523 lines
62 KiB
XML
523 lines
62 KiB
XML
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<mods:title id="591A7637DC9C4C41306B839C20E3D380">A revision of the Rutilus complex from Mediterranean Europe with description of a new genus, Sarmarutilus, and a new species, Rutilus stoumboudae (Teleostei: Cyprinidae)</mods:title>
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<mods:date id="FA71671B13DAA4808521E1D6557DC289">2014</mods:date>
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<treatment id="8E3D87DAFFDFFFBC66A3FF7CFEC6F8FB" ID-DOI="http://doi.org/10.5281/zenodo.6140523" ID-GBIF-Taxon="119512637" ID-Zenodo-Dep="6140523" LSID="urn:lsid:plazi:treatment:8E3D87DAFFDFFFBC66A3FF7CFEC6F8FB" httpUri="http://treatment.plazi.org/id/8E3D87DAFFDFFFBC66A3FF7CFEC6F8FB" lastPageId="20" lastPageNumber="399" pageId="19" pageNumber="398">
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<subSubSection id="4E8E6547FFDFFFBB66A3FF7CFF65FF67" pageId="19" pageNumber="398" type="nomenclature">
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<paragraph id="062B36CCFFDFFFBB66A3FF7CFDBBFF06" blockId="19.[151,545,151,211]" box="[151,545,151,178]" pageId="19" pageNumber="398">
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<heading id="5D6381A0FFDFFFBB66A3FF7CFDBBFF06" bold="true" box="[151,545,151,178]" fontSize="11" level="1" pageId="19" pageNumber="398" reason="1">
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<taxonomicName id="C1944D4FFFDFFFBB66A3FF7CFDBBFF06" ID-CoL="4TTHN" authority="La Cepede, 1804" authorityName="La Cepede" authorityYear="1804" box="[151,545,151,178]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="pigus">
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<emphasis id="34E0EADEFFDFFFBB66A3FF7CFDBBFF06" bold="true" box="[151,545,151,178]" pageId="19" pageNumber="398">
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<emphasis id="34E0EADEFFDFFFBB66A3FF7CFEADFF05" bold="true" box="[151,311,151,177]" italics="true" pageId="19" pageNumber="398">Rutilus pigus</emphasis>
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(La Cepède, 1804)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="062B36CCFFDFFFBB66A3FF51FF65FF67" blockId="19.[151,545,151,211]" box="[151,255,186,211]" pageId="19" pageNumber="398">
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(
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<figureCitation id="9EAF2A49FFDFFFBB66ABFF51FF78FF67" box="[159,226,186,211]" captionStart="FIGURE 2" captionStartId="4.[151,250,1514,1536]" captionTargetBox="[173,1424,552,1475]" captionTargetId="figure@4.[151,1436,519,1492]" captionTargetPageId="4" captionText="FIGURE 2. Native species of the Rutilus complex in southern Mediterranean Europe: A) Leucos aula, 115 mm SL, R. Bacchiglione, northern Italy, April 1996 (IZA 0425); B) Leucos basak, 95 mm SL, Lake Ohrid, FYROM, August 1987, (IZA 0429); C) Leucos ylikiensis, 120 mm SL, Lake Yliki, April 1987 (IZA 8733); D) Leucos panosi, 130 mm SL, canal to Lake Trichonis, May 1987 (IZA 8766); E) Leucos albus, 98 mm SL, River Moraca, Montenegro, November 1999 (IZA 0421); F) Sarmarutilus rubilio, 100 mm SL, River Ofanto, southern Italy, February 2002 (IZA 02183); G) Rutilus pigus, 230 mm SL, Lake Major, northern Italy, (IZA 0474); H) Holotype of Rutilus stoumboudae new species, 148 mm SL, October 1998, Lake Volvi, Greece (IZA 02107)." httpUri="https://zenodo.org/record/225780/files/figure.png" pageId="19" pageNumber="398">Fig. 2</figureCitation>
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G)
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||
</paragraph>
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</subSubSection>
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<subSubSection id="4E8E6547FFDFFFBB66A3FEE8FB1BFEAC" box="[151,1153,258,280]" pageId="19" pageNumber="398" type="reference_group">
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<paragraph id="062B36CCFFDFFFBB66A3FEE8FB1BFEAC" blockId="19.[151,1153,258,280]" box="[151,1153,258,280]" pageId="19" pageNumber="398">
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<taxonomicName id="C1944D4FFFDFFFBB66A3FEE8FD83FEAC" ID-CoL="33HXM" authority="La Cepede, 1803: 503" authorityName="La Cepede" authorityPageNumber="503" authorityYear="1803" box="[151,537,258,280]" class="Actinopterygii" family="Cyprinidae" genus="Cyprinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="pigus">
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<emphasis id="34E0EADEFFDFFFBB66A3FEE8FEABFEAC" box="[151,305,259,280]" italics="true" pageId="19" pageNumber="398">Cyprinus pigus</emphasis>
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La Cepède, 1803: 503
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</taxonomicName>
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, 607 (
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<typeStatus id="D92F886EFFDFFFBB6463FEE9FD1EFEAC" box="[599,644,258,280]" pageId="19" pageNumber="398">type</typeStatus>
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localities: lakes Major and Como, northern
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<collectingCountry id="7E83765CFFDFFFBB6276FEE9FBEFFEAC" box="[1090,1141,258,280]" name="Italy" pageId="19" pageNumber="398">Italy</collectingCountry>
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).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4E8E6547FFDFFFBB66A3FEAEFBF0FE12" pageId="19" pageNumber="398" type="materials_examined">
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||
<paragraph id="062B36CCFFDFFFBB66A3FEAEFBF0FE12" blockId="19.[151,1437,325,1646]" pageId="19" pageNumber="398">
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||
<emphasis id="34E0EADEFFDFFFBB66A3FEAEFE19FEEA" bold="true" box="[151,387,325,350]" pageId="19" pageNumber="398">Examined material.</emphasis>
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||
<collectionCode id="6085AE09FFDFFFBB67BDFEADFE22FEEA" box="[393,440,326,350]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="19" pageNumber="398">IZA</collectionCode>
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0 471, 12,
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<collectingCountry id="7E83765CFFDFFFBB6419FEAEFDFBFEEA" box="[557,609,325,350]" name="Italy" pageId="19" pageNumber="398">Italy</collectingCountry>
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||
, L. Como,
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<collectingCountry id="7E83765CFFDFFFBB64E8FEADFD69FEEA" box="[732,755,326,350]" name="Iceland" pageId="19" pageNumber="398">O</collectingCountry>
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. Mangione,
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<date id="722A100CFFDFFFBB6549FEAEFC62FEEA" box="[893,1016,325,350]" pageId="19" pageNumber="398" value="2004-04">April 2004</date>
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.—
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<collectionCode id="6085AE09FFDFFFBB6214FEADFBD4FEEA" box="[1056,1102,326,350]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="19" pageNumber="398">IZA</collectionCode>
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0 472, 3,
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<collectingCountry id="7E83765CFFDFFFBB6282FEAEFB70FEEA" box="[1206,1258,325,350]" name="Italy" pageId="19" pageNumber="398">Italy</collectingCountry>
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, R. Livenza, M. Povz,
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<date id="722A100CFFDFFFBB66E9FE82FEE9FE36" box="[221,371,361,386]" pageId="19" pageNumber="398" value="1996-05-22">22 May 1996</date>
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||
.—
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<collectionCode id="6085AE09FFDFFFBB67A7FE82FE71FE36" box="[403,491,361,386]" pageId="19" pageNumber="398">MSNM</collectionCode>
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||
(uncataloged), 33,
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||
<collectingCountry id="7E83765CFFDFFFBB64F2FE82FD60FE36" box="[710,762,361,386]" name="Italy" pageId="19" pageNumber="398">Italy</collectingCountry>
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, L. Maggiore, A. Mojetta,
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<date id="722A100CFFDFFFBB621DFE81FB01FE35" box="[1065,1179,361,385]" pageId="19" pageNumber="398" value="1988-06">June 1988</date>
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||
.—
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||
<collectionCode id="6085AE09FFDFFFBB6289FE82FB74FE35" box="[1213,1262,361,385]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="19" pageNumber="398">IZA</collectionCode>
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||
843,6,
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<collectingCountry id="7E83765CFFDFFFBB6377FE82FAEDFE36" box="[1347,1399,361,386]" name="Italy" pageId="19" pageNumber="398">Italy</collectingCountry>
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||
, R. Po, G. Delmastro,
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<date id="722A100CFFDFFFBB6754FE66FD96FE12" box="[352,524,397,422]" pageId="19" pageNumber="398" value="1983-03-13">13 March 1983</date>
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||
.-
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||
<collectionCode id="6085AE09FFDFFFBB642BFE65FDD7FE12" box="[543,589,398,422]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="19" pageNumber="398">IZA</collectionCode>
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||
842, 6,
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<collectingCountry id="7E83765CFFDFFFBB649DFE66FD47FE12" box="[681,733,397,422]" name="Italy" pageId="19" pageNumber="398">Italy</collectingCountry>
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||
, L. Major,
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||
<date id="722A100CFFDFFFBB656DFE66FC4CFE12" box="[857,982,397,422]" pageId="19" pageNumber="398" value="1983-04">April 1983</date>
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||
, G. Giussani.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4E8E6547FFDFFFBB66F3FE5AFBAAFD72" pageId="19" pageNumber="398" type="diagnosis">
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<paragraph id="062B36CCFFDFFFBB66F3FE5AFBAAFD72" blockId="19.[151,1437,325,1646]" pageId="19" pageNumber="398">
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||
<emphasis id="34E0EADEFFDFFFBB66F3FE5AFEDBFE7E" bold="true" box="[199,321,433,458]" pageId="19" pageNumber="398">Diagnosis.</emphasis>
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<taxonomicName id="C1944D4FFFDFFFBB677CFE5AFE40FE7D" box="[328,474,433,458]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="pigus">
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<emphasis id="34E0EADEFFDFFFBB677CFE5AFE40FE7D" box="[328,474,433,458]" italics="true" pageId="19" pageNumber="398">Rutilus pigus</emphasis>
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</taxonomicName>
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, was only recently separated from
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<taxonomicName id="C1944D4FFFDFFFBB6556FE59FC2EFE7D" box="[866,948,434,457]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="virgo">
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<emphasis id="34E0EADEFFDFFFBB6556FE59FC2EFE7D" box="[866,948,434,457]" italics="true" pageId="19" pageNumber="398">R.virgo</emphasis>
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||
</taxonomicName>
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||
(
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||
<bibRefCitation id="62054B3DFFDFFFBB65F6FE5AFB78FE7D" author="Kottelat" box="[962,1250,433,458]" pageId="19" pageNumber="398" refString="Kottelat, M. & Freyhof, J. (2007) Handbook of European Freshwater Fishes. Kottelat, Cornol, Switzerland and Frehyof, Berlin, Germany, 646 pp." type="book" year="2007">Kottelat & Frehyof, 2007</bibRefCitation>
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;
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||
<bibRefCitation id="62054B3DFFDFFFBB62DAFE5AFF4EFE5A" author="Ketmaier" pageId="19" pageNumber="398" refString="Ketmaier, V., Bianco, P. G. & Durand, J. D. (2008) Molecular systematics, phylogeny and biogeography of roaches (Rutilus, Teleostei, Cyprinidae). Molecular Phylogenetics and Evolution, 49, 362 - 367. http: // dx. doi. org / 10.1016 / j. ympev. 2008.07.012" type="journal article" year="2008">
|
||
Ketmaier
|
||
<emphasis id="34E0EADEFFDFFFBB6369FE59FA1CFE7D" box="[1373,1414,434,457]" italics="true" pageId="19" pageNumber="398">et a</emphasis>
|
||
l., 2008
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="C1944D4FFFDFFFBB66D9FE3DFECAFE59" box="[237,336,470,493]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="pigus">
|
||
<emphasis id="34E0EADEFFDFFFBB66D9FE3DFECAFE59" box="[237,336,470,493]" italics="true" pageId="19" pageNumber="398">R. pigus</emphasis>
|
||
</taxonomicName>
|
||
, is endemic of Padany-Venetian district, and differs from
|
||
<taxonomicName id="C1944D4FFFDFFFBB6221FE3DFBEFFE59" box="[1045,1141,470,493]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="virgo">
|
||
<emphasis id="34E0EADEFFDFFFBB6221FE3DFBEFFE59" box="[1045,1141,470,493]" italics="true" pageId="19" pageNumber="398">R. virgo</emphasis>
|
||
</taxonomicName>
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||
from central Europe, in squamation as it has 46–51 scales on LL, and 8.5 above and 3.5 below the LL, as against 40–46 scales on LL and 7.5 above and 3.5 below the LL in
|
||
<taxonomicName id="C1944D4FFFDFFFBB6410FDF5FDE4FD81" box="[548,638,542,565]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="virgo">
|
||
<emphasis id="34E0EADEFFDFFFBB6410FDF5FDE4FD81" box="[548,638,542,565]" italics="true" pageId="19" pageNumber="398">R. virgo</emphasis>
|
||
</taxonomicName>
|
||
. The two species show also a different position of nuptial tubercles on head (
|
||
<bibRefCitation id="62054B3DFFDFFFBB66E9FDAAFD9CFDED" author="Kottelat" box="[221,518,577,602]" pageId="19" pageNumber="398" refString="Kottelat, M. & Freyhof, J. (2007) Handbook of European Freshwater Fishes. Kottelat, Cornol, Switzerland and Frehyof, Berlin, Germany, 646 pp." type="book" year="2007">Kottelat & Freyhof, 2007</bibRefCitation>
|
||
) and also phylogenetically are very distant (Ketamaier
|
||
<emphasis id="34E0EADEFFDFFFBB62BDFDA9FB27FDEE" box="[1161,1213,577,602]" italics="true" pageId="19" pageNumber="398">et al</emphasis>
|
||
., 2008). Compared with
|
||
<taxonomicName id="C1944D4FFFDFFFBB66E5FD8DFEA0FDCA" box="[209,314,613,638]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="rutilus">
|
||
<emphasis id="34E0EADEFFDFFFBB66E5FD8DFEA0FDCA" box="[209,314,613,638]" italics="true" pageId="19" pageNumber="398">R. rutilus</emphasis>
|
||
</taxonomicName>
|
||
, has more scales on LL scales,
|
||
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|
||
<taxonomicName id="C1944D4FFFDFFFBB6524FD8DFCFFFDC9" box="[784,869,614,637]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="pigus">
|
||
<emphasis id="34E0EADEFFDFFFBB6524FD8DFCFFFDC9" box="[784,869,614,637]" italics="true" pageId="19" pageNumber="398">R pigus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
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|
||
<taxonomicName id="C1944D4FFFDFFFBB6225FD8DFBE0FDCA" box="[1041,1146,613,638]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="rutilus">
|
||
<emphasis id="34E0EADEFFDFFFBB6225FD8DFBE0FDCA" box="[1041,1146,613,638]" italics="true" pageId="19" pageNumber="398">R. rutilus</emphasis>
|
||
</taxonomicName>
|
||
; and less number of total GR,
|
||
<quantity id="C16C9B29FFDFFFBB66FFFD61FEBAFD16" box="[203,288,649,674]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="3.048" metricValueMin="2.286" pageId="19" pageNumber="398" unit="in" value="10.5" valueMax="12.0" valueMin="9.0">9–12 in</quantity>
|
||
<taxonomicName id="C1944D4FFFDFFFBB6712FD61FE1BFD15" box="[294,385,650,673]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="pigus">
|
||
<emphasis id="34E0EADEFFDFFFBB6712FD61FE1BFD15" box="[294,385,650,673]" italics="true" pageId="19" pageNumber="398">R. pigus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity id="C16C9B29FFDFFFBB6783FD62FD83FD16" box="[439,537,649,674]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="4.064" metricValueMin="3.048" pageId="19" pageNumber="398" unit="in" value="14.0" valueMax="16.0" valueMin="12.0">12–16 in</quantity>
|
||
<taxonomicName id="C1944D4FFFDFFFBB642BFD61FD1DFD16" box="[543,647,649,674]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="rutilus">
|
||
<emphasis id="34E0EADEFFDFFFBB642BFD61FD1DFD16" box="[543,647,649,674]" italics="true" pageId="19" pageNumber="398">R. rutilus</emphasis>
|
||
</taxonomicName>
|
||
. Compared to
|
||
<taxonomicName id="C1944D4FFFDFFFBB6518FD61FC45FD16" box="[812,991,649,674]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="stoumboudae" status="sp. nov.">
|
||
<emphasis id="34E0EADEFFDFFFBB6518FD61FC45FD16" box="[812,991,649,674]" italics="true" pageId="19" pageNumber="398">R. stoumboudae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2FD357A5FFDFFFBB65D2FD61FB87FD15" box="[998,1053,650,673]" pageId="19" pageNumber="398" rank="species">n.sp.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="C1944D4FFFDFFFBB621DFD61FB1EFD15" box="[1065,1156,650,673]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="pigus">
|
||
<emphasis id="34E0EADEFFDFFFBB621DFD61FB1EFD15" box="[1065,1156,650,673]" italics="true" pageId="19" pageNumber="398">R. pigus</emphasis>
|
||
</taxonomicName>
|
||
has less total GR (12–15 vs. 15–18) and more branched rays both in the dorsal and anal fins (9–11 vs. 8–9).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFDFFFBB66F3FD3AFEEEFB56" pageId="19" pageNumber="398" type="description">
|
||
<paragraph id="062B36CCFFDFFFBB66F3FD3AFEEEFB56" blockId="19.[151,1437,325,1646]" pageId="19" pageNumber="398">
|
||
<emphasis id="34E0EADEFFDFFFBB66F3FD3AFEC3FD5E" bold="true" box="[199,345,721,746]" pageId="19" pageNumber="398">Description.</emphasis>
|
||
A species of
|
||
<taxonomicName id="C1944D4FFFDFFFBB6439FD3AFDC6FD5E" box="[525,604,721,746]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="genus">
|
||
<emphasis id="34E0EADEFFDFFFBB6439FD3AFDC6FD5E" box="[525,604,721,746]" italics="true" pageId="19" pageNumber="398">Rutilus</emphasis>
|
||
</taxonomicName>
|
||
characterized by large size and the absence of mid lateral band. In reproductive males, well developed and large prominent nuptial tubercles on head, checks, and at center of the scales of the body. Absence, in reproductive males, of fan-shaped granular tubercles, on the free border of each scales of the body. Body laterally compressed, moderately elongate, dorsal profile convex; sometimes a small hump in largest specimens; snout pointed; eye diameter equal to the pre-orbital length; posterior mouth corner placed beyond the vertical crossing the anterior margin of orbital cavity; mouth opening oblique and terminal to slightly sub-terminal; lips smooth; eye small, its horizontal diameter less than or equal to the pre-orbital length; head length about 4.3–4.6 times in the SL; body quite deep, its depth about 3.2–3.4 times in the SL; origin of the D placed at same level of the insertion of the P2; free margin of dorsal and anal fins, from straight to slightly concave; caudal fin, when enlarged, moderately forked; LL complete, with 46–51 pored scales extending from the upper margin of opercular membrane, to the end of caudal peduncle; modally 3 un-branched and 10 branched rays in the dorsal and
|
||
<quantity id="C16C9B29FFDFFFBB6722FBB5FECBFBC2" box="[278,337,1117,1142]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="19" pageNumber="398" unit="in" value="11.0">11 in</quantity>
|
||
the anal fins; P1 with 1 un-branched and 15–17 branched rays; P2 with 1 un-branched ray and constantly 8 branched rays; about 8–12 total GR; color of the eye white; the color of all fins is from yellowish to dark grays; peritoneal membrane blackened by dense melanophores. For additional description and shape, see
|
||
<figureCitation id="9EAF2A49FFDFFFBB6344FB4EFF3EFB56" captionStart="FIGURE 2" captionStartId="4.[151,250,1514,1536]" captionTargetBox="[173,1424,552,1475]" captionTargetId="figure@4.[151,1436,519,1492]" captionTargetPageId="4" captionText="FIGURE 2. Native species of the Rutilus complex in southern Mediterranean Europe: A) Leucos aula, 115 mm SL, R. Bacchiglione, northern Italy, April 1996 (IZA 0425); B) Leucos basak, 95 mm SL, Lake Ohrid, FYROM, August 1987, (IZA 0429); C) Leucos ylikiensis, 120 mm SL, Lake Yliki, April 1987 (IZA 8733); D) Leucos panosi, 130 mm SL, canal to Lake Trichonis, May 1987 (IZA 8766); E) Leucos albus, 98 mm SL, River Moraca, Montenegro, November 1999 (IZA 0421); F) Sarmarutilus rubilio, 100 mm SL, River Ofanto, southern Italy, February 2002 (IZA 02183); G) Rutilus pigus, 230 mm SL, Lake Major, northern Italy, (IZA 0474); H) Holotype of Rutilus stoumboudae new species, 148 mm SL, October 1998, Lake Volvi, Greece (IZA 02107)." httpUri="https://zenodo.org/record/225780/files/figure.png" pageId="19" pageNumber="398">Fig. 2</figureCitation>
|
||
G, and Tables 4, 5.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFDFFFBB66F3FB06FC98FAFA" pageId="19" pageNumber="398" type="distribution">
|
||
<paragraph id="062B36CCFFDFFFBB66F3FB06FC98FAFA" blockId="19.[151,1437,325,1646]" pageId="19" pageNumber="398">
|
||
<emphasis id="34E0EADEFFDFFFBB66F3FB06FEFAFAB2" bold="true" box="[199,352,1261,1286]" pageId="19" pageNumber="398">Distribution.</emphasis>
|
||
Padano-Venetian district, from River Po to River Isonzo (upper Adriatic Sea drainages). Its survival is strongly affected by the introduction of two alien species: the European roach,
|
||
<taxonomicName id="C1944D4FFFDFFFBB62AAFAFAFAA5FA9E" box="[1182,1343,1297,1322]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="rutilus">
|
||
<emphasis id="34E0EADEFFDFFFBB62AAFAFAFAA5FA9E" box="[1182,1343,1297,1322]" italics="true" pageId="19" pageNumber="398">Rutilus rutilus</emphasis>
|
||
</taxonomicName>
|
||
and the European nase,
|
||
<taxonomicName id="C1944D4FFFDFFFBB677DFADEFD64FAFA" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[329,766,1333,1358]" class="Actinopterygii" family="Cyprinidae" genus="Chondrostoma" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="nasus">
|
||
<emphasis id="34E0EADEFFDFFFBB677DFADEFDAFFAF9" box="[329,565,1333,1358]" italics="true" pageId="19" pageNumber="398">Chondrostoma nasus</emphasis>
|
||
(Linnaeus, 1758)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFDFFFBB66F3FAB2FF65F94A" pageId="19" pageNumber="398" type="discussion">
|
||
<paragraph id="062B36CCFFDFFFBB66F3FAB2FB19F9DA" blockId="19.[151,1437,325,1646]" pageId="19" pageNumber="398">
|
||
<emphasis id="34E0EADEFFDFFFBB66F3FAB2FD66FAC6" bold="true" box="[199,764,1369,1394]" pageId="19" pageNumber="398">Remarks on ecology, biology and conservation.</emphasis>
|
||
A mostly lacustrine adapted species, which may reach
|
||
<quantity id="C16C9B29FFDFFFBB6346FAB2FF59FA21" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="398" unit="mm" value="500.0">500 mm</quantity>
|
||
TL and
|
||
<quantity id="C16C9B29FFDFFFBB6713FA95FEC3FA22" box="[295,345,1405,1430]" metricMagnitude="0" metricUnit="kg" metricValue="2.0" pageId="19" pageNumber="398" unit="kg" value="2.0">2 kg</quantity>
|
||
of weight, and at least 9 years of age. Sexual maturity is late and is reached at age 3+ or 4+ in males and 5+ in females at about
|
||
<quantity id="C16C9B29FFDFFFBB6426FA49FD3DFA0D" box="[530,679,1442,1466]" metricMagnitude="-1" metricUnit="m" metricValue="3.05" metricValueMax="3.3" metricValueMin="2.8" pageId="19" pageNumber="398" unit="mm" value="305.0" valueMax="330.0" valueMin="280.0">280–330 mm</quantity>
|
||
SL. Reproduction takes place between April and May, longevity is up to 9 years (
|
||
<bibRefCitation id="62054B3DFFDFFFBB6772FA2EFD10FA6A" author="Bianco" box="[326,650,1477,1502]" pageId="19" pageNumber="398" refString="Bianco, P. G. & Taraborelli, T. (1985) Contributo alla conoscenza del genere Rutilus R. in Italia e nei balcani occidentali (Pisces, Cyprinidae). Bollettino del Museo Regionale di Scienze naturali di Torino, 3, 131 - 172." type="journal article" year="1985">Bianco & Taraborelli, 1985</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="62054B3DFFDFFFBB64AEFA2EFCF5FA6A" author="Puzzi" box="[666,879,1477,1502]" pageId="19" pageNumber="398" refString="Puzzi, C. M., Trasforini, S., Bardazzi, M. A., Polisciano, N., Montonati, S. & Maggio, A. (2007) Carta Provinciale Delle Vocazioni Ittiche della Provincia Di Milano. Servizio Gestione Attivita Venatoria e Piscatoria, Milano, 447 pp." type="book" year="2007">
|
||
Puzzi
|
||
<emphasis id="34E0EADEFFDFFFBB64D7FA2DFC8DFA6A" box="[739,791,1477,1502]" italics="true" pageId="19" pageNumber="398">et al</emphasis>
|
||
., 2007
|
||
</bibRefCitation>
|
||
). Although the species was placed in the “Low Concern” category by (IUCN, 2013), according to the recent assessment made by the Italian National IUCN red list commission,
|
||
<taxonomicName id="C1944D4FFFDFFFBB6704F9E5FE0BF991" box="[304,401,1550,1573]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="pigus">
|
||
<emphasis id="34E0EADEFFDFFFBB6704F9E5FE0BF991" box="[304,401,1550,1573]" italics="true" pageId="19" pageNumber="398">R. pigus</emphasis>
|
||
</taxonomicName>
|
||
was placed in the “Critical Endangered” category (
|
||
<bibRefCitation id="62054B3DFFDFFFBB65C6F9E6FB43F992" author="Bianco" box="[1010,1241,1549,1574]" pageId="19" pageNumber="398" refString="Bianco, P. G., Caputo, V., Ferrito, V., Lorenzoni, M., Nonnis Marzano, F., Stefani, F., Sabatini, A. & Tancioni, L. (2013) Pesci d'acqua dolce. In: Rondinini, C., Battistoni, A., Peronace, V. & Teofili, C. (Compilers.), Lista Rossa IUCN dei Vertebrati Italiani. Comitato Italiano IUCN e Ministero dell'Ambiente e della Tutela del Territorio e del Mare, Roma, pp. 54." type="book chapter" year="2013">
|
||
Bianco
|
||
<emphasis id="34E0EADEFFDFFFBB627AF9E5FB18F992" box="[1102,1154,1549,1574]" italics="true" pageId="19" pageNumber="398">et al</emphasis>
|
||
., 2013
|
||
</bibRefCitation>
|
||
). This species is progressively disappearing throughout its native range, mainly as a result of competition with the invasive introduced
|
||
<taxonomicName id="C1944D4FFFDFFFBB6721F9BDFEE7F9DA" box="[277,381,1621,1646]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="rutilus">
|
||
<emphasis id="34E0EADEFFDFFFBB6721F9BDFEE7F9DA" box="[277,381,1621,1646]" italics="true" pageId="19" pageNumber="398">R. rutilus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C1944D4FFFDFFFBB6780F9BEFD3BF9D9" box="[436,673,1621,1646]" class="Actinopterygii" family="Cyprinidae" genus="Chondrostoma" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="nasus">
|
||
<emphasis id="34E0EADEFFDFFFBB6780F9BEFD3BF9D9" box="[436,673,1621,1646]" italics="true" pageId="19" pageNumber="398">Chondrostoma nasus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="62054B3DFFDFFFBB649BF9BEFCEDF9DA" author="Puzzi" box="[687,887,1621,1646]" pageId="19" pageNumber="398" refString="Puzzi, C. M., Trasforini, S., Bardazzi, M. A., Polisciano, N., Montonati, S. & Maggio, A. (2007) Carta Provinciale Delle Vocazioni Ittiche della Provincia Di Milano. Servizio Gestione Attivita Venatoria e Piscatoria, Milano, 447 pp." type="book" year="2007">
|
||
Puzzi
|
||
<emphasis id="34E0EADEFFDFFFBB64C2F9BDFCB4F9DA" box="[758,814,1621,1646]" italics="true" pageId="19" pageNumber="398">et al.</emphasis>
|
||
, 2007
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="62054B3DFFDFFFBB65B7F9BEFBEDF9DA" author="Volta" box="[899,1143,1621,1646]" pageId="19" pageNumber="398" refString="Volta, P. & Jepsen, N. (2008) The recent invasion of Rutilus rutilus (L.), (Pisces: Cyprinidae) in a large South-Alpine lake: Lago Maggiore. Journal of Limnology, 67, 163 - 170." type="journal article" year="2008">Volta & Jepsen, 2008</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="062B36CCFFDFFFBB66A3F92AFF65F94A" blockId="19.[151,786,1729,1790]" pageId="19" pageNumber="398">
|
||
<taxonomicName id="C1944D4FFFDFFFBB66A3F92AFDE1F96E" authority="Bianco & Ketmaier" authorityName="Bianco & Ketmaier" box="[151,635,1729,1754]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="stoumboudae" status="sp. nov.">
|
||
<emphasis id="34E0EADEFFDFFFBB66A3F92AFDE1F96E" bold="true" box="[151,635,1729,1754]" pageId="19" pageNumber="398">
|
||
<emphasis id="34E0EADEFFDFFFBB66A3F92AFE16F96E" bold="true" box="[151,396,1729,1754]" italics="true" pageId="19" pageNumber="398">Rutilus stoumboudae</emphasis>
|
||
Bianco & Ketmaier
|
||
</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="34E0EADEFFDFFFBB64BDF92AFC88F96E" bold="true" box="[649,786,1729,1754]" pageId="19" pageNumber="398">
|
||
<taxonomicNameLabel id="2FD357A5FFDFFFBB64BDF92AFC88F96E" box="[649,786,1729,1754]" pageId="19" pageNumber="398" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="9EAF2A49FFDFFFBB66ABF90EFF78F94A" box="[159,226,1765,1790]" captionStart="FIGURE 2" captionStartId="4.[151,250,1514,1536]" captionTargetBox="[173,1424,552,1475]" captionTargetId="figure@4.[151,1436,519,1492]" captionTargetPageId="4" captionText="FIGURE 2. Native species of the Rutilus complex in southern Mediterranean Europe: A) Leucos aula, 115 mm SL, R. Bacchiglione, northern Italy, April 1996 (IZA 0425); B) Leucos basak, 95 mm SL, Lake Ohrid, FYROM, August 1987, (IZA 0429); C) Leucos ylikiensis, 120 mm SL, Lake Yliki, April 1987 (IZA 8733); D) Leucos panosi, 130 mm SL, canal to Lake Trichonis, May 1987 (IZA 8766); E) Leucos albus, 98 mm SL, River Moraca, Montenegro, November 1999 (IZA 0421); F) Sarmarutilus rubilio, 100 mm SL, River Ofanto, southern Italy, February 2002 (IZA 02183); G) Rutilus pigus, 230 mm SL, Lake Major, northern Italy, (IZA 0474); H) Holotype of Rutilus stoumboudae new species, 148 mm SL, October 1998, Lake Volvi, Greece (IZA 02107)." httpUri="https://zenodo.org/record/225780/files/figure.png" pageId="19" pageNumber="398">Fig. 2</figureCitation>
|
||
H)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFDFFFBB66A3F8C6FB4AF83A" pageId="19" pageNumber="398" type="materials_examined">
|
||
<paragraph id="062B36CCFFDFFFBB66A3F8C6FC3AF8DE" blockId="19.[151,1436,1837,2006]" pageId="19" pageNumber="398">
|
||
<emphasis id="34E0EADEFFDFFFBB66A3F8C6FECAF8F2" bold="true" box="[151,336,1837,1862]" pageId="19" pageNumber="398">
|
||
<typeStatus id="D92F886EFFDFFFBB66A3F8C6FF48F8F2" box="[151,210,1837,1862]" pageId="19" pageNumber="398">Type</typeStatus>
|
||
materials.
|
||
</emphasis>
|
||
<typeStatus id="D92F886EFFDFFFBB6763F8C6FE25F8F2" box="[343,447,1837,1862]" pageId="19" pageNumber="398" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<figureCitation id="9EAF2A49FFDFFFBB67FBF8C6FD8BF8F2" box="[463,529,1837,1862]" captionStart="FIGURE 2" captionStartId="4.[151,250,1514,1536]" captionTargetBox="[173,1424,552,1475]" captionTargetId="figure@4.[151,1436,519,1492]" captionTargetPageId="4" captionText="FIGURE 2. Native species of the Rutilus complex in southern Mediterranean Europe: A) Leucos aula, 115 mm SL, R. Bacchiglione, northern Italy, April 1996 (IZA 0425); B) Leucos basak, 95 mm SL, Lake Ohrid, FYROM, August 1987, (IZA 0429); C) Leucos ylikiensis, 120 mm SL, Lake Yliki, April 1987 (IZA 8733); D) Leucos panosi, 130 mm SL, canal to Lake Trichonis, May 1987 (IZA 8766); E) Leucos albus, 98 mm SL, River Moraca, Montenegro, November 1999 (IZA 0421); F) Sarmarutilus rubilio, 100 mm SL, River Ofanto, southern Italy, February 2002 (IZA 02183); G) Rutilus pigus, 230 mm SL, Lake Major, northern Italy, (IZA 0474); H) Holotype of Rutilus stoumboudae new species, 148 mm SL, October 1998, Lake Volvi, Greece (IZA 02107)." httpUri="https://zenodo.org/record/225780/files/figure.png" pageId="19" pageNumber="398">Fig. 2</figureCitation>
|
||
H)
|
||
<collectionCode id="6085AE09FFDFFFBB6402F8C5FDFFF8F2" box="[566,613,1838,1862]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="19" pageNumber="398">IZA</collectionCode>
|
||
02107A,
|
||
<quantity id="C16C9B29FFDFFFBB64E1F8C5FCA8F8F1" box="[725,818,1838,1862]" metricMagnitude="-1" metricUnit="m" metricValue="1.28" pageId="19" pageNumber="398" unit="mm" value="128.0">128 mm</quantity>
|
||
<collectionCode id="6085AE09FFDFFFBB650DF8C5FCC6F8F2" box="[825,860,1838,1862]" pageId="19" pageNumber="398">SL</collectionCode>
|
||
,
|
||
<collectingCountry id="7E83765CFFDFFFBB655CF8C5FC20F8F2" box="[872,954,1838,1862]" name="Greece" pageId="19" pageNumber="398">Greece</collectingCountry>
|
||
, Lake Volvi, P.G. Bianco,
|
||
<date id="722A100CFFDFFFBB62EBF8C5FA06F8F2" box="[1247,1436,1837,1862]" pageId="19" pageNumber="398" value="1998-10-21">21 October 1998</date>
|
||
(specimen also used for molecular analysis by
|
||
<bibRefCitation id="62054B3DFFDFFFBB6496F8BAFC0EF8DE" author="Ketmaier" box="[674,916,1873,1898]" pageId="19" pageNumber="398" refString="Ketmaier, V., Bianco, P. G. & Durand, J. D. (2008) Molecular systematics, phylogeny and biogeography of roaches (Rutilus, Teleostei, Cyprinidae). Molecular Phylogenetics and Evolution, 49, 362 - 367. http: // dx. doi. org / 10.1016 / j. ympev. 2008.07.012" type="journal article" year="2008">
|
||
Ketmaier
|
||
<emphasis id="34E0EADEFFDFFFBB6525F8B9FCCAF8DE" box="[785,848,1873,1898]" italics="true" pageId="19" pageNumber="398">et al.,</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="062B36CCFFDFFFBB66A3F89DFB4AF83A" blockId="19.[151,1436,1837,2006]" box="[151,1232,1909,1934]" pageId="19" pageNumber="398">
|
||
<typeStatus id="D92F886EFFDFFFBB66A3F89DFE9EF83A" box="[151,260,1910,1934]" pageId="19" pageNumber="398" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<collectionCode id="6085AE09FFDFFFBB6720F89DFED9F83A" box="[276,323,1910,1934]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="19" pageNumber="398">IZA</collectionCode>
|
||
02107B, 11, 64–
|
||
<quantity id="C16C9B29FFDFFFBB6430F89EFDFAF839" box="[516,608,1909,1934]" metricMagnitude="-1" metricUnit="m" metricValue="1.56" pageId="19" pageNumber="398" unit="mm" value="156.0">156 mm</quantity>
|
||
<collectionCode id="6085AE09FFDFFFBB6453F89DFD10F83A" box="[615,650,1910,1934]" pageId="19" pageNumber="398">SL</collectionCode>
|
||
,
|
||
<collectingCountry id="7E83765CFFDFFFBB64A2F89DFD72F83A" box="[662,744,1910,1934]" name="Greece" pageId="19" pageNumber="398">Greece</collectingCountry>
|
||
, Lake Volvi; P.G. Bianco,
|
||
<date id="722A100CFFDFFFBB6238F89DFB56F83A" box="[1036,1228,1909,1934]" pageId="19" pageNumber="398" value="1998-10-10">10 October 1998</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFDFFFBC66F3F872FC3CFF60" lastPageId="20" lastPageNumber="399" pageId="19" pageNumber="398" type="diagnosis">
|
||
<paragraph id="062B36CCFFDFFFBC66F3F872FC3CFF60" blockId="19.[151,1436,1837,2006]" lastBlockId="20.[151,1437,151,1871]" lastPageId="20" lastPageNumber="399" pageId="19" pageNumber="398">
|
||
<emphasis id="34E0EADEFFDFFFBB66F3F872FEDBF806" bold="true" box="[199,321,1945,1970]" pageId="19" pageNumber="398">Diagnosis.</emphasis>
|
||
<taxonomicName id="C1944D4FFFDFFFBB677DF872FDACF806" box="[329,566,1945,1970]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="stoumboudae">
|
||
<emphasis id="34E0EADEFFDFFFBB677DF872FDACF806" box="[329,566,1945,1970]" italics="true" pageId="19" pageNumber="398">Rutilus stoumboudae</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<emphasis id="34E0EADEFFDFFFBB64FAF871FCA5F806" box="[718,831,1945,1970]" italics="true" pageId="19" pageNumber="398">
|
||
<taxonomicName id="C1944D4FFFDFFFBB64FAF871FCA1F806" box="[718,827,1945,1970]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="rutilus">R. rutilus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
its closest relative genetically, both in the number of total GR (16–17 vs. 13–14) and number of branched rays of the D and A (modally 9 and
|
||
<quantity id="C16C9B29FFDFFFBB62B1F855FB2AF862" box="[1157,1200,1981,2006]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="19" pageNumber="398" unit="in" value="9.0">9 in</quantity>
|
||
<taxonomicName id="C1944D4FFFDFFFBB6283F855FAF6F862" box="[1207,1388,1981,2006]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="398" phylum="Chordata" rank="species" species="stoumboudae">
|
||
<emphasis id="34E0EADEFFDFFFBB6283F855FAF6F862" box="[1207,1388,1981,2006]" italics="true" pageId="19" pageNumber="398">R. stoumboudae</emphasis>
|
||
</taxonomicName>
|
||
and 10 and
|
||
<quantity id="C16C9B29FFD8FFBC66DDFF73FEBBFF04" box="[233,289,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="399" unit="in" value="10.0">10 in</quantity>
|
||
<taxonomicName id="C1944D4FFFD8FFBC6713FF72FE15FF04" box="[295,399,151,176]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="rutilus">
|
||
<emphasis id="34E0EADEFFD8FFBC6713FF72FE15FF04" box="[295,399,151,176]" italics="true" pageId="20" pageNumber="399">R. rutilus</emphasis>
|
||
</taxonomicName>
|
||
). It differs from
|
||
<taxonomicName id="C1944D4FFFD8FFBC647EFF72FD3FFF04" box="[586,677,153,176]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="pigus">
|
||
<emphasis id="34E0EADEFFD8FFBC647EFF72FD3FFF04" box="[586,677,153,176]" italics="true" pageId="20" pageNumber="399">R. pigus</emphasis>
|
||
</taxonomicName>
|
||
in apparently lacking nuptial tubercles, in the numbers of LL scales (39–43 vs. 46–51) and of branched rays in the anal fin (8–9 vs. 9–11).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFD8FFBC66F3FF34FA0FFD1C" pageId="20" pageNumber="399" type="description">
|
||
<paragraph id="062B36CCFFD8FFBC66F3FF34FA0FFD1C" blockId="20.[151,1437,151,1871]" pageId="20" pageNumber="399">
|
||
<emphasis id="34E0EADEFFD8FFBC66F3FF34FEC3FF4C" bold="true" box="[199,345,223,248]" pageId="20" pageNumber="399">Description.</emphasis>
|
||
Apparently a small to medium sized species. Maximum SL observed,
|
||
<quantity id="C16C9B29FFD8FFBC625DFF34FB5CFF43" box="[1129,1222,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="1.56" pageId="20" pageNumber="399" unit="mm" value="156.0">156 mm</quantity>
|
||
and
|
||
<quantity id="C16C9B29FFD8FFBC62CEFF0BFACDFF43" box="[1274,1367,224,248]" metricMagnitude="-1" metricUnit="m" metricValue="1.97" pageId="20" pageNumber="399" unit="mm" value="197.0">197 mm</quantity>
|
||
of TL in our materials. Body moderately compressed, its depth about 3.1–3.3 times in SL. Head length about 3.8–4.0 times in SL. Mouth, small, terminal and slight up-turned. Pre-orbital distance less than eye diameter. Eye diameter about 2.7–3.3 times in head length. Origin of D slight behind the origin of P2. Free margins of D and A concave; dorsal profile convex, slightly humped in the
|
||
<typeStatus id="D92F886EFFD8FFBC64FCFE84FCB7FE3C" box="[712,813,367,392]" pageId="20" pageNumber="399" type="holotype">holotype</typeStatus>
|
||
; pre-dorsal distance slightly longer than pre-ventral distance. Pectoral-pelvic distance nearly equal to pelvic-anal distance. Caudal peduncle depth about 1.1–1.3 times in caudal peduncle length. Free margin of pectoral and pelvic fins rounded, caudal fin, when extended, moderately forked. D with 3 non-branched and 9 (in one case 8) branched rays. A with 3 non-branched and 9 branched rays. Fourteen circum-peduncular scales; 38–41 pored scales on the LL. 7.5 or 8.5 rows of scales above and 3.5 or 4.0 below the LL; 8–10 total GR; pharyngeal teeth on both sides, 6-5; Peritoneal membrane darkened by dense melanophores; apparently absence of tubercles. Ground color silvery; back plain dark brown. Top of head plain dark brown; sides light; ventral surface of head, belly and caudal peduncle yellowish. All fins yellow-brownish. General shape of
|
||
<typeStatus id="D92F886EFFD8FFBC676EFD64FE26FD1C" box="[346,444,655,680]" pageId="20" pageNumber="399" type="holotype">holotype</typeStatus>
|
||
in
|
||
<figureCitation id="9EAF2A49FFD8FFBC67EBFD64FDBBFD1C" box="[479,545,655,680]" captionStart="FIGURE 2" captionStartId="4.[151,250,1514,1536]" captionTargetBox="[173,1424,552,1475]" captionTargetId="figure@4.[151,1436,519,1492]" captionTargetPageId="4" captionText="FIGURE 2. Native species of the Rutilus complex in southern Mediterranean Europe: A) Leucos aula, 115 mm SL, R. Bacchiglione, northern Italy, April 1996 (IZA 0425); B) Leucos basak, 95 mm SL, Lake Ohrid, FYROM, August 1987, (IZA 0429); C) Leucos ylikiensis, 120 mm SL, Lake Yliki, April 1987 (IZA 8733); D) Leucos panosi, 130 mm SL, canal to Lake Trichonis, May 1987 (IZA 8766); E) Leucos albus, 98 mm SL, River Moraca, Montenegro, November 1999 (IZA 0421); F) Sarmarutilus rubilio, 100 mm SL, River Ofanto, southern Italy, February 2002 (IZA 02183); G) Rutilus pigus, 230 mm SL, Lake Major, northern Italy, (IZA 0474); H) Holotype of Rutilus stoumboudae new species, 148 mm SL, October 1998, Lake Volvi, Greece (IZA 02107)." httpUri="https://zenodo.org/record/225780/files/figure.png" pageId="20" pageNumber="399">Fig. 2</figureCitation>
|
||
H. Morphometric and meristic data of
|
||
<typeStatus id="D92F886EFFD8FFBC65FAFD64FBAAFD1C" box="[974,1072,655,680]" pageId="20" pageNumber="399" type="holotype">holotype</typeStatus>
|
||
and 11
|
||
<typeStatus id="D92F886EFFD8FFBC62B3FD7BFB68FD1C" box="[1159,1266,656,680]" pageId="20" pageNumber="399" type="paratype">paratypes</typeStatus>
|
||
in Tables 4, 5.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFD8FFBC66F3FD5FFBCAFA84" pageId="20" pageNumber="399" type="discussion">
|
||
<paragraph id="062B36CCFFD8FFBC66F3FD5FFECEFB70" blockId="20.[151,1437,151,1871]" pageId="20" pageNumber="399">
|
||
<emphasis id="34E0EADEFFD8FFBC66F3FD5FFEA1FD79" bold="true" box="[199,315,692,717]" pageId="20" pageNumber="399">Remarks.</emphasis>
|
||
According to
|
||
<bibRefCitation id="62054B3DFFD8FFBC67D7FD5EFD63FD78" box="[483,761,692,717]" pageId="20" pageNumber="399" refString="Larmuseau, M. H. D., Frehyof, J., Volkaert, F. A. M. & Houdt, J. K. J. van (2009) Matrilinear phylogeography and demographical patterns of Rutilus rutilus: implications for taxonomy and conservation. Journal of Fish Biology, 75, 332 - 353. http: // dx. doi. org / 10.1111 / j. 1095 - 8649.2009.02322. x" type="journal article">
|
||
Larmuseau
|
||
<emphasis id="34E0EADEFFD8FFBC645DFD5EFD3EFD79" box="[617,676,692,717]" italics="true" pageId="20" pageNumber="399">et al.</emphasis>
|
||
(2009)
|
||
</bibRefCitation>
|
||
, the species from Lake Volvi examined by
|
||
<bibRefCitation id="62054B3DFFD8FFBC62C4FD5FFF7EFD44" pageId="20" pageNumber="399" refString="Ketmaier, V., Bianco, P. G. & Durand, J. D. (2008) Molecular systematics, phylogeny and biogeography of roaches (Rutilus, Teleostei, Cyprinidae). Molecular Phylogenetics and Evolution, 49, 362 - 367. http: // dx. doi. org / 10.1016 / j. ympev. 2008.07.012" type="journal article">
|
||
Ketmaier
|
||
<emphasis id="34E0EADEFFD8FFBC6355FD5EFA06FD79" box="[1377,1436,692,717]" italics="true" pageId="20" pageNumber="399">et al.</emphasis>
|
||
(2008)
|
||
</bibRefCitation>
|
||
, should be identified as
|
||
<taxonomicName id="C1944D4FFFD8FFBC6421FD3CFD5BFD44" box="[533,705,727,752]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="heckeli">
|
||
<emphasis id="34E0EADEFFD8FFBC6421FD3CFD5BFD44" box="[533,705,727,752]" italics="true" pageId="20" pageNumber="399">Rutilus heckeli</emphasis>
|
||
</taxonomicName>
|
||
. But these authors specifically stated in their study that the identification of the alleged
|
||
<taxonomicName id="C1944D4FFFD8FFBC67D7FD17FD16FCA1" box="[483,652,764,789]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="heckeli">
|
||
<emphasis id="34E0EADEFFD8FFBC67D7FD17FD16FCA1" box="[483,652,764,789]" italics="true" pageId="20" pageNumber="399">Rutilus heckeli</emphasis>
|
||
</taxonomicName>
|
||
material they used was uncertain: the authors did not deposite the corresponding sequences in GenBank and we are thus unable to include their data in our phylogenetic analyses. According to
|
||
<bibRefCitation id="62054B3DFFD8FFBC6705FCAEFDDBFCE8" box="[305,577,836,861]" pageId="20" pageNumber="399" refString="Larmuseau, M. H. D., Frehyof, J., Volkaert, F. A. M. & Houdt, J. K. J. van (2009) Matrilinear phylogeography and demographical patterns of Rutilus rutilus: implications for taxonomy and conservation. Journal of Fish Biology, 75, 332 - 353. http: // dx. doi. org / 10.1111 / j. 1095 - 8649.2009.02322. x" type="journal article">
|
||
Larmuseau
|
||
<emphasis id="34E0EADEFFD8FFBC6780FCAEFE77FCE9" box="[436,493,836,861]" italics="true" pageId="20" pageNumber="399">et al.</emphasis>
|
||
(2009)
|
||
</bibRefCitation>
|
||
,
|
||
<taxonomicName id="C1944D4FFFD8FFBC6479FCAFFD68FCE9" box="[589,754,836,861]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="heckeli">
|
||
<emphasis id="34E0EADEFFD8FFBC6479FCAFFD68FCE9" box="[589,754,836,861]" italics="true" pageId="20" pageNumber="399">Rutilus heckeli</emphasis>
|
||
</taxonomicName>
|
||
seems the sister species of
|
||
<taxonomicName id="C1944D4FFFD8FFBC6213FCAEFB0AFCE9" box="[1063,1168,836,861]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="rutilus">
|
||
<emphasis id="34E0EADEFFD8FFBC6213FCAEFB0AFCE9" box="[1063,1168,836,861]" italics="true" pageId="20" pageNumber="399">R. rutilus</emphasis>
|
||
</taxonomicName>
|
||
. Extensive diagnosis of this species is found in
|
||
<bibRefCitation id="62054B3DFFD8FFBC6786FC83FDA5FC34" author="Berg" box="[434,575,872,896]" pageId="20" pageNumber="399" refString="Berg, L. (1949) Freshwater fish of the USSR and adjacent countries. Israel Program for Scientific Translation (1964). Vol. 2. Jerusalem, 496 pp." type="book" year="1949">Berg (1949)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="62054B3DFFD8FFBC64B7FC83FCC8FC34" author="Banarescu" box="[643,850,871,896]" pageId="20" pageNumber="399" refString="Banarescu, P. (1964) Fauna Republicii populare Romine. Pisces, Osteichthyies. Vol. 13. Bucuresti, 959 pp." type="book" year="1964">Banarescu (1964)</bibRefCitation>
|
||
: it may reach the size of about
|
||
<quantity id="C16C9B29FFD8FFBC62FAFC8CFAB5FCCB" box="[1230,1327,871,896]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="399" unit="mm" value="500.0">500 mm</quantity>
|
||
TL (size unknown in Lake Volvi, since
|
||
<taxonomicName id="C1944D4FFFD8FFBC67C5FC67FD47FC11" box="[497,733,908,933]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="stoumboudae">
|
||
<emphasis id="34E0EADEFFD8FFBC67C5FC67FD47FC11" box="[497,733,908,933]" italics="true" pageId="20" pageNumber="399">Rutilus stoumboudae</emphasis>
|
||
</taxonomicName>
|
||
belongs to a group of species of smaller size). Adult males of
|
||
<taxonomicName id="C1944D4FFFD8FFBC66A3FC44FEA6FC7C" box="[151,316,943,968]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="heckeli">
|
||
<emphasis id="34E0EADEFFD8FFBC66A3FC44FEA6FC7C" box="[151,316,943,968]" italics="true" pageId="20" pageNumber="399">Rutilus heckeli</emphasis>
|
||
</taxonomicName>
|
||
show well developed nuptial tubercles all along the dorsal area of the body (apparently not found in
|
||
<taxonomicName id="C1944D4FFFD8FFBC66A3FC3EFED6FC59" box="[151,332,980,1005]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="stoumboudae">
|
||
<emphasis id="34E0EADEFFD8FFBC66A3FC3EFED6FC59" box="[151,332,980,1005]" italics="true" pageId="20" pageNumber="399">R. stoumboudae</emphasis>
|
||
</taxonomicName>
|
||
); pharyngeal teeth formula is modally 6-
|
||
<quantity id="C16C9B29FFD8FFBC6516FC3FFCD5FC59" box="[802,847,980,1005]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="20" pageNumber="399" unit="in" value="6.0">6 in</quantity>
|
||
<taxonomicName id="C1944D4FFFD8FFBC6563FC3EFC5DFC59" box="[855,967,980,1005]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="heckeli">
|
||
<emphasis id="34E0EADEFFD8FFBC6563FC3EFC5DFC59" box="[855,967,980,1005]" italics="true" pageId="20" pageNumber="399">R. heckeli</emphasis>
|
||
</taxonomicName>
|
||
, while in our new specie is modally 6-5; the number branched rays of dorsal and anal fins are modally
|
||
<quantity id="C16C9B29FFD8FFBC6564FC13FC10FBA4" box="[848,906,1015,1040]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="399" unit="in" value="10.0">10 in</quantity>
|
||
<taxonomicName id="C1944D4FFFD8FFBC65A5FC12FB9AFBA4" box="[913,1024,1015,1040]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="heckeli">
|
||
<emphasis id="34E0EADEFFD8FFBC65A5FC12FB9AFBA4" box="[913,1024,1015,1040]" italics="true" pageId="20" pageNumber="399">R. heckeli</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<quantity id="C16C9B29FFD8FFBC620AFC13FBF0FBA4" box="[1086,1130,1015,1040]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="20" pageNumber="399" unit="in" value="9.0">9 in</quantity>
|
||
<taxonomicName id="C1944D4FFFD8FFBC6245FC12FABFFBA4" box="[1137,1317,1015,1040]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="stoumboudae">
|
||
<emphasis id="34E0EADEFFD8FFBC6245FC12FABFFBA4" box="[1137,1317,1015,1040]" italics="true" pageId="20" pageNumber="399">R. stoumboudae</emphasis>
|
||
</taxonomicName>
|
||
. Based on these differences we cannot identify samples of the new species as
|
||
<taxonomicName id="C1944D4FFFD8FFBC65ACFBF7FBA5FB81" box="[920,1087,1052,1077]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="heckeli">
|
||
<emphasis id="34E0EADEFFD8FFBC65ACFBF7FBA5FB81" box="[920,1087,1052,1077]" italics="true" pageId="20" pageNumber="399">Rutilus heckeli</emphasis>
|
||
</taxonomicName>
|
||
. Another geographically close species, described for the River
|
||
<taxonomicName id="C1944D4FFFD8FFBC6417FBD4FDE0FBEC" box="[547,634,1087,1112]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="maritza">Maritza</taxonomicName>
|
||
in
|
||
<collectingCountry id="7E83765CFFD8FFBC6498FBD4FC96FBEC" box="[684,780,1087,1112]" name="Bulgaria" pageId="20" pageNumber="399">Bulgaria</collectingCountry>
|
||
(Evros in
|
||
<collectingCountry id="7E83765CFFD8FFBC65ACFBABFC76FBEC" box="[920,1004,1088,1112]" name="Greece" pageId="20" pageNumber="399">Greece</collectingCountry>
|
||
),
|
||
<taxonomicName id="C1944D4FFFD8FFBC6231FBD4FAEDFBEC" authority="Drenski, 1926" authorityName="Drenski" authorityYear="1926" box="[1029,1399,1087,1112]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="maritza">
|
||
<emphasis id="34E0EADEFFD8FFBC6231FBD4FB22FBEC" box="[1029,1208,1087,1112]" italics="true" pageId="20" pageNumber="399">Rutilus maritza</emphasis>
|
||
, Drenski, 1926
|
||
</taxonomicName>
|
||
, is considered since a long time as a junior synonym of
|
||
<taxonomicName id="C1944D4FFFD8FFBC6515FB8EFBC3FBC8" authority="Kottelat, 1997" authorityName="Kottelat" authorityYear="1997" box="[801,1113,1124,1149]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="rutilus">
|
||
<emphasis id="34E0EADEFFD8FFBC6515FB8EFC15FBC9" box="[801,911,1124,1149]" italics="true" pageId="20" pageNumber="399">R. rutilus</emphasis>
|
||
(
|
||
<bibRefCitation id="62054B3DFFD8FFBC6590FB8FFBCBFBC8" author="Kottelat" box="[932,1105,1124,1149]" pageId="20" pageNumber="399" refString="Kottelat, M. (1997) European freshwater fish. Biologia, 52 (Supplement), 1 - 271." type="journal article" year="1997">Kottelat, 1997</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. In addition, the Bulgarian freshwater fish fauna is of Danubian origin, with the exception of a single endemic species,
|
||
<taxonomicName id="C1944D4FFFD8FFBC6299FB6CFED5FB70" authority="Drensky, 1943" authorityName="Drensky" authorityYear="1943" class="Actinopterygii" family="Cyprinidae" genus="Alburnus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="mandrensis">
|
||
<emphasis id="34E0EADEFFD8FFBC6299FB6CFA06FB14" box="[1197,1436,1159,1184]" italics="true" pageId="20" pageNumber="399">Alburnus mandrensis</emphasis>
|
||
(Drensky, 1943)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="062B36CCFFD8FFBC66F3FB24FBCAFA84" blockId="20.[151,1437,151,1871]" pageId="20" pageNumber="399">
|
||
<emphasis id="34E0EADEFFD8FFBC66F3FB24FEE8FB5C" bold="true" box="[199,370,1231,1256]" pageId="20" pageNumber="399">Color pattern.</emphasis>
|
||
In preserved specimens, dorsum brownish, sides yellowish without a longitudinal band. Scales of flanks show a reticulate pigmentation. Peritoneal membrane with more or less concentrated or scattered melanophores. Pectoral, dorsal, and caudal fins black; pelvic and anal fins yellowish.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFD8FFBC66F3FAD7FD53F9C1" pageId="20" pageNumber="399" type="distribution">
|
||
<paragraph id="062B36CCFFD8FFBC66F3FAD7FD59F9BC" blockId="20.[151,1437,151,1871]" pageId="20" pageNumber="399">
|
||
<emphasis id="34E0EADEFFD8FFBC66F3FAD7FEC3FAE1" bold="true" box="[199,345,1340,1365]" pageId="20" pageNumber="399">Distribution</emphasis>
|
||
: There are several doubts about the native origin of this species in Lake Volvi.
|
||
<bibRefCitation id="62054B3DFFD8FFBC62F3FAD7FF7BFACC" author="Kottelat" pageId="20" pageNumber="399" refString="Kottelat, M. & Freyhof, J. (2007) Handbook of European Freshwater Fishes. Kottelat, Cornol, Switzerland and Frehyof, Berlin, Germany, 646 pp." type="book" year="2007">Kottelat & Frehyof (2007)</bibRefCitation>
|
||
reported a
|
||
<emphasis id="34E0EADEFFD8FFBC6752FAB4FE42FACC" box="[358,472,1375,1400]" italics="true" pageId="20" pageNumber="399">
|
||
<taxonomicName id="C1944D4FFFD8FFBC6752FAB4FE2FFACC" box="[358,437,1375,1400]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="genus">Rutilus</taxonomicName>
|
||
sp
|
||
</emphasis>
|
||
. from River Sperchios (south-eastern
|
||
<collectingCountry id="7E83765CFFD8FFBC65BAFA8BFC79FACC" box="[910,995,1376,1400]" name="Greece" pageId="20" pageNumber="399">Greece</collectingCountry>
|
||
). According to the count methodology proposed in
|
||
<bibRefCitation id="62054B3DFFD8FFBC6716FA6FFDD8FA29" author="Bianco" box="[290,578,1412,1437]" pageId="20" pageNumber="399" refString="Bianco, P. G. & Kottelat, M. (2005) Scardinius knezevici, a new species of rudd from Lake Skadar, Montenegro (Teleostei: Cyprinidae) Ichthyological Exploration of Freshwaters, 16, 231 - 238." type="journal article" year="2005">Bianco & Kottelat (2005)</bibRefCitation>
|
||
the species shows 9 branched rays in the anal and in the dorsal fins. Among the
|
||
<taxonomicName id="C1944D4FFFD8FFBC66F4FA4CFE95FA74" box="[192,271,1447,1472]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="genus">
|
||
<emphasis id="34E0EADEFFD8FFBC66F4FA4CFE95FA74" box="[192,271,1447,1472]" italics="true" pageId="20" pageNumber="399">Rutilus</emphasis>
|
||
</taxonomicName>
|
||
species revised in this contribution, only the new species possess 9 branched rays in the dorsal and anal fins. The general shape of the specimens from the River Sperchios described by
|
||
<bibRefCitation id="62054B3DFFD8FFBC6275FA27FADEFA50" author="Kottelat" box="[1089,1348,1484,1509]" pageId="20" pageNumber="399" refString="Kottelat, M. & Freyhof, J. (2007) Handbook of European Freshwater Fishes. Kottelat, Cornol, Switzerland and Frehyof, Berlin, Germany, 646 pp." type="book" year="2007" yearSuffix="p">Kottelat (2007: p 249)</bibRefCitation>
|
||
is very similar to that of the
|
||
<typeStatus id="D92F886EFFD8FFBC67B5FA04FE79F9BC" box="[385,483,1519,1544]" pageId="20" pageNumber="399" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="C1944D4FFFD8FFBC643CFA1AFD26F9BC" box="[520,700,1519,1544]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="stoumboudae">
|
||
<emphasis id="34E0EADEFFD8FFBC643CFA1AFD26F9BC" box="[520,700,1519,1544]" italics="true" pageId="20" pageNumber="399">R. stoumboudae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="062B36CCFFD8FFBC66F3F9FFFD53F9C1" blockId="20.[151,1437,151,1871]" pageId="20" pageNumber="399">
|
||
Translocations of fishes by fishermen from southern
|
||
<collectingCountry id="7E83765CFFD8FFBC6500F9FFFC19F998" box="[820,899,1556,1580]" name="Greece" pageId="20" pageNumber="399">Greece</collectingCountry>
|
||
basins to the shore market of Lake Volvi for selling seems quite frequent.
|
||
<taxonomicName id="C1944D4FFFD8FFBC67DAF9D2FD2BF9E4" box="[494,689,1591,1616]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="ylikiensis">
|
||
<emphasis id="34E0EADEFFD8FFBC67DAF9D2FD2BF9E4" box="[494,689,1591,1616]" italics="true" pageId="20" pageNumber="399">Leucos ylikiensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C1944D4FFFD8FFBC64DBF9DCFC61F9E4" box="[751,1019,1591,1616]" class="Actinopterygii" family="Cyprinidae" genus="Scardinius" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="399" phylum="Chordata" rank="species" species="acarnanicus">
|
||
<emphasis id="34E0EADEFFD8FFBC64DBF9DCFC61F9E4" box="[751,1019,1591,1616]" italics="true" pageId="20" pageNumber="399">Scardinius acarnanicus</emphasis>
|
||
</taxonomicName>
|
||
, endemic to Lake Yliki, have been transplanted in Lake Volvi following this practice.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFD8FFBC66F3F994FB28F908" pageId="20" pageNumber="399" type="etymology">
|
||
<paragraph id="062B36CCFFD8FFBC66F3F994FB28F908" blockId="20.[151,1437,151,1871]" pageId="20" pageNumber="399">
|
||
<emphasis id="34E0EADEFFD8FFBC66F3F994FED4F92C" bold="true" box="[199,334,1663,1688]" pageId="20" pageNumber="399">Etymology.</emphasis>
|
||
The species is named after the Greek ichthyologist, colleague and friend, Maria Stoumboudi, in honor of her research activity on the ecology and conservation of freshwater fishes of
|
||
<collectingCountry id="7E83765CFFD8FFBC626FF94FFB34F908" box="[1115,1198,1700,1724]" name="Greece" pageId="20" pageNumber="399">Greece</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFD8FFBC66F3F92CFEC6F8FB" pageId="20" pageNumber="399" type="discussion">
|
||
<paragraph id="062B36CCFFD8FFBC66F3F92CFEC6F8FB" blockId="20.[151,1437,151,1871]" pageId="20" pageNumber="399">
|
||
<emphasis id="34E0EADEFFD8FFBC66F3F92CFC95F954" bold="true" box="[199,783,1735,1760]" pageId="20" pageNumber="399">Remarks on ecology biology and conservation:</emphasis>
|
||
a still water adapted species. No data available on its biology. According to the IUCN (2012) category, it satisfies point B and potentially should be placed in the “Endangered category” as the area of occupancy is less than
|
||
<quantity id="C16C9B29FFD8FFBC6576F8F9FC0DF89F" box="[834,919,1810,1835]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="399" unit="km" value="500.0">500 km</quantity>
|
||
2 and the species keeps declining as a result of habitat reduction.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |