153 lines
18 KiB
XML
153 lines
18 KiB
XML
<document id="1EC91986C3A5109399772C1B4830F4B9" ID-DOI="10.11646/zootaxa.5497.1.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="13618068" ID-ZooBank="7B226163-8E94-413A-9549-E0B74502878C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725025544867" checkinUser="plazi" docAuthor="Azar, Dany, Maksoud, Sibelle, Abi-Saad, Pierre & Nel, Andre" docDate="2024" docId="03CD87ECD23F1905FF1EF18DFA863B82" docLanguage="en" docName="zootaxa.5497.1.9.pdf" docOrigin="Zootaxa 5497 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5497.1.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lebanoaeshna mikhaeli Azar, Maksoud, Abi-Saad & Nel, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="146" masterDocId="FFF4FF94D23B1900FF89F579FF973907" masterDocTitle="A new, to date endemic, family of dragonfly in the mid-Cretaceous fossil fish Konservat-Lagerstätte of Haqel, Lebanon (Odonata: Anisoptera)" masterLastPageNumber="150" masterPageNumber="142" pageNumber="146" updateTime="1725026673843" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="965C7A81427E812DCBB8D14F186AECA1">
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<mods:title id="F6E06B94A03AC0341D532EF99942963A">A new, to date endemic, family of dragonfly in the mid-Cretaceous fossil fish Konservat-Lagerstätte of Haqel, Lebanon (Odonata: Anisoptera)</mods:title>
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<mods:name id="8149170398C25766048DC9A0D7920846" type="personal">
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<mods:roleTerm id="1419DA061829779BAB2E0039E7B62571">Author</mods:roleTerm>
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<mods:namePart id="C7997389168953E2022E51ECA1F4F574">Azar, Dany</mods:namePart>
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<mods:nameIdentifier id="F62DDFDF63F6AE53B5074C1E7A4CFCCA" type="ORCID">0000-0002-4485-197X</mods:nameIdentifier>
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<mods:affiliation id="5685AC0395CF2ED945D7660C1914C3C8">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, Jiangsu, 210008, China & Department of Natural Sciences, Faculty of Sciences II, Lebanese University, P. O. Box: 26110217, Fanar, Matn, Lebanon</mods:affiliation>
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<mods:nameIdentifier id="04F9D663022FA5314EB2CA450251F255" type="email">danyazar@ul.edu.lb</mods:nameIdentifier>
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</mods:name>
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<mods:name id="62240833E2378A7E4101071487D2B4A0" type="personal">
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<mods:role id="3DFEE93D629B97512F1E96768FE6778B">
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<mods:roleTerm id="34F65ED09AFF589715B05676B802E4F9">Author</mods:roleTerm>
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<mods:namePart id="88D54D890548A1856AED7ABDC31EFE80">Maksoud, Sibelle</mods:namePart>
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<mods:nameIdentifier id="504422468B67DBD95672D300DBAF5DFD" type="ORCID">0000-0003-4004-6735</mods:nameIdentifier>
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<mods:affiliation id="FB70406E55DF1359B0FF5AB21F2A10F0">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, Jiangsu, 210008, China & Department of Natural Sciences, Faculty of Sciences II, Lebanese University, P. O. Box: 26110217, Fanar, Matn, Lebanon</mods:affiliation>
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<mods:nameIdentifier id="4CB46D6914CC3215CCD4EE4E3BF1451A" type="email">sibelle.maksoud@ul.edu.lb</mods:nameIdentifier>
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</mods:name>
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<mods:name id="4D6C25DE33C8D8CC2D6C1C27448D28A4" type="personal">
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<mods:namePart id="96306D68448B5B8BFFEB98740086AE1C">Abi-Saad, Pierre</mods:namePart>
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<mods:affiliation id="E9114D665410ED445CC7B3D02991AD8B">Memory of Time, Byblos old sook, Byblos, Lebanon</mods:affiliation>
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<mods:name id="CB2DFB808AB7D684FD24FC6DEABD036C" type="personal">
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<mods:namePart id="B7E4D786D623F60A1B451DDC8EEA69E4">Nel, Andre</mods:namePart>
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<mods:affiliation id="4815804486C1CEAC2434BD2FAB37C790">Institut de Systématique, Evolution, Biodiversité, ISYEB-UMR 7205 - CNRS, MNHN, UPMC, EPHE, Muséum National d’Histoire Naturelle, Sorbonne Universités, 57 Rue Cuvier, CP 50, Entomologie, F- 75005, Paris, France</mods:affiliation>
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<mods:title id="7E08D028B2555378E73E498CA50861A2">Zootaxa</mods:title>
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<mods:part id="928A40BCB6F2A3D5BE461753861DF448">
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<mods:date id="ECC377CFFCB1F70149754FDDE1F28B19">2024</mods:date>
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<mods:number id="D98C98DAC406C1C76C96C8C0BA6013C9">2024-08-22</mods:number>
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<mods:number id="F8288EFECD17D3983849A5A7B4A3BC02">5497</mods:number>
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<mods:number id="1800078ACD3A7D7EA783E2662FCD511F">1</mods:number>
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<mods:url id="E03926E490FF5B3D8A842CDA86F26585">http://dx.doi.org/10.11646/zootaxa.5497.1.9</mods:url>
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<mods:classification id="20D65887E0B8DB5BC17AFDF4D3017D1D">journal article</mods:classification>
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<mods:identifier id="9ED23190111223891732217CD2889CB9" type="DOI">10.11646/zootaxa.5497.1.9</mods:identifier>
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<mods:identifier id="51706A7FA92B353A9CF92B10B137218E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="2CB310786D11A0DC7C9A64C2B2BDBB80" type="Zenodo-Dep">13618068</mods:identifier>
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<treatment id="03CD87ECD23F1905FF1EF18DFA863B82" LSID="urn:lsid:plazi:treatment:03CD87ECD23F1905FF1EF18DFA863B82" httpUri="http://treatment.plazi.org/id/03CD87ECD23F1905FF1EF18DFA863B82" lastPageId="5" lastPageNumber="146" pageId="4" pageNumber="146">
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<subSubSection id="C37E6571D23F1904FF1EF18DFBAE3C33" pageId="4" pageNumber="146" type="nomenclature">
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<paragraph id="8BDB36FAD23F1904FF1EF18DFD883C08" blockId="4.[151,1082,1268,1403]" box="[151,543,1268,1295]" pageId="4" pageNumber="146">
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<heading id="D0938196D23F1904FF1EF18DFD883C08" bold="true" box="[151,543,1268,1295]" fontSize="11" level="1" pageId="4" pageNumber="146" reason="1">
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<emphasis id="B910EAE8D23F1904FF1EF18DFD883C08" bold="true" box="[151,543,1268,1295]" pageId="4" pageNumber="146">
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<taxonomicName id="4C644D79D23F1904FF1EF18DFE2D3C08" authorityName="Azar & Maksoud & Abi-Saad & Nel" authorityYear="2024" box="[151,442,1268,1295]" class="Insecta" family="Liupanshaniidae" genus="Lebanoaeshna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="146" phylum="Arthropoda" rank="species" species="mikhaeli" status="sp. nov.">
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<emphasis id="B910EAE8D23F1904FF1EF18DFE2D3C08" bold="true" box="[151,442,1268,1295]" italics="true" pageId="4" pageNumber="146">Lebanoaeshna mikhaeli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2235793D23F1904FE48F18CFD883C08" box="[449,543,1269,1295]" pageId="4" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BDB36FAD23F1904FF1EF060FBAE3C33" blockId="4.[151,1082,1268,1403]" box="[151,1081,1305,1332]" pageId="4" pageNumber="146">
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Genus:
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<uri id="FFF53AF8D23F1904FF67F060FBAE3C33" box="[238,1081,1305,1332]" pageId="4" pageNumber="146">
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urn:lsid:zoobank.org:act:
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<uuid id="FFC20C2FD23F1904FE77F060FBAE3C33" box="[510,1081,1305,1332]" pageId="4" pageNumber="146">2AAF9E86-B9CA-4B6D-A25E-EC9D430B529A</uuid>
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</uri>
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</paragraph>
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<subSubSection id="C37E6571D23F1904FF1EF044FF4E3C7C" pageId="4" pageNumber="146" type="description">
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<paragraph id="8BDB36FAD23F1904FF1EF044FBAD3C5F" blockId="4.[151,1082,1268,1403]" box="[151,1082,1341,1368]" pageId="4" pageNumber="146">
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Species:
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<uri id="FFF53AF8D23F1904FF75F044FBAD3C5F" box="[252,1082,1341,1368]" pageId="4" pageNumber="146">
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urn:lsid:zoobank.org:act:
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<uuid id="FFC20C2FD23F1904FD85F044FBAD3C5F" box="[524,1082,1341,1368]" pageId="4" pageNumber="146">5AD24D5E-B77D-4F29-9CFF-D690B84F7DC4</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BDB36FAD23F1904FF1EF018FF4E3C7C" blockId="4.[151,1082,1268,1403]" box="[151,217,1377,1403]" pageId="4" pageNumber="146">
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<figureCitation id="135F2A7FD23F1904FF1EF018FF4E3C7C" box="[151,217,1377,1403]" captionStart="FIGURE 3" captionStartId="4.[151,250,1091,1115]" captionTargetBox="[151,1435,181,1056]" captionTargetId="figure-17@4.[151,1435,181,1067]" captionTargetPageId="4" captionText="FIGURE 3. Lebanoaeshna mikhaeli gen et sp. nov., Holotype I-26247/1A and B, part and counterpart. A, Counterpart, specimen I-26247/1B. B, Part, specimen I-26247/1A. C, Drawing of the specimen I-26247/1A completed by parts preserved on specimen I-26247/1B." figureDoi="http://doi.org/10.5281/zenodo.13618074" httpUri="https://zenodo.org/record/13618074/files/figure.png" pageId="4" pageNumber="146">Fig. 3</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37E6571D23F1904FF1EF0D0FD7C3CE0" pageId="4" pageNumber="146" type="materials_examined">
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<paragraph id="8BDB36FAD23F1904FF1EF0D0FD7C3CE0" blockId="4.[151,1437,1449,2016]" pageId="4" pageNumber="146">
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<emphasis id="B910EAE8D23F1904FF1EF0D0FE913CC4" bold="true" box="[151,262,1449,1475]" pageId="4" pageNumber="146">Material.</emphasis>
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<typeStatus id="54DF8858D23F1904FE86F0D0FEE03CC4" box="[271,375,1449,1475]" pageId="4" pageNumber="146" type="holotype">Holotype</typeStatus>
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I-26247/1A and B (part and counterpart of a rather poorly preserved forewing), stored in the Natural History Museum of the Lebanese University.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37E6571D23F1904FF4EF088FE963F70" pageId="4" pageNumber="146" type="etymology">
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<paragraph id="8BDB36FAD23F1904FF4EF088FEBA3F37" blockId="4.[151,1437,1449,2016]" pageId="4" pageNumber="146">
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<emphasis id="B910EAE8D23F1904FF4EF088FED93F0C" bold="true" box="[199,334,1521,1547]" pageId="4" pageNumber="146">Etymology.</emphasis>
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After Mr Mikhael Abi Saad, late father of one of us PAS, who worked several years in Haqel fossil fish’ outcrop.
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</paragraph>
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<paragraph id="8BDB36FAD23F1904FF4EF340FE963F70" blockId="4.[151,1437,1449,2016]" pageId="4" pageNumber="146">
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<emphasis id="B910EAE8D23F1904FF4EF340FE043F54" bold="true" box="[199,403,1593,1619]" pageId="4" pageNumber="146">Age and outcrop.</emphasis>
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Early late Cenomanian (
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<figureCitation id="135F2A7FD23F1904FD2EF340FD7D3F54" box="[679,746,1593,1619]" captionStart="FIGURE 2" captionStartId="3.[151,250,1839,1863]" captionTargetBox="[158,1430,181,1816]" captionTargetId="figure-19@3.[158,1430,181,1816]" captionTargetPageId="3" captionText="FIGURE 2. Mid-Cretaceous fossil fish Konservat-Lagerstätte of Haqel, Lebanon. A, Geological map of outcrop (modified from Dubertret, 1945; Dubertret & Wetzel, 1945). J6 = uppermost Jurassic; C2a = lower Barremian ‘Grès du Liban’ sandstone; C2b = Barremian clay and oolitic deposition of upper part of ‘Grès du Liban’ and oolitic deposition of lower part of Jezzinian; C3 = micritic part of Jezzinian (uppermost Barremian-lowermost Aptian); C4 = Albian; C5 = Cenomanian; Q = Quaternary scree; βJ6 = Kimmeridgian volcanic deposition; βC3 = Jezzinian volcanic deposition. Thick lines represent faults. Scale bar = 1 km. B, General view of outcrop. C & D, Details of stratigraphic layers of outcrop." figureDoi="http://doi.org/10.5281/zenodo.13618072" httpUri="https://zenodo.org/record/13618072/files/figure.png" pageId="4" pageNumber="146">Fig. 2</figureCitation>
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), Haqel, Caza Byblos, Governorate Jbeil-Kesserouan, Central
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<collectingCountry id="F373766AD23F1904FF1EF324FF6A3F70" box="[151,253,1629,1655]" name="Lebanon" pageId="4" pageNumber="146">Lebanon</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37E6571D23F1904FF4EF3F8FCA93F9C" box="[199,830,1665,1692]" pageId="4" pageNumber="146" type="diagnosis">
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<paragraph id="8BDB36FAD23F1904FF4EF3F8FCA93F9C" blockId="4.[151,1437,1449,2016]" box="[199,830,1665,1692]" pageId="4" pageNumber="146">
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<emphasis id="B910EAE8D23F1904FF4EF3F8FED53F9C" bold="true" box="[199,322,1665,1691]" pageId="4" pageNumber="146">Diagnosis.</emphasis>
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As for the genus and the family (
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<emphasis id="B910EAE8D23F1904FD31F3F8FCB93F9C" box="[696,814,1665,1691]" italics="true" pageId="4" pageNumber="146">vide supra</emphasis>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C37E6571D23F1905FF4EF3DCFA863B82" lastPageId="5" lastPageNumber="147" pageId="4" pageNumber="146" type="description">
|
||
<paragraph id="8BDB36FAD23F1905FF4EF3DCFA863B82" blockId="4.[151,1437,1449,2016]" lastBlockId="5.[151,1437,151,645]" lastPageId="5" lastPageNumber="147" pageId="4" pageNumber="146">
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<emphasis id="B910EAE8D23F1904FF4EF3DCFECE3FB8" bold="true" box="[199,345,1701,1727]" pageId="4" pageNumber="146">Description.</emphasis>
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Forewing hyaline, incomplete, with distal third missing and partly wrinkled, wing
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<emphasis id="B910EAE8D23F1904FA84F3DFFAB83FB8" box="[1293,1327,1702,1727]" italics="true" pageId="4" pageNumber="146">ca.</emphasis>
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50.0 mm long,
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<quantity id="4C9C9B1FD23F1904FF53F3B0FEA33FE3" box="[218,308,1737,1764]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="4" pageNumber="146" unit="mm" value="8.8">8.8 mm</quantity>
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wide, distance between base and arculus 5.0 mm, between arculus and nodus
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<emphasis id="B910EAE8D23F1904FB44F3B3FB793FE4" box="[1229,1262,1738,1763]" italics="true" pageId="4" pageNumber="146">ca.</emphasis>
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||
<quantity id="4C9C9B1FD23F1904FB70F3B0FAF43FE3" box="[1273,1379,1737,1764]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="4" pageNumber="146" unit="mm" value="18.7">18.7 mm</quantity>
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; two rows of cells between posterior wing margin and AA, defining an intercalary longitudinal vein parallel to AP; AA nearly parallel to MP+CuA; median area free of crossveins, six crossveins (including CuP) in submedian space; subdiscoidal space in continuity with submedian space, not transverse and with a distal angle acute; discoidal triangle very long and narrow, divided into seven smaller cells, with vein MAb having a very strong angle and a strong curvature; arculus with RP separated from MA; a strong trigonal planate in postdiscoidal area, emerging at angle of MAb, two rows of cells between MAb and MP basad angle of MAb; two rows of cells between MAa and trigonal planate and one row between trigonal planate and MP; Mspl rudimentary; distal part of postdiscodal area widened with at least five of cells between MAba and MP; CuA making a very strong double curve, very close to posterior margin of wing in its basal half, without well-defined posterior branches distally; cubito-anal area with at least three-four rows of cells in its broadest part; area between CuA and MP with two rows of large cells basally; area between MP and posterior margin very broad with numerous rows of cells and intercalary veins; two strong primary antenodal crossveins, Ax1 1.0 mm basal of arculus and Ax2
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<quantity id="4C9C9B1FD23E1905FC2FF45EFC6A3846" box="[934,1021,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="5" pageNumber="147" unit="mm" value="3.7">3.7 mm</quantity>
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distal of arculus, opposite middle of discoidal hypertriangle;
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<emphasis id="B910EAE8D23E1905FE2CF432FE503863" box="[421,455,331,356]" italics="true" pageId="5" pageNumber="147">ca.</emphasis>
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28 secondary antenodal crossveins of first row between C and ScP distad Ax2 and three-four between Ax1 and Ax2; secondary antenodal crossveins of second row between ScP and RA present but poorly preserved; nodus in a very distal position; bases of RP3/4 and IR2 very far distad arculus, closer to nodus than to arculus; a supplementary longitudinal vein aligned with ScP distad nodus,
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<emphasis id="B910EAE8D23E1905FC63F4CEFB9B38D7" box="[1002,1036,439,464]" italics="true" pageId="5" pageNumber="147">ca.</emphasis>
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||
<quantity id="4C9C9B1FD23E1905FB9CF4CFFBFB38D6" box="[1045,1132,438,465]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="147" unit="mm" value="2.5">2.5 mm</quantity>
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long; base of RP2 aligned with subnodus; area between RP1 and RP2 very narrow up to pterostigma; postnodal crossveins numerous, more than 22, not well-aligned with the postsubnodal crossveins; part of areas between C and RA and area between RA and RP1 distad pterostigma very long (possibly due to taphonomy deformation); pterostigma in a basal position, elongate, 4.0 mm long,
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||
<quantity id="4C9C9B1FD23E1905FE29F73EFE603B66" box="[416,503,583,609]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="147" unit="mm" value="0.7">0.7 mm</quantity>
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wide, covering eight cells; Rspl rudimentary; IR2 probably simple, closely parallel to RP2 at least in their basal halves; cells and intercalary crossveins in area below IR2 very numerous.
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</paragraph>
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</subSubSection>
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</treatment>
|
||
</document> |