251 lines
18 KiB
XML
251 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.695.19906" ID-GBIF-Dataset="07675d9d-03d7-4cc3-9846-352b910e0add" ID-PMC="PMC5673837" ID-Pensoft-Pub="1313-2970-695-111" ID-PubMed="29134010" ID-ZBK="091F6C649E8D45DA9F31F5127DFA10CB" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-695-111" ModsDocOrigin="ZooKeys 695" ModsDocTitle="Nornalup, a new genus of pselaphine beetle from southwestern Australia (Coleoptera, Staphylinidae, Pselaphinae, Faronitae)" checkinTime="1504626281574" checkinUser="pensoft" docAuthor="Park, Jong-Seok & Chandler, Donald S." docDate="2017" docId="7659369AC8FE3AF12D96F9F902CC55C2" docLanguage="en" docName="ZooKeys 695: 111-121" docOrigin="ZooKeys 695" docSource="http://dx.doi.org/10.3897/zookeys.695.19906" docTitle="Nornalup Park & Chandler, gen. n." docType="treatment" docUuid="691083BF-7137-48CF-BB87-7F735392CFCD" docUuidSource="ZooBank" docVersion="5" lastPageNumber="114" masterDocId="FFB6292D5839FF89FFD83B67B214FF92" masterDocTitle="Nornalup, a new genus of pselaphine beetle from southwestern Australia (Coleoptera, Staphylinidae, Pselaphinae, Faronitae)" masterLastPageNumber="121" masterPageNumber="111" pageNumber="112" updateTime="1668164759149" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Nornalup, a new genus of pselaphine beetle from southwestern Australia (Coleoptera, Staphylinidae, Pselaphinae, Faronitae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Park, Jong-Seok</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chandler, Donald S.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>695</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>111</mods:start>
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<mods:end>121</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.695.19906</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.695.19906</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-695-111</mods:identifier>
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<mods:identifier type="ZBK">091F6C649E8D45DA9F31F5127DFA10CB</mods:identifier>
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<mods:identifier type="ZooBank">091F6C649E8D45DA9F31F5127DFA10CB</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="133186111" LSID="urn:lsid:zoobank.org:act:691083BF-7137-48CF-BB87-7F735392CFCD" httpUri="http://treatment.plazi.org/id/7659369AC8FE3AF12D96F9F902CC55C2" lastPageId="3" lastPageNumber="114" pageId="1" pageNumber="112">
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<subSubSection pageId="1" pageNumber="112" type="nomenclature">
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<paragraph pageId="1" pageNumber="112">
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<taxonomicName LSID="http://zoobank.org/691083BF-7137-48CF-BB87-7F735392CFCD" authority="Park & Chandler" class="Insecta" family="Staphylinidae" genus="Nornalup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nornalup" order="Coleoptera" pageId="1" pageNumber="112" phylum="Arthropoda" rank="genus">Nornalup Park & Chandler</taxonomicName>
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<taxonomicNameLabel pageId="1" pageNumber="112">gen. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="112" type="type species">
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<paragraph pageId="1" pageNumber="112">Type species.</paragraph>
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<paragraph pageId="1" pageNumber="112">
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Nornalup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nornalup afoveatus" order="Coleoptera" pageId="1" pageNumber="112" phylum="Arthropoda" rank="species" species="afoveatus">Nornalup afoveatus</taxonomicName>
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Park and Chandler, sp. n., herein designated.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="112" type="diagnosis">
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<paragraph pageId="1" pageNumber="112">Diagnosis.</paragraph>
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<paragraph pageId="1" pageNumber="112">
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Members of this genus are easily separated from other faronite genera by the following combination of characters: rostrum separated by distinct frontal sulcus (Fig. 3a); ventral surface of head swollen (Fig. 3b); eyes extremely large, longer than length of temples (Fig. 2
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<normalizedToken originalValue="g–l">g-l</normalizedToken>
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); frontal sulcus deep and wide, open anteriorly (Fig. 3a); mesoventrite with lateral mesosternal fovea and promesocoxal fovea (Fig. 3d); metaventrite with or without median metasternal fovea (Fig. 3d: arrow); abdominal length of tergite and sternite VI approximately 1.5 times longer than V (Fig. 1); female sternite IX bearing two pairs of long setae (Fig. 4
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<normalizedToken originalValue="a–c">a-c</normalizedToken>
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); species only known from Western Australia (Fig. 5).
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</paragraph>
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<caption pageId="1" pageNumber="112">
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<paragraph pageId="1" pageNumber="112">
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Figure 1. Habiti, dorsal view. a
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Nornalup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nornalup afoveatus" order="Coleoptera" pageId="1" pageNumber="112" phylum="Arthropoda" rank="species" species="afoveatus">Nornalup afoveatus</taxonomicName>
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sp. n. b
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<taxonomicName lsidName="N. quadratus" pageId="1" pageNumber="112" rank="species" species="quadratus">N. quadratus</taxonomicName>
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sp. n. c
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<taxonomicName lsidName="N. minusculus" pageId="1" pageNumber="112" rank="species" species="minusculus">N. minusculus</taxonomicName>
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sp. n. Scale bars: 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="113" type="description">
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<paragraph pageId="2" pageNumber="113">
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<pageBreakToken pageId="2" pageNumber="113" start="start">Description</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="2" pageNumber="113">
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Small body size, 1.1-2.5 mm (Fig. 1). Body yellowish to reddish-brown (Fig. 1). Head. Triangular with extremely large eyes, widest across eyes (Fig. 2
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<normalizedToken originalValue="g–l">g-l</normalizedToken>
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). Gular area convex (Fig. 3b). Male antennomeres longer than those of female. Male and female antennomeres with tubercles on 4-11 and 8-11, respectively (Fig. 2
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<normalizedToken originalValue="a–f">a-f</normalizedToken>
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). Frontal sulcus deep and wide, open anteriorly (Fig. 3a). Thorax. Prosternum as long as wide, widest at midpoint of prosternum (Fig. 3c). Prosternum with lateral procoxal and median procoxal fovea (Fig. 3c). Meso- and metathorax trapezoidal, longer than wide (Fig. 3d). Mesoventrite with lateral mesosternal, promesocoxal and lateral mesocoxal foveae (Fig. 3d). Metaventrite with lateral metasternal foveae (Fig. 3d). Abdomen. Length of tergite and sternite VI approximately 1.5 times longer than V (Fig. 1). Aedeagus. Median lobe longer than parameres (Fig. 4
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<normalizedToken originalValue="d–i">d-i</normalizedToken>
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). Phallobase rounded (Fig. 4
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<normalizedToken originalValue="d–i">d-i</normalizedToken>
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). Parameres symmetrical, as wide as median lobe, bearing setae at apex (Fig. 4
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<normalizedToken originalValue="d–i">d-i</normalizedToken>
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).
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</paragraph>
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<caption pageId="2" pageNumber="113">
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<paragraph pageId="2" pageNumber="113">
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Figure 2. Antennae of
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Nornalup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nornalup afoveatus" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="afoveatus">Nornalup afoveatus</taxonomicName>
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sp. n. a male b female. Antennae of
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<taxonomicName lsidName="N. quadratus" pageId="2" pageNumber="113" rank="species" species="quadratus">N. quadratus</taxonomicName>
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sp. n. c male d female. Antennae of
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<taxonomicName lsidName="N. minusculus" pageId="2" pageNumber="113" rank="species" species="minusculus">N. minusculus</taxonomicName>
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sp. n. e male f female. Male heads, dorsal view g
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<taxonomicName lsidName="N. afoveatus" pageId="2" pageNumber="113" rank="species" species="afoveatus">N. afoveatus</taxonomicName>
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sp. n. h
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<taxonomicName lsidName="N. quadratus" pageId="2" pageNumber="113" rank="species" species="quadratus">N. quadratus</taxonomicName>
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sp. n. i
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<taxonomicName lsidName="N. minusculus" pageId="2" pageNumber="113" rank="species" species="minusculus">N. minusculus</taxonomicName>
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sp. n. Female heads, dorsal view j
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<taxonomicName lsidName="N. afoveatus" pageId="2" pageNumber="113" rank="species" species="afoveatus">N. afoveatus</taxonomicName>
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sp. n. k
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<taxonomicName lsidName="N. quadratus" pageId="2" pageNumber="113" rank="species" species="quadratus">N. quadratus</taxonomicName>
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sp. n. l
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<taxonomicName lsidName="N. minusculus" pageId="2" pageNumber="113" rank="species" species="minusculus">N. minusculus</taxonomicName>
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sp. n. Scale bars: 0.1 mm.
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</paragraph>
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</caption>
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<caption pageId="2" pageNumber="113">
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<paragraph pageId="2" pageNumber="113">
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Figure 3. Heads of
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Nornalup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nornalup afoveatus" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="afoveatus">Nornalup afoveatus</taxonomicName>
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sp. n. a dorsal view b ventral view.
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<taxonomicName lsidName="N. quadratus" pageId="2" pageNumber="113" rank="species" species="quadratus">N. quadratus</taxonomicName>
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sp. n. c prosternum, ventral view d meso- and metaventrite, ventral view. Scale bars: 0.1 mm.
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</paragraph>
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</caption>
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<caption pageId="2" pageNumber="113">
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<paragraph pageId="2" pageNumber="113">
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Figure 4. Female abdominal sternites IX, ventral view. a
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Nornalup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nornalup afoveatus" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="afoveatus">Nornalup afoveatus</taxonomicName>
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sp. n. b
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<taxonomicName lsidName="N. quadratus" pageId="2" pageNumber="113" rank="species" species="quadratus">N. quadratus</taxonomicName>
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sp. n. c
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<taxonomicName lsidName="N. minusculus" pageId="2" pageNumber="113" rank="species" species="minusculus">N. minusculus</taxonomicName>
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sp. n. Aedeagi of
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<taxonomicName lsidName="N. afoveatus" pageId="2" pageNumber="113" rank="species" species="afoveatus">N. afoveatus</taxonomicName>
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sp. n. d dorsal view e lateral view. Aedeagi of
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<taxonomicName lsidName="N. quadratus" pageId="2" pageNumber="113" rank="species" species="quadratus">N. quadratus</taxonomicName>
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sp. n. f dorsal view g lateral view. Aedeagi of
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<taxonomicName lsidName="N. minusculus" pageId="2" pageNumber="113" rank="species" species="minusculus">N. minusculus</taxonomicName>
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sp. n. h dorsal view i lateral view. Scale bars: 0.1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="114" type="etymology">
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<paragraph pageId="3" pageNumber="114">
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<pageBreakToken pageId="3" pageNumber="114" start="start">Etymology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="3" pageNumber="114">
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Nornalup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nornalup" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="genus">Nornalup</taxonomicName>
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gen. n. is named for
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Nornalup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nornalup" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="genus">Nornalup</taxonomicName>
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, one of the national parks where one of the species was collected.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="114" type="distribution">
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<paragraph pageId="3" pageNumber="114">Distribution.</paragraph>
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<paragraph pageId="3" pageNumber="114">Australia.</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="114" type="comments">
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<paragraph pageId="3" pageNumber="114">Comments about secondary sexual characters.</paragraph>
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<paragraph pageId="3" pageNumber="114">
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Male specimens possess tubercles on antennomeres 4-11, but females have the tubercles on antennomeres 8-11 (Fig. 2
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<normalizedToken originalValue="a–f">a-f</normalizedToken>
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). Males have slightly larger eyes (Fig. 2
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<normalizedToken originalValue="g–l">g-l</normalizedToken>
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). Male abdominal sternite IX is usually fragile and is partially concealed by sternite VIII, rendering it simple and reduced in appearance. Females possess a more robust, rectangular abdominal sternite IX bearing two pairs of long setae (Fig. 4
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<normalizedToken originalValue="a–c">a-c</normalizedToken>
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) that are usually visible in ventral view.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="114" type="comments">
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<paragraph pageId="3" pageNumber="114">Comments about biotic region.</paragraph>
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<paragraph pageId="3" pageNumber="114">
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Nornalup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nornalup" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="genus">Nornalup</taxonomicName>
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gen. n. is found at the very southwestern corner of Australia, which is known as a global biodiversity hotspot (Hopper and Gioia 2004). This region has a higher average annual rainfall (300-1200 mm) than the surrounding more internal deserts of the mainland, and is mostly covered by
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<taxonomicName genus="Eucalytus" lsidName="Eucalytus" pageId="3" pageNumber="114" rank="genus">Eucalytus</taxonomicName>
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forests (Hopper and Gioia 2004). Approximately 740 native vascular plants are known from this area, half of which are endemic (Hopper and Gioia 2004). All species are found in the karri (
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<taxonomicName lsidName="E. diversicolor" pageId="3" pageNumber="114" rank="species" species="diversicolor">E. diversicolor</taxonomicName>
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F.Muell.), tingle (
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<taxonomicName lsidName="E. jacksonii" pageId="3" pageNumber="114" rank="species" species="jacksonii">E. jacksonii</taxonomicName>
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Maiden), and jarrah (
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<taxonomicName lsidName="E. marginata" pageId="3" pageNumber="114" rank="species" species="marginata">E. marginata</taxonomicName>
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Donn ex Sm.) forests unique to this area, where the distributions of three species do not overlap (Fig. 5).
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</paragraph>
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<caption pageId="3" pageNumber="114">
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<paragraph pageId="3" pageNumber="114">
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Figure 5. Known collection localities of
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Nornalup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nornalup" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="genus">Nornalup</taxonomicName>
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gen. n.
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<taxonomicName lsidName="N. afoveatus" pageId="3" pageNumber="114" rank="species" species="afoveatus">N. afoveatus</taxonomicName>
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sp. n.: squares;
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<taxonomicName lsidName="N. quadratus" pageId="3" pageNumber="114" rank="species" species="quadratus">N. quadratus</taxonomicName>
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sp. n.: triangle;
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<taxonomicName lsidName="N. minusculus" pageId="3" pageNumber="114" rank="species" species="minusculus">N. minusculus</taxonomicName>
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sp. n.: circles.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="114" type="comments">
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<paragraph pageId="3" pageNumber="114">Comments about related taxa.</paragraph>
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<paragraph pageId="3" pageNumber="114">
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Based on thoracic foveal system,
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Nornalup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nornalup" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="genus">Nornalup</taxonomicName>
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gen. n. is closest to the genus
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sagola" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="genus">Sagola</taxonomicName>
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Sharp. However, the frontal sulcus (Fig. 3a), abdominal length of tergite and sternite VI (Fig. 1), and form of the male aedeagus are not shared with any species of
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sagola" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="genus">Sagola</taxonomicName>
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or other faronite genera. To understand the specific relationship with other faronites, phylogenetic analysis based on morphology and molecular data is needed.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="114" type="key">
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<paragraph pageId="3" pageNumber="114">
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Key to species of the genus
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Nornalup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nornalup" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="genus">Nornalup</taxonomicName>
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gen. n.
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</paragraph>
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<paragraph pageId="3" pageNumber="114">
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<table pageId="3" pageNumber="114">
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<tr pageId="3" pageNumber="114">
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<td colspan="1" pageId="3" pageNumber="114" rowspan="1">1b3d4b</td>
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<td colspan="1" pageId="3" pageNumber="114" rowspan="1">
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Nornalup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nornalup quadratus" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="quadratus">Nornalup quadratus</taxonomicName>
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sp. n.
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</td>
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</tr>
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<tr pageId="3" pageNumber="114">
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<td colspan="1" pageId="3" pageNumber="114" rowspan="1">1a, c4a, c</td>
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<td colspan="1" pageId="3" pageNumber="114" rowspan="1">2</td>
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</tr>
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<tr pageId="3" pageNumber="114">
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<td colspan="1" pageId="3" pageNumber="114" rowspan="1">1a4d4a</td>
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<td colspan="1" pageId="3" pageNumber="114" rowspan="1">
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<taxonomicName lsidName="N. afoveatus" pageId="3" pageNumber="114" rank="species" species="afoveatus">N. afoveatus</taxonomicName>
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sp. n.
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</td>
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</tr>
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<tr pageId="3" pageNumber="114">
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<td colspan="1" pageId="3" pageNumber="114" rowspan="1">1c4h4c</td>
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<td colspan="1" pageId="3" pageNumber="114" rowspan="1">
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<taxonomicName lsidName="N. minusculus" pageId="3" pageNumber="114" rank="species" species="minusculus">N. minusculus</taxonomicName>
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sp. n.
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</td>
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</tr>
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</table>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |