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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1175.107636" ID-Pensoft-Pub="1313-2970-1175-163" ID-Pensoft-UUID="CE03B1B394CF56B48E8F7E5026CD639C" ID-ZooBank="E88FF072E8C945AEA4944B2A5F7DD7E8" ModsDocID="1313-2970-1175-163" checkinTime="1692265381537" checkinUser="pensoft" docAuthor="Yin, Wenqi, Zhu, Pingzhou &amp; Shi, Hongliang" docDate="2023" docId="346E35B1437A5730BA307B66F4002F56" docLanguage="en" docName="ZooKeys 1175: 163-185" docOrigin="ZooKeys 1175" docPubDate="2023-08-16" docSource="http://dx.doi.org/10.3897/zookeys.1175.107636" docTitle="Pterostichus (Orientostichus) liyuani Yin &amp; Zhu &amp; Shi 2023, sp. nov." docType="treatment" docUuid="48C49F07-DDA3-418B-A284-EAA52596104F" docUuidSource="ZooBank" docVersion="1" id="CE03B1B394CF56B48E8F7E5026CD639C" lastPageNumber="163" masterDocId="CE03B1B394CF56B48E8F7E5026CD639C" masterDocTitle="Taxonomic study on the subgenus Orientostichus, the Pterostichus pulcher species group (Coleoptera, Carabidae, Pterostichus)" masterLastPageNumber="185" masterPageNumber="163" pageNumber="163" updateTime="1692265381537" updateUser="pensoft">
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<mods:title id="7466A89D89BE41F67D8EE13F7A2CD8FA">Taxonomic study on the subgenus Orientostichus, the Pterostichus pulcher species group (Coleoptera, Carabidae, Pterostichus)</mods:title>
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<mods:namePart id="AC2B488C680C7E3836E5B01B856F4053">Yin, Wenqi</mods:namePart>
<mods:nameIdentifier id="733DEC2B3DB6BE71B29E569153114474" type="ORCID">https://orcid.org/0000-0003-2903-1399</mods:nameIdentifier>
<mods:affiliation id="AE1C42F700F9EC98EEA0AA733B6D90C0">College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
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<mods:namePart id="C699284E38A98AB916B8081E657AE504">Zhu, Pingzhou</mods:namePart>
<mods:nameIdentifier id="74632720B21278E0CF91B65BB44274AD" type="ORCID">https://orcid.org/0000-0003-2139-6764</mods:nameIdentifier>
<mods:affiliation id="602325D2880DD9E697CECBB2E878AEA6">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing 100093, China</mods:affiliation>
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<mods:namePart id="1C032B1919F05E3E9118231E377C5BFB">Shi, Hongliang</mods:namePart>
<mods:nameIdentifier id="DC222C1B6DFEB8465D58BDE3DE29A87C" type="ORCID">https://orcid.org/0000-0002-9989-5830</mods:nameIdentifier>
<mods:affiliation id="1918FC2ADC41BBDBD4F3A21E62FD236A">College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
<mods:nameIdentifier id="C4DF4DF9D1DD35DE7A70F40A658C583D" type="email">shihl@bjfu.edu.cn</mods:nameIdentifier>
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<mods:date id="A2176ED46FFFB9BE5C6D4B76934653CC">2023</mods:date>
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<mods:number id="F559A2C41CD7C75A5E2D7DDD36448FB2">2023-08-16</mods:number>
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<treatment id="346E35B1437A5730BA307B66F4002F56" LSID="urn:lsid:zoobank.org:act:48C49F07-DDA3-418B-A284-EAA52596104F" httpUri="http://treatment.plazi.org/id/346E35B1437A5730BA307B66F4002F56" lastPageNumber="163" pageId="0" pageNumber="163">
<subSubSection id="35102BA9A08413ACF2FE4983F627F47A" pageId="0" pageNumber="163" type="nomenclature">
<paragraph id="058DFA64F22D60A73F073CC6B582F5AD" pageId="0" pageNumber="163">
<taxonomicName id="99EF2BEDA72ABC6BE37C29225E859F23" LSID="https://zoobank.org/48C49F07-DDA3-418B-A284-EAA52596104F" authority="Yin &amp; Zhu &amp; Shi, 2023" authorityName="Yin &amp; Zhu &amp; Shi" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) liyuani" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="liyuani" status="sp. nov." subGenus="Orientostichus">Pterostichus (Orientostichus) liyuani</taxonomicName>
<taxonomicNameLabel id="6DAD1B42984EF9B7CF77562959ED8C27" pageId="0" pageNumber="163">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="A27CB28A747D57E99795CE06F63D0BC9" pageId="0" pageNumber="163" type="description">
<paragraph id="A1F9C68E4BF3CAEB25238270D817F301" pageId="0" pageNumber="163">
<figureCitation id="9A3F79744D5712321C4B0B69B2D4C2B4" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus and labels of Pterostichus pulcher species group 1 P. pemphis sp. nov., holotype 2 P. orbicollis sp. nov., holotype 3 P. leo sp. nov., holotype 4 P. liyuani sp. nov., holotype 5 P. condylus sp. nov., holotype 6 P. jialini sp. nov., holotype. Scale bar: 5 mm." figureDoi="10.3897/zookeys.1175.107636.figures1-6" httpUri="https://binary.pensoft.net/fig/893313" pageId="0" pageNumber="163">Figs 4</figureCitation>
<figureCitation id="0B05B66F22FF003A9E615553B9DB8119" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Pronota of Pterostichus pulcher species group 7 P. pemphis sp. nov., holotype 8 P. orbicollis sp. nov., holotype 9 P. leo sp. nov., holotype 10 P. liyuani sp. nov., holotype 11 P. condylus sp. nov., holotype 12 P. jialini sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures7-12" httpUri="https://binary.pensoft.net/fig/893314" pageId="0" pageNumber="163">, 10</figureCitation>
<figureCitation id="BBFEC9A8E3DDE1C8C1596BB734BF793A" captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus pulcher species group, left lateral view and dorsal view of median lobe of aedeagus 13 P. pemphis sp. nov., holotype 14 P. orbicollis sp. nov., holotype 15 P. leo sp. nov., holotype 16 P. liyuani sp. nov., holotype 17 P. condylus sp. nov., holotype 18 P. jialini sp. nov., holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures13-18" httpUri="https://binary.pensoft.net/fig/893315" pageId="0" pageNumber="163">, 16</figureCitation>
<figureCitation id="462FD9D3444049D3D01B45C625097CB4" captionStart="Figures 19–24" captionStartId="F4" captionText="Figures 19 - 24. Endophallus of Pterostichus pulcher species group, left lateral view and right lateral view 19 P. pemphis sp. nov., paratype from Zhaojue county 20 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 21 P. leo sp. nov., paratype from Shizishan mountain, Jinyang county 22 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 23 P. condylus sp. nov., paratype from Mianning county 24 P. pulcher Sciaky &amp; Allegro, paratype from Yizi Yakou, E'bian county to Meigu county. Scale bar: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures19-24" httpUri="https://binary.pensoft.net/fig/893316" pageId="0" pageNumber="163">, 22</figureCitation>
<figureCitation id="BF3EEBA9B18F7A829FACC4DD66EBD557" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. 25 - 28 endophallus bsp of Pterostichus pulcher species group, left lateral view 25 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 26 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 27 P. condylus sp. nov., paratype from Mianning county 28 P. pulcher Sciaky &amp; Allegro, paratype from Yizi Yakou, E'bian county to Meigu county 29, 30 Endophallus of Pterostichus (Orientostichus) spp., left lateral view and right lateral view 29 P. prattii Bates, 1890, a male from Sanming city, Fujian province 30 P. curtatus Fairmaire, a male from Heqing county, Yunnan province. Scale bars: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures25-30" httpUri="https://binary.pensoft.net/fig/893317" pageId="0" pageNumber="163">, 26</figureCitation>
<figureCitation id="E7FF8D830BB2FF3B5A35799639EEE7B9" captionStart="Figures 34–39" captionStartId="F7" captionText="Figures 34 - 39. Male sternum VII of Pterostichus pulcher species group spp. 34 P. pemphis sp. nov., holotype 35 P. orbicollis sp. nov., holotype 36 P. liyuani sp. nov., holotype 37 P. condylus sp. nov., holotype 38 P. jialini sp. nov., holotype 39 P. pulcher Sciaky &amp; Allegro, holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures34-39" httpUri="https://binary.pensoft.net/fig/893319" pageId="0" pageNumber="163">
, 36 Chinese vernacular name:
<normalizedToken id="25FA847E6B9328D615B2AEA15D6EA4D0" originalValue="李圆通缘步甲">李圆通缘步甲</normalizedToken>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="43B94AB2ACC1864CBBE806B91D488635" pageId="0" pageNumber="163" type="type locality">
<paragraph id="70412C728C1E77D0328DA41A26C55B16" pageId="0" pageNumber="163">Type locality.</paragraph>
<paragraph id="3258A89BF78D0BC4DFBFC19E3C4E1904" pageId="0" pageNumber="163">
China, Sichuan province: Liangshan Yi Autonomous Prefecture, Puge county, Luojishan mountain (
<geoCoordinate id="541AC1CCD70767C29C7E512766F12C22" degrees="27.33" direction="north" orientation="latitude" precision="555" value="27.33">27.33N</geoCoordinate>
,
<geoCoordinate id="2D7C78ED594698E618233A8F08FA9953" degrees="102.44" direction="east" orientation="longitude" precision="555" value="102.44">102.44E</geoCoordinate>
, alt 3500 m).
</paragraph>
</subSubSection>
<subSubSection id="A5793C5DC4944F13C119CA645BA5A5BB" pageId="0" pageNumber="163" type="type material">
<paragraph id="49B822E735946B0749E418F768993360" pageId="0" pageNumber="163">Type material.</paragraph>
<paragraph id="A8A7C500224AD7C983EF3513A4A36D1E" pageId="0" pageNumber="163">
<emphasis id="70F87A0EF30C3B9D87E55F3046A39F83" bold="true" italics="true" pageId="0" pageNumber="163">Holotype</emphasis>
: ♂: &quot;China, Sichuan province, Liangshan autonomous prefecture, Puge county, Luojishan mountain&quot;, &quot;N27.3355, E102.4461, 3500 m, pitfall trap by Li Yuan, 2022.VII.26&quot;, &quot;HOLOTYPE ♂
<taxonomicName id="A459549B535E05478A8C24C33498793A" authorityName="Yin &amp; Zhu &amp; Shi" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) liyuani" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="liyuani" subGenus="Orientostichus">Pterostichus (Orientostichus) liyuani</taxonomicName>
sp. nov. det. Yin &amp; Shi, 2022&quot; [red label];
<emphasis id="E415160501F4B0F42874B695DFD78491" bold="true" italics="true" pageId="0" pageNumber="163">Paratypes</emphasis>
(a total of 64 ♂ and 72 ♀): 18 ♂ and 15 ♀ (CHYL): the same data as holotype but labeled as paratype; 8 ♂ and 7 ♀: &quot;China: Sichuan, Liangshan Dist., Puge county, Luojishan, tourist path, mixed forest, 2805 m, N27.58253 E102.39497&quot;, &quot;by pitfall trap, 2012.VI.10, Huang Hao lgt., Institute of Zoology, CAS&quot;, &quot;PARATYPE of
<taxonomicName id="E741910BDAB7D86A32D27B97E9CD0987" authorityName="Yin &amp; Zhu &amp; Shi" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) liyuani" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="liyuani" subGenus="Orientostichus">Pterostichus (Orientostichus) liyuani</taxonomicName>
sp. nov. det. Yin &amp; Shi, 2022&quot; [red label]; 6 ♂ and 29 ♀: &quot;China: Sichuan prov., Puge County, Shuihaizi wind power station, meadow + rhodo + fir. 3515 m, N27.3355, E102.4461&quot;, &quot;2018.VII.21, pitfall trap. Shi HL, Yan WF, Zhu PZ &amp; Jiang ZY lgt., IZAS &amp; BJFU exp. 2018&quot;, &quot;PARATYPE of
<taxonomicName id="A16621C57937397549ED7DDCCA36FF66" authorityName="Yin &amp; Zhu &amp; Shi" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) liyuani" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="liyuani" subGenus="Orientostichus">Pterostichus (Orientostichus) liyuani</taxonomicName>
sp. nov. det. Yin &amp; Shi, 2022&quot; [red label]; 1 ♂: &quot;China: Sichuan prov, Zhaojue county, Mufoshan mountain. peak. Rhododendron shrub + alpine meadow.&quot;, &quot;N28.0685, E102.7870, 3485 m. pitfall trap. 2015.VII.27. Shi HL, Liu B &amp; Ma YL lgt.&quot;, &quot;PARATYPE of
<taxonomicName id="D121E00A177622EAD6D81FFB7D4AED3C" authorityName="Yin &amp; Zhu &amp; Shi" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) liyuani" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="liyuani" subGenus="Orientostichus">Pterostichus (Orientostichus) liyuani</taxonomicName>
sp. nov. det. Yin &amp; Shi, 2022&quot; [red label]; 26 ♂ and 18 ♀: &quot;China: Sichuan, Liangshan Dist., Puge county, Luojishan, near No. 1 glacial groove, mixed forest, 3470 m, N27.58445, E103.7913&quot;, &quot;by pitfall trap, 2012.VI.10, Huang Hao lgt., Institute of Zoology, CAS&quot;, &quot;PARATYPE of
<taxonomicName id="1BC0153AAEE4C23FC00253B5BFE42550" authorityName="Yin &amp; Zhu &amp; Shi" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) liyuani" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="liyuani" subGenus="Orientostichus">Pterostichus (Orientostichus) liyuani</taxonomicName>
sp. nov. det. Yin &amp; Shi, 2022&quot; [red label]; 5 ♂ and 3 ♀: &quot;China, Sichuan, Puge, Luojishan Mt., near ropeway upper station, 3595 m, N27.58108, E102.38012&quot;, &quot;2012.VI.10, pitfall trap, under rhododendra forest, Shi Hongliang &amp; Liu Ye leg. Institute of Zoology, CAS.&quot;, &quot;PARATYPE of
<taxonomicName id="E5FAD8B832B47DA74F6301092539CF60" authorityName="Yin &amp; Zhu &amp; Shi" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) liyuani" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="liyuani" subGenus="Orientostichus">Pterostichus (Orientostichus) liyuani</taxonomicName>
sp. nov. det. Yin &amp; Shi, 2022&quot; [red label].
</paragraph>
</subSubSection>
<subSubSection id="9CD357909BCDA97AE00BE5F3A8714338" pageId="0" pageNumber="163" type="diagnosis">
<paragraph id="6E8DFFD09A700E037F6961238B04DAFA" pageId="0" pageNumber="163">Diagnosis.</paragraph>
<paragraph id="5417761A7B92E5B1C6EE706B833B9F07" pageId="0" pageNumber="163">
Elytral intervals 3 and 5 each with ≥ 3 large foveate discal pores, interval 7 without discal pore. Antennomere 3 without accessory setae. Pronotum subcordate, with lateral margins nearly straight before posterior angles (Fig.
<figureCitation id="3D4E72CCA56FF33A0902111EC60B1B07" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Pronota of Pterostichus pulcher species group 7 P. pemphis sp. nov., holotype 8 P. orbicollis sp. nov., holotype 9 P. leo sp. nov., holotype 10 P. liyuani sp. nov., holotype 11 P. condylus sp. nov., holotype 12 P. jialini sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures7-12" httpUri="https://binary.pensoft.net/fig/893314" pageId="0" pageNumber="163">10</figureCitation>
); lateral margins with 2-4 mid-lateral setae; pronotal disc with shallow but distinct transverse wrinkles. Male sternite VII with a very small smooth tubercle (Fig.
<figureCitation id="BB8A951779CDDEB118AB7F2F133D143B" captionStart="Figures 34–39" captionStartId="F7" captionText="Figures 34 - 39. Male sternum VII of Pterostichus pulcher species group spp. 34 P. pemphis sp. nov., holotype 35 P. orbicollis sp. nov., holotype 36 P. liyuani sp. nov., holotype 37 P. condylus sp. nov., holotype 38 P. jialini sp. nov., holotype 39 P. pulcher Sciaky &amp; Allegro, holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures34-39" httpUri="https://binary.pensoft.net/fig/893319" pageId="0" pageNumber="163">36</figureCitation>
). Apical lamella of aedeagus short, apex rounded, LL/LW = 1.9-2.1 (Fig.
<figureCitation id="FFE7A48821E2C4820CFF834014CE7CF7" captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus pulcher species group, left lateral view and dorsal view of median lobe of aedeagus 13 P. pemphis sp. nov., holotype 14 P. orbicollis sp. nov., holotype 15 P. leo sp. nov., holotype 16 P. liyuani sp. nov., holotype 17 P. condylus sp. nov., holotype 18 P. jialini sp. nov., holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures13-18" httpUri="https://binary.pensoft.net/fig/893315" pageId="0" pageNumber="163">16</figureCitation>
); endophallus with spiral-shaped bsp (Figs
<figureCitation id="C9E8008417B7E289FFC5B5EEEBDA1D21" captionStart="Figures 19–24" captionStartId="F4" captionText="Figures 19 - 24. Endophallus of Pterostichus pulcher species group, left lateral view and right lateral view 19 P. pemphis sp. nov., paratype from Zhaojue county 20 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 21 P. leo sp. nov., paratype from Shizishan mountain, Jinyang county 22 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 23 P. condylus sp. nov., paratype from Mianning county 24 P. pulcher Sciaky &amp; Allegro, paratype from Yizi Yakou, E'bian county to Meigu county. Scale bar: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures19-24" httpUri="https://binary.pensoft.net/fig/893316" pageId="0" pageNumber="163">22</figureCitation>
,
<figureCitation id="1BFE57D0198C6F1A49591291A54688C4" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. 25 - 28 endophallus bsp of Pterostichus pulcher species group, left lateral view 25 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 26 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 27 P. condylus sp. nov., paratype from Mianning county 28 P. pulcher Sciaky &amp; Allegro, paratype from Yizi Yakou, E'bian county to Meigu county 29, 30 Endophallus of Pterostichus (Orientostichus) spp., left lateral view and right lateral view 29 P. prattii Bates, 1890, a male from Sanming city, Fujian province 30 P. curtatus Fairmaire, a male from Heqing county, Yunnan province. Scale bars: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures25-30" httpUri="https://binary.pensoft.net/fig/893317" pageId="0" pageNumber="163">26</figureCitation>
).
</paragraph>
<caption id="C51AD351F4C98DED390A93EC0BB25C01" doi="10.3897/zookeys.1175.107636.figures34-39" httpUri="https://binary.pensoft.net/fig/893319" pageId="0" pageNumber="163" start="Figures 34–39" startId="F7">
<paragraph id="0E82249088FB724F332079FAB9774E00" pageId="0" pageNumber="163">
<emphasis id="5DAFEE10B1ABFCAAB9F7D45316FF84AC" bold="true" pageId="0" pageNumber="163">Figures 34-39.</emphasis>
Male sternum VII of
<taxonomicName id="B32B5BC9CD488DDBD0FB2ED709B761DC" authorityName="Sciaky &amp; Allegro" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus pulcher" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="AB651E55F36EB5BA6600477E17A9A335" italics="true" pageId="0" pageNumber="163">Pterostichus pulcher</emphasis>
</taxonomicName>
species group spp.
<emphasis id="004FF249F6E91495348C7A02AC53AF14" bold="true" pageId="0" pageNumber="163">34</emphasis>
<taxonomicName id="C7DE38FB88B9561167D2F84E7BEB9925" lsidName="P. pemphis" pageId="0" pageNumber="163" rank="species" species="pemphis">
<emphasis id="54ED7EC0B24468CDDD0CA54D72F14576" italics="true" pageId="0" pageNumber="163">P. pemphis</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis id="27116445B01B22FFF9EBA0AC1F0B425A" bold="true" pageId="0" pageNumber="163">35</emphasis>
<taxonomicName id="0DA9066449C237753A0CF750996B2E07" lsidName="P. orbicollis" pageId="0" pageNumber="163" rank="species" species="orbicollis">
<emphasis id="E9557932125F86EA62A9999EFBC24192" italics="true" pageId="0" pageNumber="163">P. orbicollis</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis id="FDC21CE932285D5584BEB3B9526AEC56" bold="true" pageId="0" pageNumber="163">36</emphasis>
<taxonomicName id="C16FA912F4E4BCC75D1AF9890A615B6B" lsidName="P. liyuani" pageId="0" pageNumber="163" rank="species" species="liyuani">
<emphasis id="B150C9C47CAD0B6A66A1F728710D4C9C" italics="true" pageId="0" pageNumber="163">P. liyuani</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis id="C909E4CDAC4BB8CF7DCABE29A04EF435" bold="true" pageId="0" pageNumber="163">37</emphasis>
<taxonomicName id="E3165FE2C204F0EE517559F6E692FD9D" lsidName="P. condylus" pageId="0" pageNumber="163" rank="species" species="condylus">
<emphasis id="F71A151C8FB97EFE5008DD286F602715" italics="true" pageId="0" pageNumber="163">P. condylus</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis id="A4DA0AA1B32D43081C8729520C52CF42" bold="true" pageId="0" pageNumber="163">38</emphasis>
<taxonomicName id="0E9972D262BAA162E0C5CC73338A994E" lsidName="P. jialini" pageId="0" pageNumber="163" rank="species" species="jialini">
<emphasis id="9844264A773DA4275C1B6C36F754E5F5" italics="true" pageId="0" pageNumber="163">P. jialini</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis id="4D7B13C0476A576314310927D1D508AE" bold="true" pageId="0" pageNumber="163">39</emphasis>
<taxonomicName id="4945363786D389A6D97C66B479B4C21E" lsidName="P. pulcher" pageId="0" pageNumber="163" rank="species" species="pulcher">
<emphasis id="B0397181D8EA777A8167EC6DFB5333D7" italics="true" pageId="0" pageNumber="163">P. pulcher</emphasis>
</taxonomicName>
Sciaky &amp; Allegro, holotype. Scale bar: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4EA116F340E882C841134B9B1B719C96" pageId="0" pageNumber="163" type="comparisons">
<paragraph id="C968B0FA3C023C76814A13C84370C5BE" pageId="0" pageNumber="163">Comparison.</paragraph>
<paragraph id="4B5AE43DE901AFDC501CD6C8077C5F60" pageId="0" pageNumber="163">
From the external features,
<taxonomicName id="2C0B60ABF7979D860351AC560B21F1FB" lsidName="P. liyuani" pageId="0" pageNumber="163" rank="species" species="liyuani">
<emphasis id="AC66B725E54B11A320C0ABDBCE4559AC" italics="true" pageId="0" pageNumber="163">P. liyuani</emphasis>
</taxonomicName>
sp. nov. is most similar to another new species
<taxonomicName id="BEE65D204D488AA8726CFC67EA510778" lsidName="P. condylus" pageId="0" pageNumber="163" rank="species" species="condylus">
<emphasis id="C967497D9AE7DEBF7DB195739ACEF5ED" italics="true" pageId="0" pageNumber="163">P. condylus</emphasis>
</taxonomicName>
sp. nov. These two species can be easily distinguished from
<taxonomicName id="03FDE399DCF7358B0A3B690AD27CAE53" lsidName="P. pemphis" pageId="0" pageNumber="163" rank="species" species="pemphis">
<emphasis id="CDDF8DF9DA3D1DF1531603EBA08288AB" italics="true" pageId="0" pageNumber="163">P. pemphis</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName id="2DA6E5170888B2A94FEE9BEE9C86CF76" lsidName="P. leo" pageId="0" pageNumber="163" rank="species" species="leo">
<emphasis id="D92FF267D73BCD7EDD0A0A11F9E4D6BF" italics="true" pageId="0" pageNumber="163">P. leo</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="2FE6D63019FB814CB6D5A1399B8FD642" lsidName="P. orbicollis" pageId="0" pageNumber="163" rank="species" species="orbicollis">
<emphasis id="09A8BC5E3328B8F5295A54CF6B0E8628" italics="true" pageId="0" pageNumber="163">P. orbicollis</emphasis>
</taxonomicName>
sp. nov. by antennomere 3 without accessory setae and quite different pronotal shape, and from
<taxonomicName id="A9A2B1F689AD8E3EB22CF689CCD5EC62" lsidName="P. pulcher" pageId="0" pageNumber="163" rank="species" species="pulcher">
<emphasis id="527259DDDD9DDFA02217A3EE184439A7" italics="true" pageId="0" pageNumber="163">P. pulcher</emphasis>
</taxonomicName>
Sciaky &amp; Allegro and
<taxonomicName id="AAC315D7E98C777B73E642F8FC8B869B" lsidName="P. jialini" pageId="0" pageNumber="163" rank="species" species="jialini">
<emphasis id="8E7DAAEFFA7FFBF9A51DCBDC918EDD48" italics="true" pageId="0" pageNumber="163">P. jialini</emphasis>
</taxonomicName>
sp. nov. by interval 7 without discal pores.
</paragraph>
<paragraph id="5064C44F4CFCAC1B289CC068B359A3F9" pageId="0" pageNumber="163">
Although these two new species are very similar to each other,
<taxonomicName id="01B1AABE144491D24BFC8BACE6D46127" lsidName="P. liyuani" pageId="0" pageNumber="163" rank="species" species="liyuani">
<emphasis id="D761F67FF55390E5D874381A5FAFC29F" italics="true" pageId="0" pageNumber="163">P. liyuani</emphasis>
</taxonomicName>
sp. nov. is distinguishable from
<taxonomicName id="F63B8F909CB5F9BBCA825D2C71EA0982" lsidName="P. condylus" pageId="0" pageNumber="163" rank="species" species="condylus">
<emphasis id="67F16DB947C529F4FF901C0422EB15E0" italics="true" pageId="0" pageNumber="163">P. condylus</emphasis>
</taxonomicName>
sp. nov. by: (1) the sexual modification on male sternite VII is quite small and inconspicuous of
<taxonomicName id="A21B8F92212E203B5F06F775301F3B07" lsidName="P. liyuani" pageId="0" pageNumber="163" rank="species" species="liyuani">
<emphasis id="866A215534E7946912077C496E6ACD60" italics="true" pageId="0" pageNumber="163">P. liyuani</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="8873C30349386C01B4878A005EB5DE85" captionStart="Figures 34–39" captionStartId="F7" captionText="Figures 34 - 39. Male sternum VII of Pterostichus pulcher species group spp. 34 P. pemphis sp. nov., holotype 35 P. orbicollis sp. nov., holotype 36 P. liyuani sp. nov., holotype 37 P. condylus sp. nov., holotype 38 P. jialini sp. nov., holotype 39 P. pulcher Sciaky &amp; Allegro, holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures34-39" httpUri="https://binary.pensoft.net/fig/893319" pageId="0" pageNumber="163">36</figureCitation>
), while the latter species has larger and more well-defined tubercle on sternite VII (Fig.
<figureCitation id="7FCFF2899BF4460E4F4F9DDE54193C06" captionStart="Figures 34–39" captionStartId="F7" captionText="Figures 34 - 39. Male sternum VII of Pterostichus pulcher species group spp. 34 P. pemphis sp. nov., holotype 35 P. orbicollis sp. nov., holotype 36 P. liyuani sp. nov., holotype 37 P. condylus sp. nov., holotype 38 P. jialini sp. nov., holotype 39 P. pulcher Sciaky &amp; Allegro, holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures34-39" httpUri="https://binary.pensoft.net/fig/893319" pageId="0" pageNumber="163">37</figureCitation>
); (2) the pronotal disc is more evidently wrinkled in
<taxonomicName id="A2A61672BFDF58478CE936197E9B5485" lsidName="P. liyuani" pageId="0" pageNumber="163" rank="species" species="liyuani">
<emphasis id="F87D9095C144E5FDA6F4D8478DF54C69" italics="true" pageId="0" pageNumber="163">P. liyuani</emphasis>
</taxonomicName>
sp. nov., but the latter species at most has very faint wrinkles; (3) the pronotal lateral margin is hardly sinuate in
<taxonomicName id="95AE4A340B109AE436F41C9E865AEB0A" lsidName="P. liyuani" pageId="0" pageNumber="163" rank="species" species="liyuani">
<emphasis id="AB46E554ADFC9CE192E31C29CAFB6206" italics="true" pageId="0" pageNumber="163">P. liyuani</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="8C3868BC8D89066C6E4C43D33C8A623E" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Pronota of Pterostichus pulcher species group 7 P. pemphis sp. nov., holotype 8 P. orbicollis sp. nov., holotype 9 P. leo sp. nov., holotype 10 P. liyuani sp. nov., holotype 11 P. condylus sp. nov., holotype 12 P. jialini sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures7-12" httpUri="https://binary.pensoft.net/fig/893314" pageId="0" pageNumber="163">10</figureCitation>
), but evidently sinuate in the latter species (Fig.
<figureCitation id="10A536480C69E8F989C3E2429CA83C8A" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Pronota of Pterostichus pulcher species group 7 P. pemphis sp. nov., holotype 8 P. orbicollis sp. nov., holotype 9 P. leo sp. nov., holotype 10 P. liyuani sp. nov., holotype 11 P. condylus sp. nov., holotype 12 P. jialini sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures7-12" httpUri="https://binary.pensoft.net/fig/893314" pageId="0" pageNumber="163">11</figureCitation>
); (4) comparing to
<taxonomicName id="1ECCD92B56DCC597FA5A595647254107" lsidName="P. liyuani" pageId="0" pageNumber="163" rank="species" species="liyuani">
<emphasis id="BD0C195EB943EA3503734B7DB0F79CCD" italics="true" pageId="0" pageNumber="163">P. liyuani</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="5B284608CDE058988BAF4C5006C716ED" captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus pulcher species group, left lateral view and dorsal view of median lobe of aedeagus 13 P. pemphis sp. nov., holotype 14 P. orbicollis sp. nov., holotype 15 P. leo sp. nov., holotype 16 P. liyuani sp. nov., holotype 17 P. condylus sp. nov., holotype 18 P. jialini sp. nov., holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures13-18" httpUri="https://binary.pensoft.net/fig/893315" pageId="0" pageNumber="163">16</figureCitation>
), in
<taxonomicName id="F9F1FD916123ED403437C6E32018578B" lsidName="P. condylus" pageId="0" pageNumber="163" rank="species" species="condylus">
<emphasis id="2D4987A7C3A2CF5B76CCAEF21367C955" italics="true" pageId="0" pageNumber="163">P. condylus</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="91D9E61B08CCB81BE432A9208C164FCB" captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus pulcher species group, left lateral view and dorsal view of median lobe of aedeagus 13 P. pemphis sp. nov., holotype 14 P. orbicollis sp. nov., holotype 15 P. leo sp. nov., holotype 16 P. liyuani sp. nov., holotype 17 P. condylus sp. nov., holotype 18 P. jialini sp. nov., holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures13-18" httpUri="https://binary.pensoft.net/fig/893315" pageId="0" pageNumber="163">17</figureCitation>
) the apical lamella of male genitalia is a little narrower, slightly narrowed near base and more truncated at apex; (5) the endophallus of
<taxonomicName id="208C74AEDB6050E8479450917D4A3D0B" lsidName="P. liyuani" pageId="0" pageNumber="163" rank="species" species="liyuani">
<emphasis id="15C85F41871B7413D696784A52068599" italics="true" pageId="0" pageNumber="163">P. liyuani</emphasis>
</taxonomicName>
sp. nov. has spiral shaped bsp with apex of right branch extended and bent reversely (Figs
<figureCitation id="EE3FDBE25E327C0A79BA7D7F46129DDA" captionStart="Figures 19–24" captionStartId="F4" captionText="Figures 19 - 24. Endophallus of Pterostichus pulcher species group, left lateral view and right lateral view 19 P. pemphis sp. nov., paratype from Zhaojue county 20 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 21 P. leo sp. nov., paratype from Shizishan mountain, Jinyang county 22 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 23 P. condylus sp. nov., paratype from Mianning county 24 P. pulcher Sciaky &amp; Allegro, paratype from Yizi Yakou, E'bian county to Meigu county. Scale bar: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures19-24" httpUri="https://binary.pensoft.net/fig/893316" pageId="0" pageNumber="163">22</figureCitation>
,
<figureCitation id="B240768377B0E479D4A28CA4B98E2511" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. 25 - 28 endophallus bsp of Pterostichus pulcher species group, left lateral view 25 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 26 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 27 P. condylus sp. nov., paratype from Mianning county 28 P. pulcher Sciaky &amp; Allegro, paratype from Yizi Yakou, E'bian county to Meigu county 29, 30 Endophallus of Pterostichus (Orientostichus) spp., left lateral view and right lateral view 29 P. prattii Bates, 1890, a male from Sanming city, Fujian province 30 P. curtatus Fairmaire, a male from Heqing county, Yunnan province. Scale bars: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures25-30" httpUri="https://binary.pensoft.net/fig/893317" pageId="0" pageNumber="163">26</figureCitation>
) which is unique among this species group, while the latter species has a U-shaped bsp (Figs
<figureCitation id="0D8B4E5AB665096F1F58265E2CF50205" captionStart="Figures 19–24" captionStartId="F4" captionText="Figures 19 - 24. Endophallus of Pterostichus pulcher species group, left lateral view and right lateral view 19 P. pemphis sp. nov., paratype from Zhaojue county 20 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 21 P. leo sp. nov., paratype from Shizishan mountain, Jinyang county 22 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 23 P. condylus sp. nov., paratype from Mianning county 24 P. pulcher Sciaky &amp; Allegro, paratype from Yizi Yakou, E'bian county to Meigu county. Scale bar: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures19-24" httpUri="https://binary.pensoft.net/fig/893316" pageId="0" pageNumber="163">23</figureCitation>
,
<figureCitation id="01AB4947FF797B02D366ABF320FBE3FE" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. 25 - 28 endophallus bsp of Pterostichus pulcher species group, left lateral view 25 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 26 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 27 P. condylus sp. nov., paratype from Mianning county 28 P. pulcher Sciaky &amp; Allegro, paratype from Yizi Yakou, E'bian county to Meigu county 29, 30 Endophallus of Pterostichus (Orientostichus) spp., left lateral view and right lateral view 29 P. prattii Bates, 1890, a male from Sanming city, Fujian province 30 P. curtatus Fairmaire, a male from Heqing county, Yunnan province. Scale bars: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures25-30" httpUri="https://binary.pensoft.net/fig/893317" pageId="0" pageNumber="163">27</figureCitation>
).
</paragraph>
<caption id="3DB2CB59CD17BE99634FDCC110B28857" doi="10.3897/zookeys.1175.107636.figures40-41" httpUri="https://binary.pensoft.net/fig/893320" pageId="0" pageNumber="163" start="Figures 40, 41" startId="F8">
<paragraph id="78C0F109EF8E5CF09B632419264E97B0" pageId="0" pageNumber="163">
<emphasis id="633421F7B8BD617FAA669E7C932117AF" bold="true" pageId="0" pageNumber="163">Figures 40, 41.</emphasis>
Female genitalia of
<taxonomicName id="1CB44F8E0C145308E22BB285BED58A8C" authorityName="Yin &amp; Zhu &amp; Shi" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) pemphis" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="pemphis" subGenus="Orientostichus">Pterostichus (Orientostichus) pemphis</taxonomicName>
sp. nov., paratype from Shuihaizi, Puge county
<emphasis id="A532062E95338EA91CEA670A659A140D" bold="true" pageId="0" pageNumber="163">40</emphasis>
ventral view and inner lateral view of ovipositor.
<emphasis id="67FA4DF9D34FECCE2DC4C224855D6C9C" bold="true" pageId="0" pageNumber="163">41</emphasis>
. Female reproductive system. Scale bars: 0.5 mm (
<emphasis id="478FA0D532D31454F8C015C4EA2A22B1" bold="true" pageId="0" pageNumber="163">40</emphasis>
); 1 mm (
<emphasis id="B045FFADD2693A076EC7CFAA3688A34B" bold="true" pageId="0" pageNumber="163">41</emphasis>
). Abbreviations: g1: gonocoxite 1, g2: gonocoxite 2, es: ensiform setae, ns, nematiform setae, bc: bursa copulatrix, co: common oviduct, sc: seminal canal, rc: receptaculum, sg: spermathecal gland, spc: spermathecal canal.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="88EF61E9E2053E3867811434F6155825" pageId="0" pageNumber="163" type="description">
<paragraph id="9BF2F4CD76D093FC0E5EA88E801EED6B" pageId="0" pageNumber="163">Description.</paragraph>
<paragraph id="D4AA9C45D0621C5101822159AC5C2415" pageId="0" pageNumber="163">
BL = 14.7-18.5 mm, BW = 5.3-6.5 mm, dorsal surface and appendages black, elytra often a little reddish brown. Antennomere 3 without accessory setae, only with primary setae forming apical ring. Pronotum subcordate, PW/PL = 1.24-1.28, widest near anterior 1/3; anterior margin wider than posterior margin; slightly narrowed to base (PBW/PW = 0.71-0.75); lateral margins evenly arched from anterior angles to middle, hardly sinuate before posterior angles; posterior angles nearly rectangular, apex not dentate; lateral margins with 2-4 mid-lateral setae near maximum width; basal foveae often with slightly transverse wrinkles, inner and outer grooves straight, partly fused, outer groove slightly longer than 1/2 length of inner groove, area between them slightly depressed; disc with shallow but distinct transverse wrinkles aside median line. Elytra oblong; parascutellar pore usually absent, but present in one specimen from Zhaojue; intervals 3 and 5 each with ≥ 3 foveate discal pores; interval 7 without discal pore. Male sternite VII with an inconspicuous tubercle on middle, very small and smooth (Fig.
<figureCitation id="381D42B4E94560A94E15DB899973A169" captionStart="Figures 34–39" captionStartId="F7" captionText="Figures 34 - 39. Male sternum VII of Pterostichus pulcher species group spp. 34 P. pemphis sp. nov., holotype 35 P. orbicollis sp. nov., holotype 36 P. liyuani sp. nov., holotype 37 P. condylus sp. nov., holotype 38 P. jialini sp. nov., holotype 39 P. pulcher Sciaky &amp; Allegro, holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures34-39" httpUri="https://binary.pensoft.net/fig/893319" pageId="0" pageNumber="163">36</figureCitation>
). Median lobe of aedeagus stout, strongly curved near basal 1/3; apical lamella gradually dilated and deflected ventrally; apical lamella slightly twisted longitudinally, relatively long (LL/LW = 1.9-2.1), apex rounded (Fig.
<figureCitation id="B307D7B9B206D47E9CD37B725E0BEC80" captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus pulcher species group, left lateral view and dorsal view of median lobe of aedeagus 13 P. pemphis sp. nov., holotype 14 P. orbicollis sp. nov., holotype 15 P. leo sp. nov., holotype 16 P. liyuani sp. nov., holotype 17 P. condylus sp. nov., holotype 18 P. jialini sp. nov., holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures13-18" httpUri="https://binary.pensoft.net/fig/893315" pageId="0" pageNumber="163">16</figureCitation>
). Endophallus long, straightly directed ventrally, gonopore opened to ventral-basal direction of aedeagus;
<emphasis id="32C53803434449EEB0D76C53FE9F52E1" bold="true" pageId="0" pageNumber="163">vb</emphasis>
distinct, oblate, rounded at both apexes; bsp spiral-shaped, deeply grooved, apex of right branch strongly extended and bent reversely (Figs
<figureCitation id="01B99A90A8729187D1F520DF6B60BD80" captionStart="Figures 19–24" captionStartId="F4" captionText="Figures 19 - 24. Endophallus of Pterostichus pulcher species group, left lateral view and right lateral view 19 P. pemphis sp. nov., paratype from Zhaojue county 20 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 21 P. leo sp. nov., paratype from Shizishan mountain, Jinyang county 22 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 23 P. condylus sp. nov., paratype from Mianning county 24 P. pulcher Sciaky &amp; Allegro, paratype from Yizi Yakou, E'bian county to Meigu county. Scale bar: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures19-24" httpUri="https://binary.pensoft.net/fig/893316" pageId="0" pageNumber="163">22</figureCitation>
,
<figureCitation id="0B198A04A25951BADB27A37A9AB797AF" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. 25 - 28 endophallus bsp of Pterostichus pulcher species group, left lateral view 25 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 26 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 27 P. condylus sp. nov., paratype from Mianning county 28 P. pulcher Sciaky &amp; Allegro, paratype from Yizi Yakou, E'bian county to Meigu county 29, 30 Endophallus of Pterostichus (Orientostichus) spp., left lateral view and right lateral view 29 P. prattii Bates, 1890, a male from Sanming city, Fujian province 30 P. curtatus Fairmaire, a male from Heqing county, Yunnan province. Scale bars: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures25-30" httpUri="https://binary.pensoft.net/fig/893317" pageId="0" pageNumber="163">26</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7D4CE2F8E57F7EA96CD45B380E54C9E1" pageId="0" pageNumber="163" type="remarks">
<paragraph id="3D44CA5DE5FC7F38320CC8E231D87241" pageId="0" pageNumber="163">Remarks.</paragraph>
<paragraph id="C0831158376CF8A23EE965740B707577" pageId="0" pageNumber="163">
The only examined specimen from Zhaojue county is different from other ones from Puge county for its elytral parascutellar pores present. It is interesting because most specimens of this species group have no parascutellar pores, but these pores are present only in some specimens collected from Zhaojue of two not closely related species,
<taxonomicName id="1EB2447CEAFA5D4307933E4185F7E236" lsidName="P. liyuani" pageId="0" pageNumber="163" rank="species" species="liyuani">
<emphasis id="2EA6063BE5056DF94DB4B9E7A80C3A31" italics="true" pageId="0" pageNumber="163">P. liyuani</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="96BC7ABE71C21E1F154113CB723D27A6" lsidName="P. pemphis" pageId="0" pageNumber="163" rank="species" species="pemphis">
<emphasis id="A72933387353F4DA955EADDC3C7589E0" italics="true" pageId="0" pageNumber="163">P. pemphis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection id="A52FCEFA142C1B39EF7C4E013415645E" pageId="0" pageNumber="163" type="distribution">
<paragraph id="95E3F2F6E15777507B3D10DFE3D684A7" pageId="0" pageNumber="163">Distribution.</paragraph>
<paragraph id="90E890F04798FF480FF3FAEE1B0DD761" pageId="0" pageNumber="163">
This species is known from two localities in Puge county and Zhaojue county, Liangshan Yi Autonomous Prefecture (Fig.
<figureCitation id="69BFAA5D640D7EC1428E4F5BC6E366E2" captionStart="Figure 42" captionStartId="F9" captionText="Figure 42. Distribution of the taxa of Pterostichus pulcher species group in Sichuan province, China. Brown: P. pemphis sp. nov. Yellow: P. orbicollis sp. nov. Red: P. leo sp. nov. Purple: P. liyuani sp. nov. Blue: P. condylus sp. nov. Green: P. jialini sp. nov. Orange: P. pulcher Sciaky &amp; Allegro." figureDoi="10.3897/zookeys.1175.107636.figure42" httpUri="https://binary.pensoft.net/fig/893321" pageId="0" pageNumber="163">42</figureCitation>
).
</paragraph>
<caption id="E300734415EC03156E205A9018EA450E" doi="10.3897/zookeys.1175.107636.figure42" httpUri="https://binary.pensoft.net/fig/893321" pageId="0" pageNumber="163" start="Figure 42" startId="F9">
<paragraph id="8F91E9876295631029DF94A4150F64D2" pageId="0" pageNumber="163">
<emphasis id="7D8C54E4BED82589F605BE4F65B61980" bold="true" pageId="0" pageNumber="163">Figure 42.</emphasis>
Distribution of the taxa of
<taxonomicName id="6A9F8DC9C1E7B42212A2BCB5CD5B95B7" authorityName="Sciaky &amp; Allegro" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus pulcher" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="744467B69D11C2DBEBA07A2BEA878D44" italics="true" pageId="0" pageNumber="163">Pterostichus pulcher</emphasis>
</taxonomicName>
species group in Sichuan province, China. Brown:
<taxonomicName id="1C4F8373F70EB5B494F0FFCC65A073AD" lsidName="P. pemphis" pageId="0" pageNumber="163" rank="species" species="pemphis">
<emphasis id="1AF111F060C887199C79C2E2A9710BCD" italics="true" pageId="0" pageNumber="163">P. pemphis</emphasis>
</taxonomicName>
sp. nov. Yellow:
<taxonomicName id="0D5FA9E1E6BBE93A02D17D2A2911C14F" lsidName="P. orbicollis" pageId="0" pageNumber="163" rank="species" species="orbicollis">
<emphasis id="7B9046CA6386DA0746F8999E707D2DFF" italics="true" pageId="0" pageNumber="163">P. orbicollis</emphasis>
</taxonomicName>
sp. nov. Red:
<taxonomicName id="A7866BFDF8A5060A0BF2C9B7C247AC04" lsidName="P. leo" pageId="0" pageNumber="163" rank="species" species="leo">
<emphasis id="DDA0C940F95EE40B41B83950CAC0225C" italics="true" pageId="0" pageNumber="163">P. leo</emphasis>
</taxonomicName>
sp. nov. Purple:
<taxonomicName id="309ACD1E9FC01612E16DA90CCD17C1CD" lsidName="P. liyuani" pageId="0" pageNumber="163" rank="species" species="liyuani">
<emphasis id="3A3C9C2AF91827DD24978A7F3A1E8FF5" italics="true" pageId="0" pageNumber="163">P. liyuani</emphasis>
</taxonomicName>
sp. nov. Blue:
<taxonomicName id="82BC8EEC8776D383069A6F6C5D6F381E" lsidName="P. condylus" pageId="0" pageNumber="163" rank="species" species="condylus">
<emphasis id="010A15E0A42FE3F9B2448F0664E55D00" italics="true" pageId="0" pageNumber="163">P. condylus</emphasis>
</taxonomicName>
sp. nov. Green:
<taxonomicName id="2180F03FE5E4C7884ED08CCEFB3203A7" lsidName="P. jialini" pageId="0" pageNumber="163" rank="species" species="jialini">
<emphasis id="9A1F6CDB13C7150C4EC6D6D718672663" italics="true" pageId="0" pageNumber="163">P. jialini</emphasis>
</taxonomicName>
sp. nov. Orange:
<taxonomicName id="2C1A85067311A9F98836C0D8930D4131" lsidName="P. pulcher" pageId="0" pageNumber="163" rank="species" species="pulcher">
<emphasis id="24EC3D5AAAA9996B83077DC0A094E850" italics="true" pageId="0" pageNumber="163">P. pulcher</emphasis>
</taxonomicName>
Sciaky &amp; Allegro.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="F42887FF0C3DF5830D5E3229F49457CD" pageId="0" pageNumber="163" type="etymology">
<paragraph id="AC84A724852B8208538A77E8DC546F64" pageId="0" pageNumber="163">Etymology.</paragraph>
<paragraph id="6340A9CC911E8CBB3EDAB907B3E8EE49" pageId="0" pageNumber="163">This new species is named for Mr. Yuan Li, who contributed large number of specimens for the present study including many ones of this new species.</paragraph>
</subSubSection>
</treatment>
</document>
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