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<document id="BC31E434590D2783ADE25AE9AEDB435E" ID-DOI="10.11646/zootaxa.3841.3.4" ID-GBIF-Dataset="fd4f8193-3005-4d9b-ae7f-304b847215a5" ID-ISSN="1175-5326" ID-Zenodo-Dep="225778" ID-ZooBank="E52D2F6B-631D-45FE-97E0-DD26137757FE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460462027068" checkinUser="plazi" docAuthor="Bianco, Pier Giorgio &amp; Ketmaier, Valerio" docDate="2014" docId="8E3D87DAFFDCFFB866A3FEA0FD88F8D0" docLanguage="en" docName="zt03841p402.pdf" docOrigin="Zootaxa 3841 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rutilus rutilus Linnaeus 1758" docType="treatment" docVersion="8" lastPageNumber="395" masterDocId="7204FFA2FFCCFFA86634FFEBFF9AFFB4" masterDocTitle="A revision of the Rutilus complex from Mediterranean Europe with description of a new genus, Sarmarutilus, and a new species, Rutilus stoumboudae (Teleostei: Cyprinidae)" masterLastPageNumber="402" masterPageNumber="379" pageNumber="395" updateTime="1698304242431" updateUser="plazi">
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<mods:title id="5C0E1F50009A70E12AF61ECEDCB94E59">A revision of the Rutilus complex from Mediterranean Europe with description of a new genus, Sarmarutilus, and a new species, Rutilus stoumboudae (Teleostei: Cyprinidae)</mods:title>
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<mods:namePart id="CF975273E896EA09FCBD998455C4CC21">Bianco, Pier Giorgio</mods:namePart>
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<mods:namePart id="B7F43B327E0DAE776853ADB0D4FC83DB">Ketmaier, Valerio</mods:namePart>
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<mods:date id="2C9D0622DC26233A5FB00760BCBF0774">2014</mods:date>
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<treatment id="8E3D87DAFFDCFFB866A3FEA0FD88F8D0" ID-DOI="http://doi.org/10.5281/zenodo.6140519" ID-GBIF-Taxon="119512638" ID-Zenodo-Dep="6140519" LSID="urn:lsid:plazi:treatment:8E3D87DAFFDCFFB866A3FEA0FD88F8D0" httpUri="http://treatment.plazi.org/id/8E3D87DAFFDCFFB866A3FEA0FD88F8D0" lastPageNumber="395" pageId="16" pageNumber="395">
<subSubSection id="4E8E6547FFDCFFB866A3FEA0FE7BFE33" pageId="16" pageNumber="395" type="nomenclature">
<paragraph id="062B36CCFFDCFFB866A3FEA0FD87FED2" blockId="16.[151,541,331,391]" box="[151,541,331,358]" pageId="16" pageNumber="395">
<heading id="5D6381A0FFDCFFB866A3FEA0FD87FED2" bold="true" box="[151,541,331,358]" fontSize="11" level="1" pageId="16" pageNumber="395" reason="1">
<taxonomicName id="C1944D4FFFDCFFB866A3FEA0FD87FED2" ID-CoL="4TTHT" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[151,541,331,358]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="rutilus">
<emphasis id="34E0EADEFFDCFFB866A3FEA0FD87FED2" bold="true" box="[151,541,331,358]" pageId="16" pageNumber="395">
<emphasis id="34E0EADEFFDCFFB866A3FEA0FEDCFED1" bold="true" box="[151,326,331,357]" italics="true" pageId="16" pageNumber="395">Rutilus rutilus</emphasis>
(Linnaeus, 1758)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="062B36CCFFDCFFB866A3FE85FE7BFE33" blockId="16.[151,541,331,391]" box="[151,481,366,391]" pageId="16" pageNumber="395">(introduced in the study area)</paragraph>
</subSubSection>
<subSubSection id="4E8E6547FFDCFFB866A3FE5CFBE2FE5F" pageId="16" pageNumber="395" type="reference_group">
<paragraph id="062B36CCFFDCFFB866A3FE5CFD67FE78" blockId="16.[151,1144,438,491]" box="[151,765,438,460]" pageId="16" pageNumber="395">
<taxonomicName id="C1944D4FFFDCFFB866A3FE5CFD88FE78" ID-CoL="643L5" authority="Linnaeus, 1758: 234" authorityName="Linnaeus" authorityPageNumber="234" authorityYear="1758" box="[151,530,438,460]" class="Actinopterygii" family="Cyprinidae" genus="Cyprinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="rutilus">
<emphasis id="34E0EADEFFDCFFB866A3FE5CFEA6FE78" box="[151,316,438,460]" italics="true" pageId="16" pageNumber="395">Cyprinus rutilus</emphasis>
Linnaeus, 1758: 234
</taxonomicName>
(
<typeStatus id="D92F886EFFDCFFB8642BFE5DFDD6FE78" box="[543,588,438,460]" pageId="16" pageNumber="395">type</typeStatus>
locality: Europa)
</paragraph>
<paragraph id="062B36CCFFDCFFB866A3FE3EFBE2FE5F" blockId="16.[151,1144,438,491]" box="[151,1144,469,491]" pageId="16" pageNumber="395">
<taxonomicName id="C1944D4FFFDCFFB866A3FE3EFD16FE5F" ID-CoL="4TTJ2" authority="Stephanidis, 1950: 203" authorityName="Stephanidis" authorityPageNumber="203" authorityYear="1950" box="[151,652,469,491]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="subSpecies" species="rutilus" subSpecies="vegariticus">
<emphasis id="34E0EADEFFDCFFB866A3FE3EFE04FE5F" box="[151,414,469,491]" italics="true" pageId="16" pageNumber="395">Rutilus rutilus vegariticus</emphasis>
Stephanidis, 1950: 203
</taxonomicName>
(
<typeStatus id="D92F886EFFDCFFB864AEFE3EFD5DFE5F" box="[666,711,469,491]" pageId="16" pageNumber="395">type</typeStatus>
locality, Lake Vegoritis, northern
<collectingCountry id="7E83765CFFDCFFB86215FE3EFBF7FE5F" box="[1057,1133,469,491]" name="Greece" pageId="16" pageNumber="395">Greece</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="4E8E6547FFDCFFB866A3FDFCFAA1FD74" pageId="16" pageNumber="395" type="materials_examined">
<paragraph id="062B36CCFFDCFFB866A3FDFCFAA1FD74" blockId="16.[151,1437,535,1893]" pageId="16" pageNumber="395">
<emphasis id="34E0EADEFFDCFFB866A3FDFCFE19FD84" bold="true" box="[151,387,535,560]" pageId="16" pageNumber="395">Examined material.</emphasis>
<collectionCode id="6085AE09FFDCFFB867BDFDF3FE22FD84" box="[393,440,536,560]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="16" pageNumber="395">IZA</collectionCode>
83108A, 107,
<collectingCountry id="7E83765CFFDCFFB8646AFDF3FD2AFD84" box="[606,688,536,560]" name="Greece" pageId="16" pageNumber="395">Greece</collectingCountry>
, Epirus, L. Joannina, P.G. Bianco,
<date id="722A100CFFDCFFB86205FDF3FB6DFD84" box="[1073,1271,535,560]" pageId="16" pageNumber="395" value="1976-02-23">23 February 1976</date>
.—
<collectionCode id="6085AE09FFDCFFB8632CFDFCFAD3FD84" box="[1304,1353,535,560]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="16" pageNumber="395">IZA</collectionCode>
88124, 6,
<collectingCountry id="7E83765CFFDCFFB8668CFDD7FE82FDE1" box="[184,280,572,597]" name="United Kingdom" pageId="16" pageNumber="395">England</collectingCountry>
, R. Thames, A. Wheeler,
<date id="722A100CFFDCFFB86464FDD7FD75FDE1" box="[592,751,572,597]" pageId="16" pageNumber="395" value="1986-06-18">18 June 1986</date>
.—
<collectionCode id="6085AE09FFDCFFB86524FDD7FCDBFDE1" box="[784,833,572,597]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="16" pageNumber="395">IZA</collectionCode>
78115, 11,
<collectingCountry id="7E83765CFFDCFFB865E6FDD7FBA0FDE1" box="[978,1082,572,597]" name="Romania" pageId="16" pageNumber="395">Romania</collectingCountry>
, R. Danube, P. Banarescu,
<date id="722A100CFFDCFFB863B4FDD7FECAFDCC" pageId="16" pageNumber="395" value="1975-09-23">23 September 1975</date>
.—
<collectionCode id="6085AE09FFDCFFB86745FDB4FE38FDCC" box="[369,418,607,632]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="16" pageNumber="395">IZA</collectionCode>
0 475, 15,
<collectingCountry id="7E83765CFFDCFFB86416FD8BFDEEFDCC" box="[546,628,608,632]" name="Greece" pageId="16" pageNumber="395">Greece</collectingCountry>
, L. Vegoritis (
<typeStatus id="D92F886EFFDCFFB8652FFD8BFCD7FDCC" box="[795,845,608,632]" pageId="16" pageNumber="395">type</typeStatus>
locality of
<taxonomicName id="C1944D4FFFDCFFB865E2FD8AFBEFFDCC" box="[982,1141,609,632]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="vegariticus">
<emphasis id="34E0EADEFFDCFFB865E2FD8AFBEFFDCC" box="[982,1141,609,632]" italics="true" pageId="16" pageNumber="395">R. vegariticus</emphasis>
</taxonomicName>
), P.G. Bianco,
<date id="722A100CFFDCFFB86311FD8BFF54FD28" pageId="16" pageNumber="395" value="1998-08-21">21 August 1998</date>
.—
<collectionCode id="6085AE09FFDCFFB866DBFD6FFEBAFD28" box="[239,288,644,668]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="16" pageNumber="395">IZA</collectionCode>
0 452, 5,
<collectionCode id="6085AE09FFDCFFB867B8FD6FFE69FD28" box="[396,499,644,668]" pageId="16" pageNumber="395">FYROM</collectionCode>
, L. Dojran, R. Gruptché,
<date id="722A100CFFDCFFB86526FD6FFC56FD28" box="[786,972,644,669]" pageId="16" pageNumber="395" value="1971-01-14">14 January 1971</date>
.—0477, 7,
<collectingCountry id="7E83765CFFDCFFB86267FD6FFB1DFD29" box="[1107,1159,644,669]" name="Italy" pageId="16" pageNumber="395">Italy</collectingCountry>
L. Como,
<collectingCountry id="7E83765CFFDCFFB86335FD6FFA82FD28" box="[1281,1304,644,668]" name="Iceland" pageId="16" pageNumber="395">O</collectingCountry>
. Mangione,
<date id="722A100CFFDCFFB866A3FD4CFE88FD74" box="[151,274,679,704]" pageId="16" pageNumber="395" value="2004-04">April 2004</date>
.—
<collectionCode id="6085AE09FFDCFFB86707FD43FEF9FD74" box="[307,355,680,704]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="16" pageNumber="395">IZA</collectionCode>
0 478, 5,
<collectingCountry id="7E83765CFFDCFFB867F8FD4CFDABFD74" box="[460,561,679,704]" name="Slovenia" pageId="16" pageNumber="395">Slovenia</collectingCountry>
, L. Pernica (R. Danube basin) at Mari-Bor, M. Povz,
<date id="722A100CFFDCFFB862BCFD43FAACFD74" box="[1160,1334,679,704]" pageId="16" pageNumber="395" value="1999-03-29">29 March 1999</date>
.
</paragraph>
</subSubSection>
<subSubSection id="4E8E6547FFDCFFB866F3FD27FD88FCE4" pageId="16" pageNumber="395" type="diagnosis">
<paragraph id="062B36CCFFDCFFB866F3FD27FD88FCE4" blockId="16.[151,1437,535,1893]" pageId="16" pageNumber="395">
<emphasis id="34E0EADEFFDCFFB866F3FD27FEA0FD51" bold="true" box="[199,314,716,741]" pageId="16" pageNumber="395">Diagnosis</emphasis>
.
<taxonomicName id="C1944D4FFFDCFFB8677CFD27FE7DFD51" box="[328,487,716,741]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="rutilus">
<emphasis id="34E0EADEFFDCFFB8677CFD27FE7DFD51" box="[328,487,716,741]" italics="true" pageId="16" pageNumber="395">Rutilus rutilus</emphasis>
</taxonomicName>
compared to
<taxonomicName id="C1944D4FFFDCFFB864B4FD26FD46FD50" box="[640,732,717,740]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="pigus">
<emphasis id="34E0EADEFFDCFFB864B4FD26FD46FD50" box="[640,732,717,740]" italics="true" pageId="16" pageNumber="395">R. pigus</emphasis>
</taxonomicName>
has less scales on LL (3944 vs 4651), and higher number of total GR (1216 vs. 912). Compared to
<taxonomicName id="C1944D4FFFDCFFB86450FD04FCD5FCBC" box="[612,847,751,776]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="stoumboudae" status="sp. nov.">
<emphasis id="34E0EADEFFDCFFB86450FD04FCD5FCBC" box="[612,847,751,776]" italics="true" pageId="16" pageNumber="395">Rutilus stoumboudae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FD357A5FFDCFFB86563FD1BFC14FCB3" box="[855,910,752,775]" pageId="16" pageNumber="395" rank="species">n.sp.</taxonomicNameLabel>
,
<taxonomicName id="C1944D4FFFDCFFB865AEFD1AFB99FCBC" box="[922,1027,751,776]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="rutilus">
<emphasis id="34E0EADEFFDCFFB865AEFD1AFB99FCBC" box="[922,1027,751,776]" italics="true" pageId="16" pageNumber="395">R. rutilus</emphasis>
</taxonomicName>
has less total GR (1215 vs. 1518) and more branched rays both in the dorsal and anal fins (911 vs. 89). Proportional measurements and meristic counts are reported in Tables 4, 5.
</paragraph>
</subSubSection>
<subSubSection id="4E8E6547FFDCFFB866F3FCB7FBAFFAB4" pageId="16" pageNumber="395" type="description">
<paragraph id="062B36CCFFDCFFB866F3FCB7FBAFFAB4" blockId="16.[151,1437,535,1893]" pageId="16" pageNumber="395">
<emphasis id="34E0EADEFFDCFFB866F3FCB7FEC3FCC1" bold="true" box="[199,345,860,885]" pageId="16" pageNumber="395">Description.</emphasis>
A species of
<taxonomicName id="C1944D4FFFDCFFB867CDFCB7FDD2FCC1" box="[505,584,860,885]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="genus">
<emphasis id="34E0EADEFFDCFFB867CDFCB7FDD2FCC1" box="[505,584,860,885]" italics="true" pageId="16" pageNumber="395">Rutilus</emphasis>
</taxonomicName>
of medium-large size, characterized by the absence of a mid lateral band; body slightly compressed; and moderately elongate, dorsal profile convex; sometimes humped in larger specimens; posterior mouth corner placed above the vertical crossing the anterior margin of orbital cavity; mouth terminal, opening small, slightly downturned; lips smooth; horizontal diameter of the eye more than or equal to the preorbital length; head small, with a length about 4.24.6 times the SL; body depth, about 3.23.6 times the SL; origin of the D slightly below the insertion of the P2; free margin of dorsal and anal fins slightly concave; caudal fin deeply forked; LL complete, with 3944 pored scales, placed below the middle of the body, slightly concave and extending from the upper margin of the opercular membrane to the end of caudal peduncle; modally 3 un-branched and modally 10 branched rays both in the dorsal and anal fins; P1 with 1 un-branched and 1517 branched rays; P2 with 1 un-branched ray and constantly 8 branched rays; usually 1214 total GR; color in life: eye from yellowish to a brilliant red; fins yellowish with some red; dorsal and caudal fins dark brown; peritoneal membrane silvery sometimes with few scattered melanophores. Additional description in Tables 4, 5.
</paragraph>
</subSubSection>
<subSubSection id="4E8E6547FFDCFFB866F3FAE7FA0CF904" pageId="16" pageNumber="395" type="discussion">
<paragraph id="062B36CCFFDCFFB866F3FAE7FA0CF904" blockId="16.[151,1437,535,1893]" pageId="16" pageNumber="395">
<emphasis id="34E0EADEFFDCFFB866F3FAE7FE5CFA91" bold="true" box="[199,454,1292,1317]" pageId="16" pageNumber="395">Remark on synonym.</emphasis>
The status of
<taxonomicName id="C1944D4FFFDCFFB86455FAE7FCAEFA90" box="[609,820,1292,1317]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="vegariticus">
<emphasis id="34E0EADEFFDCFFB86455FAE7FCAEFA90" box="[609,820,1292,1317]" italics="true" pageId="16" pageNumber="395">Rutilus vegariticus</emphasis>
</taxonomicName>
was still confused and inquirenda. It was considered as a valid species (
<bibRefCitation id="62054B3DFFDCFFB8677AFAC4FD21FAFC" author="Economidis" box="[334,699,1327,1352]" pageId="16" pageNumber="395" refString="Economidis, P. S. &amp; Banarescu, P. (1991) The distribution and origins of freshwater fish in the Balkan peninsula, especially in Greece. Internationale Revue gesamten Hydrobiologie, 76, 257 - 283. http: // dx. doi. org / 10.1002 / iroh. 19910760209" type="journal article" year="1991">Economidis &amp; Banarescu, 1991</bibRefCitation>
), or a subspecies of
<taxonomicName id="C1944D4FFFDCFFB86593FAC4FB78FAFC" authority="Economidis, 1995" authorityName="Economidis" authorityYear="1995" box="[935,1250,1327,1352]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="genus">
<emphasis id="34E0EADEFFDCFFB86593FAC4FC6CFAFC" box="[935,1014,1327,1352]" italics="true" pageId="16" pageNumber="395">Rutilus</emphasis>
(
<bibRefCitation id="62054B3DFFDCFFB86232FAC4FB43FAFC" author="Economidis" box="[1030,1241,1327,1352]" pageId="16" pageNumber="395" refString="Economidis, P. S. (1995) Endangered freshwater fishes of Greece. Biological conservation, 72, 201 - 211. http: // dx. doi. org / 10.1016 / 0006 - 3207 (94) 00083 - 3" type="journal article" year="1995">Economidis, 1995</bibRefCitation>
)
</taxonomicName>
.
<bibRefCitation id="62054B3DFFDCFFB862DAFAC4FA06FAFC" author="Kottelat" box="[1262,1436,1327,1352]" pageId="16" pageNumber="395" refString="Kottelat, M. (1997) European freshwater fish. Biologia, 52 (Supplement), 1 - 271." type="journal article" year="1997">Kottelat (1997)</bibRefCitation>
still considered questionable its validity as distinct taxon. Following its original description, no recent diagnosis and description of this nominal taxon were performed. According to molecular data (
<bibRefCitation id="62054B3DFFDCFFB86260FA9CFADDFA24" author="Ketmaier" box="[1108,1351,1399,1424]" pageId="16" pageNumber="395" refString="Ketmaier, V., Bianco, P. G. &amp; Durand, J. D. (2008) Molecular systematics, phylogeny and biogeography of roaches (Rutilus, Teleostei, Cyprinidae). Molecular Phylogenetics and Evolution, 49, 362 - 367. http: // dx. doi. org / 10.1016 / j. ympev. 2008.07.012" type="journal article" year="2008">
Ketmaier
<emphasis id="34E0EADEFFDCFFB862F2FA92FA9CFA24" box="[1222,1286,1399,1424]" italics="true" pageId="16" pageNumber="395">et al.,</emphasis>
2008
</bibRefCitation>
) and to the morphological analyses presented here (Tables 4, 5),
<taxonomicName id="C1944D4FFFDCFFB8652CFA77FC71FA00" box="[792,1003,1436,1461]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="vegariticus">
<emphasis id="34E0EADEFFDCFFB8652CFA77FC71FA00" box="[792,1003,1436,1461]" italics="true" pageId="16" pageNumber="395">Rutilus vegariticus</emphasis>
</taxonomicName>
cannot be consistently separated from
<taxonomicName id="C1944D4FFFDCFFB866A3FA54FF62FA6C" box="[151,248,1471,1496]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="rutilus">
<emphasis id="34E0EADEFFDCFFB866A3FA54FF62FA6C" box="[151,248,1471,1496]" italics="true" pageId="16" pageNumber="395">R.rutilus</emphasis>
</taxonomicName>
as all morphometric and meristic characters overlap (Tables 4, 5).
<taxonomicName id="C1944D4FFFDCFFB865C9FA54FB49FA6C" box="[1021,1235,1471,1496]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="vegariticus">
<emphasis id="34E0EADEFFDCFFB865C9FA54FB49FA6C" box="[1021,1235,1471,1496]" italics="true" pageId="16" pageNumber="395">Rutilus vegariticus</emphasis>
</taxonomicName>
(Lake Vegoritis, eastern
<collectingCountry id="7E83765CFFDCFFB866C1FA0FFEDCFA48" box="[245,326,1508,1532]" name="Greece" pageId="16" pageNumber="395">Greece</collectingCountry>
) is genetically very close to a number of central European and introduced northern Italian populations of
<taxonomicName id="C1944D4FFFDCFFB86777F9E2FD25F994" authority="Ketmaier et al., 2008" authorityName="Ketmaier et al." authorityYear="2008" box="[323,703,1543,1568]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="rutilus">
<emphasis id="34E0EADEFFDCFFB86777F9E2FE37F994" box="[323,429,1543,1568]" italics="true" pageId="16" pageNumber="395">R. rutilus</emphasis>
(
<bibRefCitation id="62054B3DFFDCFFB86789F9ECFD2DF994" author="Ketmaier" box="[445,695,1543,1568]" pageId="16" pageNumber="395" refString="Ketmaier, V., Bianco, P. G. &amp; Durand, J. D. (2008) Molecular systematics, phylogeny and biogeography of roaches (Rutilus, Teleostei, Cyprinidae). Molecular Phylogenetics and Evolution, 49, 362 - 367. http: // dx. doi. org / 10.1016 / j. ympev. 2008.07.012" type="journal article" year="2008">
Ketmaier
<emphasis id="34E0EADEFFDCFFB86404F9E2FDEBF994" box="[560,625,1543,1568]" italics="true" pageId="16" pageNumber="395">et al.,</emphasis>
2008
</bibRefCitation>
)
</taxonomicName>
.
<bibRefCitation id="62054B3DFFDCFFB864FFF9E2FC45F994" box="[715,991,1543,1568]" pageId="16" pageNumber="395" refString="Larmuseau, M. H. D., Frehyof, J., Volkaert, F. A. M. &amp; Houdt, J. K. J. van (2009) Matrilinear phylogeography and demographical patterns of Rutilus rutilus: implications for taxonomy and conservation. Journal of Fish Biology, 75, 332 - 353. http: // dx. doi. org / 10.1111 / j. 1095 - 8649.2009.02322. x" type="journal article">
Larmuseau
<emphasis id="34E0EADEFFDCFFB86565F9E2FC11F994" box="[849,907,1543,1568]" italics="true" pageId="16" pageNumber="395">et al.</emphasis>
(2009)
</bibRefCitation>
claimed that this population should be regarded as a distinct
<taxonomicName id="C1944D4FFFDCFFB867A1F9C7FDADF9F1" box="[405,567,1580,1605]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="rutilus">
<emphasis id="34E0EADEFFDCFFB867A1F9C7FDADF9F1" box="[405,567,1580,1605]" italics="true" pageId="16" pageNumber="395">Rutilus rutilus</emphasis>
</taxonomicName>
lineage, possibly a junior synonym for the Ponto-Caspian
<taxonomicName id="C1944D4FFFDCFFB862D5F9C6FAC0F9F1" box="[1249,1370,1580,1605]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="heckelii">
<emphasis id="34E0EADEFFDCFFB862D5F9C6FAC0F9F1" box="[1249,1370,1580,1605]" italics="true" pageId="16" pageNumber="395">R. heckelii</emphasis>
</taxonomicName>
. This seems not realistic biogeographically (
<bibRefCitation id="62054B3DFFDCFFB8647BF9A4FD77F9DC" author="Bianco" box="[591,749,1615,1640]" pageId="16" pageNumber="395" refString="Bianco, P. G. (1990) Potential role of the palaeohistory of the Mediterranean and Parathetys basin on the early dispersal of Europe-Mediterranean freshwater fishes. Ichthyological Exploration of Freshwaters, 1, 167 - 184." type="journal article" year="1990">Bianco, 1990</bibRefCitation>
), since the Ponto-Caspian region is well differentiated from the circum-Mediterranean one; we believe that the most parsimonious explanation for the occurrence of
<taxonomicName id="C1944D4FFFDCFFB86378F99FFF7AF904" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="rutilus">
<emphasis id="34E0EADEFFDCFFB86378F99FFF7AF904" italics="true" pageId="16" pageNumber="395">Rutilus rutilus</emphasis>
</taxonomicName>
in the area is introduction by humans and, therefore, we consider this taxon as junior synonym of
<emphasis id="34E0EADEFFDCFFB86312F972FA0CF904" box="[1318,1430,1687,1712]" italics="true" pageId="16" pageNumber="395">
<taxonomicName id="C1944D4FFFDCFFB86312F972FA0BF904" box="[1318,1425,1687,1712]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="rutilus">R. rutilus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="4E8E6547FFDCFFB866F3F957FD88F8D0" pageId="16" pageNumber="395" type="distribution">
<paragraph id="062B36CCFFDCFFB866F3F957FD88F8D0" blockId="16.[151,1437,535,1893]" pageId="16" pageNumber="395">
<emphasis id="34E0EADEFFDCFFB866F3F957FEC3F961" bold="true" box="[199,345,1724,1749]" pageId="16" pageNumber="395">Distribution</emphasis>
. Native to central Europe, the species has been introduced in several circum-Mediterranean ichthyogeographical districts. In northern
<collectingCountry id="7E83765CFFDCFFB864B5F934FD2FF94C" box="[641,693,1759,1784]" name="Italy" pageId="16" pageNumber="395">Italy</collectingCountry>
, the species has been introduced in the Po basin and in many subalpine lakes where it compete with the native
<taxonomicName id="C1944D4FFFDCFFB864FDF8EFFCC7F8A8" box="[713,861,1796,1821]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="pigus">
<emphasis id="34E0EADEFFDCFFB864FDF8EFFCC7F8A8" box="[713,861,1796,1821]" italics="true" pageId="16" pageNumber="395">Rutilus pigus</emphasis>
</taxonomicName>
and
<taxonomicName id="C1944D4FFFDCFFB865A2F8EEFBBAF8A9" box="[918,1056,1796,1821]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="aula">
<emphasis id="34E0EADEFFDCFFB865A2F8EEFBBAF8A9" box="[918,1056,1796,1821]" italics="true" pageId="16" pageNumber="395">Leucos aula</emphasis>
</taxonomicName>
; in central
<collectingCountry id="7E83765CFFDCFFB86296F8EFFB4FF8A9" box="[1186,1237,1796,1821]" name="Italy" pageId="16" pageNumber="395">Italy</collectingCountry>
introductions are known for the Tiber and Arno rivers and for lakes Montedoglio and Corbara, where it caused the disappearance of the endemic
<taxonomicName id="C1944D4FFFDCFFB86712F8A7FD91F8D1" box="[294,523,1868,1893]" class="Actinopterygii" family="Cyprinidae" genus="Sarmarutilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="395" phylum="Chordata" rank="species" species="rubilio">
<emphasis id="34E0EADEFFDCFFB86712F8A7FD91F8D1" box="[294,523,1868,1893]" italics="true" pageId="16" pageNumber="395">Sarmarutilus rubilio</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>