398 lines
56 KiB
XML
398 lines
56 KiB
XML
<document id="FAE8FB10BB3379B6753700183F7DB105" ID-DOI="10.11646/zootaxa.3994.1.1" ID-GBIF-Dataset="1334b429-62a4-4505-9e05-d68b39058e5c" ID-ISSN="1175-5326" ID-Zenodo-Dep="289137" ID-ZooBank="093B124E-58AE-4303-8C07-2D7B27E6AC38" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461271356316" checkinUser="plazi" docAuthor="Nogueira, João Miguel De Matos, Carrerette, Orlemir & Hutchings, Pat" docDate="2015" docId="DD7687BBFFACFF8FFF66FF62D9C6FE71" docLanguage="en" docName="zt03994p052.pdf" docOrigin="Zootaxa 3994 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amaeana apheles Hutchings 1974" docType="treatment" docVersion="6" lastPageNumber="24" masterDocId="214FFFC3FFBFFF98FFF1FFF5D92AFF84" masterDocTitle="Review of Amaeana Hartman, 1959 (Annelida, Terebelliformia, Polycirridae), with descriptions of seven new species" masterLastPageNumber="52" masterPageNumber="1" pageNumber="20" updateTime="1698611839755" updateUser="plazi">
|
||
<mods:mods id="0D39D2CE2A5F177B8CC257DF2CCC84D7" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="25B04BB5FC4E8388B541542553FDFA93">
|
||
<mods:title id="A596A53EEC938F1130EAD6AC55F36C49">Review of Amaeana Hartman, 1959 (Annelida, Terebelliformia, Polycirridae), with descriptions of seven new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="879E3EB0949D0743FC472C9EF8D575A6" type="personal">
|
||
<mods:role id="1F65CE51317F473897EE8942913471FB">
|
||
<mods:roleTerm id="E16E7357C5AA4DB5BBB31F4F62125DE1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="85BB7F23E8F48C726D757A0DE38ACC95">Nogueira, João Miguel De Matos</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="FAD16656C7EAF753260EAEFE46B12CA0" type="personal">
|
||
<mods:role id="B266FCB238957646B633971B0688B78F">
|
||
<mods:roleTerm id="0C18D8DF9842C0C54C4F5CDDB72050DE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="05DB7F7C5AF4D619539D529B8D40485B">Carrerette, Orlemir</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="E504BF18D289B9E1E41032FD4A2F6C9A" type="personal">
|
||
<mods:role id="3E217900B8BFD88D21D4A382507A1A4C">
|
||
<mods:roleTerm id="9D03ECF174A07BAF3A6E0F65A7CFFD84">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="11A68029E902026DA4C6B7549F470A48">Hutchings, Pat</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="11E3E25FD71F2670E8A620CB8EB5C9F0">text</mods:typeOfResource>
|
||
<mods:relatedItem id="730E6599857C7211797CE60CF0E27B39" type="host">
|
||
<mods:titleInfo id="B9F957F6689430AB1F5E45F3B1BEAB08">
|
||
<mods:title id="A2F330D4000A8E4E00A4B93114E9DC43">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="8EBF6CECF856503C9DA440AE0A338D45">
|
||
<mods:date id="749E0756E6268BE9F74917081EE269C0">2015</mods:date>
|
||
<mods:detail id="14E1977E9DF00668A8A6ED867BDFC02C" type="volume">
|
||
<mods:number id="6AAA3E8B3F747287F2C1E2EB8A2261FE">3994</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="C6BFB8B98A1997F3991AF101E7F7EE50" type="issue">
|
||
<mods:number id="90A719A7F46F8A2FF4D2A3C652F826BF">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="6669F4A44E467A3DD3588B950BD45693" unit="page">
|
||
<mods:start id="36AAF4676A1A5A3E8D0701B8023A0485">1</mods:start>
|
||
<mods:end id="FC042FD667CB7E623F298537C88A1FB3">52</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="F3CFDBB5BFD7A2F23C382BD6E81C9898">journal article</mods:classification>
|
||
<mods:identifier id="85B0536A1B99435A94526191B045DF13" type="DOI">10.11646/zootaxa.3994.1.1</mods:identifier>
|
||
<mods:identifier id="94B6D52E5C42FA670F4B771203610ED6" type="GBIF-Dataset">1334b429-62a4-4505-9e05-d68b39058e5c</mods:identifier>
|
||
<mods:identifier id="9DD5B855EF83B82810C44DDEF68AABCB" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="DAF2568AB62AF629A628C74A51DAB045" type="Zenodo-Dep">289137</mods:identifier>
|
||
<mods:identifier id="9CF05B244E21B6FC6D8F7967203D5703" type="ZooBank">093B124E-58AE-4303-8C07-2D7B27E6AC38</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="DD7687BBFFACFF8FFF66FF62D9C6FE71" ID-DOI="http://doi.org/10.5281/zenodo.6094904" ID-GBIF-Taxon="127692608" ID-Zenodo-Dep="6094904" LSID="urn:lsid:plazi:treatment:DD7687BBFFACFF8FFF66FF62D9C6FE71" httpUri="http://treatment.plazi.org/id/DD7687BBFFACFF8FFF66FF62D9C6FE71" lastPageId="23" lastPageNumber="24" pageId="19" pageNumber="20">
|
||
<subSubSection id="1DC56526FFACFF8BFF66FF62D85FFF57" pageId="19" pageNumber="20" type="nomenclature">
|
||
<paragraph id="556036ADFFACFF8BFF66FF62DB62FF36" blockId="19.[151,584,151,211]" box="[151,584,151,178]" pageId="19" pageNumber="20">
|
||
<heading id="0E2881C1FFACFF8BFF66FF62DB62FF36" bold="true" box="[151,584,151,178]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
|
||
<taxonomicName id="92DF4D2EFFACFF8BFF66FF62DB62FF36" ID-CoL="5TXMF" authority="Hutchings, 1974" authorityName="Hutchings" authorityYear="1974" box="[151,584,151,178]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="19" pageNumber="20" phylum="Annelida" rank="species" species="apheles">
|
||
<emphasis id="67ABEABFFFACFF8BFF66FF62DB62FF36" bold="true" box="[151,584,151,178]" pageId="19" pageNumber="20">
|
||
<emphasis id="67ABEABFFFACFF8BFF66FF62D842FF35" bold="true" box="[151,360,151,177]" italics="true" pageId="19" pageNumber="20">Amaeana apheles</emphasis>
|
||
(
|
||
<bibRefCitation id="314E4B5CFFACFF8BFE86FF6DDB6AFF36" author="Hutchings" box="[375,576,152,178]" pageId="19" pageNumber="20" refString="Hutchings, P. A. (1974) Polychaeta of WalIis Lake, New South Wales. Proceedings of the Linnean Society of New South Wales, 98 (4), 175 - 195." type="journal article" year="1974">Hutchings, 1974</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="556036ADFFACFF8BFF66FF4FD85FFF57" blockId="19.[151,584,151,211]" box="[151,373,186,211]" pageId="19" pageNumber="20">
|
||
<figureCitation id="CDE42A28FFACFF8BFF66FF4FD9D5FF57" box="[151,255,186,211]" captionStart="FIGURE 9" captionStartId="20.[151,250,1851,1873]" captionTargetBox="[193,1404,197,1818]" captionTargetId="figure@20.[187,1408,193,1826]" captionTargetPageId="20" captionText="FIGURE 9. Amaeana apheles, holotype AM W. 5239. A – C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D – F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial / genital papillae. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W. 29203: I – J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.7 mm; B – C = 1 mm; D – F = 0.5 mm; G – H = 0.3 mm; I – K = 20 µm." httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="19" pageNumber="20">Figures 9</figureCitation>
|
||
,
|
||
<figureCitation id="CDE42A28FFACFF8BFEFAFF4ED80DFF57" box="[267,295,187,211]" captionStart="FIGURE 10" captionStartId="21.[151,250,1882,1904]" captionTargetBox="[204,1384,181,1856]" captionTargetId="figure@21.[198,1388,177,1861]" captionTargetPageId="21" captionText="FIGURE 10. Amaeana apheles, photos D and G from specimen AM W. 10864, all other photos from specimen AM W. 19333. A – C, thorax, right lateral, ventral and dorsal views, respectively. D – E, close ups of the anterior end, left and right lateral views, respectively. F – G, close ups of the anterior end, dorsal view. H, detail of the oral area. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A – B = 0.7 mm; C = 0.6 mm; D = 0.15 mm; E – F = 0.2 mm; G – H = 0.1 mm." httpUri="https://zenodo.org/record/289147/files/figure.png" pageId="19" pageNumber="20">10</figureCitation>
|
||
and
|
||
<figureCitation id="CDE42A28FFACFF8BFEAAFF4ED85FFF57" box="[347,373,187,211]" captionStart="FIGURE 11" captionStartId="22.[151,250,1897,1919]" captionTargetBox="[171,1413,196,1851]" captionTargetId="figure@22.[164,1417,192,1859]" captionTargetPageId="22" captionText="FIGURE 11. Amaeana apheles, photos A – E from specimen AM W. 19333, photos F – I from specimen AM W. 10864. A, notopodia, segments 9 – 10; B – E, notochaetae, segment 10; F – G, posterior neuropodia; H – I, close up of posterior neuropodia with protruding spines. Scale bars: A = 70 µm; B, I = 10 µm; C – D = 5 µm; E = 4 µm; F = 200 µm; G = 8 µm; H = 20 µm." httpUri="https://zenodo.org/record/289148/files/figure.png" pageId="19" pageNumber="20">11</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1DC56526FFACFF8BFF66FF08DA66FEB0" pageId="19" pageNumber="20" type="reference_group">
|
||
<paragraph id="556036ADFFACFF8BFF66FF08DA66FEB0" blockId="19.[151,1253,253,308]" pageId="19" pageNumber="20">
|
||
<treatmentCitationGroup id="75CF1183FFACFF8BFF66FF08DA66FEB0" pageId="19" pageNumber="20">
|
||
<taxonomicName id="92DF4D2EFFACFF8BFF66FF08DB28FE97" ID-CoL="6R2ZP" authority="Hutchings 1974: 190" authorityName="Hutchings" authorityPageNumber="190" authorityYear="1974" box="[151,514,253,275]" class="Polychaeta" family="Terebellidae" genus="Lysilla" kingdom="Animalia" order="Terebellida" pageId="19" pageNumber="20" phylum="Annelida" rank="species" species="apheles">
|
||
<emphasis id="67ABEABFFFACFF8BFF66FF08D801FE97" box="[151,299,253,275]" italics="true" pageId="19" pageNumber="20">Lysilla apheles</emphasis>
|
||
<treatmentCitation id="D47E10BCFFACFF8BFEC0FF08DB28FE97" author="Hutchings" box="[305,514,253,275]" page="190" pageId="19" pageNumber="20" year="1974">
|
||
<bibRefCitation id="314E4B5CFFACFF8BFEC0FF08D8F8FE97" author="Hutchings" box="[305,466,253,275]" pageId="19" pageNumber="20" refString="Hutchings, P. A. (1974) Polychaeta of WalIis Lake, New South Wales. Proceedings of the Linnean Society of New South Wales, 98 (4), 175 - 195." type="journal article" year="1974">Hutchings 1974</bibRefCitation>
|
||
: 190
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
–191,
|
||
<figureCitation id="CDE42A28FFACFF8BFDCFFF08DB5CFE97" box="[574,630,253,275]" captionStart="FIGURE 5" captionStartId="15.[151,250,1791,1813]" captionTargetBox="[236,1361,191,1760]" captionTargetId="figure@15.[230,1364,187,1773]" captionTargetPageId="15" captionText="FIGURE 5. Amaeana hsiehae sp. nov., ASIZW 0000935. A – D, entire worm, right ventro-lateral, right lateral, left lateral and left dorso-lateral views, respectively; arrows point to beginning of neuropodia; E – F, progressively closer views of the anterior end, ventral view; G – H, thorax, right and left lateral views, respectively; I – K, close ups of the anterior end, left dorso-lateral, ventral and right lateral views, respectively; L, posterior end, left dorso-lateral view; M, close up of the anterior end, left dorsolateral view; N, abdominal segments; arrows point to neuropodia; O, posterior end, right ventro-lateral view. Paratype AM W. 47363: P, entire worm, left lateral view (anteriorly); Q, thorax, left lateral view. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip. Scale bars: A = 0.6 mm; B – D, H = 0.5 mm; E, G = 0.4 mm; F, I – L, N = 0.2 mm; M, O = 0.15 mm; P = 1.5 mm; Q = 1 mm." httpUri="https://zenodo.org/record/289142/files/figure.png" pageId="19" pageNumber="20">Fig. 5</figureCitation>
|
||
A;—
|
||
<treatmentCitation id="D47E10BCFFACFF8BFD57FF08DA41FE97" author="Hutchings" box="[678,875,253,275]" page="10" pageId="19" pageNumber="20" year="1977">
|
||
<bibRefCitation id="314E4B5CFFACFF8BFD57FF08DA60FE97" author="Hutchings" box="[678,842,253,275]" pageId="19" pageNumber="20" refString="Hutchings, P. A. (1977) TerebelIiform Polychaeta of the families Ampharetidae, Terebellidae and Trichobranchidae from Australia, chiefly from Moreton Bay, Queensland. Records of the Australian Museum, 31, 1 - 38. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.31.1977.232" type="journal article" year="1977">Hutchings 1977</bibRefCitation>
|
||
: 10
|
||
</treatmentCitation>
|
||
–11;—
|
||
<treatmentCitation id="D47E10BCFFACFF8BFC5DFF08DDCAFE97" author="Hutchings" box="[940,1248,253,275]" page="90" pageId="19" pageNumber="20" year="1984">
|
||
<bibRefCitation id="314E4B5CFFACFF8BFC5DFF08DD96FE97" author="Hutchings" box="[940,1212,253,275]" pageId="19" pageNumber="20" refString="Hutchings, P. A. & Murray, A. (1984) Taxonomy of Polychaetes from the Hawkesbury River and the southern estuaries of New South Wales, Australia. Records of the Australian Museum, 3 (Supplement), 1 - 118. http: // dx. doi. org / 10.3853 / j. 0812 - 7387.3.1984.101" type="journal article" year="1984">Hutchings & Murray 1984</bibRefCitation>
|
||
: 90
|
||
</treatmentCitation>
|
||
.
|
||
<emphasis id="67ABEABFFFACFF8BFF66FEEAD848FEB0" box="[151,354,286,308]" italics="true" pageId="19" pageNumber="20">
|
||
<taxonomicName id="92DF4D2EFFACFF8BFF66FEEAD86CFEB0" ID-CoL="5TXMF" box="[151,326,286,308]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="19" pageNumber="20" phylum="Annelida" rank="species" species="apheles">Amaeana apheles</taxonomicName>
|
||
.—
|
||
</emphasis>
|
||
<treatmentCitation id="D47E10BCFFACFF8BFE92FEEBDBB0FEB0" author="Hutchings" box="[355,666,286,308]" page="321" pageId="19" pageNumber="20" year="1986">
|
||
<bibRefCitation id="314E4B5CFFACFF8BFE92FEEBDB43FEB0" author="Hutchings" box="[355,617,286,308]" pageId="19" pageNumber="20" refString="Hutchings, P. A. & Glasby, C. J. (1986) The Polycirrinae (F. Terebellidae) of Australia. Records of the Australian Museum, 38, 319 - 350. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.38.1986.185" type="journal article" year="1986">Hutchings & Glasby 1986</bibRefCitation>
|
||
: 321
|
||
</treatmentCitation>
|
||
,
|
||
<figureCitation id="CDE42A28FFACFF8BFD55FEEBDBF7FEB0" box="[676,733,286,308]" captionStart="FIGURE 1" captionStartId="8.[151,250,1538,1560]" captionTargetBox="[157,1431,805,1488]" captionTargetId="figure@8.[151,1435,801,1491]" captionTargetPageId="8" captionText="FIGURE 1. Amaeana ellobophora, paratype 1 AM W. 10341, with egg cocoons: A, entire worm, dorsal view. B, anterior end, dorsal view. Scale bars: A = 0.5 mm; B = 0.3 mm." httpUri="https://zenodo.org/record/289138/files/figure.png" pageId="19" pageNumber="20">Figs 1</figureCitation>
|
||
C–E; 12A.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1DC56526FFACFF8BFF66FEA9DDE2FD5B" pageId="19" pageNumber="20" type="materials_examined">
|
||
<paragraph id="556036ADFFACFF8BFF66FEA9DC0FFE7C" blockId="19.[151,1437,348,1993]" pageId="19" pageNumber="20">
|
||
<emphasis id="67ABEABFFFACFF8BFF66FEA9D8AAFEF1" bold="true" box="[151,384,348,373]" pageId="19" pageNumber="20">Material examined.</emphasis>
|
||
<typeStatus id="8A64880FFFACFF8BFE76FEA9D8DEFEF1" box="[391,500,348,373]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
|
||
: AM W.5239: incomplete specimen, with 11 abdominal segments, in excellent state of preservation,
|
||
<quantity id="92279B48FFACFF8BFEBEFE88D8B5FE11" box="[335,415,381,405]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="20" unit="mm" value="12.0">12 mm</quantity>
|
||
long, ~
|
||
<quantity id="92279B48FFACFF8BFE04FE89DB67FE11" box="[501,589,380,405]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="20" unit="mm" value="2.5">2.5 mm</quantity>
|
||
wide at posterior thorax.
|
||
<typeStatus id="8A64880FFFACFF8BFC81FE8BDAC8FE12" box="[880,994,382,406]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
|
||
: AM W.5237: incomplete specimen, in good state of preservation, ~
|
||
<quantity id="92279B48FFACFF8BFE17FE6BDB13FE32" box="[486,569,414,438]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="19" pageNumber="20" unit="mm" value="14.0">14 mm</quantity>
|
||
long, ~
|
||
<quantity id="92279B48FFACFF8BFD66FE6BDBDBFE32" box="[663,753,414,438]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="19" pageNumber="20" unit="mm" value="1.8">1.8 mm</quantity>
|
||
wide. AM W.5238: incomplete specimen in good state of preservation, ~
|
||
<quantity id="92279B48FFACFF8BFEB1FE4AD8BBFE53" box="[320,401,447,472]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="20" unit="mm" value="15.0">15 mm</quantity>
|
||
long, ~
|
||
<quantity id="92279B48FFACFF8BFE18FE4ADB06FE53" box="[489,556,447,471]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="mm" value="2.0">2 mm</quantity>
|
||
wide. All
|
||
<typeStatus id="8A64880FFFACFF8BFD56FE35DBFDFE5C" box="[679,727,448,472]" pageId="19" pageNumber="20">type</typeStatus>
|
||
material collected from—Australia, New South
|
||
<collectingCountry id="2DC8763DFFACFF8BFAF3FE4ADC63FE5C" box="[1282,1353,447,472]" name="United Kingdom" pageId="19" pageNumber="20">Wales</collectingCountry>
|
||
,
|
||
<collectingCountry id="2DC8763DFFACFF8BFAA4FE4ADCB6FE5C" box="[1365,1436,447,472]" name="United Kingdom" pageId="19" pageNumber="20">Wallis</collectingCountry>
|
||
Lake,
|
||
<geoCoordinate id="30EB506AFFACFF8BFF2CFE15D81DFE7C" box="[221,311,480,504]" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="925" value="-32.283333">32°17'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="30EB506AFFACFF8BFEB2FE15D886FE7D" box="[323,428,480,505]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="925" value="152.48334">152°29'E</geoCoordinate>
|
||
,
|
||
<date id="2161106DFFACFF8BFE49FE15DB0CFE7C" box="[440,550,480,504]" pageId="19" pageNumber="20" value="1970-12">Dec.1970</date>
|
||
, coll. by P. Dixon & K. O’Gower, in seagrass beds in shallow water.
|
||
</paragraph>
|
||
<paragraph id="556036ADFFACFF8BFF36FDF4DD99FD3B" blockId="19.[151,1437,348,1993]" pageId="19" pageNumber="20">
|
||
<emphasis id="67ABEABFFFACFF8BFF36FDF4DB11FD9E" bold="true" box="[199,571,513,538]" pageId="19" pageNumber="20">Additional material examined</emphasis>
|
||
. AM W.5384 (Queensland, Mary River, Kangaroo Island, Hervey Bay,
|
||
<geoCoordinate id="30EB506AFFACFF8BFF66FDD7D9DAFDBF" box="[151,240,546,571]" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="925" value="-25.416666">25°25'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="30EB506AFFACFF8BFEF2FDD7D847FDBF" box="[259,365,546,571]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="925" value="152.93333">152°56'E</geoCoordinate>
|
||
,
|
||
<date id="2161106DFFACFF8BFE8EFDD6D8DAFDBF" box="[383,496,547,571]" pageId="19" pageNumber="20" value="1971-12">Dec.1971</date>
|
||
):
|
||
<specimenCount id="43D9FD24FFACFF8BFDFBFDD6DBB7FDBF" box="[522,669,546,571]" pageId="19" pageNumber="20" type="generic">2 specimens</specimenCount>
|
||
, 1 of which complete. AM W.29203 (Queensland, Mary River, Kangaroo Island, Hervey Bay,
|
||
<geoCoordinate id="30EB506AFFACFF8BFE09FDB6DB7BFDD8" box="[504,593,579,604]" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="925" value="-25.416666">25°25'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="30EB506AFFACFF8BFDAEFDB6DBE3FDD8" box="[607,713,579,604]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="925" value="152.93333">152°56'E</geoCoordinate>
|
||
,
|
||
<date id="2161106DFFACFF8BFD27FDB1DA6DFDD8" box="[726,839,580,604]" pageId="19" pageNumber="20" value="1971-12">Dec.1971</date>
|
||
): complete, in excellent state of preservation; slides: notopodium, segment 9; anterior and posterior neuropodia. AM W.10864 (
|
||
<collectingCountry id="2DC8763DFFACFF8BFC1EFD91DD70FDF9" box="[1007,1114,612,637]" name="Australia" pageId="19" pageNumber="20">Australia</collectingCountry>
|
||
, Queensland, Nerang River,
|
||
<collectingCountry id="2DC8763DFFACFF8BFF66FD70D80AFD1A" box="[151,288,645,670]" name="Ghana" pageId="19" pageNumber="20">Gold Coast</collectingCountry>
|
||
,
|
||
<geoCoordinate id="30EB506AFFACFF8BFEC3FD70D854FD1A" box="[306,382,645,670]" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="925" value="-28.1">28°6'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="30EB506AFFACFF8BFE61FD70D8D0FD1A" box="[400,506,645,670]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="925" value="153.3">153°18'E</geoCoordinate>
|
||
, 1975): mounted on SEM pin. AM W.19333 (
|
||
<collectingCountry id="2DC8763DFFACFF8BFBB3FD70DD87FD1A" box="[1090,1197,645,670]" name="Australia" pageId="19" pageNumber="20">Australia</collectingCountry>
|
||
, New South
|
||
<collectingCountry id="2DC8763DFFACFF8BFAA3FD70DCB3FD1A" box="[1362,1433,645,670]" name="United Kingdom" pageId="19" pageNumber="20">Wales</collectingCountry>
|
||
, Hawkesbury River, near Juno Head,
|
||
<geoCoordinate id="30EB506AFFACFF8BFDCBFD52DBB9FD3B" box="[570,659,679,703]" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="925" value="-33.566666">33°34'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="30EB506AFFACFF8BFD51FD53DA3AFD3A" box="[672,784,678,703]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="925" value="151.25">151°15' E</geoCoordinate>
|
||
,
|
||
<date id="2161106DFFACFF8BFCEDFD52DA82FD3B" box="[796,936,679,703]" pageId="19" pageNumber="20" value="1977-01-12">12.Jan.1977</date>
|
||
): mounted on SEM pin.
|
||
</paragraph>
|
||
<paragraph id="556036ADFFACFF8BFF36FD32DDE2FD5B" blockId="19.[151,1437,348,1993]" box="[199,1224,711,736]" pageId="19" pageNumber="20">
|
||
<emphasis id="67ABEABFFFACFF8BFF36FD32D84FFD64" bold="true" box="[199,357,711,736]" pageId="19" pageNumber="20">
|
||
<typeStatus id="8A64880FFFACFF8BFF36FD32D828FD64" box="[199,258,711,736]" pageId="19" pageNumber="20">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="2DC8763DFFACFF8BFE9DFD32D8FCFD64" box="[364,470,711,736]" name="Australia" pageId="19" pageNumber="20">Australia</collectingCountry>
|
||
, New South
|
||
<collectingCountry id="2DC8763DFFACFF8BFD94FD32DB86FD64" box="[613,684,711,736]" name="United Kingdom" pageId="19" pageNumber="20">Wales</collectingCountry>
|
||
,
|
||
<collectingCountry id="2DC8763DFFACFF8BFD46FD32DBD4FD64" box="[695,766,711,736]" name="United Kingdom" pageId="19" pageNumber="20">Wallis</collectingCountry>
|
||
Lake,
|
||
<geoCoordinate id="30EB506AFFACFF8BFCBBFD3DDA8FFD64" box="[842,933,712,736]" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="925" value="-32.283333">32°17'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="30EB506AFFACFF8BFC41FD32DD30FD64" box="[944,1050,711,736]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="925" value="152.48334">152°29'E</geoCoordinate>
|
||
, shallow water.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1DC56526FFACFF8BFF36FD1DDBD2F965" pageId="19" pageNumber="20" type="description">
|
||
<paragraph id="556036ADFFACFF8BFF36FD1DDA58FCA6" blockId="19.[151,1437,348,1993]" pageId="19" pageNumber="20">
|
||
<emphasis id="67ABEABFFFACFF8BFF36FD1DD87BFC85" bold="true" box="[199,337,744,769]" pageId="19" pageNumber="20">Description</emphasis>
|
||
:
|
||
<typeStatus id="8A64880FFFACFF8BFE91FD1DD8E2FC85" box="[352,456,744,769]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
|
||
incomplete specimen,
|
||
<quantity id="92279B48FFACFF8BFD3BFD1CDA33FC84" box="[714,793,745,769]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="20" unit="mm" value="12.0">12 mm</quantity>
|
||
long, ~
|
||
<quantity id="92279B48FFACFF8BFC9FFD1DDAEFFC84" box="[878,965,744,769]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="20" unit="mm" value="2.5">2.5 mm</quantity>
|
||
wide at posterior thorax, maximum width of body; longest specimen examined ~
|
||
<quantity id="92279B48FFACFF8BFDB9FCFCDBBDFCA5" box="[584,663,777,802]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="20" unit="mm" value="15.0">15 mm</quantity>
|
||
long, ~
|
||
<quantity id="92279B48FFACFF8BFD1DFCFFDA07FCA5" box="[748,813,778,802]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="mm" value="2.0">2 mm</quantity>
|
||
wide.
|
||
</paragraph>
|
||
<paragraph id="556036ADFFACFF8BFF36FCDEDDBBFC6C" blockId="19.[151,1437,348,1993]" pageId="19" pageNumber="20">
|
||
Prostomium at base of upper lip, both basal and distal parts developed, basal part as thickened crest, distal part with large, flaring lobes and also short, rounded to squared mid-dorsal process; prostomium covering segment 1 laterally and terminating laterally to lower lip, near mouth (
|
||
<figureCitation id="CDE42A28FFACFF8BFCCDFC99DAA9FC01" box="[828,899,876,901]" captionStart="FIGURE 9" captionStartId="20.[151,250,1851,1873]" captionTargetBox="[193,1404,197,1818]" captionTargetId="figure@20.[187,1408,193,1826]" captionTargetPageId="20" captionText="FIGURE 9. Amaeana apheles, holotype AM W. 5239. A – C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D – F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial / genital papillae. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W. 29203: I – J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.7 mm; B – C = 1 mm; D – F = 0.5 mm; G – H = 0.3 mm; I – K = 20 µm." httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="19" pageNumber="20">Figs 9</figureCitation>
|
||
A–H; 10A–H). Three
|
||
<typeStatus id="8A64880FFFACFF8BFB8EFC9BDD90FC02" box="[1151,1210,878,902]" pageId="19" pageNumber="20">types</typeStatus>
|
||
of buccal tentacles, short ones thin, uniformly cylindrical; intermediate tentacles distally spatulate, with convolute margins; long buccal tentacles progressively widening towards subdistal cylindrical inflation, collar-like, with relatively long pointed to distally blunt tip, as wide as tentacle basally to collar-like inflation (
|
||
<figureCitation id="CDE42A28FFACFF8BFBF6FC25DD60FC6C" box="[1031,1098,976,1001]" captionStart="FIGURE 9" captionStartId="20.[151,250,1851,1873]" captionTargetBox="[193,1404,197,1818]" captionTargetId="figure@20.[187,1408,193,1826]" captionTargetPageId="20" captionText="FIGURE 9. Amaeana apheles, holotype AM W. 5239. A – C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D – F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial / genital papillae. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W. 29203: I – J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.7 mm; B – C = 1 mm; D – F = 0.5 mm; G – H = 0.3 mm; I – K = 20 µm." httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
|
||
A–H).
|
||
</paragraph>
|
||
<paragraph id="556036ADFFACFF8BFF36FC04DA0BFBAF" blockId="19.[151,1437,348,1993]" pageId="19" pageNumber="20">
|
||
Peristomium restricted to lips, upper lip almost circular, usually folded into three lobes; lower lip short, buttonlike, rectangular to crescent-shaped (
|
||
<figureCitation id="CDE42A28FFACFF8BFDC3FBE7DB52FBAF" box="[562,632,1042,1067]" captionStart="FIGURE 9" captionStartId="20.[151,250,1851,1873]" captionTargetBox="[193,1404,197,1818]" captionTargetId="figure@20.[187,1408,193,1826]" captionTargetPageId="20" captionText="FIGURE 9. Amaeana apheles, holotype AM W. 5239. A – C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D – F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial / genital papillae. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W. 29203: I – J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.7 mm; B – C = 1 mm; D – F = 0.5 mm; G – H = 0.3 mm; I – K = 20 µm." httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="19" pageNumber="20">Figs 9</figureCitation>
|
||
A–H; 10A–H).
|
||
</paragraph>
|
||
<paragraph id="556036ADFFACFF8BFF36FBC6DC1AFB54" blockId="19.[151,1437,348,1993]" pageId="19" pageNumber="20">
|
||
Body anteriorly swollen, progressively broader until segments 10–11, then of uniform width until segments 19–20, tapering on segments 20–23, and again of relatively uniform width through posterior body, distinctly narrower than anterior segments (
|
||
<figureCitation id="CDE42A28FFACFF8BFDE2FB80DB70FB0A" box="[531,602,1141,1166]" captionStart="FIGURE 9" captionStartId="20.[151,250,1851,1873]" captionTargetBox="[193,1404,197,1818]" captionTargetId="figure@20.[187,1408,193,1826]" captionTargetPageId="20" captionText="FIGURE 9. Amaeana apheles, holotype AM W. 5239. A – C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D – F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial / genital papillae. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W. 29203: I – J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.7 mm; B – C = 1 mm; D – F = 0.5 mm; G – H = 0.3 mm; I – K = 20 µm." httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="19" pageNumber="20">Figs 9</figureCitation>
|
||
A–H; 10A–F; 11G); achaetous gap between termination of notopodia and beginning of neuropodia, corresponding to segments
|
||
<date id="2161106DFFACFF8BFD19FB62DAB2FB2B" box="[744,920,1175,1199]" pageId="19" pageNumber="20" value="1922-12-13" valueMax="1922-12-21" valueMin="1922-12-13">12–13 to 21–22</date>
|
||
(9–10 segments), twice as long as region with notopodia, with poorly marked segmentation and fragile, with thin body wall dorsally (
|
||
<figureCitation id="CDE42A28FFACFF8BFB97FB42DD84FB54" box="[1126,1198,1207,1232]" captionStart="FIGURE 9" captionStartId="20.[151,250,1851,1873]" captionTargetBox="[193,1404,197,1818]" captionTargetId="figure@20.[187,1408,193,1826]" captionTargetPageId="20" captionText="FIGURE 9. Amaeana apheles, holotype AM W. 5239. A – C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D – F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial / genital papillae. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W. 29203: I – J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.7 mm; B – C = 1 mm; D – F = 0.5 mm; G – H = 0.3 mm; I – K = 20 µm." httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="19" pageNumber="20">Figs 9</figureCitation>
|
||
B; 10A–C).
|
||
</paragraph>
|
||
<paragraph id="556036ADFFACFF8BFF36FB2CDD5DFA13" blockId="19.[151,1437,348,1993]" pageId="19" pageNumber="20">
|
||
Segments biannulated, segment 1 short, visible dorsally and ventrally, laterally covered by expanded prostomium; segment 2 narrower and shorter than following segments, with small, pentagonal mid-ventral shield at beginning of mid-ventral groove, extending anteriorly through segment 1 until near ventral edge of lower lip (
|
||
<figureCitation id="CDE42A28FFACFF8BFA9BFAEED98EFAD0" captionStart="FIGURE 9" captionStartId="20.[151,250,1851,1873]" captionTargetBox="[193,1404,197,1818]" captionTargetId="figure@20.[187,1408,193,1826]" captionTargetPageId="20" captionText="FIGURE 9. Amaeana apheles, holotype AM W. 5239. A – C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D – F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial / genital papillae. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W. 29203: I – J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.7 mm; B – C = 1 mm; D – F = 0.5 mm; G – H = 0.3 mm; I – K = 20 µm." httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="19" pageNumber="20">Figs 9</figureCitation>
|
||
A–H; 10A–H). Ventrum highly glandular, covered on small papillae, arranged on paired ventro-lateral pads on segments 2–12; papillae larger and more numerous on anterior segments, progressively less numerous on segments 11–12, still conspicuous on both parts of each segment (
|
||
<figureCitation id="CDE42A28FFACFF8BFCFBFA8BDA7AFA13" box="[778,848,1406,1431]" captionStart="FIGURE 9" captionStartId="20.[151,250,1851,1873]" captionTargetBox="[193,1404,197,1818]" captionTargetId="figure@20.[187,1408,193,1826]" captionTargetPageId="20" captionText="FIGURE 9. Amaeana apheles, holotype AM W. 5239. A – C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D – F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial / genital papillae. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W. 29203: I – J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.7 mm; B – C = 1 mm; D – F = 0.5 mm; G – H = 0.3 mm; I – K = 20 µm." httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="19" pageNumber="20">Figs 9</figureCitation>
|
||
A–F, H; 10A–B, D–E, H).
|
||
</paragraph>
|
||
<paragraph id="556036ADFFACFF8BFF36FA6ADAD7F9B8" blockId="19.[151,1437,348,1993]" pageId="19" pageNumber="20">
|
||
Notopodia starting from segment 3 and extending through 10 segments, until segment 12, in
|
||
<typeStatus id="8A64880FFFACFF8BFB1EFA6ADC7EFA3C" box="[1263,1364,1439,1464]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
, some specimens with 9 pairs of notopodia only; elongate, cylindrical notopodia, with equal sized lobes and elongate, distally blunt tip (
|
||
<figureCitation id="CDE42A28FFACFF8BFE91FA14D88DFA7E" box="[352,423,1505,1530]" captionStart="FIGURE 9" captionStartId="20.[151,250,1851,1873]" captionTargetBox="[193,1404,197,1818]" captionTargetId="figure@20.[187,1408,193,1826]" captionTargetPageId="20" captionText="FIGURE 9. Amaeana apheles, holotype AM W. 5239. A – C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D – F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial / genital papillae. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W. 29203: I – J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.7 mm; B – C = 1 mm; D – F = 0.5 mm; G – H = 0.3 mm; I – K = 20 µm." httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="19" pageNumber="20">Figs 9</figureCitation>
|
||
A–I; 10A–G; 11A–B). Notochaetae as nearly alimbate capillaries in both rows, wings not visible under higher magnifications of light microscopy (
|
||
<figureCitation id="CDE42A28FFACFF8BFCDAF9F7DA5AF99F" box="[811,880,1538,1563]" captionStart="FIGURE 9" captionStartId="20.[151,250,1851,1873]" captionTargetBox="[193,1404,197,1818]" captionTargetId="figure@20.[187,1408,193,1826]" captionTargetPageId="20" captionText="FIGURE 9. Amaeana apheles, holotype AM W. 5239. A – C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D – F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial / genital papillae. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W. 29203: I – J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.7 mm; B – C = 1 mm; D – F = 0.5 mm; G – H = 0.3 mm; I – K = 20 µm." httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="19" pageNumber="20">
|
||
Fig.
|
||
<date id="2161106DFFACFF8BFC93F9F6DA5AF99F" box="[866,880,1539,1563]" pageId="19" pageNumber="20">9</date>
|
||
</figureCitation>
|
||
I–J), as short and fine hairs under SEM, closely resembling pinnate chaetae, but with finer hairs in oblique tiers (
|
||
<figureCitation id="CDE42A28FFACFF8BFC9BF9D6DA9DF9B8" box="[874,951,1571,1596]" captionStart="FIGURE 11" captionStartId="22.[151,250,1897,1919]" captionTargetBox="[171,1413,196,1851]" captionTargetId="figure@22.[164,1417,192,1859]" captionTargetPageId="22" captionText="FIGURE 11. Amaeana apheles, photos A – E from specimen AM W. 19333, photos F – I from specimen AM W. 10864. A, notopodia, segments 9 – 10; B – E, notochaetae, segment 10; F – G, posterior neuropodia; H – I, close up of posterior neuropodia with protruding spines. Scale bars: A = 70 µm; B, I = 10 µm; C – D = 5 µm; E = 4 µm; F = 200 µm; G = 8 µm; H = 20 µm." httpUri="https://zenodo.org/record/289148/files/figure.png" pageId="19" pageNumber="20">Fig. 11</figureCitation>
|
||
A–E).
|
||
</paragraph>
|
||
<paragraph id="556036ADFFACFF8BFF36F9B1D9C3F91B" blockId="19.[151,1437,348,1993]" pageId="19" pageNumber="20">
|
||
Neuropodia present from segments 23–24, laterally to mid-ventral groove, on outer margins of longitudinal crests; neurochaetae as 1–2 thick, distally sharp spines, tips somewhat narrower and slightly hooked (
|
||
<figureCitation id="CDE42A28FFACFF8BFAC7F990DCAAF9FA" box="[1334,1408,1637,1662]" captionStart="FIGURE 9" captionStartId="20.[151,250,1851,1873]" captionTargetBox="[193,1404,197,1818]" captionTargetId="figure@20.[187,1408,193,1826]" captionTargetPageId="20" captionText="FIGURE 9. Amaeana apheles, holotype AM W. 5239. A – C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D – F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial / genital papillae. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W. 29203: I – J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.7 mm; B – C = 1 mm; D – F = 0.5 mm; G – H = 0.3 mm; I – K = 20 µm." httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="19" pageNumber="20">Figs 9</figureCitation>
|
||
K; 11F–I).
|
||
</paragraph>
|
||
<paragraph id="556036ADFFACFF8BFF36F952DBD2F965" blockId="19.[151,1437,348,1993]" pageId="19" pageNumber="20">
|
||
Nephridial and genital papillae at anterior bases of all notopodia, larger on segments 5–10 (
|
||
<figureCitation id="CDE42A28FFACFF8BFB02F952DC14F944" box="[1267,1342,1703,1728]" captionStart="FIGURE 9" captionStartId="20.[151,250,1851,1873]" captionTargetBox="[193,1404,197,1818]" captionTargetId="figure@20.[187,1408,193,1826]" captionTargetPageId="20" captionText="FIGURE 9. Amaeana apheles, holotype AM W. 5239. A – C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D – F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial / genital papillae. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W. 29203: I – J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.7 mm; B – C = 1 mm; D – F = 0.5 mm; G – H = 0.3 mm; I – K = 20 µm." httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="19" pageNumber="20">Figs 9</figureCitation>
|
||
B, D–E; 10A–B, D–E). Pygidium with rounded ventral papilla.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1DC56526FFACFF8FFF36F91CD9C6FE71" lastPageId="23" lastPageNumber="24" pageId="19" pageNumber="20" type="discussion">
|
||
<paragraph id="556036ADFFACFF8BFF36F91CD81CF8E1" blockId="19.[151,1437,348,1993]" pageId="19" pageNumber="20">
|
||
<emphasis id="67ABEABFFFACFF8BFF36F91CD811F886" bold="true" box="[199,315,1769,1794]" pageId="19" pageNumber="20">Remarks.</emphasis>
|
||
This taxon was originally described as a species of
|
||
<taxonomicName id="92DF4D2EFFACFF8BFC66F91FDAC9F886" box="[919,995,1770,1794]" class="Polychaeta" family="Terebellidae" genus="Lysilla" kingdom="Animalia" order="Terebellida" pageId="19" pageNumber="20" phylum="Annelida" rank="genus">
|
||
<emphasis id="67ABEABFFFACFF8BFC66F91FDAC9F886" box="[919,995,1770,1794]" italics="true" pageId="19" pageNumber="20">Lysilla</emphasis>
|
||
</taxonomicName>
|
||
, due to the
|
||
<typeStatus id="8A64880FFFACFF8BFB81F91CDDF8F886" box="[1136,1234,1769,1794]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
being posteriorly incomplete, with few segments after termination of notopodia, not presenting neuropodia (
|
||
<bibRefCitation id="314E4B5CFFACFF8BFB23F8FEDCBAF8A7" author="Hutchings" box="[1234,1424,1803,1828]" pageId="19" pageNumber="20" refString="Hutchings, P. A. (1974) Polychaeta of WalIis Lake, New South Wales. Proceedings of the Linnean Society of New South Wales, 98 (4), 175 - 195." type="journal article" year="1974">Hutchings 1974</bibRefCitation>
|
||
). Subsequently, complete specimens were found and the species was correctly transferred to
|
||
<taxonomicName id="92DF4D2EFFACFF8BFB63F8D9D818F8E1" authority="Hutchings & Glasby 1986" authorityName="Hutchings & Glasby" authorityYear="1986" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="19" pageNumber="20" phylum="Annelida" rank="genus">
|
||
<emphasis id="67ABEABFFFACFF8BFB63F8D9DDD6F8C7" box="[1170,1276,1836,1859]" italics="true" pageId="19" pageNumber="20">Amaeana</emphasis>
|
||
(
|
||
<bibRefCitation id="314E4B5CFFACFF8BFAFBF8D9D803F8E1" author="Hutchings" pageId="19" pageNumber="20" refString="Hutchings, P. A. & Glasby, C. J. (1986) The Polycirrinae (F. Terebellidae) of Australia. Records of the Australian Museum, 38, 319 - 350. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.38.1986.185" type="journal article" year="1986">Hutchings & Glasby 1986</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="556036ADFFACFF8BFF36F89BD8C0F84D" blockId="19.[151,1437,348,1993]" pageId="19" pageNumber="20">
|
||
Members of
|
||
<taxonomicName id="92DF4D2EFFACFF8BFE91F89BD8F3F802" box="[352,473,1902,1926]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="19" pageNumber="20" phylum="Annelida" rank="species" species="apheles">
|
||
<emphasis id="67ABEABFFFACFF8BFE91F89BD8F3F802" box="[352,473,1902,1926]" italics="true" pageId="19" pageNumber="20">A. apheles</emphasis>
|
||
</taxonomicName>
|
||
are characterised by having 1–2 stout, distally sharp spines per neuropodium. In contrast to the other species in this genus, members of
|
||
<taxonomicName id="92DF4D2EFFACFF8BFD0DF87ADA5BF823" box="[764,881,1935,1959]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="19" pageNumber="20" phylum="Annelida" rank="species" species="apheles">
|
||
<emphasis id="67ABEABFFFACFF8BFD0DF87ADA5BF823" box="[764,881,1935,1959]" italics="true" pageId="19" pageNumber="20">A. apheles</emphasis>
|
||
</taxonomicName>
|
||
present slight variation on the number of pairs of notopodia present, 9–10 pairs.
|
||
</paragraph>
|
||
<caption id="01A06625FFABFF8CFF66F8CEDBEBF86E" httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="20" pageNumber="21" targetBox="[193,1404,197,1818]" targetPageId="20">
|
||
<paragraph id="556036ADFFABFF8CFF66F8CEDBEBF86E" blockId="20.[151,1437,1851,2026]" pageId="20" pageNumber="21">
|
||
<emphasis id="67ABEABFFFABFF8CFF66F8CED83DF8D4" bold="true" box="[151,279,1851,1873]" pageId="20" pageNumber="21">FIGURE 9.</emphasis>
|
||
<taxonomicName id="92DF4D2EFFABFF8CFED2F8CED8F0F8D5" box="[291,474,1851,1873]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="20" pageNumber="21" phylum="Annelida" rank="species" species="apheles">
|
||
<emphasis id="67ABEABFFFABFF8CFED2F8CED8F0F8D5" box="[291,474,1851,1873]" italics="true" pageId="20" pageNumber="21">Amaeana apheles</emphasis>
|
||
</taxonomicName>
|
||
, holotype AM W.5239. A–C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D–F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial/genital papillae. G–H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W.29203: I–J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P* = distal part of prostomium. Scale bars: A = 0.7 mm; B–C = 1 mm; D–F= 0.5 mm; G–H = 0.3 mm; I–K = 20 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="01A06625FFAAFF8DFF66F8AFDAB2F86F" httpUri="https://zenodo.org/record/289147/files/figure.png" pageId="21" pageNumber="22" targetBox="[204,1384,181,1856]" targetPageId="21">
|
||
<paragraph id="556036ADFFAAFF8DFF66F8AFDAB2F86F" blockId="21.[151,1437,1882,2027]" pageId="21" pageNumber="22">
|
||
<emphasis id="67ABEABFFFAAFF8DFF66F8AFD80AF8EB" bold="true" box="[151,288,1882,1904]" pageId="21" pageNumber="22">FIGURE 10.</emphasis>
|
||
<taxonomicName id="92DF4D2EFFAAFF8DFED6F8AED8F0F8F4" box="[295,474,1882,1904]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="apheles">
|
||
<emphasis id="67ABEABFFFAAFF8DFED6F8AED8F0F8F4" box="[295,474,1882,1904]" italics="true" pageId="21" pageNumber="22">Amaeana apheles</emphasis>
|
||
</taxonomicName>
|
||
, photos D and G from specimen AM W.10864, all other photos from specimen AM W.19333. A–C, thorax, right lateral, ventral and dorsal views, respectively. D–E, close ups of the anterior end, left and right lateral views, respectively. F–G, close ups of the anterior end, dorsal view. H, detail of the oral area. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P* = distal part of prostomium. Scale bars: A–B = 0.7 mm; C = 0.6 mm; D = 0.15 mm; E–F= 0.2 mm; G–H = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="01A06625FFA9FF8EFF66F89CDC47F838" httpUri="https://zenodo.org/record/289148/files/figure.png" pageId="22" pageNumber="23" targetBox="[171,1413,196,1851]" targetPageId="22">
|
||
<paragraph id="556036ADFFA9FF8EFF66F89CDC47F838" blockId="22.[151,1436,1897,1980]" pageId="22" pageNumber="23">
|
||
<emphasis id="67ABEABFFFA9FF8EFF66F89CD808F8FA" bold="true" box="[151,290,1897,1919]" pageId="22" pageNumber="23">FIGURE 11.</emphasis>
|
||
<taxonomicName id="92DF4D2EFFA9FF8EFEDCF89FD8CEF8FB" box="[301,484,1897,1919]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="22" pageNumber="23" phylum="Annelida" rank="species" species="apheles">
|
||
<emphasis id="67ABEABFFFA9FF8EFEDCF89FD8CEF8FB" box="[301,484,1897,1919]" italics="true" pageId="22" pageNumber="23">Amaeana apheles</emphasis>
|
||
</taxonomicName>
|
||
, photos A–E from specimen AM W.19333, photos F–I from specimen AM W.10864. A, notopodia, segments 9–10; B–E, notochaetae, segment 10; F–G, posterior neuropodia; H–I, close up of posterior neuropodia with protruding spines. Scale bars: A = 70 µm; B, I = 10 µm; C–D = 5 µm; E = 4 µm; F = 200 µm; G = 8 µm; H = 20 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="556036ADFFA8FF8FFF36FF62DD9AFE98" blockId="23.[151,1437,151,501]" pageId="23" pageNumber="24">
|
||
Members of
|
||
<taxonomicName id="92DF4D2EFFA8FF8FFEA7FF6DD8EDFF2B" box="[342,455,152,175]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="yirrarn">
|
||
<emphasis id="67ABEABFFFA8FF8FFEA7FF6DD8EDFF2B" box="[342,455,152,175]" italics="true" pageId="23" pageNumber="24">A. yirrarn</emphasis>
|
||
</taxonomicName>
|
||
also have 1–2 spines per neuropodium, but these species are distinguished by the latter having 12 pairs of notopodia, instead of 9–10, as in
|
||
<taxonomicName id="92DF4D2EFFA8FF8FFD1BFF49DA4AFF50" box="[746,864,188,212]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="apheles">
|
||
<emphasis id="67ABEABFFFA8FF8FFD1BFF49DA4AFF50" box="[746,864,188,212]" italics="true" pageId="23" pageNumber="24">A. apheles</emphasis>
|
||
</taxonomicName>
|
||
. There is also a difference on the extension of the achaetous gap between termination of notopodia and beginning of neuropodia between these species, although with some overlapping, as this gap extends for 7–9 segments in
|
||
<taxonomicName id="92DF4D2EFFA8FF8FFCDAFEF1DAB1FE98" box="[811,923,260,284]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="yirrarn">
|
||
<emphasis id="67ABEABFFFA8FF8FFCDAFEF1DAB1FE98" box="[811,923,260,284]" italics="true" pageId="23" pageNumber="24">A. yirrarn</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<quantity id="92279B48FFA8FF8FFC29FEF1DD07FE99" box="[984,1069,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.54" metricValueMin="2.286" pageId="23" pageNumber="24" unit="in" value="9.5" valueMax="10.0" valueMin="9.0">9–10 in</quantity>
|
||
<emphasis id="67ABEABFFFA8FF8FFBC5FEF1DD9AFE98" box="[1076,1200,260,284]" italics="true" pageId="23" pageNumber="24">
|
||
<taxonomicName id="92DF4D2EFFA8FF8FFBC5FEF1DD86FE98" box="[1076,1196,260,284]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="apheles">A. apheles</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="556036ADFFA8FF8FFF36FED2D9C6FE71" blockId="23.[151,1437,151,501]" pageId="23" pageNumber="24">
|
||
Besides
|
||
<taxonomicName id="92DF4D2EFFA8FF8FFED4FEDDD8BCFEBB" box="[293,406,296,319]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="yirrarn">
|
||
<emphasis id="67ABEABFFFA8FF8FFED4FEDDD8BCFEBB" box="[293,406,296,319]" italics="true" pageId="23" pageNumber="24">A. yirrarn</emphasis>
|
||
</taxonomicName>
|
||
, three other species have a number of spines per neuropodium similar to that of
|
||
<taxonomicName id="92DF4D2EFFA8FF8FFAD1FEDDDCBEFEC4" box="[1312,1428,296,320]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="apheles">
|
||
<emphasis id="67ABEABFFFA8FF8FFAD1FEDDDCBEFEC4" box="[1312,1428,296,320]" italics="true" pageId="23" pageNumber="24">A. apheles</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="92DF4D2EFFA8FF8FFF66FEB9D84EFEE0" box="[151,356,332,356]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="crassispinulata" status="sp. nov.">
|
||
<emphasis id="67ABEABFFFA8FF8FFF66FEB9D84EFEE0" box="[151,356,332,356]" italics="true" pageId="23" pageNumber="24">A. crassispinulata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="67ABEABFFFA8FF8FFE9FFEB9D8E0FEE1" bold="true" box="[366,458,332,357]" pageId="23" pageNumber="24">
|
||
<taxonomicNameLabel id="7C9857C4FFA8FF8FFE9FFEB9D8E0FEE1" box="[366,458,332,357]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="92DF4D2EFFA8FF8FFE2AFEB9DB79FEE0" box="[475,595,332,356]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="hsiehae" status="sp. nov.">
|
||
<emphasis id="67ABEABFFFA8FF8FFE2AFEB9DB79FEE0" box="[475,595,332,356]" italics="true" pageId="23" pageNumber="24">A. hsiehae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="67ABEABFFFA8FF8FFDADFEB9DB93FEE1" bold="true" box="[604,697,332,357]" pageId="23" pageNumber="24">
|
||
<taxonomicNameLabel id="7C9857C4FFA8FF8FFDADFEB9DB93FEE1" box="[604,697,332,357]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="92DF4D2EFFA8FF8FFD0DFEB9DAB4FEE0" box="[764,926,332,356]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="brasiliensis" status="sp. nov.">
|
||
<emphasis id="67ABEABFFFA8FF8FFD0DFEB9DAB4FEE0" box="[764,926,332,356]" italics="true" pageId="23" pageNumber="24">A. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="67ABEABFFFA8FF8FFC58FEB9DD2FFEE1" bold="true" box="[937,1029,332,357]" pageId="23" pageNumber="24">
|
||
<taxonomicNameLabel id="7C9857C4FFA8FF8FFC58FEB9DD2FFEE1" box="[937,1029,332,357]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, all these species described herein. However, those spines are basally wider and distally truncate in
|
||
<taxonomicName id="92DF4D2EFFA8FF8FFC86FE85DD69FE0C" box="[887,1091,368,392]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="crassispinulata" status="sp. nov.">
|
||
<emphasis id="67ABEABFFFA8FF8FFC86FE85DD69FE0C" box="[887,1091,368,392]" italics="true" pageId="23" pageNumber="24">A. crassispinulata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="67ABEABFFFA8FF8FFBBDFE9ADD8DFE0C" bold="true" box="[1100,1191,367,392]" pageId="23" pageNumber="24">
|
||
<taxonomicNameLabel id="7C9857C4FFA8FF8FFBBDFE9ADD8DFE0C" box="[1100,1191,367,392]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(see below), tapered, but not as sharp in
|
||
<taxonomicName id="92DF4D2EFFA8FF8FFE9DFE61D8C8FE28" box="[364,482,404,428]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="hsiehae" status="sp. nov.">
|
||
<emphasis id="67ABEABFFFA8FF8FFE9DFE61D8C8FE28" box="[364,482,404,428]" italics="true" pageId="23" pageNumber="24">A. hsiehae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="67ABEABFFFA8FF8FFE19FE61DB68FE29" bold="true" box="[488,578,404,429]" pageId="23" pageNumber="24">
|
||
<taxonomicNameLabel id="7C9857C4FFA8FF8FFE19FE61DB68FE29" box="[488,578,404,429]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, described above, and distally blunt or with oblique tip in
|
||
<taxonomicName id="92DF4D2EFFA8FF8FFB22FE61DC59FE28" box="[1235,1395,404,428]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="brasiliensis" status="sp. nov.">
|
||
<emphasis id="67ABEABFFFA8FF8FFB22FE61DC59FE28" box="[1235,1395,404,428]" italics="true" pageId="23" pageNumber="24">A. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7C9857C4FFA8FF8FFA8BFE61D9EDFE4B" pageId="23" pageNumber="24" rank="species">
|
||
<emphasis id="67ABEABFFFA8FF8FFA8BFE61D9EBFE54" bold="true" pageId="23" pageNumber="24">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
; in addition, this latter. In addition, this latter species also has a different number of pairs of notopodia (see below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |