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<document id="B2F5A3AA975FBD7881DA9EB07A375A3C" ID-DOI="10.5252/z2011n1a5" ID-ISSN="1638-9387" ID-Zenodo-Dep="4546704" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613600601004" checkinUser="felipe" docAuthor="Montreuil, Olivier" docDate="2011" docId="03DC9E64DA5F3A6E3AF4FB2A8750E30E" docLanguage="fr" docName="Zoosystema.33.1.101-122.pdf" docOrigin="Zoosystema 33 (1)" docSource="http://dx.doi.org/10.5252/z2011n1a5" docStyle="DocumentStyle:EB3C2C73672D2E883259A050BD27978B.6:Zoosystema.2011-2012.journal_article" docStyleId="EB3C2C73672D2E883259A050BD27978B" docStyleName="Zoosystema.2011-2012.journal_article" docStyleVersion="6" docTitle="Epilissus fantamattii Montreuil 2011, n. sp." docType="treatment" docVersion="3" lastPageNumber="114" masterDocId="FFE5E61CDA533A633A1FFFA48336E764" masterDocTitle="Monophylie du « groupe Epilissus » et refonte des genres Epilissus Reiche, 1841, Arachnodes Westwood, 1847 et Apterepilissus n. gen. de Madagascar (Insecta, Coleoptera, Scarabaeidae, Epilissini)" masterLastPageNumber="122" masterPageNumber="101" pageNumber="113" updateTime="1698872838645" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FD3057957E96A13773C0F88B2EEB2CE8">Monophylie du « groupe Epilissus » et refonte des genres Epilissus Reiche, 1841, Arachnodes Westwood, 1847 et Apterepilissus n. gen. de Madagascar (Insecta, Coleoptera, Scarabaeidae, Epilissini)</mods:title>
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<mods:namePart id="49E8F8BF9BD6AF699140E21B507C7D8C">Montreuil, Olivier</mods:namePart>
<mods:affiliation id="DC233A687EB251FC0B4DEB54C5B32233">Muséum national dHistoire naturelle, UMR 7205, Département Systématique et Évolution, Entomologie, case postale 50, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) o. montr. @ mnhn. fr</mods:affiliation>
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<mods:title id="643BA803627EC69BBCC84DB2FD65B689">Zoosystema</mods:title>
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<mods:date id="319B1B94A41D805C03A6220EC67D6EEA">2011</mods:date>
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<mods:number id="CE616243E00C550B1F149BC0030DCC57">2011-03-31</mods:number>
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<mods:identifier id="83F44F72DD1E89C711CADF93F7C779B4" type="ISSN">1638-9387</mods:identifier>
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<treatment id="03DC9E64DA5F3A6E3AF4FB2A8750E30E" ID-DOI="http://doi.org/10.5281/zenodo.4546741" ID-Zenodo-Dep="4546741" LSID="urn:lsid:plazi:treatment:03DC9E64DA5F3A6E3AF4FB2A8750E30E" httpUri="http://treatment.plazi.org/id/03DC9E64DA5F3A6E3AF4FB2A8750E30E" lastPageId="13" lastPageNumber="114" pageId="12" pageNumber="113">
<subSubSection id="C36F7CF9DA5F3A6F3AF4FB2A813BE3CD" box="[235,525,1166,1193]" pageId="12" pageNumber="113" type="nomenclature">
<paragraph id="8BCA2F72DA5F3A6F3AF4FB2A813BE3CD" blockId="12.[235,525,1166,1193]" box="[235,525,1166,1193]" pageId="12" pageNumber="113">
<heading id="D082981EDA5F3A6F3AF4FB2A813BE3CD" box="[235,525,1166,1193]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="113" reason="2">
<taxonomicName id="4C7554F1DA5F3A6F3AF4FB2A82FDE3CC" ID-CoL="6G4VJ" authority="Montreuil, 2011" authorityName="Montreuil" authorityYear="2011" box="[235,459,1166,1193]" class="Insecta" family="Scarabaeidae" genus="Epilissus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="fantamattii" status="sp. nov.">
<emphasis id="B901F360DA5F3A6F3AF4FB2A82FDE3CC" bold="true" box="[235,459,1166,1193]" italics="true" pageId="12" pageNumber="113">Epilissus fantamattii</emphasis>
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<taxonomicNameLabel id="A2324E1BDA5F3A6F3BCDFB2A813BE3CD" box="[466,525,1166,1193]" pageId="12" pageNumber="113" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C36F7CF9DA5F3A6F3A6EFB638176E2F7" pageId="12" pageNumber="113" type="materials_examined">
<paragraph id="8BCA2F72DA5F3A6F3A6EFB638124E270" blockId="12.[112,648,1223,1300]" pageId="12" pageNumber="113">
<materialsCitation id="3B1D252FDA5F3A6F3A6EFB638124E270" collectingDate="2005-12-01" collectionCode="MNHN" collectorName="Andranobe &amp; Masoala NP &amp; Ilkka Hanski" pageId="12" pageNumber="113" specimenCode="EC1910" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CE91D0DA5F3A6F3A6EFB6383EDE3BA" box="[113,219,1223,1247]" pageId="12" pageNumber="113" type="holotype">HOLOTYPE</typeStatus>
. —
<taxonomicName id="4C7554F1DA5F3A6F3B15FB6382B6E3BB" authority="Montreuil, 2011" authorityName="Montreuil" authorityYear="2011" box="[266,384,1223,1247]" genus="Madagascar" pageId="12" pageNumber="102" rank="genus" status="gen. nov.">
<emphasis id="B901F360DA5F3A6F3B15FB6382B6E3BB" bold="true" box="[266,384,1223,1247]" pageId="12" pageNumber="113">Madagascar</emphasis>
</taxonomicName>
.
<collectorName id="26804AA4DA5F3A6F3B90FB6382C9E3BB" box="[399,511,1223,1247]" pageId="12" pageNumber="113">Andranobe</collectorName>
,
<collectorName id="26804AA4DA5F3A6F3815FB6381B5E3BB" box="[522,643,1223,1247]" pageId="12" pageNumber="113">Masoala NP</collectorName>
,
<date id="FFCB09B2DA5F3A6F3A6CFB4683D7E39E" box="[115,225,1250,1274]" pageId="12" pageNumber="113" value="2005-12-01">
<collectingDate id="EF8FF05ADA5F3A6F3A6CFB4683D7E39E" box="[115,225,1250,1274]" pageId="12" pageNumber="113" value="2005-12-01">1.XII.2005</collectingDate>
</date>
, littoral forest, 0 m,
<collectingMethod id="52345765DA5F3A6F3B82FB4681B2E39E" box="[413,644,1250,1274]" pageId="12" pageNumber="113">human dung baited trap</collectingMethod>
,
<collectorName id="26804AA4DA5F3A6F3A6FFB5883D8E270" box="[112,238,1276,1300]" pageId="12" pageNumber="113">Ilkka Hanski</collectorName>
leg.,1 ơ (
<collectionCode id="ED64B7B7DA5F3A6F3B49FB58829FE270" box="[342,425,1276,1300]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="113">MNHN</collectionCode>
<specimenCode id="DBD38709DA5F3A6F3BB0FB588130E270" box="[431,518,1276,1300]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="113">EC1910</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCA2F72DA5F3A6F3A6EFA888176E2F7" blockId="12.[111,648,1324,1427]" pageId="12" pageNumber="113">
<typeStatus id="54CE91D0DA5F3A6F3A6EFA8883E5E226" box="[113,211,1324,1348]" pageId="12" pageNumber="113" type="paratype">PARATYPES</typeStatus>
. — mêmes données que lholotype,13 ơơ et
<specimenCount id="9D73E4FBDA5F3A6F387AFA8981B1E220" box="[613,647,1325,1348]" pageId="12" pageNumber="113" type="female">♀♀</specimenCount>
(
<collectionCode id="ED64B7B7DA5F3A6F3A67FAE283F1E23A" box="[120,199,1350,1374]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="113">MNHN</collectionCode>
et
<collectionCode id="ED64B7B7DA5F3A6F3AC2FAE2821EE23A" box="[221,296,1350,1374]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="113" type="Museum">FMNH</collectionCode>
),XI-XII.2005.
<taxonomicName id="4C7554F1DA5F3A6F3BB5FAE28123E23A" authority="Montreuil, 2011" authorityName="Montreuil" authorityYear="2011" box="[426,533,1350,1374]" genus="Madagascar" pageId="12" pageNumber="102" rank="genus" status="gen. nov.">
<emphasis id="B901F360DA5F3A6F3BB5FAE28123E23A" bold="true" box="[426,533,1350,1374]" pageId="12" pageNumber="113">Madagascar</emphasis>
</taxonomicName>
. Andranobe, Masoala, wet forest,
<date id="FFCB09B2DA5F3A6F3B2FFAC5824FE21D" box="[304,377,1377,1401]" pageId="12" pageNumber="113" value="2004-02">II.2004</date>
, littoral forest, alt.
<quantity id="4C8D8297DA5F3A6F3834FAC581B6E21D" box="[555,640,1377,1401]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="12" pageNumber="113" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0-100 m</quantity>
, fish baited trap, Ilkka Hanski leg.,
<specimenCount id="9D73E4FBDA5F3A6F3BACFADF82E0E2F7" box="[435,470,1403,1427]" pageId="12" pageNumber="113" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED64B7B7DA5F3A6F3BF9FADF8103E2F7" box="[486,565,1403,1427]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="113" type="Museum">FMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C36F7CF9DA5F3A6E3A6EFA0F8750E30E" lastPageId="13" lastPageNumber="114" pageId="12" pageNumber="113" type="description">
<paragraph id="8BCA2F72DA5F3A6F3A6EFA0F8178E29C" blockId="12.[113,647,1451,1528]" pageId="12" pageNumber="113">ÉTYMOLOGIE. — Cette espèce est amicalement dédiée à Matti Hanski, fils dIlkka Hanski, agréable compagnon de terrain et grand amateur dorangeade gazeuse.</paragraph>
<paragraph id="8BCA2F72DA5F3A6F3A6EF98E8234E127" blockId="12.[113,646,1578,1700]" box="[113,258,1578,1605]" pageId="12" pageNumber="113">DESCRIPTION</paragraph>
<paragraph id="8BCA2F72DA5F3A6F3A6EF9EE8229E1C0" blockId="12.[113,646,1578,1700]" pageId="12" pageNumber="113">
Longueur
<quantity id="4C8D8297DA5F3A6F3AF7F9EE824DE101" box="[232,379,1610,1637]" metricMagnitude="-2" metricUnit="m" metricValue="1.275" metricValueMax="1.4" metricValueMin="1.15" pageId="12" pageNumber="113" unit="mm" value="12.75" valueMax="14.0" valueMin="11.5">11,5-14 mm</quantity>
. Noir mat, à très légers reflets olive-foncé, face sternale et pattes à légers reflets verdâtres.
</paragraph>
<paragraph id="8BCA2F72DA5F3A6F38BAFB2A81F9E3CC" blockId="12.[674,1210,1166,1383]" box="[677,719,1166,1192]" pageId="12" pageNumber="113">
<emphasis id="B901F360DA5F3A6F38BAFB2A81F9E3CC" box="[677,719,1166,1192]" italics="true" pageId="12" pageNumber="113">Tête</emphasis>
</paragraph>
<paragraph id="8BCA2F72DA5F3A6F38BAFB0A8723E203" blockId="12.[674,1210,1166,1383]" pageId="12" pageNumber="113">Inerme, clypéus bidenté. Côtés de la tête anguleux au niveau de la suture clypéo-génale. Front, joues et arrière du clypéus très finement microréticulés, à ponctuation imperceptible. Partie antérieure du clypéus légèrement déprimée, lisse, laissant apparaître une fi ne ponctuation superficielle.</paragraph>
<paragraph id="8BCA2F72DA5F3A6F38BAFA28803AE2C1" blockId="12.[675,1210,1420,1700]" box="[677,780,1420,1445]" pageId="12" pageNumber="113">
<emphasis id="B901F360DA5F3A6F38BAFA28803AE2C1" box="[677,780,1420,1445]" italics="true" pageId="12" pageNumber="113">Pronotum</emphasis>
</paragraph>
<paragraph id="8BCA2F72DA5F3A6E38BAFA0F8132E4AF" blockId="12.[675,1210,1420,1700]" lastBlockId="13.[112,646,754,971]" lastPageId="13" lastPageNumber="114" pageId="12" pageNumber="113">Marge basale à rebord extrêmement fin. Marges latérales et antérieure bien rebordées. Tégument finement microréticulé, avec une ponctuation très fine, dense et régulière sur toute la surface, un peu plus superficielle près de la marge antérieure. Côtés plongeant dans leur partie postérieure, les angles postérieurs ainsi pratiquement invisibles et les côtés apparaissant fortement divergents en vue dorsale; convergents et droits avant les angles antérieurs, qui sont peu saillants, droits. La plus grande largeur du pronotum semble située à peu près au milieu en vue dorsale alors quelle est au niveau des angles postérieurs en vue latérale. Un profond sillon longe intérieurement les marges latérales sur leur moitié postérieure.</paragraph>
<caption id="DF0A7FFADA5E3A6E3A6EFDDF8283E5C7" ID-DOI="http://doi.org/10.5281/zenodo.4546720" ID-Zenodo-Dep="4546720" httpUri="https://zenodo.org/record/4546720/files/figure.png" pageId="13" pageNumber="114" startId="13.[113,124,635,652]" targetBox="[210,551,215,579]" targetPageId="13">
<paragraph id="8BCA2F72DA5E3A6E3A6EFDDF8283E5C7" blockId="13.[112,646,635,675]" pageId="13" pageNumber="114">
FIG. 5. — Paramères d
<taxonomicName id="4C7554F1DA5E3A6E3B34FDDF82F8E5E8" authority="Montreuil, 2011" authorityName="Montreuil" authorityYear="2011" box="[299,462,635,652]" class="Insecta" family="Scarabaeidae" genus="Epilissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="fantamattii" status="sp. nov.">
<emphasis id="B901F360DA5E3A6E3B34FDDF82F8E5E8" box="[299,462,635,652]" italics="true" pageId="13" pageNumber="114">Epilissus fantamattii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2324E1BDA5E3A6E3BCDFDDF8136E5E8" box="[466,512,635,652]" pageId="13" pageNumber="114" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B901F360DA5E3A6E3815FDD88121E5E9" bold="true" box="[522,535,636,653]" pageId="13" pageNumber="114">A</emphasis>
, vue dorsale;
<emphasis id="B901F360DA5E3A6E3A6FFD36834BE5C7" bold="true" box="[112,125,658,675]" pageId="13" pageNumber="114">B</emphasis>
, vue latérale gauche. Échelle: 1 mm.
</paragraph>
</caption>
<paragraph id="8BCA2F72DA5E3A6E3A6EFC548385E36F" blockId="13.[113,648,1008,1130]" box="[113,179,1008,1035]" pageId="13" pageNumber="114">
<emphasis id="B901F360DA5E3A6E3A6EFC548385E36F" box="[113,179,1008,1035]" italics="true" pageId="13" pageNumber="114">Élytres</emphasis>
</paragraph>
<paragraph id="8BCA2F72DA5E3A6E3A6EFBB483E4E30E" blockId="13.[113,648,1008,1130]" pageId="13" pageNumber="114">Stries élytrales peu marquées, à ponctuation peu nette. Interstries imponctués, fortement microréticulés.</paragraph>
<paragraph id="8BCA2F72DA5E3A6E38BAFF7C81C9E796" blockId="13.[675,1210,216,370]" box="[677,767,216,242]" pageId="13" pageNumber="114">
<emphasis id="B901F360DA5E3A6E38BAFF7C81C9E796" box="[677,767,216,242]" italics="true" pageId="13" pageNumber="114">
8
<superScript id="7C00823ADA5E3A6E38ACFF7E818CE78E" attach="left" box="[691,698,218,234]" fontSize="7" pageId="13" pageNumber="114">e</superScript>
tergite
</emphasis>
</paragraph>
<paragraph id="8BCA2F72DA5E3A6E38BAFF5C8016E616" blockId="13.[675,1210,216,370]" pageId="13" pageNumber="114">Déprimé dans sa moitié antérieure, présentant à cet endroit une petite ponctuation nette et régulière. Partie postérieure bombée et lisse, à ponctuation indistincte.</paragraph>
<paragraph id="8BCA2F72DA5E3A6E38BAFE3381D6E6D4" blockId="13.[674,1212,407,750]" box="[677,736,407,432]" pageId="13" pageNumber="114">
<emphasis id="B901F360DA5E3A6E38BAFE3381D6E6D4" box="[677,736,407,432]" italics="true" pageId="13" pageNumber="114">Pattes</emphasis>
</paragraph>
<paragraph id="8BCA2F72DA5E3A6E38BAFE12873BE58A" blockId="13.[674,1212,407,750]" pageId="13" pageNumber="114">Protibia tronqué droit à lapex, faiblement élargi à partir du milieu, apparaissant ainsi légèrement courbe en dedans. Arête externe tridentée, la dent médiane plus proche de lapicale, avec une rangée de petites dents tout le long côté externe y compris entre les dents principales. Angle apical interne en fort éperon recourbé vers le bas chez le mâle, en éperon plus petit chez la femelle. Méso- et métatibias recourbés en dedans, progressivement élargis de la base à lapex côté arête inférieure.</paragraph>
<paragraph id="8BCA2F72DA5E3A6E38BAFCB681DAE449" blockId="13.[677,826,786,845]" box="[677,748,786,813]" pageId="13" pageNumber="114">
<emphasis id="B901F360DA5E3A6E38BAFCB681DAE449" box="[677,748,786,813]" italics="true" pageId="13" pageNumber="114">Édéage</emphasis>
</paragraph>
<paragraph id="8BCA2F72DA5E3A6E38BAFC96800CE429" blockId="13.[677,826,786,845]" box="[677,826,818,845]" pageId="13" pageNumber="114">
Voir
<figureCitation id="134E33F7DA5E3A6E38C5FC96800CE429" box="[730,826,818,845]" captionStart="FIG" captionStartId="13.[113,124,635,652]" captionTargetBox="[210,551,215,579]" captionTargetId="figure-432@13.[210,550,220,579]" captionTargetPageId="13" captionText="FIG. 5. — Paramères dEpilissus fantamattii n. sp.: A, vue dorsale; B, vue latérale gauche. Échelle: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4546720" httpUri="https://zenodo.org/record/4546720/files/figure.png" pageId="13" pageNumber="114">Figure 5.</figureCitation>
</paragraph>
<paragraph id="8BCA2F72DA5E3A6E38BAFCD58011E4EE" blockId="13.[674,1210,881,1130]" box="[677,807,881,908]" pageId="13" pageNumber="114">REMARQUES</paragraph>
<paragraph id="8BCA2F72DA5E3A6E38BAFC358750E30E" blockId="13.[674,1210,881,1130]" pageId="13" pageNumber="114">
Par sa couleur entièrement noire, cette espèce ressemble à
<taxonomicName id="4C7554F1DA5E3A6E38ABFC148075E4AE" authorityName="Harold" authorityYear="1879" box="[692,835,944,970]" class="Insecta" family="Scarabaeidae" genus="Epilissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B901F360DA5E3A6E38ABFC148075E4AE" box="[692,835,944,970]" italics="true" pageId="13" pageNumber="114">Epilissus morio</emphasis>
</taxonomicName>
et
<taxonomicName id="4C7554F1DA5E3A6E3941FC1480D7E4AE" authorityName="Montreuil" authorityYear="2011" box="[862,993,944,970]" class="Insecta" family="Scarabaeidae" genus="Epilissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="hova" status="stat. nov.">
<emphasis id="B901F360DA5E3A6E3941FC1480D7E4AE" box="[862,993,944,970]" italics="true" pageId="13" pageNumber="114">Epilissus hova</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2324E1BDA5E3A6E39FAFC14871DE4AF" box="[997,1067,944,971]" pageId="13" pageNumber="114">n. stat.</taxonomicNameLabel>
, mais la forme des paramères la rapproche davantage d
<taxonomicName id="4C7554F1DA5E3A6E3E7DFC7481CCE36E" authorityName="Montreuil" authorityYear="2011" baseAuthorityName="Montreuil" baseAuthorityYear="2006" class="Insecta" family="Scarabaeidae" genus="Epilissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="planatus">
<emphasis id="B901F360DA5E3A6E3E7DFC7481CCE36E" italics="true" pageId="13" pageNumber="114">Epilissus planatus</emphasis>
</taxonomicName>
. Le lustre très luisant de sa face supérieure, dû à la très forte microréticulation de son tégument, est unique au sein de ce genre et permet de séparer facilement cette espèce de toutes les autres.
</paragraph>
</subSubSection>
</treatment>
</document>