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<document id="AEBF68CDD56252F5BDE730F5F40438ED" ID-DOI="10.11646/zootaxa.5377.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10208826" ID-ZooBank="2BA30329-4BDF-4595-9864-3CA23D56A659" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1701074315795" checkinUser="plazi" docAuthor="González-Obando, Ranulfo, Calderón-Martínez, Nadia R. &amp; Carrejo-Gironza, Nancy S." docDate="2023" docId="345F87BEFB2DFFDFD9C83898AF1FFBB8" docLanguage="en" docName="zootaxa.5377.1.1.pdf" docOrigin="Zootaxa 5377 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5377.1.1/52352" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Myopsocus caliensis González-Obando &amp; Calderón-Martínez &amp; Carrejo-Gironza 2023, n. sp." docType="treatment" docVersion="3" lastPageNumber="34" masterDocId="C866FFC6FB0CFFFED95F380FAA27FFE1" masterDocTitle="New species of Myopsocidae (Insecta: Psocodea: “ Psocoptera ”) from Colombia" masterLastPageNumber="61" masterPageNumber="1" pageNumber="34" updateTime="1701260917186" updateUser="juliana">
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<mods:title id="3BD50C8FEB0E5E6DE97E2CD75BAEE422">New species of Myopsocidae (Insecta: Psocodea: “ Psocoptera ”) from Colombia</mods:title>
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<mods:namePart id="864E98948450E129CF005F05F86A6DAD">González-Obando, Ranulfo</mods:namePart>
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<mods:affiliation id="4A16D77B929BA7BB18E7945B8C5A73E5">Grupo de Investigaciones Entomológicas (GIE), Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, Colombia &amp; ranulfo. gonzalez @ correounivalle. edu. co; https: // orcid. org / 0000 - 0003 - 4047 - 8677</mods:affiliation>
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<mods:namePart id="75B952CC5B9C46939525A3D0742E9B44">Calderón-Martínez, Nadia R.</mods:namePart>
<mods:nameIdentifier id="D7F11F7D494B59BDEEC2ECB26AC08566" type="ORCID">0009-0005-3222-2403</mods:nameIdentifier>
<mods:affiliation id="DE7E04D736018AB57A254EEAC6AFE3F5">Grupo de Investigaciones Entomológicas (GIE), Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, Colombia &amp; calderonmartinez. nadia @ gmail. com; https: // orcid. org / 0009 - 0005 - 3222 - 2403</mods:affiliation>
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<mods:namePart id="CE62E4C64145478D99E8BC4D32751EC9">Carrejo-Gironza, Nancy S.</mods:namePart>
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<mods:affiliation id="C8BD297A2CAF09F9C67410568042E6BE">Grupo de Investigaciones Entomológicas (GIE), Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, Colombia &amp; nancy. carrejo @ correounivalle. edu. co; https: // orcid. org / 0000 - 0002 - 2395 - 6887</mods:affiliation>
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<treatment id="345F87BEFB2DFFDFD9C83898AF1FFBB8" LSID="urn:lsid:plazi:treatment:345F87BEFB2DFFDFD9C83898AF1FFBB8" httpUri="http://treatment.plazi.org/id/345F87BEFB2DFFDFD9C83898AF1FFBB8" lastPageNumber="34" pageId="33" pageNumber="34">
<subSubSection id="F4EC6523FB2DFFDFD9C83898ABFFFF50" box="[151,472,151,178]" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="BC4936A8FB2DFFDFD9C83898ABFFFF50" blockId="33.[151,472,151,213]" box="[151,472,151,178]" pageId="33" pageNumber="34">
<heading id="E70181C4FB2DFFDFD9C83898ABFFFF50" bold="true" box="[151,472,151,178]" fontSize="11" level="2" pageId="33" pageNumber="34" reason="4">
<emphasis id="8E82EABAFB2DFFDFD9C83898ABFFFF50" bold="true" box="[151,472,151,178]" pageId="33" pageNumber="34">
<taxonomicName id="7BF64D2BFB2DFFDFD9C83898ABA9FF53" authority="González-Obando &amp; Calderón-Martínez &amp; Carrejo-Gironza, 2023" authorityName="González-Obando &amp; Calderón-Martínez &amp; Carrejo-Gironza" authorityYear="2023" box="[151,398,151,178]" class="Insecta" family="Myopsocidae" genus="Myopsocus" kingdom="Animalia" order="Psocodea" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="caliensis" status="sp. nov.">
<emphasis id="8E82EABAFB2DFFDFD9C83898ABA9FF53" bold="true" box="[151,398,151,178]" italics="true" pageId="33" pageNumber="34">Myopsocus caliensis</emphasis>
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<taxonomicNameLabel id="95B157C1FB2DFFDFD8CA3898ABFFFF50" box="[405,472,151,177]" pageId="33" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="F4EC6523FB2DFFDFD9C838B4AB00FF34" box="[151,295,187,213]" pageId="33" pageNumber="34" type="description">
<paragraph id="BC4936A8FB2DFFDFD9C838B4AB00FF34" blockId="33.[151,472,151,213]" box="[151,295,187,213]" pageId="33" pageNumber="34">
(
<figureCitation id="24CD2A2DFB2DFFDFD9C038B4AB38FF34" box="[159,287,187,213]" captionStart="FIGURES 9196" captionStartId="32.[152,270,1943,1967]" captionTargetBox="[158,1427,188,1919]" captionTargetId="figure-24@32.[151,1436,181,1919]" captionTargetPageId="32" captionText="FIGURES 9196. Myopsocus caliensis n. sp. Male. 91. Forewing. 92. Hindwing. 93. Head (frontal view). 94. Epiproct and paraproct. 95. Hypandrium. 96. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10208862" httpUri="https://zenodo.org/record/10208862/files/figure.png" pageId="33" pageNumber="34">Figs 9196</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F4EC6523FB2DFFDFD9C8390CA83FFC63" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="BC4936A8FB2DFFDFD9C8390CA88AFE68" blockId="33.[151,1437,259,1114]" pageId="33" pageNumber="34">
<emphasis id="8E82EABAFB2DFFDFD9C8390CAB35FEFC" bold="true" box="[151,274,259,285]" pageId="33" pageNumber="34">Diagnosis.</emphasis>
Belonging to the assemblage of species that do not present the forewing with the scalloped apical margin, unlike these it presents a pyriform phallosome, with narrow and fused phallobase, wider distally (
<figureCitation id="24CD2A2DFB2DFFDFDD853928AF09FEA0" box="[1242,1326,295,321]" captionStart="FIGURES 9196" captionStartId="32.[152,270,1943,1967]" captionTargetBox="[158,1427,188,1919]" captionTargetId="figure-24@32.[151,1436,181,1919]" captionTargetPageId="32" captionText="FIGURES 9196. Myopsocus caliensis n. sp. Male. 91. Forewing. 92. Hindwing. 93. Head (frontal view). 94. Epiproct and paraproct. 95. Hypandrium. 96. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10208862" httpUri="https://zenodo.org/record/10208862/files/figure.png" pageId="33" pageNumber="34">Fig. 96</figureCitation>
), external parameres flattened, widened and curved inward, apically separated by short membrane; endophallus elongated, flattened, distally and apically more broadened.
</paragraph>
<paragraph id="BC4936A8FB2DFFDFD998399CAE3BFD80" blockId="33.[151,1437,259,1114]" pageId="33" pageNumber="34">
<emphasis id="8E82EABAFB2DFFDFD998399CAB79FE4C" bold="true" box="[199,350,403,429]" pageId="33" pageNumber="34">Male. Color.</emphasis>
Head (
<figureCitation id="24CD2A2DFB2DFFDFD8E8399CA82BFE4C" box="[439,524,403,429]" captionStart="FIGURES 9196" captionStartId="32.[152,270,1943,1967]" captionTargetBox="[158,1427,188,1919]" captionTargetId="figure-24@32.[151,1436,181,1919]" captionTargetPageId="32" captionText="FIGURES 9196. Myopsocus caliensis n. sp. Male. 91. Forewing. 92. Hindwing. 93. Head (frontal view). 94. Epiproct and paraproct. 95. Hypandrium. 96. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10208862" httpUri="https://zenodo.org/record/10208862/files/figure.png" pageId="33" pageNumber="34">Fig. 93</figureCitation>
) dark brown, darker genae, clypeus and labrum. Compound eyes black, ocelli hyaline, with ochre centripetal crescents. Forewing (
<figureCitation id="24CD2A2DFB2DFFDFDBBF39B8A916FE30" box="[736,817,439,465]" captionStart="FIGURES 9196" captionStartId="32.[152,270,1943,1967]" captionTargetBox="[158,1427,188,1919]" captionTargetId="figure-24@32.[151,1436,181,1919]" captionTargetPageId="32" captionText="FIGURES 9196. Myopsocus caliensis n. sp. Male. 91. Forewing. 92. Hindwing. 93. Head (frontal view). 94. Epiproct and paraproct. 95. Hypandrium. 96. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10208862" httpUri="https://zenodo.org/record/10208862/files/figure.png" pageId="33" pageNumber="34">Fig. 91</figureCitation>
) with dark brown spots, with well-marked basal fascia, pterostigma filled with small purple spots that extends outside the margin, while retaining its shape, alar margin dotted, venation poorly differentiated basally and distally marked. Hindwing (
<figureCitation id="24CD2A2DFB2DFFDFDD5239F0AE47FDF8" box="[1037,1120,511,537]" captionStart="FIGURES 9196" captionStartId="32.[152,270,1943,1967]" captionTargetBox="[158,1427,188,1919]" captionTargetId="figure-24@32.[151,1436,181,1919]" captionTargetPageId="32" captionText="FIGURES 9196. Myopsocus caliensis n. sp. Male. 91. Forewing. 92. Hindwing. 93. Head (frontal view). 94. Epiproct and paraproct. 95. Hypandrium. 96. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10208862" httpUri="https://zenodo.org/record/10208862/files/figure.png" pageId="33" pageNumber="34">Fig. 92</figureCitation>
) smoked, with veins brown and dotted alar margin. Legs: coxae and trochanter dark brown, femur dark brown with subapical band white-creamy, tibia and tarsi brown. Hypandrium, paraprocts and epiproct dark brown.
</paragraph>
<paragraph id="BC4936A8FB2DFFDFD9983A64A83FFC63" blockId="33.[151,1437,259,1114]" pageId="33" pageNumber="34">
<emphasis id="8E82EABAFB2DFFDFD9983A64AB46FD64" bold="true" box="[199,353,619,645]" pageId="33" pageNumber="34">Morphology.</emphasis>
As in diagnosis, plus the following: Head (
<figureCitation id="24CD2A2DFB2DFFDFDA393A64A99CFD64" box="[870,955,619,645]" captionStart="FIGURES 9196" captionStartId="32.[152,270,1943,1967]" captionTargetBox="[158,1427,188,1919]" captionTargetId="figure-24@32.[151,1436,181,1919]" captionTargetPageId="32" captionText="FIGURES 9196. Myopsocus caliensis n. sp. Male. 91. Forewing. 92. Hindwing. 93. Head (frontal view). 94. Epiproct and paraproct. 95. Hypandrium. 96. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10208862" httpUri="https://zenodo.org/record/10208862/files/figure.png" pageId="33" pageNumber="34">Fig. 93</figureCitation>
) H/MxW: 1.25; compound eyes grandes, H/D: 1.74, with interonmatidial setae; vertex with emargination V-shaped, with slightly pronounced bulge before the epicranial suture. IO/MxW: 0.50. Outer cusp of lacinial tips little wide, with four denticles; Mx4/Mx2: 1.38. Forewings (
<figureCitation id="24CD2A2DFB2DFFDFD87F3AD8AB50FD10" box="[288,375,727,753]" captionStart="FIGURES 9196" captionStartId="32.[152,270,1943,1967]" captionTargetBox="[158,1427,188,1919]" captionTargetId="figure-24@32.[151,1436,181,1919]" captionTargetPageId="32" captionText="FIGURES 9196. Myopsocus caliensis n. sp. Male. 91. Forewing. 92. Hindwing. 93. Head (frontal view). 94. Epiproct and paraproct. 95. Hypandrium. 96. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10208862" httpUri="https://zenodo.org/record/10208862/files/figure.png" pageId="33" pageNumber="34">Fig. 91</figureCitation>
): L/W: 2.68. Pterostigma: lp/wp: 2.67, areola postica wider than tall, apically angled: al/ah: 1.67. Hindwings (
<figureCitation id="24CD2A2DFB2DFFDFD83C3AF4AB9FFCF4" box="[355,440,763,789]" captionStart="FIGURES 9196" captionStartId="32.[152,270,1943,1967]" captionTargetBox="[158,1427,188,1919]" captionTargetId="figure-24@32.[151,1436,181,1919]" captionTargetPageId="32" captionText="FIGURES 9196. Myopsocus caliensis n. sp. Male. 91. Forewing. 92. Hindwing. 93. Head (frontal view). 94. Epiproct and paraproct. 95. Hypandrium. 96. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10208862" httpUri="https://zenodo.org/record/10208862/files/figure.png" pageId="33" pageNumber="34">Fig. 92</figureCitation>
): l/w: 2.62. Paraprocts (
<figureCitation id="24CD2A2DFB2DFFDFDB973AF4A93DFCF4" box="[712,794,763,789]" captionStart="FIGURES 9196" captionStartId="32.[152,270,1943,1967]" captionTargetBox="[158,1427,188,1919]" captionTargetId="figure-24@32.[151,1436,181,1919]" captionTargetPageId="32" captionText="FIGURES 9196. Myopsocus caliensis n. sp. Male. 91. Forewing. 92. Hindwing. 93. Head (frontal view). 94. Epiproct and paraproct. 95. Hypandrium. 96. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10208862" httpUri="https://zenodo.org/record/10208862/files/figure.png" pageId="33" pageNumber="34">Fig. 94</figureCitation>
) with well-sclerotized conical lateral projection hiding a small membranous tubercle covered setae, sensory fields with 23 trichobothria on basal rosettes. Epiproct (
<figureCitation id="24CD2A2DFB2DFFDFDC1C3B10AFB3FCD8" box="[1347,1428,799,825]" captionStart="FIGURES 9196" captionStartId="32.[152,270,1943,1967]" captionTargetBox="[158,1427,188,1919]" captionTargetId="figure-24@32.[151,1436,181,1919]" captionTargetPageId="32" captionText="FIGURES 9196. Myopsocus caliensis n. sp. Male. 91. Forewing. 92. Hindwing. 93. Head (frontal view). 94. Epiproct and paraproct. 95. Hypandrium. 96. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10208862" httpUri="https://zenodo.org/record/10208862/files/figure.png" pageId="33" pageNumber="34">Fig. 94</figureCitation>
) crescent-shaped, with a transverse dorsal protuberance with superficial small tubercles, distal edge rounded and an apical bulge with four distal setae.
</paragraph>
</subSubSection>
<subSubSection id="F4EC6523FB2DFFDFD9983B84A97EFC28" pageId="33" pageNumber="34" type="description">
<paragraph id="BC4936A8FB2DFFDFD9983B84A97EFC28" blockId="33.[151,1437,259,1114]" pageId="33" pageNumber="34">
<emphasis id="8E82EABAFB2DFFDFD9983B84AB59FC44" bold="true" box="[199,382,907,933]" pageId="33" pageNumber="34">Measurements.</emphasis>
FW: 2907.5, HW: 2175, F: 525, T: 1075, t1: 400, t2: 65, t3: 70, ctt1: 19, f1: 675, f2: 462.5, f3: 382.5, Mx4: 165, IO: 266, d: 300, D: 382, IO/d: 0.89, PO: 0.79
</paragraph>
</subSubSection>
<subSubSection id="F4EC6523FB2DFFDFD9983BDCA9F4FBD4" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="BC4936A8FB2DFFDFD9983BDCA9F4FBD4" blockId="33.[151,1437,259,1114]" pageId="33" pageNumber="34">
<emphasis id="8E82EABAFB2DFFDFD9983BDCABB3FC0C" bold="true" box="[199,404,979,1005]" pageId="33" pageNumber="34">Material studied.</emphasis>
<materialsCitation id="0C9E3CF5FB2DFFDFD8C43BDCAFBEFBF0" collectingDate="2009-04-08" collectingMethod="Manual collection" collectionCode="MUSENUV" collectorName="R. Gonzalez" country="Colombia" elevation="1153" latitude="3.3723612" location="Santiago de Cali" longLatPrecision="1" longitude="-76.5698" pageId="33" pageNumber="34" specimenCode="31328" specimenCount="1" stateProvince="Valle del Cauca" typeStatus="holotype">
<typeStatus id="634D880AFB2DFFDFD8C43BDCA824FC0C" box="[411,515,979,1005]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
<specimenCount id="AAF0FD21FB2DFFDFDB553BDCA863FC0C" box="[522,580,979,1005]" pageId="33" pageNumber="34" type="male">male</specimenCount>
.
<collectingCountry id="C4E17638FB2DFFDFDB113BDCA8C0FC0C" box="[590,743,979,1005]" name="Colombia" pageId="33" pageNumber="34">COLOMBIA</collectingCountry>
.
<collectingRegion id="7E32F84AFB2DFFDFDBAF3BDCA985FC0C" box="[752,930,979,1005]" country="Colombia" name="Valle del Cauca" pageId="33" pageNumber="34">Valle del Cauca</collectingRegion>
,
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<location id="B9296073FB2DFFDFDD2C3BDBAED1FC0C" LSID="urn:lsid:plazi:treatment:345F87BEFB2DFFDFD9C83898AF1FFBB8:B9296073FB2DFFDFDD2C3BDBAED1FC0C" box="[1139,1270,979,1006]" country="Colombia" latitude="3.3723612" longLatPrecision="1" longitude="-76.5698" name="La Buitrera" pageId="33" pageNumber="34" stateProvince="Valle del Cauca">La Buitrera</location>
,
<geoCoordinate id="D9C2506FFB2DFFDFDC5F3BDCAFBEFC0F" box="[1280,1433,979,1006]" degrees="3" direction="north" minutes="22" orientation="latitude" pageId="33" pageNumber="34" precision="1" seconds="20.5" value="3.3723612">3°2220.5N</geoCoordinate>
:
<geoCoordinate id="D9C2506FFB2DFFDFD9C83BF8AB1BFBF3" box="[151,316,1015,1042]" degrees="76" direction="west" minutes="34" orientation="longitude" pageId="33" pageNumber="34" precision="1" seconds="11.3" value="-76.5698">76°3411.3W</geoCoordinate>
.
<quantity id="7B0E9B4DFB2DFFDFD80E3BF8AB86FBF3" box="[337,417,1015,1042]" metricMagnitude="3" metricUnit="m" metricValue="1.153" pageId="33" pageNumber="34" unit="m" value="1153.0">
<elevation id="37DBD19BFB2DFFDFD80E3BF8AB86FBF3" box="[337,417,1015,1042]" metricMagnitude="3" metricUnit="m" metricValue="1.153" pageId="33" pageNumber="34" unit="m" value="1153.0">1153m</elevation>
</quantity>
.
<date id="C8481068FB2DFFDFD8ED3BF8A80BFBF0" box="[434,556,1015,1041]" pageId="33" pageNumber="34" value="2009-04-08">
<collectingDate id="D80CE980FB2DFFDFD8ED3BF8A80BFBF0" box="[434,556,1015,1041]" pageId="33" pageNumber="34" value="2009-04-08">08.iv.2009</collectingDate>
</date>
.
<collectionCode id="DAE7AE6DFB2DFFDFDB623BF8A8E0FBF0" box="[573,711,1015,1041]" pageId="33" pageNumber="34">MUSENUV</collectionCode>
slide code:
<specimenCode id="EC509ED3FB2DFFDFDA3E3BF8A98CFBF0" box="[865,939,1015,1041]" pageId="33" pageNumber="34">31328</specimenCode>
.
<collectorName id="1103537EFB2DFFDFDAE33BF7AE76FBF0" box="[956,1105,1015,1042]" pageId="33" pageNumber="34">R. González</collectorName>
.
<collectingMethod id="65B74EBFFB2DFFDFDD3D3BF8AF14FBF0" box="[1122,1331,1015,1041]" pageId="33" pageNumber="34">Manual collection</collectingMethod>
method
</materialsCitation>
.
<materialsCitation id="0C9E3CF5FB2DFFDFD9C83C13A9F7FBD4" box="[151,976,1051,1078]" collectingDate="2009-04-08" collectingMethod="Manual collection" collectionCode="MUSENUV" collectorName="La Buitrera" country="Colombia" elevation="1153" latitude="3.3723612" location="Santiago de Cali" longLatPrecision="1" longitude="-76.5698" pageId="33" pageNumber="34" specimenCode="31329" specimenCount="1" specimenCount-male="1" stateProvince="Valle del Cauca" typeStatus="paratype">
<typeStatus id="634D880AFB2DFFDFD9C83C13AB2FFBD7" box="[151,264,1052,1078]" pageId="33" pageNumber="34" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="AAF0FD21FB2DFFDFD84C3C14AB45FBD4" box="[275,354,1051,1077]" count="1" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
. Same data as holotype.
<collectionCode id="DAE7AE6DFB2DFFDFDB2A3C14A8D8FBD4" box="[629,767,1051,1077]" pageId="33" pageNumber="34">MUSENUV</collectionCode>
slide code:
<specimenCode id="EC509ED3FB2DFFDFDAD93C14A9F7FBD4" box="[902,976,1051,1077]" pageId="33" pageNumber="34">31329</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F4EC6523FB2DFFDFD9983C30AF1FFBB8" box="[199,1336,1087,1114]" pageId="33" pageNumber="34" type="etymology">
<paragraph id="BC4936A8FB2DFFDFD9983C30AF1FFBB8" blockId="33.[151,1437,259,1114]" box="[199,1336,1087,1114]" pageId="33" pageNumber="34">
<emphasis id="8E82EABAFB2DFFDFD9983C30AB69FBB8" bold="true" box="[199,334,1087,1113]" pageId="33" pageNumber="34">Etymology.</emphasis>
The specific epithet refers to Santiago de Cali,
<collectingCountry id="C4E17638FB2DFFDFDA3C3C30A9F3FBB8" box="[867,980,1087,1113]" name="Colombia" pageId="33" pageNumber="34">Colombia</collectingCountry>
, where the
<typeStatus id="634D880AFB2DFFDFDD0B3C30AE91FBB8" box="[1108,1206,1087,1113]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
was found.
</paragraph>
</subSubSection>
</treatment>
</document>