184 lines
21 KiB
XML
184 lines
21 KiB
XML
<document id="7E51400A92AA8B980184EBB7DFA119FE" ID-CLB-Dataset="55847" ID-DOI="10.11646/zootaxa.5128.2.4" ID-GBIF-Dataset="a9b6e575-57e1-4fe3-90ee-c4a375f2958e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6479814" ID-ZooBank="5C65C12C-7943-47C8-B08E-01B84B4B0ED0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1650721128229" checkinUser="plazi" docAuthor="Viraktamath, C. A., Webb, M. D. & Yeshwanth, H. M." docDate="2022" docId="844C5277FFA7FFBA15DBF947B796FBE1" docLanguage="en" docName="zootaxa.5128.2.4.pdf" docOrigin="Zootaxa 5128 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Drabescus nervosopunctatus Signoret 1880" docType="treatment" docVersion="7" lastPageNumber="238" masterDocId="78752A0FFFABFFB7154CFF81B637FFD4" masterDocTitle="Review of the leafhopper genus Drabescus Stål (Hemiptera: Cicadellidae: Deltocephalinae) in the Indian subcontinent with description of one new species" masterLastPageNumber="247" masterPageNumber="225" pageNumber="237" updateTime="1699329088942" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="78D312798C74CF8A0AAAD81A2DEB90A0">Review of the leafhopper genus Drabescus Stål (Hemiptera: Cicadellidae: Deltocephalinae) in the Indian subcontinent with description of one new species</mods:title>
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<mods:roleTerm id="0BDC91EAFCA50471037F5201849CB461">Author</mods:roleTerm>
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<mods:namePart id="B0AB6BAB806D7A8BDF34B73BCA2958DD">Viraktamath, C. A.</mods:namePart>
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<mods:affiliation id="46A2953361F738A0AB2208482085B88C">Department of Entomology, University of Agricultural Sciences, GKVK, Bengaluru 560065, India.</mods:affiliation>
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<mods:nameIdentifier id="9F8B6CA95595DABAB266B2A8D1FCC935" type="email">chandrashekara.viraktamath@gmail.com</mods:nameIdentifier>
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<mods:namePart id="8A4213AE3637A9CEDE6ABB4E4FB213B0">Webb, M. D.</mods:namePart>
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<mods:nameIdentifier id="F0EC636EC5A65C053442975CFFFE4292" type="ORCID">0000-0002-1312-6142</mods:nameIdentifier>
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<mods:affiliation id="CAA23A1666F38DDB55E216DEC967710D">Department of Life Science, The Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom. & m. webb @ nhm. ac. uk; https: // orcid. org / 0000 - 0002 - 1312 - 6142</mods:affiliation>
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<mods:nameIdentifier id="3E951515E6AB39F897D9D7AE9AC2D4E7" type="email">m.webb@nhm.ac.uk</mods:nameIdentifier>
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<mods:namePart id="4DBEDCD47F15DE5AD9374BF59A099F43">Yeshwanth, H. M.</mods:namePart>
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<mods:affiliation id="60F608EC9CC65FFBC55F53DE230E8F35">Department of Entomology, University of Agricultural Sciences, GKVK, Bengaluru 560065, India. & hmyeshwanth @ gmail. com; https: // orcid. org / 0000 - 0002 - 5104 - 3401</mods:affiliation>
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<mods:nameIdentifier id="32C8F5E0CFEF7F5BF114C54AF71AC082" type="email">chandrashekara.viraktamath@gmail.com</mods:nameIdentifier>
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<mods:title id="AF138FD8509C03EEEAC69EF5C7D70C73">Zootaxa</mods:title>
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<mods:date id="C6A52841D39AA1CC40847E311F33BA76">2022</mods:date>
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<mods:number id="BB81BC8A10243882FBC9434B0DFE2D92">2022-04-20</mods:number>
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<mods:number id="DF4F307ACE3958D0DFF476A289393CAE">5128</mods:number>
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<treatment id="844C5277FFA7FFBA15DBF947B796FBE1" ID-DOI="http://doi.org/10.5281/zenodo.6479834" ID-GBIF-Taxon="194848000" ID-Zenodo-Dep="6479834" LSID="urn:lsid:plazi:treatment:844C5277FFA7FFBA15DBF947B796FBE1" httpUri="http://treatment.plazi.org/id/844C5277FFA7FFBA15DBF947B796FBE1" lastPageId="13" lastPageNumber="238" pageId="12" pageNumber="237">
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<subSubSection id="44FFB0EAFFA7FFBB15DBF947B454F934" box="[151,611,1734,1761]" pageId="12" pageNumber="237" type="nomenclature">
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<paragraph id="0C5AE361FFA7FFBB15DBF947B454F934" blockId="12.[151,611,1734,1797]" box="[151,611,1734,1761]" pageId="12" pageNumber="237">
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<heading id="5712540DFFA7FFBB15DBF947B454F934" bold="true" box="[151,611,1734,1761]" fontSize="11" level="1" pageId="12" pageNumber="237" reason="1">
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<taxonomicName id="CBE598E2FFA7FFBB15DBF947B454F934" ID-CoL="37JMG" authority="Signoret" authorityName="Signoret" authorityYear="1880" box="[151,611,1734,1761]" class="Insecta" family="Cicadellidae" genus="Drabescus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="237" phylum="Arthropoda" rank="species" species="nervosopunctatus">
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<emphasis id="3E913F73FFA7FFBB15DBF947B454F934" bold="true" box="[151,611,1734,1761]" pageId="12" pageNumber="237">
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<emphasis id="3E913F73FFA7FFBB15DBF947B7C3F935" bold="true" box="[151,500,1734,1761]" italics="true" pageId="12" pageNumber="237">Drabescus nervosopunctatus</emphasis>
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Signoret
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="44FFB0EAFFA7FFBB15DBF96AB7E4F8D1" box="[151,467,1770,1797]" pageId="12" pageNumber="237" type="description">
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<paragraph id="0C5AE361FFA7FFBB15DBF96AB7E4F8D1" blockId="12.[151,611,1734,1797]" box="[151,467,1770,1797]" pageId="12" pageNumber="237">
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<figureCitation id="94DEFFE4FFA7FFBB15DBF96AB6EBF8D1" box="[151,220,1771,1797]" captionStart="FIGURES 2" captionStartId="5.[151,264,1900,1924]" captionTargetBox="[200,1388,193,1866]" captionTargetId="figure-18@5.[187,1401,181,1876]" captionTargetPageId="5" captionText="FIGURES 2 A–L. Species of Drabescus from Indian subcontinent. Habitus, dorsal and lateral views (A–F) and female sternite VII (G–L). A–B, D. nervosopunctatus Signoret; C–D, D. nigrofacialis (Distant); E–F, D. shillongensis Rao; G, D. angulatus Signoret; H, D. kempi Pruthi; I, D. limbaticeps Signoret; J, D. nervosopunctatus Signoret; K, D. nigrofacialis (Distant); L, D. shillongensis Rao." figureDoi="http://doi.org/10.5281/zenodo.6479818" httpUri="https://zenodo.org/record/6479818/files/figure.png" pageId="12" pageNumber="237">Figs 2</figureCitation>
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ABJ, 3G, 5E, 10A–H.
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</paragraph>
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</subSubSection>
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<subSubSection id="44FFB0EAFFA7FFBB15DBF8AEB45AF87B" pageId="12" pageNumber="237" type="reference_group">
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<paragraph id="0C5AE361FFA7FFBB15DBF8AEB6C3F8B2" blockId="12.[151,1437,1839,1894]" pageId="12" pageNumber="237">
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<treatmentCitationGroup id="2CF5C44FFFA7FFBB15DBF8AEB6C3F8B2" pageId="12" pageNumber="237">
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<treatmentCitation id="8D44C570FFA7FFBB15DBF8AEB4B5F893" author="Signoret, V." box="[151,642,1839,1863]" page="209" pageId="12" pageNumber="237" year="1880">
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<taxonomicName id="CBE598E2FFA7FFBB15DBF8AEB4B5F893" ID-CoL="37JMG" authority="Signoret 1880: 209" authorityName="Signoret" authorityPageNumber="209" authorityYear="1880" box="[151,642,1839,1863]" class="Insecta" family="Cicadellidae" genus="Drabescus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="237" phylum="Arthropoda" rank="species" species="nervosopunctatus">
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<emphasis id="3E913F73FFA7FFBB15DBF8AEB78BF893" box="[151,444,1839,1863]" italics="true" pageId="12" pageNumber="237">Drabescus nervosopunctatus</emphasis>
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<bibRefCitation id="68749E90FFA7FFBB148DF8AEB4B5F893" author="Signoret, V." box="[449,642,1839,1863]" pageId="12" pageNumber="237" pagination="189 - 212" refId="ref11552" refString="Signoret, V. (1880) Essai sur les Jassides Stal, Fieb. et plus particulierement sur les Acocephalides Puton. Annates de la Societe Entomologique de France 10, 189 - 212." type="journal article" year="1880">Signoret 1880: 209</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, pl.23, fig. 72;
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<treatmentCitation id="8D44C570FFA7FFBB1654F8AEB23FF893" author="Distant, W. L." box="[792,1032,1839,1863]" page="304" pageId="12" pageNumber="237" year="1908">
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<bibRefCitation id="68749E90FFA7FFBB1654F8AEB23FF893" author="Distant, W. L." box="[792,1032,1839,1863]" pageId="12" pageNumber="237" pagination="1 - 501" refId="ref11088" refString="Distant, W. L. (1908) Rhynchota-Homoptera. In: Bingham, C. T. (Ed.), The Fauna of British India including Ceylon and Burma, IV, pp. i - xv + 1 - 501." type="book chapter" year="1908">Distant, 1908: 304–305</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="8D44C570FFA7FFBB115CF8AEB33FF893" author="Zhang, Y. L. & Webb, M. D." box="[1040,1288,1839,1863]" page="25" pageId="12" pageNumber="237" year="1996">
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<bibRefCitation id="68749E90FFA7FFBB115CF8AEB33FF893" author="Zhang, Y. L. & Webb, M. D." box="[1040,1288,1839,1863]" pageId="12" pageNumber="237" pagination="1 - 103" refId="ref11914" refString="Zhang, Y. L. & Webb, M. D. (1996) A revised classification of the Asian and Pacific Selenocephalinae leafhoppers (Homoptera: Cicadellidae). Bulletin of the Natural History Museum, Entomology Series, 65, 1 - 103." type="journal article" year="1996">Zhang & Webb 1996: 25</bibRefCitation>
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</treatmentCitation>
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, figs 413–417, 527.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="0C5AE361FFA7FFBB15DBF815B45AF87B" blockId="12.[151,1436,1940,2039]" box="[151,621,1940,1967]" pageId="12" pageNumber="237">
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Coloration similar to that in
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<taxonomicName id="CBE598E2FFA7FFBB1499F814B452F87A" ID-CoL="37JLR" authorityName="Signoret" authorityYear="1880" box="[469,613,1940,1966]" class="Insecta" family="Cicadellidae" genus="Drabescus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="237" phylum="Arthropoda" rank="species" species="angulatus">
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<emphasis id="3E913F73FFA7FFBB1499F814B452F87A" box="[469,613,1940,1966]" italics="true" pageId="12" pageNumber="237">D. angulatus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="44FFB0EAFFA7FFBA158BF839B74EFD25" lastPageId="13" lastPageNumber="238" pageId="12" pageNumber="237" type="description">
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<paragraph id="0C5AE361FFA7FFBA158BF839B326FF01" blockId="12.[151,1436,1940,2039]" lastBlockId="13.[151,1437,151,1077]" lastPageId="13" lastPageNumber="238" pageId="12" pageNumber="237">Crown medially little shorter than next to eyes, more than 7× as wide between eyes as long medially, transversely depressed, transition to face rimmed. Face with lateral frontal sutures slightly convex in basal half; distance between antennae about 3.8 as wide as apex of frontoclypeus. Clypellus widened apically. Pronotum about 2.6× as wide as median length. Forewings without cross veins between claval veins. Fore tibiae not expanded.</paragraph>
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<paragraph id="0C5AE361FFA6FFBA158BFF5EB4B1FDE9" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="238">
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<emphasis id="3E913F73FFA6FFBA158BFF5EB75AFF2D" box="[199,365,223,249]" italics="true" pageId="13" pageNumber="238">Male genitalia</emphasis>
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. Pygofer about 1.5× as long as basal height in lateral view, basal half broad then abruptly narrowed then gradually to conically rounded apex, with spine-like process arising from ventral margin, dorsal half of ventral margin serrated before the origin of the process; a few macrosetae present in distal 0.33. Valve with anterior margin convex, about 3.3 times as wide as long medially. Subgenital plate, elongate, triangular, about 2.7 × as long as basal width, distal half a membranous appendix. Style with well-developed subapical lobe, apophysis curved laterally, apex narrowed and surface transversely striate. Connective with broad, quadrangular stem, about as long as arms, anterior margin without median lobe. Aedeagus with shaft more or less of uniform width in lateral view with lateral processes at base, widely separated at base, about as wide as shaft at midlength with their apices divergent in dorsal and ventral views, slightly shorter than shaft; dorsal apodeme well-developed, slightly more than half length of shaft; gonopore apical on ventral surface.
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</paragraph>
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<paragraph id="0C5AE361FFA6FFBA158BFDC6B23EFD51" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="238">
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<emphasis id="3E913F73FFA6FFBA158BFDC6B7C3FDB4" box="[199,500,583,609]" italics="true" pageId="13" pageNumber="238">Female pregenital sternite</emphasis>
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. Sternite VII about 1.75 as wide as long medially, Posterior margin sinuate, with lateral angles slightly produced posteriorly; medially dark brown in distal half.
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</paragraph>
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<paragraph id="0C5AE361FFA6FFBA158BFD0EB74EFD25" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="238">
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<emphasis id="3E913F73FFA6FFBA158BFD0EB75EFD7C" box="[199,361,655,680]" italics="true" pageId="13" pageNumber="238">Measurements</emphasis>
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. Male
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<quantity id="CB1D4E84FFA6FFBA14F1FD0EB471FD7D" box="[445,582,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="8.4" metricValueMin="8.2" pageId="13" pageNumber="238" unit="mm" value="8.3" valueMax="8.4" valueMin="8.2">8.2–8.4 mm</quantity>
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long,
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<quantity id="CB1D4E84FFA6FFBA17DDFD0EB52DFD7D" box="[657,794,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="2.9" metricValueMin="2.8" pageId="13" pageNumber="238" unit="mm" value="2.8499999999999996" valueMax="2.9" valueMin="2.8">2.8–2.9 mm</quantity>
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wide across eyes and
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<quantity id="CB1D4E84FFA6FFBA1153FD0FB29FFD7D" box="[1055,1192,654,681]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="13" pageNumber="238" unit="mm" value="2.55" valueMax="2.6" valueMin="2.5">2.5–2.6 mm</quantity>
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wide across posterolateral angles of pronotum. Female
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<quantity id="CB1D4E84FFA6FFBA1765FD32B448FD19" box="[553,639,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="13" pageNumber="238" unit="mm" value="8.8">8.8 mm</quantity>
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long,
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<quantity id="CB1D4E84FFA6FFBA178BFD32B529FD19" box="[711,798,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="238" unit="mm" value="3.2">3.2 mm</quantity>
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wide across eyes and
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<quantity id="CB1D4E84FFA6FFBA1156FD32B246FD19" box="[1050,1137,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="13" pageNumber="238" unit="mm" value="2.9">2.9 mm</quantity>
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wide across posterolateral angles of pronotum.
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</paragraph>
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</subSubSection>
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<subSubSection id="44FFB0EAFFA6FFBA158BFD7BB773FCED" pageId="13" pageNumber="238" type="materials_examined">
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<paragraph id="0C5AE361FFA6FFBA158BFD7BB773FCED" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="238">
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<emphasis id="3E913F73FFA6FFBA158BFD7BB781FCC0" bold="true" box="[199,438,762,788]" pageId="13" pageNumber="238">Material examined.</emphasis>
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<materialsCitation id="BC8DE93CFFA6FFBA148EFD7AB777FCED" ID-GBIF-Occurrence="3756540309" collectingDate="2015-06-10" collectionCode="IARI, UASB" country="India" county="Dehradun" location="Chakrata" municipality="Mukta" pageId="13" pageNumber="238" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Uttarakhand">
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<collectingCountry id="74F2A3F1FFA6FFBA148EFD7AB421FCC1" box="[450,534,763,789]" name="India" pageId="13" pageNumber="238">INDIA</collectingCountry>
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:
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<collectingRegion id="CE212D83FFA6FFBA176AFD7AB482FCC1" box="[550,693,763,789]" country="India" name="Uttarakhand" pageId="13" pageNumber="238">Uttarakhand</collectingRegion>
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:
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<specimenCount id="1AE328E8FFA6FFBA1788FD7BB4DEFCC1" box="[708,745,762,789]" pageId="13" pageNumber="238" type="male">5♂</specimenCount>
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,
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<specimenCount id="1AE328E8FFA6FFBA17B7FD7AB52AFCC1" box="[763,797,763,789]" pageId="13" pageNumber="238" type="female">1♀</specimenCount>
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,
|
||
<collectingCounty id="E53B9BEDFFA6FFBA167DFD7AB595FCC1" box="[817,930,763,789]" pageId="13" pageNumber="238">Dehradun</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="EC3E791BFFA6FFBA16FFFD7AB5CBFCC1" box="[947,1020,763,789]" pageId="13" pageNumber="238">Mukta</collectingMunicipality>
|
||
pokhari, new
|
||
<location id="093AB5BAFFA6FFBA11E0FD7AB323FCC1" LSID="urn:lsid:plazi:treatment:844C5277FFA7FFBA15DBF947B796FBE1:093AB5BAFFA6FFBA11E0FD7AB323FCC1" box="[1196,1300,763,789]" country="India" county="Dehradun" municipality="Mukta" name="Chakrata" pageId="13" pageNumber="238" stateProvince="Uttarakhand">Chakrata</location>
|
||
,
|
||
<date id="785BC5A1FFA6FFBA1068FD7BB3AAFCC1" box="[1316,1437,762,789]" pageId="13" pageNumber="238" value="2015-06-10">
|
||
<collectingDate id="681F3C49FFA6FFBA1068FD7BB3AAFCC1" box="[1316,1437,762,789]" pageId="13" pageNumber="238" value="2015-06-10">10.vi.2015</collectingDate>
|
||
</date>
|
||
(
|
||
<collectionCode id="6AF47BA4FFA6FFBA15D3FC9EB6EAFCED" box="[159,221,799,825]" country="India" lsid="urn:lsid:biocol.org:col:13272" name="Indian Agricultural Research Institute" pageId="13" pageNumber="238">IARI</collectionCode>
|
||
,
|
||
<collectionCode id="6AF47BA4FFA6FFBA15A4FC9EB70FFCED" box="[232,312,799,825]" country="India" httpUri="http://grbio.org/cool/yr7i-jnjm" name="University of Agricultural Sciences" pageId="13" pageNumber="238">UASB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="44FFB0EAFFA6FFBA158BFCC2B796FBE1" pageId="13" pageNumber="238" type="discussion">
|
||
<paragraph id="0C5AE361FFA6FFBA158BFCC2B796FBE1" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="238">
|
||
<emphasis id="3E913F73FFA6FFBA158BFCC2B703FC89" bold="true" box="[199,308,835,861]" pageId="13" pageNumber="238">Remarks</emphasis>
|
||
. This species was described from an unknown number of female specimens (
|
||
<typeStatus id="D35E5DC3FFA6FFBA11FAFCC2B325FC89" box="[1206,1298,835,861]" pageId="13" pageNumber="238" type="syntype">syntype</typeStatus>
|
||
(s)) with the following data: “Indes nord” in “coll. Signoret”. The extant type was examined by
|
||
<bibRefCitation id="68749E90FFA6FFBA1103FCE6B35CFC55" author="Zhang, Y. L. & Webb, M. D." box="[1103,1387,870,897]" pageId="13" pageNumber="238" pagination="1 - 103" refId="ref11914" refString="Zhang, Y. L. & Webb, M. D. (1996) A revised classification of the Asian and Pacific Selenocephalinae leafhoppers (Homoptera: Cicadellidae). Bulletin of the Natural History Museum, Entomology Series, 65, 1 - 103." type="journal article" year="1996">Zhang & Webb 1996: 25</bibRefCitation>
|
||
and designated as
|
||
<typeStatus id="D35E5DC3FFA6FFBA1479FC0AB7A8FC71" box="[309,415,907,933]" pageId="13" pageNumber="238" type="lectotype">lectotype</typeStatus>
|
||
. Male of this species can be readily recognised among species of
|
||
<taxonomicName id="CBE598E2FFA6FFBA11C9FC0AB2CAFC71" authorityName="Stal" authorityYear="1870" box="[1157,1277,907,933]" class="Insecta" family="Cicadellidae" genus="Drabescus" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="238" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3E913F73FFA6FFBA11C9FC0AB2CAFC71" box="[1157,1277,907,933]" italics="true" pageId="13" pageNumber="238">Drabescus</emphasis>
|
||
</taxonomicName>
|
||
from the subcontinent by the short somewhat compressed aedeagal shaft with the basal processes divergent and wider than shaft at midlength in either ventral or dorsal view and also shorter. Also, the pygofer ventral margin is serrated before the apex. The female can be recognised by the sternite VII being sinuate and neither excavated nor incised medially and nor with a median lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |