id="74378CB4C30922A527E88210FE507153" type="email">joseamf@ufpa.br</mods:nameIdentifier>
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<mods:title id="703123BBC4E787EEB8A764FEB97FA7A4">Zootaxa</mods:title>
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<mods:part id="913B1FE8B4D5B1BB51E8F95BB15B6779">
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<mods:date id="52F9AFB35875993288BF48E03FC2A030">2023</mods:date>
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<mods:number id="CB935DB3E3CACF4AE5ACA1155636F7F3">2023-11-14</mods:number>
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<mods:detail id="48FCE49E0D630283CDE3AB21F3131FB0" type="volume">
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<mods:number id="66BCF170324CCD48050017D0DBF6CC1E">5372</mods:number>
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<mods:url id="279E8593FC6E304D3C2B97ED7E50A461">https://www.mapress.com/zt/article/download/zootaxa.5372.1.1/52272</mods:url>
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</mods:location>
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<mods:classification id="3C1A046BFE55D36DB42D9AA7454EE9AF">journal article</mods:classification>
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<mods:identifier id="9C4C9CEEB1ACC4D803E9ACF96937E8E1" type="DOI">10.11646/zootaxa.5372.1.1</mods:identifier>
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<mods:identifier id="7A6FA3F750F6B14035A01950784660D6" type="ISSN">1175-5334</mods:identifier>
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<mods:identifier id="64482D85C518FA924779F0C2D2E7235E" type="Zenodo-Dep">10146342</mods:identifier>
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<mods:identifier id="6E9E295C7BBE2EACC49B62568F77D158" type="ZooBank">1300D562-3E0B-4F72-933F-8FA7D28F6853</mods:identifier>
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</mods:mods>
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<treatment id="414B87B5FF94980DFF63DC0177B4F83A" LSID="urn:lsid:plazi:treatment:414B87B5FF94980DFF63DC0177B4F83A" httpUri="http://treatment.plazi.org/id/414B87B5FF94980DFF63DC0177B4F83A" lastPageNumber="72" pageId="71" pageNumber="72">
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<subSubSection id="81F86528FF94980DFF63DC01775CFDA0" box="[151,591,582,609]" pageId="71" pageNumber="72" type="nomenclature">
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<paragraph id="C95D36A3FF94980DFF63DC01775CFDA0" blockId="71.[151,591,582,645]" box="[151,591,582,609]" pageId="71" pageNumber="72">
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<heading id="921581CFFF94980DFF63DC01775CFDA0" bold="true" box="[151,591,582,609]" fontSize="11" level="2" pageId="71" pageNumber="72" reason="4">
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<emphasis id="FB96EAB1FF94980DFF63DC01775CFDA0" bold="true" box="[151,591,582,609]" pageId="71" pageNumber="72">
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<taxonomicName id="0EE24D20FF94980DFF63DC017715FDA0" authority="Mendonça & Silva & Fernandes, 2023" authorityName="Mendonça & Silva & Fernandes" authorityYear="2023" box="[151,518,582,609]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="nigroacantha" status="sp. nov." subGenus="Edessa">
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<emphasis id="FB96EAB1FF94980DFF63DC017715FDA0" bold="true" box="[151,518,582,609]" italics="true" pageId="71" pageNumber="72">Edessa (Edessa) nigroacantha</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E0A557CAFF94980DFDF9DC00775CFDA0" box="[525,591,583,609]" pageId="71" pageNumber="72" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="81F86528FF94980DFF63DC2C7480FD44" box="[151,403,618,645]" pageId="71" pageNumber="72" type="description">
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<paragraph id="C95D36A3FF94980DFF63DC2C7480FD44" blockId="71.[151,591,582,645]" box="[151,403,618,645]" pageId="71" pageNumber="72">
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(
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<figureCitation id="51D92A26FF94980DFF6BDC2C75EEFD44" box="[159,253,619,645]" captionStart="FIGURE 34" captionStartId="72.[152,255,1943,1967]" captionTargetBox="[184,1409,187,1913]" captionTargetId="figure-22@72.[178,1409,181,1919]" captionTargetPageId="72" captionText="FIGURE 34. A–F. Edessa (E.) nigroacantha sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146415" httpUri="https://zenodo.org/record/10146415/files/figure.png" pageId="71" pageNumber="72">Figs. 34</figureCitation>
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,
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<figureCitation id="51D92A26FF94980DFEFDDC2D7471FD44" box="[265,354,618,645]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">65 A–B</figureCitation>
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,
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<figureCitation id="51D92A26FF94980DFE99DC2C7499FD44" box="[365,394,619,645]" captionStart="FIGURE 72" captionStartId="125.[152,255,1071,1095]" captionTargetBox="[195,1391,181,1040]" captionTargetId="figure-20@125.[195,1391,181,1050]" captionTargetPageId="125" captionText="FIGURE 72. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.10146495" httpUri="https://zenodo.org/record/10146495/files/figure.png" pageId="71" pageNumber="72">72</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="81F86528FF94980DFF63DCEB71ABFD07" box="[151,1208,684,711]" pageId="71" pageNumber="72" type="etymology">
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<paragraph id="C95D36A3FF94980DFF63DCEB71ABFD07" blockId="71.[151,1437,684,2043]" box="[151,1208,684,711]" pageId="71" pageNumber="72">
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<emphasis id="FB96EAB1FF94980DFF63DCEB740BFD07" bold="true" box="[151,280,684,710]" pageId="71" pageNumber="72">Etymology</emphasis>
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. The name refers to the black humeral angles (L.
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<emphasis id="FB96EAB1FF94980DFCB2DCEA769BFD07" box="[838,904,685,710]" italics="true" pageId="71" pageNumber="72">nigro,</emphasis>
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black; L.
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<emphasis id="FB96EAB1FF94980DFC0EDCEB7142FD07" box="[1018,1105,684,710]" italics="true" pageId="71" pageNumber="72">canthus</emphasis>
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, corner).
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</paragraph>
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</subSubSection>
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<subSubSection id="81F86528FF94980DFF33DC977441FCCF" pageId="71" pageNumber="72" type="materials_examined">
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<paragraph id="C95D36A3FF94980DFF33DC977441FCCF" blockId="71.[151,1437,684,2043]" pageId="71" pageNumber="72">
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<emphasis id="FB96EAB1FF94980DFF33DC9774A3FD2B" bold="true" box="[199,432,720,746]" pageId="71" pageNumber="72">Material examined.</emphasis>
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<materialsCitation id="798A3CFEFF94980DFE41DC977441FCCF" collectingDate="1921-12" collectionCode="USNM" collectorName="W. M. Manm" country="Bolivia" location="Mulford BioExpl" municipality="Tumupasa" pageId="71" pageNumber="72" specimenCount="1" stateProvince="La Paz" typeStatus="holotype">
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<typeStatus id="16598801FF94980DFE41DC97770EFD2B" box="[437,541,720,746]" pageId="71" pageNumber="72" type="holotype">Holotype</typeStatus>
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male.
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<emphasis id="FB96EAB1FF94980DFD93DC97765AFD2B" bold="true" box="[615,841,720,746]" pageId="71" pageNumber="72">
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<collectingCountry id="B1F57633FF94980DFD93DC9777F1FD2B" box="[615,738,720,746]" name="Bolivia" pageId="71" pageNumber="72">BOLIVIA</collectingCountry>
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,
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<collectingRegion id="0B26F841FF94980DFD19DC977657FD2B" box="[749,836,720,746]" country="Bolivia" name="La Paz" pageId="71" pageNumber="72">La Paz</collectingRegion>
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:
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</emphasis>
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<collectingMunicipality id="2939ACD9FF94980DFCBBDC9676D5FD2A" box="[847,966,721,747]" pageId="71" pageNumber="72">Tumupasa</collectingMunicipality>
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,
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<location id="CC3D6078FF94980DFC3BDC977180FD2B" LSID="urn:lsid:plazi:treatment:414B87B5FF94980DFF63DC0177B4F83A:CC3D6078FF94980DFC3BDC977180FD2B" box="[975,1171,720,746]" country="Bolivia" municipality="Tumupasa" name="Mulford BioExpl" pageId="71" pageNumber="72" stateProvince="La Paz">Mulford BioExpl</location>
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,
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<date id="BD5C1063FF94980DFB69DC977016FD2B" box="[1181,1285,720,746]" pageId="71" pageNumber="72" value="1921-12">
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<collectingDate id="AD18E98BFF94980DFB69DC977016FD2B" box="[1181,1285,720,746]" pageId="71" pageNumber="72" value="1921-12">XII-1921</collectingDate>
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</date>
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–1922,
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<collectorName id="64175375FF94980DFAA3DC9675F3FCCE" pageId="71" pageNumber="72">W. M. Manm</collectorName>
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(
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<collectionCode id="AFF3AE66FF94980DFF1BDCB37455FCCF" box="[239,326,756,782]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="71" pageNumber="72" type="Museum">USNM</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="81F86528FF94980DFF33DD5F745BFCBB" pageId="71" pageNumber="72" type="description">
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<paragraph id="C95D36A3FF94980DFF33DD5F745BFCBB" blockId="71.[151,1437,684,2043]" pageId="71" pageNumber="72">
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<emphasis id="FB96EAB1FF94980DFF33DD5F7465FCF3" bold="true" box="[199,374,792,818]" pageId="71" pageNumber="72">Measurements</emphasis>
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(n= 1). Total length: 19.8; head length: 1.8; head width: 3.7; pronotum length: 4.1; pronotum width: 13.6; scutellum length: 9.4; scutellum width: 7.0; abdominal width: 12.2; length antennomeres: I: 1.0; II: 1.9; III: 2.2; IV: 4.4.
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</paragraph>
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</subSubSection>
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<subSubSection id="81F86528FF94980DFF33DDC37675F91B" pageId="71" pageNumber="72" type="diagnosis">
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<paragraph id="C95D36A3FF94980DFF33DDC37675F91B" blockId="71.[151,1437,684,2043]" pageId="71" pageNumber="72">
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<emphasis id="FB96EAB1FF94980DFF33DDC37451FC5F" bold="true" box="[199,322,900,926]" pageId="71" pageNumber="72">Diagnosis.</emphasis>
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Large (
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<quantity id="0E1A9B46FF94980DFE6FDDC37715FC5F" box="[411,518,900,926]" metricMagnitude="-2" metricUnit="m" metricValue="1.98" pageId="71" pageNumber="72" unit="mm" value="19.8">19.8 mm</quantity>
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). Dorsal body surface green, densely punctured (
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<figureCitation id="51D92A26FF94980DFBC7DDC371B0FC5F" box="[1075,1187,900,927]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 A</figureCitation>
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). Ventral surface dark yellow with transversal black lines on thorax and abdomen (
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<figureCitation id="51D92A26FF94980DFCC6DDEF768EFC03" box="[818,925,936,963]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 B</figureCitation>
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). Antennae, with first segment yellow, second and thrid reddish brown and fourth yellow (
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<figureCitation id="51D92A26FF94980DFD8BDD8B761DFC27" box="[639,782,972,999]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 A–B</figureCitation>
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). Pronotum coarse, with concolorous with the surrounding surface punctures (
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<figureCitation id="51D92A26FF94980DFE9EDDB774CBFBCB" box="[362,472,1008,1035]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 A</figureCitation>
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). Humeral angles short (1.25 times wider than long); apex with black spot restricted to the angles in dorsal view and restricted to the margin of the angles in ventral view; bent backward (
|
||
<figureCitation id="51D92A26FF94980DFB0BDA537082FBEF" box="[1279,1425,1044,1071]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 A–B</figureCitation>
|
||
). Scutellum with concolorous with the surrounding surface punctures; apex not reaching the end of coria (
|
||
<figureCitation id="51D92A26FF94980DFAD7DA7F7083FB93" box="[1315,1424,1080,1107]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 A</figureCitation>
|
||
). Coria with all veins concolorous with the surrounding surface (
|
||
<figureCitation id="51D92A26FF94980DFC91DA1B76C5FBB7" box="[869,982,1116,1143]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 A</figureCitation>
|
||
). Connexival segments with concavities almost entirely covered by subrectangular black spots separated by a large yellow median spot (
|
||
<figureCitation id="51D92A26FF94980DFB29DAC7705EFB5B" box="[1245,1357,1152,1179]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 A</figureCitation>
|
||
), spots extending ventrally, triangular (
|
||
<figureCitation id="51D92A26FF94980DFDF7DAE37764FB7F" box="[515,631,1188,1215]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 B</figureCitation>
|
||
). Posterolateral angles of connexivum with apices black (
|
||
<figureCitation id="51D92A26FF94980DFAEEDAE37083FB7F" box="[1306,1424,1188,1215]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 A</figureCitation>
|
||
). Ventral surface, thorax with brown lines; dark line of the propleura covering 2/3 of the width of the sclerite (
|
||
<figureCitation id="51D92A26FF94980DFAB8DA8F75BFFAC7" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 B</figureCitation>
|
||
). Evaporatorium concolorous with the surrounding surface (
|
||
<figureCitation id="51D92A26FF94980DFCB8DAAB76A4FAC7" box="[844,951,1260,1287]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 B</figureCitation>
|
||
). Metasternal process (
|
||
<figureCitation id="51D92A26FF94980DFB4CDAAB700CFAC7" box="[1208,1311,1260,1286]" captionStart="FIGURE 34" captionStartId="72.[152,255,1943,1967]" captionTargetBox="[184,1409,187,1913]" captionTargetId="figure-22@72.[178,1409,181,1919]" captionTargetPageId="72" captionText="FIGURE 34. A–F. Edessa (E.) nigroacantha sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146415" httpUri="https://zenodo.org/record/10146415/files/figure.png" pageId="71" pageNumber="72">Fig. 34 F</figureCitation>
|
||
) with arms of anterior bifurcation straight and laterally well expanded at apex; anterior bifurcation broadly excavated receiving fourth rostral segment. Legs with tibiae and tarsi reddish, remaining yellow (
|
||
<figureCitation id="51D92A26FF94980DFBFDDB73716AFA8F" box="[1033,1145,1332,1359]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 B</figureCitation>
|
||
). Abdomen with spine of segment III rounded (
|
||
<figureCitation id="51D92A26FF94980DFE70DB1F74FEFAB3" box="[388,493,1368,1395]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 B</figureCitation>
|
||
). Intersegmental areas covered by very narrow black lines with well-defined margins, reaching the lateral spots (
|
||
<figureCitation id="51D92A26FF94980DFE4EDB3B7737FA57" box="[442,548,1404,1431]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 B</figureCitation>
|
||
). Pseudosutures concolorous with the surrounding surface (
|
||
<figureCitation id="51D92A26FF94980DFB4EDB3B7036FA56" box="[1210,1317,1404,1431]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 B</figureCitation>
|
||
, dark lines are a deterioration of the exoskeleton). Median longitudinal brown band incomplete, restricted to the segment VII (
|
||
<figureCitation id="51D92A26FF94980DFF6BDB837401FA1F" box="[159,274,1476,1503]" captionStart="FIGURE 65" captionStartId="118.[152,255,1943,1967]" captionTargetBox="[210,1377,193,1914]" captionTargetId="figure-22@118.[204,1383,181,1920]" captionTargetPageId="118" captionText="FIGURE 65. A–F. Dorsal and ventral surface: A–B, Edessa (E.) nigroacantha sp. n.; C–D, Edessa (E.) nigroantennata sp. n.; E–F, Edessa (E.) nigrocrocata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146481" httpUri="https://zenodo.org/record/10146481/files/figure.png" pageId="71" pageNumber="72">Fig. 65 B</figureCitation>
|
||
). Trichobothria both parallel to the spiracle. Male genitalia, posterolateral angles of pygophore very developed, well projected laterally, apices with spots black (
|
||
<figureCitation id="51D92A26FF94980DFCCEDBAF76F8F9C3" box="[826,1003,1512,1538]" captionStart="FIGURE 34" captionStartId="72.[152,255,1943,1967]" captionTargetBox="[184,1409,187,1913]" captionTargetId="figure-22@72.[178,1409,181,1919]" captionTargetPageId="72" captionText="FIGURE 34. A–F. Edessa (E.) nigroacantha sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146415" httpUri="https://zenodo.org/record/10146415/files/figure.png" pageId="71" pageNumber="72">Fig. 34 A–B, D</figureCitation>
|
||
). Superior processes of the genital cup subtriangular, narrow, continuing ventrally in a smooth high carina, ending in a dentiform projection (
|
||
<figureCitation id="51D92A26FF94980DFAF8D84B7084F9E6" box="[1292,1431,1548,1575]" captionStart="FIGURE 34" captionStartId="72.[152,255,1943,1967]" captionTargetBox="[184,1409,187,1913]" captionTargetId="figure-22@72.[178,1409,181,1919]" captionTargetPageId="72" captionText="FIGURE 34. A–F. Edessa (E.) nigroacantha sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146415" httpUri="https://zenodo.org/record/10146415/files/figure.png" pageId="71" pageNumber="72">Fig. 34 B, E</figureCitation>
|
||
, dentiform projection barely visible). Diaphragm with contrasting yellow spot (
|
||
<figureCitation id="51D92A26FF94980DFBEFD87771BCF98B" box="[1051,1199,1584,1610]" captionStart="FIGURE 34" captionStartId="72.[152,255,1943,1967]" captionTargetBox="[184,1409,187,1913]" captionTargetId="figure-22@72.[178,1409,181,1919]" captionTargetPageId="72" captionText="FIGURE 34. A–F. Edessa (E.) nigroacantha sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146415" httpUri="https://zenodo.org/record/10146415/files/figure.png" pageId="71" pageNumber="72">Fig. 34 D–E</figureCitation>
|
||
). Parameres (
|
||
<figureCitation id="51D92A26FF94980DFABED87775DFF9AF" captionStart="FIGURE 34" captionStartId="72.[152,255,1943,1967]" captionTargetBox="[184,1409,187,1913]" captionTargetId="figure-22@72.[178,1409,181,1919]" captionTargetPageId="72" captionText="FIGURE 34. A–F. Edessa (E.) nigroacantha sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146415" httpUri="https://zenodo.org/record/10146415/files/figure.png" pageId="71" pageNumber="72">Fig. 34 D–E</figureCitation>
|
||
) with black margins; anterior lobe subrectangular; dorsal lobe rounded with very curved apex; posterior lobe subrectangular. Proctiger, posterior face triangular (
|
||
<figureCitation id="51D92A26FF94980DFD28D83F767DF953" box="[732,878,1656,1682]" captionStart="FIGURE 34" captionStartId="72.[152,255,1943,1967]" captionTargetBox="[184,1409,187,1913]" captionTargetId="figure-22@72.[178,1409,181,1919]" captionTargetPageId="72" captionText="FIGURE 34. A–F. Edessa (E.) nigroacantha sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146415" httpUri="https://zenodo.org/record/10146415/files/figure.png" pageId="71" pageNumber="72">Fig. 34 D–E</figureCitation>
|
||
). Ventral surface sparsely punctuated (
|
||
<figureCitation id="51D92A26FF94980DFAD4D83F709CF953" box="[1312,1423,1656,1682]" captionStart="FIGURE 34" captionStartId="72.[152,255,1943,1967]" captionTargetBox="[184,1409,187,1913]" captionTargetId="figure-22@72.[178,1409,181,1919]" captionTargetPageId="72" captionText="FIGURE 34. A–F. Edessa (E.) nigroacantha sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146415" httpUri="https://zenodo.org/record/10146415/files/figure.png" pageId="71" pageNumber="72">Fig. 34 C</figureCitation>
|
||
). Ventral rim setose, with medial tuft (
|
||
<figureCitation id="51D92A26FF94980DFDC8D8DB77BEF977" box="[572,685,1692,1718]" captionStart="FIGURE 34" captionStartId="72.[152,255,1943,1967]" captionTargetBox="[184,1409,187,1913]" captionTargetId="figure-22@72.[178,1409,181,1919]" captionTargetPageId="72" captionText="FIGURE 34. A–F. Edessa (E.) nigroacantha sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146415" httpUri="https://zenodo.org/record/10146415/files/figure.png" pageId="71" pageNumber="72">Fig. 34 C</figureCitation>
|
||
); with expansions little developed, rounded, concolorous with the surrounding surface (
|
||
<figureCitation id="51D92A26FF94980DFE72D8877706F91B" box="[390,533,1728,1754]" captionStart="FIGURE 34" captionStartId="72.[152,255,1943,1967]" captionTargetBox="[184,1409,187,1913]" captionTargetId="figure-22@72.[178,1409,181,1919]" captionTargetPageId="72" captionText="FIGURE 34. A–F. Edessa (E.) nigroacantha sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146415" httpUri="https://zenodo.org/record/10146415/files/figure.png" pageId="71" pageNumber="72">Fig. 34 A–C</figureCitation>
|
||
). Female genitalia. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="81F86528FF94980DFF33D8A37636F817" pageId="71" pageNumber="72" type="discussion">
|
||
<paragraph id="C95D36A3FF94980DFF33D8A37636F817" blockId="71.[151,1437,684,2043]" pageId="71" pageNumber="72">
|
||
<emphasis id="FB96EAB1FF94980DFF33D8A3745CF93F" bold="true" box="[199,335,1764,1790]" pageId="71" pageNumber="72">Comments.</emphasis>
|
||
<taxonomicName id="0EE24D20FF94980DFEACD8A3776AF93F" authority="Mendonça & Silva & Fernandes, 2023" authorityName="Mendonça & Silva & Fernandes" authorityYear="2023" box="[344,633,1764,1790]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="nigroacantha" status="sp. nov." subGenus="Edessa">
|
||
<emphasis id="FB96EAB1FF94980DFEACD8A3776AF93F" box="[344,633,1764,1790]" italics="true" pageId="71" pageNumber="72">Edessa (E.) nigroacantha</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FB96EAB1FF94980DFD76D8A377D0F93F" bold="true" box="[642,707,1764,1790]" pageId="71" pageNumber="72">
|
||
<taxonomicNameLabel id="E0A557CAFF94980DFD76D8A377D0F93F" box="[642,707,1764,1790]" pageId="71" pageNumber="72" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resembles
|
||
<taxonomicName id="0EE24D20FF94980DFCB0D8A27111F93F" authority="Mendonça & Silva & Fernandes, 2023" authorityName="Mendonça & Silva & Fernandes" authorityYear="2023" box="[836,1026,1764,1790]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="omopalla" status="sp. n." subGenus="Edessa">
|
||
<emphasis id="FB96EAB1FF94980DFCB0D8A27111F93F" box="[836,1026,1764,1790]" italics="true" pageId="71" pageNumber="72">E. (E.) omopalla</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FB96EAB1FF94980DFBFFD8A3715FF93F" bold="true" box="[1035,1100,1764,1790]" pageId="71" pageNumber="72">
|
||
<taxonomicNameLabel id="E0A557CAFF94980DFBFFD8A3715FF93F" box="[1035,1100,1764,1790]" pageId="71" pageNumber="72" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
due to humeral angles black, globose and curved; connexival segments black with subrectangular spots; abdomen ventrally with intersegmental areas brown reaching the lateral spots; and the shape of superior processes of the genital cup. On the other hand, both species can be easily separated by the pygophore, the superior processes of the genital cup with smooth carina ending in a dentiform projection (
|
||
<taxonomicName id="0EE24D20FF94980DFDD2D93277FAF84F" authority="Mendonça & Silva & Fernandes, 2023" authorityName="Mendonça & Silva & Fernandes" authorityYear="2023" box="[550,745,1908,1934]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="omopalla" status="sp. n." subGenus="Edessa">
|
||
<emphasis id="FB96EAB1FF94980DFDD2D93277FAF84F" box="[550,745,1908,1934]" italics="true" pageId="71" pageNumber="72">E. (E.) omopalla</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FB96EAB1FF94980DFD00D9337624F84F" bold="true" box="[756,823,1908,1934]" pageId="71" pageNumber="72">
|
||
<taxonomicNameLabel id="E0A557CAFF94980DFD00D9337624F84F" box="[756,823,1908,1934]" pageId="71" pageNumber="72" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
with crenulate carina without dentiform projection); diaphragm with contrasting yellow spot (
|
||
<taxonomicName id="0EE24D20FF94980DFD96D9DE760FF873" authority="Mendonça & Silva & Fernandes, 2023" authorityName="Mendonça & Silva & Fernandes" authorityYear="2023" box="[610,796,1944,1970]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="omopalla" status="sp. n." subGenus="Edessa">
|
||
<emphasis id="FB96EAB1FF94980DFD96D9DE760FF873" box="[610,796,1944,1970]" italics="true" pageId="71" pageNumber="72">E. (E.) omopalla</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FB96EAB1FF94980DFCD7D9DF7671F873" bold="true" box="[803,866,1944,1970]" pageId="71" pageNumber="72">
|
||
<taxonomicNameLabel id="E0A557CAFF94980DFCD7D9DF7671F873" box="[803,866,1944,1970]" pageId="71" pageNumber="72" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
without spot) and ventral rim with lateral subdistal tuft of setae (
|
||
<taxonomicName id="0EE24D20FF94980DFED8D9FA74F4F817" authority="Mendonça & Silva & Fernandes, 2023" authorityName="Mendonça & Silva & Fernandes" authorityYear="2023" box="[300,487,1980,2006]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="omopalla" status="sp. n." subGenus="Edessa">
|
||
<emphasis id="FB96EAB1FF94980DFED8D9FA74F4F817" box="[300,487,1980,2006]" italics="true" pageId="71" pageNumber="72">E. (E.) omopalla</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FB96EAB1FF94980DFE19D9FB773EF817" bold="true" box="[493,557,1980,2006]" pageId="71" pageNumber="72">
|
||
<taxonomicNameLabel id="E0A557CAFF94980DFE19D9FB773EF817" box="[493,557,1980,2006]" pageId="71" pageNumber="72" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
without tuft of setae).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="81F86528FF94980DFF33D9A777B4F83A" box="[199,679,2016,2043]" pageId="71" pageNumber="72" type="distribution">
|
||
<paragraph id="C95D36A3FF94980DFF33D9A777B4F83A" blockId="71.[151,1437,684,2043]" box="[199,679,2016,2043]" pageId="71" pageNumber="72">
|
||
<emphasis id="FB96EAB1FF94980DFF33D9A7744AF83B" bold="true" box="[199,345,2016,2042]" pageId="71" pageNumber="72">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="51D92A26FF94980DFE9DD9A774A9F83B" box="[361,442,2016,2042]" captionStart="FIGURE 72" captionStartId="125.[152,255,1071,1095]" captionTargetBox="[195,1391,181,1040]" captionTargetId="figure-20@125.[195,1391,181,1050]" captionTargetPageId="125" captionText="FIGURE 72. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.10146495" httpUri="https://zenodo.org/record/10146495/files/figure.png" pageId="71" pageNumber="72">Fig. 72</figureCitation>
|
||
).
|
||
<collectingCountry id="B1F57633FF94980DFE24D9A7775BF83B" box="[464,584,2016,2042]" name="Bolivia" pageId="71" pageNumber="72">BOLIVIA</collectingCountry>
|
||
:
|
||
<collectingRegion id="0B26F841FF94980DFDA6D9A677B0F83A" box="[594,675,2017,2043]" country="Bolivia" name="La Paz" pageId="71" pageNumber="72">La Paz</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |