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<document id="938A70185ECAB2CD9BB85DB25C8D715C" ID-CLB-Dataset="29643" ID-DOI="http://dx.doi.org/10.3897/dez.65.27033" ID-GBIF-Dataset="8916b689-6eca-4121-9066-01e3ccfdbb56" ID-Pensoft-Pub="1860-1324-2-117" ID-ZooBank="B1A8523CA4634FC4A0C3072C2E78BA02" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1860-1324-2-117" ModsDocOrigin="Deutsche Entomologische Zeitschrift 65 (2)" ModsDocTitle="Revision of the Quedius fauna of Middle Asia (Coleoptera, Staphylinidae, Staphylininae)" checkinTime="1533122252714" checkinUser="pensoft" docAuthor="Salnitska, Maria &amp; Solodovnikov, Alexey" docDate="2018" docId="AABA22035174DA6C68181C79CD408A0C" docLanguage="en" docName="DeutEntomolZeit 65(2): 117-159" docOrigin="Deutsche Entomologische Zeitschrift 65 (2)" docSource="http://dx.doi.org/10.3897/dez.65.27033" docTitle="Quedius (Microsaurus) capitalis Eppelsheim 1892" docType="treatment" docVersion="6" lastPageNumber="124" masterDocId="FFF6FFC22708A131FF960567FFF2FFB9" masterDocTitle="Revision of the Quedius fauna of Middle Asia (Coleoptera, Staphylinidae, Staphylininae)" masterLastPageNumber="159" masterPageNumber="117" pageNumber="123" updateTime="1701380021094" updateUser="ExternalLinkService">
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<mods:title id="51E633CD898EFC8EACC8736E8B18A0F4">Revision of the Quedius fauna of Middle Asia (Coleoptera, Staphylinidae, Staphylininae)</mods:title>
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<treatment id="AABA22035174DA6C68181C79CD408A0C" ID-GBIF-Taxon="146754235" LSID="urn:lsid:plazi:treatment:AABA22035174DA6C68181C79CD408A0C" httpUri="http://treatment.plazi.org/id/AABA22035174DA6C68181C79CD408A0C" lastPageId="7" lastPageNumber="124" pageId="6" pageNumber="123">
<subSubSection id="2909CD7AD879DBD551B8DF68E0D9C171" pageId="6" pageNumber="123" type="nomenclature">
<paragraph id="A8FC7105DD4FCA30BF26841BD425B375" pageId="6" pageNumber="123">
<taxonomicName id="8BD9C1875B972DF0744C7502A999BCEB" ID-CoL="4R2LF" authority="Eppelsheim, 1892" authorityName="Eppelsheim" authorityYear="1892" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Microsaurus) capitalis" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="capitalis" subGenus="Microsaurus">Quedius (Microsaurus) capitalis Eppelsheim, 1892</taxonomicName>
Fig. 7
</paragraph>
</subSubSection>
<subSubSection id="30EC53C1FAAA1C7C3515FE161BE92903" pageId="6" pageNumber="123" type="reference_group">
<paragraph id="54624A77690986F3A60762E13F2AC908" pageId="6" pageNumber="123">
<taxonomicName id="C5704A13922D01F5080BE856985F13FB" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius capitalis" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="capitalis">Quedius capitalis</taxonomicName>
<bibRefCitation id="F0C0121F84757011F09FE54260B0EB90" author="Eppelsheim, E" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="25" pageNumber="142" pagination="321 - 346" title="Zur Staphylinenfauna Turkestan's." url="https://doi.org/10.1002/mmnd.48018920111" volume="1892" year="1892">Eppelsheim 1892</bibRefCitation>
, 329 (original description);
<bibRefCitation id="D66FE26E4007A03FF3AFE707D32D47CC" author="Gridelli, E" journalOrPublisher="Memorie della Societa Entomologica Italiana" pageId="25" pageNumber="142" pagination="5 - 180" title="Studi sul genere Quedius Steph. (Coleoptera: Staphylinidae). Secondo contributo. Specie della regione paleartica." volume="3" year="1924">Gridelli 1924</bibRefCitation>
, 40 (characters);
<bibRefCitation id="D271E8670E17EABE6EFF539B91E775F8" pageId="6" pageNumber="123">Coiffait 1978</bibRefCitation>
, 148 (characters and illustration of aedeagus);
<bibRefCitation id="3AD74E6C96D85D6968FE6E9594671BDD" pageId="6" pageNumber="123">Kadyrov et al. 2014a</bibRefCitation>
, 31 and 2014b, 49 (distributional records).
</paragraph>
</subSubSection>
<subSubSection id="41EE8422C803AC260397E455A4F6AC05" pageId="6" pageNumber="123" type="materials_examined">
<paragraph id="20453AA72A64B591855D45B0167BA003" pageId="6" pageNumber="123">Material examined.</paragraph>
<paragraph id="F1410E06F049C52545D6156EEBBC1CC5" pageId="6" pageNumber="123">Type material: Uzbekistan: Syntypes: 1 ♂, &quot;♂/ c.Epplsh. Steind. d. [printed]/ Qu. capitalis Epp. Type Taschkent, Leder [handwritten]/ Typus&quot;; 1 ♂, &quot;♂/ capitalis Epp. Taschkent Leder. [handwritten]/ Typus&quot; (Fig. 7E, F) (NMW).</paragraph>
</subSubSection>
<subSubSection id="3367B4EBBDD4FBF28DF8607E910289E5" pageId="6" pageNumber="123" type="materials_examined">
<paragraph id="1D3CCFA76C3488D0BDB819FB8921E503" pageId="6" pageNumber="123">Additional material.</paragraph>
<paragraph id="37D1AF6948BC991947CBB3EAAEEADA20" pageId="6" pageNumber="123">
Uzbekistan: 2 ♀, Tien Shan, Aktasch, near Taschkent, 2000 m a.s.l., 13.VII.1984, D.W. Wrase leg. (cSch); Kazakhstan: 3 ♂, Karatau Mts, Khantagi River, 570 m a.s.l., 43°33'
<geoCoordinate id="1F14C91C87FDF588EE2C14FC3A0F818A" direction="north" orientation="latitude" precision="5555" value="32.4">32.4N</geoCoordinate>
, 68°40'
<geoCoordinate id="C293DF054DA12F6D5A768A491AECEEE8" direction="east" orientation="longitude" precision="5555" value="52.7">52.7E</geoCoordinate>
, 25.VI.2011, V.A. Kastcheev leg. (ZIN); Tajikistan: 3 ♂, Mountains near Kuljab, 1500-2000 m a.s.l., 20.VI.1963, A.V. Bogachev leg. (ZMMU).
</paragraph>
</subSubSection>
<subSubSection id="0F00F48D2C3929DB1CE411B585DF277D" pageId="6" pageNumber="123" type="type material">
<paragraph id="4C4EB9B5E5037F514EA419C262D85511" pageId="6" pageNumber="123">Comments on taxonomy and type material.</paragraph>
<paragraph id="68596BCDA114FF3F0AA27586E6AA6B20" pageId="6" pageNumber="123">
In the original description,
<bibRefCitation id="827444835B445A784E40E9D04914D076" author="Eppelsheim, E" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="25" pageNumber="142" pagination="321 - 346" title="Zur Staphylinenfauna Turkestan's." url="https://doi.org/10.1002/mmnd.48018920111" volume="1892" year="1892">Eppelsheim (1892)</bibRefCitation>
mentioned morphological characters of both males and females and stated that the species was known to him from a few specimens from
<normalizedToken id="FA70325126EE917D4D3EF08329AEC366" originalValue="Tashkent">''Tashkent''</normalizedToken>
. As we learn from the introduction in his paper, specimens were collected by Hans Leder. Both male specimens from NMW labeled as
<normalizedToken id="36DB49B4C8B3A059D9B7B7A6EF3DDF8A" originalValue="“types”">&quot;types&quot;</normalizedToken>
of
<taxonomicName id="9A312F7CDEE77F7CD8FA42FE8692CDBA" lsidName="Q. capitalis" pageId="6" pageNumber="123" rank="species" species="capitalis">Q. capitalis</taxonomicName>
originally come from
<normalizedToken id="9A7F6BFEB5A1A762F4924F4F4300674A" originalValue="Eppelsheims">Eppelsheim's</normalizedToken>
collection and their morphology and label information fit the original description; therefore they are syntypes.
</paragraph>
<paragraph id="795F2E424489A3430B684BD74C79D852" pageId="6" pageNumber="123">
We have examined aedeagi of both syntypes and confirm they are conspecific.
<bibRefCitation id="58513DB287AAEFC176BBBCB6B69F568A" author="Eppelsheim, E" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="25" pageNumber="142" pagination="321 - 346" title="Zur Staphylinenfauna Turkestan's." url="https://doi.org/10.1002/mmnd.48018920111" volume="1892" year="1892">Eppelsheim (1892)</bibRefCitation>
compared
<taxonomicName id="1388886D674DEE9B794E53AB5192ED9D" lsidName="Q. capitalis" pageId="6" pageNumber="123" rank="species" species="capitalis">Q. capitalis</taxonomicName>
with
<taxonomicName id="53BC97392176166A343B411225BD72F9" lsidName="Q. ragusai" pageId="6" pageNumber="123" rank="species" species="ragusai">Q. ragusai</taxonomicName>
Eppelsheim, 1889.
<bibRefCitation id="280251D05338C8E55D6BA180538ABCBB" author="Gridelli, E" journalOrPublisher="Memorie della Societa Entomologica Italiana" pageId="25" pageNumber="142" pagination="5 - 180" title="Studi sul genere Quedius Steph. (Coleoptera: Staphylinidae). Secondo contributo. Specie della regione paleartica." volume="3" year="1924">Gridelli (1924)</bibRefCitation>
, based on the examination of a syntype, provided additional morphological details for the species including verbal description of its aedeagus (but no illustration) and placed it near
<taxonomicName id="2E177E6CC4ACC64DB15ABE5FEC15C16E" lsidName="Q. ochripennis" pageId="6" pageNumber="123" rank="species" species="ochripennis">Q. ochripennis</taxonomicName>
. Based on a syntype male,
<bibRefCitation id="92C7B0417A7B09F3E791570CB6A74C49" pageId="6" pageNumber="123">Coiffait (1978)</bibRefCitation>
again redescribed this species and provided its first and hitherto the only available illustration of the aedeagus. Our examination of the syntypes of
<taxonomicName id="70896A35BB55D7920806C142868423ED" lsidName="Q. capitalis" pageId="6" pageNumber="123" rank="species" species="capitalis">Q. capitalis</taxonomicName>
confirms the correct identification of this species by both
<bibRefCitation id="8891A0790CD7E92D7040BAE5576A2FBD" author="Gridelli, E" journalOrPublisher="Memorie della Societa Entomologica Italiana" pageId="25" pageNumber="142" pagination="5 - 180" title="Studi sul genere Quedius Steph. (Coleoptera: Staphylinidae). Secondo contributo. Specie della regione paleartica." volume="3" year="1924">Gridelli (1924)</bibRefCitation>
and
<bibRefCitation id="1C04B04FD56535F6247D26F872D65657" pageId="6" pageNumber="123">Coiffait (1978)</bibRefCitation>
. In
<bibRefCitation id="E396EC59FEF8AD79AB6ECB8AFD3F7CC2" author="Schuelke, M" journalOrPublisher="Brill, Leiden, Boston" pageId="26" pageNumber="143" title="Staphylinidae, 304 - 1134. In: Loebl I, Loebl D (Eds) Catalogue of Palaearctic Coleoptera. Volume 2. Hydrophiloidea-Staphylinoidea, Revised and updated edition." year="2015">
<normalizedToken id="A1D07B7C590AF8ADD5BA063707F2171C" originalValue="Schülke">Schuelke</normalizedToken>
and Smetana (2015)
</bibRefCitation>
<taxonomicName id="D49E5648DCFA780BDCE0165E1EABBF36" lsidName="Q. capitalis" pageId="6" pageNumber="123" rank="species" species="capitalis">Q. capitalis</taxonomicName>
was erroneously placed in the Subgenus
<taxonomicName id="8BF62F1338303E91DF68B93CC349EF24" genus="Amblyopinina" lsidName="Amblyopinina (Raphirus)" pageId="6" pageNumber="123" rank="subGenus" subGenus="Raphirus">Raphirus</taxonomicName>
. Here we redescribe this poorly known species and provide further data on its distribution.
</paragraph>
</subSubSection>
<subSubSection id="032785466EEAF1BDCD7A84B4EC7BAEA2" lastPageId="7" lastPageNumber="124" pageId="6" pageNumber="123" type="redescription">
<paragraph id="2A2CB9F60A9F00764CC31050A699A139" pageId="6" pageNumber="123">Redescription.</paragraph>
<paragraph id="CF691A1CC11184B024983718DC67080D" pageId="6" pageNumber="123">Measurements and ratios (range, arithmetic mean; n = 10): HL: 0.8-1.3 (1.0); HW: 0.8-1.5 (1.1); PL: 0.9-1.6 (1.3); PW: 1.1-1.8 (1.4); EL: 1.2-2.0 (1.6); EW: 1.2-1.9 (1.5); FB: 2.9-4.7 (3.9); TL: 6.5-9.3 (7.8); HL/HW: 0.8-1.1 (1.0); PL/PW: 0.8-1.0 (0.9); EL/EW: 1.0-1.2 (1.1).</paragraph>
<paragraph id="1E306869C1BD59A8671D8C2517DD5DBB" pageId="6" pageNumber="123">Body black to dark brown, hind margins of abdominal tergites slightly paler; elytra reddish; palpi and other appendages slightly lighter; body glossy (Fig. 7A, B).</paragraph>
<paragraph id="AAB1627EB94C416D8B432ED4414B2176" pageId="6" pageNumber="123">Head approximately as wide as long or slightly longer; eyes small, not convex; temples as long as longitudinal diameter of eye; posterior frontal puncture closer to posterior margin of head than to anterior frontal puncture; temporal puncture closer to posterior margin of head than to posterior margin of eye; two vertical punctures behind posterior frontal puncture arranged as slightly oblique line between posterior margin of eye and dorsal part of neck; microsculpture of head with transverse distinct wavelines.</paragraph>
<paragraph id="77D9FF49D984A632E6D3A5A5ECA13AFE" pageId="6" pageNumber="123">
Antennae moderately long, antennal segments: 3rd longer than 2nd, 4
<normalizedToken id="B76CB3703009710D892794F45D56A552" originalValue="th">th-</normalizedToken>
10th gradually widening towards apex of antenna.
</paragraph>
<paragraph id="79DD349E0422931E15B164D5C2158295" pageId="6" pageNumber="123">Pronotum slightly wider than long, widest at about middle to posterior third; hind angles rounded but distinct; dorsal and sublateral rows each with three punctures; microsculpture with transverse waves as on posterior part of head. Scutellum impunctate with microsculpture slightly coarser than on pronotum. Elytra parallel-sided, slightly longer than wide, longer than pronotum, their punctation dense, interspaces shiny with distinct minute irregularities.</paragraph>
<paragraph id="065F9D199995CDA9DF2E79C4A33175FD" pageId="6" pageNumber="123">Abdomen: punctation fine and dense; interspaces with minute irregularities; posterior margin of tergite VII with palisade fringe.</paragraph>
<paragraph id="E33F9744AC9B387B6A8F141C6D446378" lastPageId="7" lastPageNumber="124" pageId="6" pageNumber="123">
Male: protarsi with tarsomeres 1-4 dilated stronger than in females. Sternite VIII with weak triangular medio-apical emargination; tergite X triangular with setae; sternite IX elongate, gradually narrowed apically, with moderately wide and long basal portion and obtusely rounded apical margin with numerous setae. Aedeagus (Fig. 7C, D): median lobe parallel-sided with broad and obtuse apex and tooth located near apex (Fig. 7 C). Paramere rhomboid sharply narrowing apicad; its apex almost reaching apex of median lobe, with two pairs of apical setae and two pairs of lateral setae below apex;
<pageBreakToken id="E50AB6FC9BC6E8CCD3B0E5B67A3A93C9" pageId="7" pageNumber="124" start="start">paramere</pageBreakToken>
(underside) with ca. 4-8 sensory peg setae in each of two sinuate lateral rows that extend basad over pairs of lateral setae (Fig. 7D).
</paragraph>
</subSubSection>
<subSubSection id="4E16A22351E4974870C7925140CF3C3C" pageId="7" pageNumber="124" type="comparisons">
<paragraph id="5F35B18AAE44557B5A57A7FDB53141B4" pageId="7" pageNumber="124">Comparison.</paragraph>
<paragraph id="CB18EF2AD5EEB0BE063AD9554AFF15B8" pageId="7" pageNumber="124">
<taxonomicName id="FD4F4A9C6EAFABA706DC727BB9AE3687" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius capitalis" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="capitalis">Quedius capitalis</taxonomicName>
seems to be closely related to
<taxonomicName id="5FBAE183B6B64391B0DDEED38272ED12" lsidName="Q. fusicornis" pageId="7" pageNumber="124" rank="species" species="fusicornis">Q. fusicornis</taxonomicName>
and
<taxonomicName id="24E4FCCD0988315ADB6C105C9DB0054B" lsidName="Q. ochripennis" pageId="7" pageNumber="124" rank="species" species="ochripennis">Q. ochripennis</taxonomicName>
from which it can be easily distinguished externally by smaller body size and proportions, and by the structure of paramere with two sinuate lateral rows of peg setae (ca.4-8 in each row) extending basad over pairs of lateral setae.
</paragraph>
</subSubSection>
<subSubSection id="7CD6B10DD34C2FD88A80F98FCE9332A1" pageId="7" pageNumber="124" type="distribution">
<paragraph id="92847709C5E4D08CDE2A57E1521B6288" pageId="7" pageNumber="124">Distribution.</paragraph>
<paragraph id="A2DF2B17C3AC916C85A05313AAD22705" pageId="7" pageNumber="124">
Based on the literature data (Table 1) and newly examined material,
<taxonomicName id="58C0B2B460CAD545B401498CF039E3A4" lsidName="Q. capitalis" pageId="7" pageNumber="124" rank="species" species="capitalis">Q. capitalis</taxonomicName>
is known from several localities near Tashkent (Uzbekistan), Karatau Mountains (southwestern Kazakhstan) and Hazratisho Mountains (southern Tajikistan).
</paragraph>
<caption id="AAAB37F99EBF2D01F5EA30635CDC0E94" pageId="7" pageNumber="124">
<paragraph id="BE22C5853F1D46503B74686756A11C6B" pageId="7" pageNumber="124">
Figure 7.
<taxonomicName id="64A26C9D32B8D183E5F8A3E4A177954A" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius capitalis" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="capitalis">Quedius capitalis</taxonomicName>
, syntypes, males. A, B, habitus; C, D, aedeagus of the syntype in the photo A. E (of the syntype in the photo A), F (of the syntype in the photo B), labels. C, median lobe, lateral view; D, paramere, underside. Scale bars: A, B = 1 mm; C = 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="087F79B34CB46DB888E1CE3234032930" pageId="7" pageNumber="124" type="bionomics">
<paragraph id="0BED97170182ED7E60B3816D6CC6508F" pageId="7" pageNumber="124">Bionomics.</paragraph>
<paragraph id="6B0D0158AB67F8D99D216A1CA20822D8" pageId="7" pageNumber="124">Unknown.</paragraph>
</subSubSection>
</treatment>
</document>