499 lines
79 KiB
XML
499 lines
79 KiB
XML
<document id="5DBB35D63210D5E24E55FBF7A5E696B6" ID-CLB-Dataset="24014" ID-DOI="10.1080/00222933.2019.1660429" ID-GBIF-Dataset="95d9643a-ed68-42cf-a00c-fbaaa113c71f" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670379" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581942039985" checkinUser="valdenar" docAuthor="Molano-Rendón, Arturo González-Alvarado ı Fredy & Vaz-de- Mello, Fernando Z." docDate="2019" docId="8D45C315FFAF1242FE9CEFB7FE74FE9F" docLanguage="en" docName="JNATHIST.53.27–28.1751–1765.pdf.imf" docOrigin="Journal of Natural History 53 (27)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Atlantemolanum Gonzalez-Alvarado 2019, new genus" docType="treatment" docVersion="9" lastPageNumber="1757" masterDocId="717CBB6DFFAC124AFFADEF76FFFDFF91" masterDocTitle="A new genus of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) endemic to the Brazilian Atlantic Forest" masterLastPageNumber="1765" masterPageNumber="1751" pageNumber="1753" updateTime="1698783642502" updateUser="ExternalLinkService">
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<mods:mods id="38B04617D15307EB6D0B841DA13453AE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9ECD61E469D261B82F0B70E995D49150">
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<mods:title id="A7A469AFD872B823861CFA23FBB002E5">A new genus of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) endemic to the Brazilian Atlantic Forest</mods:title>
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<mods:name id="335B53AE9AFF7250E7ECBE95A95840C4" type="personal">
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<mods:namePart id="41A1525BACA4E62B6427B91AAE4372F5">Molano-Rendón, Arturo González-Alvarado ı Fredy</mods:namePart>
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<mods:name id="91B90D0D50E0B640991EF24DFFBCFE67" type="personal">
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<mods:roleTerm id="8054F7756DF7B44EDEEC01EE1E839CC1">Author</mods:roleTerm>
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<mods:namePart id="9E061482EE3624E23EDD8D9532BA20D9">Vaz-de- Mello, Fernando Z.</mods:namePart>
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<mods:title id="A4C1979000A8CE3F1DC696A155B613D3">Journal of Natural History</mods:title>
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<mods:part id="ED1EF5E4021428EF4EE04801D2CD7CDA">
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<mods:date id="A256C87F3B587776C5428AD5C062B02E">2019</mods:date>
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<mods:number id="FF484F127E436B18016773D56E38FCE7">2019-09-27</mods:number>
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<mods:number id="B126D616E6C63D84ED2912B78D6D1D79">53</mods:number>
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<mods:number id="8C766A84408984467ABE604342B1B824">27</mods:number>
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<mods:identifier id="BD384BC7A77936E3C1DEEC1090466445" type="DOI">10.1080/00222933.2019.1660429</mods:identifier>
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<treatment id="8D45C315FFAF1242FE9CEFB7FE74FE9F" ID-DOI="http://doi.org/10.5281/zenodo.3671063" ID-GBIF-Taxon="162146594" ID-Zenodo-Dep="3671063" LSID="urn:lsid:plazi:treatment:8D45C315FFAF1242FE9CEFB7FE74FE9F" httpUri="http://treatment.plazi.org/id/8D45C315FFAF1242FE9CEFB7FE74FE9F" lastPageId="8" lastPageNumber="1757" pageId="3" pageNumber="1753">
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<subSubSection id="4DF62188FFAF1249FE9CEFB7FCE1FF6A" pageId="3" pageNumber="1753" type="nomenclature">
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<paragraph id="05537203FFAF1249FE9CEFB7FCE1FF6A" blockId="3.[305,1011,193,251]" pageId="3" pageNumber="1753">
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||
<emphasis id="3798AE11FFAF1249FE9CEFB7FCE1FF6A" bold="true" pageId="3" pageNumber="1753">
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||
<heading id="5E1BC56FFFAF1249FE9CEFB7FC0EFF4D" box="[305,1011,193,220]" pageId="3" pageNumber="1753">
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||
<taxonomicName id="C2EC0980FFAF1249FE9CEFB7FCA5FF4D" ID-CoL="354F" authority="Gonzalez-Alvarado et al." authorityName="Gonzalez-Alvarado" authorityYear="2019" box="[305,856,193,220]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="3798AE11FFAF1249FE9CEFB7FDF4FF4A" bold="true" box="[305,521,193,219]" italics="true" pageId="3" pageNumber="1753">Atlantemolanum</emphasis>
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González-Alvarado et al.
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</taxonomicName>
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<taxonomicNameLabel id="2CAB136AFFAF1249FCCFEFB4FC0EFF4D" box="[866,1011,194,220]" pageId="3" pageNumber="1753" rank="genus">new genus</taxonomicNameLabel>
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</heading>
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(
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<figureCitation id="9DD76E86FFAF1249FD82EF95FD66FF6A" box="[559,667,227,251]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="3" pageNumber="1753">Figures 1</figureCitation>
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(a–e)ı 2–5)
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="4DF62188FFAF1249FF0DEE5AFDF1FDAA" pageId="3" pageNumber="1753" type="type_taxon">
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<subSubSection id="4DF62188FFAF1249FF0DEE5AFDACFE1B" pageId="3" pageNumber="1753" type="type_taxon">
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<paragraph id="05537203FFAF1249FF0DEE5AFEC9FED4" blockId="3.[160,1155,300,394]" box="[160,308,300,325]" pageId="3" pageNumber="1753">
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<emphasis id="3798AE11FFAF1249FF0DEE5AFEC9FED4" bold="true" box="[160,308,300,325]" italics="true" pageId="3" pageNumber="1753">
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<typeStatus id="DA57CCA1FFAF1249FF0DEE5AFF25FED4" box="[160,216,300,325]" pageId="3" pageNumber="1753">Type</typeStatus>
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species
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</emphasis>
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</paragraph>
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<paragraph id="05537203FFAF1249FF0DEE39FDACFE1B" blockId="3.[160,1155,300,394]" pageId="3" pageNumber="1753">
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<taxonomicName id="C2EC0980FFAF1249FF0DEE39FE06FEF6" ID-CoL="6CJ4Z" authority="Harold 1868" authorityName="Harold" authorityYear="1868" box="[160,507,335,359]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="species" species="riehli">
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<emphasis id="3798AE11FFAF1249FF0DEE39FE9EFEF6" box="[160,355,335,359]" italics="true" pageId="3" pageNumber="1753">Deltochilum riehli</emphasis>
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<bibRefCitation id="617D0FF2FFAF1249FEC3EE39FE06FEF6" author="Harold E" box="[366,507,335,359]" pageId="3" pageNumber="1753" pagination="79 - 86" refId="ref6497" refString="Harold E. 1868. Diagnosen Neuer Coprophagen. Coleopterol Hefte. 1: 79 - 86." type="journal article" year="1868">Harold 1868</bibRefCitation>
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</taxonomicName>
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(
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<emphasis id="3798AE11FFAF1249FDA0EE39FCFCFEF6" bold="true" box="[525,769,335,359]" pageId="3" pageNumber="1753">present designation</emphasis>
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) =
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<taxonomicName id="C2EC0980FFAF1249FC83EE39FF22FE1B" ID-CoL="JGW5" authority="(Harold 1868)" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" baseAuthorityName="Harold" baseAuthorityYear="1868" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="species" species="riehli" status="comb. nov.">
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<emphasis id="3798AE11FFAF1249FC83EE39FBDBFEF6" box="[814,1062,335,359]" italics="true" pageId="3" pageNumber="1753">Atlantemolanum riehli</emphasis>
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(
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<bibRefCitation id="617D0FF2FFAF1249FB9AEE39FF2AFE1B" author="Harold E" pageId="3" pageNumber="1753" pagination="79 - 86" refId="ref6497" refString="Harold E. 1868. Diagnosen Neuer Coprophagen. Coleopterol Hefte. 1: 79 - 86." type="journal article" year="1868">Harold 1868</bibRefCitation>
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)
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</taxonomicName>
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<emphasis id="3798AE11FFAF1249FF4AEE04FE44FE1B" bold="true" box="[231,441,370,394]" pageId="3" pageNumber="1753">
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<taxonomicNameLabel id="2CAB136AFFAF1249FF4AEE04FE44FE1B" box="[231,441,370,394]" pageId="3" pageNumber="1753" rank="species">new combination</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="9DD76E86FFAF1249FE67EE04FDDAFE1B" box="[458,551,370,394]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="3" pageNumber="1753">Figure 1</figureCitation>
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(a)).
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</paragraph>
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</subSubSection>
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<subSubSection id="4DF62188FFAF1249FF0DEECCFDF1FDAA" pageId="3" pageNumber="1753" type="discussion">
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<paragraph id="05537203FFAF1249FF0DEECCFECBFE42" blockId="3.[160,1156,442,571]" box="[160,310,442,467]" pageId="3" pageNumber="1753">
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||
<emphasis id="3798AE11FFAF1249FF0DEECCFECBFE42" bold="true" box="[160,310,442,467]" italics="true" pageId="3" pageNumber="1753">Composition</emphasis>
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</paragraph>
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<paragraph id="05537203FFAF1249FF0DEEA8FDF1FDAA" blockId="3.[160,1156,442,571]" pageId="3" pageNumber="1753">
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Besides the
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<typeStatus id="DA57CCA1FFAF1249FE83EEA8FE9DFE67" box="[302,352,478,502]" pageId="3" pageNumber="1753">type</typeStatus>
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speciesı
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<taxonomicName id="C2EC0980FFAF1249FE61EEA8FD7EFE67" ID-CoL="354F" authority="Molano-Rendón & Vaz-de- Mello, 2019" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" box="[460,643,478,502]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="3798AE11FFAF1249FE61EEA8FD7EFE67" box="[460,643,478,502]" italics="true" pageId="3" pageNumber="1753">Atlantemolanum</emphasis>
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</taxonomicName>
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<emphasis id="3798AE11FFAF1249FD22EEA8FCEEFE67" bold="true" box="[655,787,478,502]" pageId="3" pageNumber="1753">
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<taxonomicNameLabel id="2CAB136AFFAF1249FD22EEA8FCEEFE67" box="[655,787,478,502]" pageId="3" pageNumber="1753" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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includes
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<taxonomicName id="C2EC0980FFAF1249FC2BEEA8FF10FD88" ID-CoL="34KBC" authority="Pereira" authorityName="Pereira" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="species" species="costalimai">
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<emphasis id="3798AE11FFAF1249FC2BEEA8FB7EFE67" box="[902,1155,478,502]" italics="true" pageId="3" pageNumber="1753">Deltochilum costalimai</emphasis>
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Pereira
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</taxonomicName>
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and d
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<emphasis id="3798AE11FFAF1249FE9AED76FEC1FD89" box="[311,316,512,536]" italics="true" pageId="3" pageNumber="1753">’</emphasis>
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Andretta 1955 =
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<taxonomicName id="C2EC0980FFAF1249FE52ED77FCD0FD88" ID-CoL="JGW4" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" baseAuthorityName="Pereira and d' Andretta" baseAuthorityYear="1955" box="[511,813,513,537]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="species" species="costalimai">
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<emphasis id="3798AE11FFAF1249FE52ED77FCD0FD88" box="[511,813,513,537]" italics="true" pageId="3" pageNumber="1753">Atlantemolanum costalimai</emphasis>
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</taxonomicName>
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(Pereira and d
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<emphasis id="3798AE11FFAF1249FC78ED76FC27FD89" box="[981,986,512,536]" italics="true" pageId="3" pageNumber="1753">’</emphasis>
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Andretta 1955)
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<emphasis id="3798AE11FFAF1249FF0DED55FE8FFDAA" bold="true" box="[160,370,547,571]" pageId="3" pageNumber="1753">new combination</emphasis>
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(
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<figureCitation id="9DD76E86FFAF1249FE2EED55FE1DFDAA" box="[387,480,547,571]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="3" pageNumber="1753">Figure 1</figureCitation>
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(b)).
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</paragraph>
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</subSubSection>
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||
</subSubSection>
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<subSubSection id="4DF62188FFAF1249FF0DED1AFB84FB3E" pageId="3" pageNumber="1753" type="diagnosis">
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||
<paragraph id="05537203FFAF1249FF0DED1AFEE9FD14" blockId="3.[160,1156,620,1199]" box="[160,276,620,645]" pageId="3" pageNumber="1753">
|
||
<emphasis id="3798AE11FFAF1249FF0DED1AFEE9FD14" bold="true" box="[160,276,620,645]" italics="true" pageId="3" pageNumber="1753">Diagnosis</emphasis>
|
||
</paragraph>
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||
<paragraph id="05537203FFAF1249FF0DEDF9FB84FB3E" blockId="3.[160,1156,620,1199]" pageId="3" pageNumber="1753">
|
||
<taxonomicName id="C2EC0980FFAF1249FF0DEDF9FEA9FD36" authority="Molano-Rendón & Vaz-de- Mello, 2019" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" box="[160,340,655,679]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1754" phylum="Arthropoda" rank="genus" status="gen. nov.">
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||
<emphasis id="3798AE11FFAF1249FF0DEDF9FEA9FD36" box="[160,340,655,679]" italics="true" pageId="3" pageNumber="1753">Atlantemolanum</emphasis>
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</taxonomicName>
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can be separated from all other genera of Scarabaeinae of the New Word by the following combination of characters: (1) mesotibiae with only one spur (
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<figureCitation id="9DD76E86FFAF1249FBA5EDC4FB9CFD5B" box="[1032,1121,690,714]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="3" pageNumber="1753">Figure 1</figureCitation>
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(c)) and (2) Mesosternum with a transversal sulcus anteriorly at the meso-metasternal suture. The two species included in
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<taxonomicName id="C2EC0980FFAF1249FDB5ED81FD32FC9E" authority="Molano-Rendón & Vaz-de- Mello, 2019" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" box="[536,719,759,783]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1754" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="3798AE11FFAF1249FDB5ED81FD32FC9E" box="[536,719,759,783]" italics="true" pageId="3" pageNumber="1753">Atlantemolanum</emphasis>
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</taxonomicName>
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were originally described within the genus
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<taxonomicName id="C2EC0980FFAF1249FF5BEC6CFE84FCA3" authorityName="Eschscholtz" authorityYear="1822" box="[246,377,794,818]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="genus">
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<emphasis id="3798AE11FFAF1249FF5BEC6CFE84FCA3" box="[246,377,794,818]" italics="true" pageId="3" pageNumber="1753">Deltochilum</emphasis>
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</taxonomicName>
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by having apical elytral tubercles in some of the interstriaeı a character present in all species of the genus
|
||
<taxonomicName id="C2EC0980FFAF1249FD46EC4BFC93FCC4" authorityName="Eschscholtz" authorityYear="1822" box="[747,878,829,853]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3798AE11FFAF1249FD46EC4BFC93FCC4" box="[747,878,829,853]" italics="true" pageId="3" pageNumber="1753">Deltochilum</emphasis>
|
||
</taxonomicName>
|
||
; however
|
||
<taxonomicName id="C2EC0980FFAF1249FC5FEC4BFBB7FCC4" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[1010,1098,829,853]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="species" species="riehli">
|
||
<emphasis id="3798AE11FFAF1249FC5FEC4BFBB7FCC4" box="[1010,1098,829,853]" italics="true" pageId="3" pageNumber="1753">A. riehli</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C2EC0980FFAF1249FF0DEC29FED3FCE6" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" baseAuthorityName="Pereira and d' Andretta" baseAuthorityYear="1955" box="[160,302,863,887]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="species" species="costalimai">
|
||
<emphasis id="3798AE11FFAF1249FF0DEC29FED3FCE6" box="[160,302,863,887]" italics="true" pageId="3" pageNumber="1753">A. costalimai</emphasis>
|
||
</taxonomicName>
|
||
can be separated from all other species of
|
||
<taxonomicName id="C2EC0980FFAF1249FC9FEC29FC48FCE6" authorityName="Eschscholtz" authorityYear="1822" box="[818,949,863,887]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3798AE11FFAF1249FC9FEC29FC48FCE6" box="[818,949,863,887]" italics="true" pageId="3" pageNumber="1753">Deltochilum</emphasis>
|
||
</taxonomicName>
|
||
ı by the following combination of characters: (1) mesotibiae with only one spur (
|
||
<figureCitation id="9DD76E86FFAF1249FCC5ECF4FC3BFC0B" box="[872,966,898,922]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="3" pageNumber="1753">Figure 1</figureCitation>
|
||
(c))ı two spurs in the species of
|
||
<taxonomicName id="C2EC0980FFAF1249FEE0ECD3FE2DFC2C" authorityName="Eschscholtz" authorityYear="1822" box="[333,464,933,957]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3798AE11FFAF1249FEE0ECD3FE2DFC2C" box="[333,464,933,957]" italics="true" pageId="3" pageNumber="1753">Deltochilum</emphasis>
|
||
</taxonomicName>
|
||
; (2) hypomeron divided by a strong transverse carinaı and with a deep excavation anteriorly (
|
||
<figureCitation id="9DD76E86FFAF1249FD9EECB1FD6CFC4E" box="[563,657,967,991]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="3" pageNumber="1753">Figure 1</figureCitation>
|
||
(d) black arrow) to the carina; in the genus
|
||
<taxonomicName id="C2EC0980FFAF1249FF0DEC9CFEDEFB93" authorityName="Eschscholtz" authorityYear="1822" box="[160,291,1002,1026]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3798AE11FFAF1249FF0DEC9CFEDEFB93" box="[160,291,1002,1026]" italics="true" pageId="3" pageNumber="1753">Deltochilum</emphasis>
|
||
</taxonomicName>
|
||
ı only the species of the subgenus
|
||
<taxonomicName id="C2EC0980FFAF1249FD0AEC9CFCDEFB93" authorityName="Kolbe" authorityYear="1893" box="[679,803,1002,1026]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="subGenus" subGenus="Calhyboma">
|
||
<emphasis id="3798AE11FFAF1249FD0AEC9CFCDEFB93" box="[679,803,1002,1026]" italics="true" pageId="3" pageNumber="1753">Calhyboma</emphasis>
|
||
</taxonomicName>
|
||
have the transverse carinaı but anteriorly to the carina is weakly excavated; (3) pronotumı in dorsal viewı with a carina (that separates it from the hypomeron) not visible between the medial and posterior angles (
|
||
<figureCitation id="9DD76E86FFAF1249FF55EB24FEA9FBFB" box="[248,340,1106,1130]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="3" pageNumber="1753">Figure 1</figureCitation>
|
||
(e) arrow)ı carina visible (in dorsal view) (
|
||
<figureCitation id="9DD76E86FFAF1249FCB2EB24FC81FBFB" box="[799,892,1106,1130]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="3" pageNumber="1753">Figure 1</figureCitation>
|
||
(f) arrow) in the species of
|
||
<taxonomicName id="C2EC0980FFAF1249FF6DEB03FEBEFB1C" authorityName="Eschscholtz" authorityYear="1822" box="[192,323,1141,1165]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3798AE11FFAF1249FF6DEB03FEBEFB1C" box="[192,323,1141,1165]" italics="true" pageId="3" pageNumber="1753">Deltochilum</emphasis>
|
||
</taxonomicName>
|
||
; (4) meso- and metafemora marginated anteriorly and posteriorly on its ventral face (
|
||
<figureCitation id="9DD76E86FFAF1249FE98EBE1FE60FB3E" box="[309,413,1175,1199]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="3" pageNumber="1753">Figures 1</figureCitation>
|
||
(c)ı 2(g
|
||
<emphasis id="3798AE11FFAF1249FE47EBE1FE0AFB3E" box="[490,503,1175,1199]" italics="true" pageId="3" pageNumber="1753">–</emphasis>
|
||
h)ı 4(h
|
||
<emphasis id="3798AE11FFAF1249FDEDEBE1FDB0FB3E" box="[576,589,1175,1199]" italics="true" pageId="3" pageNumber="1753">–</emphasis>
|
||
i))ı not marginated in the species of
|
||
<taxonomicName id="C2EC0980FFAF1249FC5DEBE1FB8EFB3E" authorityName="Eschscholtz" authorityYear="1822" box="[1008,1139,1175,1199]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1753" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3798AE11FFAF1249FC5DEBE1FB8EFB3E" box="[1008,1139,1175,1199]" italics="true" pageId="3" pageNumber="1753">Deltochilum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4DF62188FFAF124DFF0DEB96FC11FE2D" lastPageId="7" lastPageNumber="1757" pageId="3" pageNumber="1753" type="description">
|
||
<paragraph id="05537203FFAF1249FF0DEB96FEDBFB68" blockId="3.[160,1155,1248,1411]" box="[160,294,1248,1273]" pageId="3" pageNumber="1753">
|
||
<emphasis id="3798AE11FFAF1249FF0DEB96FEDBFB68" bold="true" box="[160,294,1248,1273]" italics="true" pageId="3" pageNumber="1753">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="05537203FFAF1249FF0DEA75FE23FA12" blockId="3.[160,1155,1248,1411]" pageId="3" pageNumber="1753">
|
||
Sizeı length
|
||
<quantity id="C214DFE6FFAF1249FE80EA75FE51FA8A" box="[301,428,1283,1307]" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="1.25" metricValueMin="0.9" pageId="3" pageNumber="1753" unit="mm" value="10.75" valueMax="12.5" valueMin="9.0">
|
||
9
|
||
<emphasis id="3798AE11FFAF1249FE96EA75FEB5FA8A" box="[315,328,1283,1307]" italics="true" pageId="3" pageNumber="1753">–</emphasis>
|
||
12.5 mm
|
||
</quantity>
|
||
(from anterior margin of pronotum to apex of elytra)ı humeral width
|
||
<quantity id="C214DFE6FFAF1249FF43EA50FE84FAAF" box="[238,377,1318,1342]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.5" metricValueMin="6.5" pageId="3" pageNumber="1753" unit="mm" value="8.0" valueMax="9.5" valueMin="6.5">
|
||
6.5
|
||
<emphasis id="3798AE11FFAF1249FEA2EA50FEE1FAAF" box="[271,284,1318,1342]" italics="true" pageId="3" pageNumber="1753">–</emphasis>
|
||
9.5 mm
|
||
</quantity>
|
||
. Body oval; colour black or black with coppery reflections. Body covered by circuları ocellate punctures; body with setose puncturesı setae longest dorsally and on the femora.
|
||
</paragraph>
|
||
<paragraph id="05537203FFAF1249FF0DEAC2FE4DF92E" blockId="3.[160,1156,1460,1727]" pageId="3" pageNumber="1753">
|
||
<emphasis id="3798AE11FFAF1249FF0DEAC2FE36FA5C" bold="true" box="[160,459,1460,1485]" italics="true" pageId="3" pageNumber="1753">
|
||
Head (
|
||
<figureCitation id="9DD76E86FFAF1249FF5DEAC2FEA3FA5C" box="[240,350,1460,1485]" captionStart="Figure 2" captionStartId="5.[160,225,1645,1668]" captionTargetBox="[167,1153,441,1612]" captionTargetId="figure@5.[163,1153,437,1614]" captionTargetPageId="5" captionText="Figure 2. Male Atlantemolanum riehli. (a) head, dorsal view; (b) right elytron; (c) elytral apex; (d) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (e) metaesternum; (f) ventrites; (g) mesoleg; (h) metaleg." figureDoi="http://doi.org/10.5281/zenodo.3670385" httpUri="https://zenodo.org/record/3670385/files/figure.png" pageId="3" pageNumber="1753">Figures 2</figureCitation>
|
||
(a)ı 4(a)).
|
||
</emphasis>
|
||
Wider than long; covered with punctures separated by one diameter on the centre and the genae; vertex and between the clypealı punctures separated by less than one diameter. Posterior margin of head without carina. Clypeus bidentateı teeth acuteı each tooth separated by a U-shaped emargination; clypeo-genal suture visible; clypeus almost straight between tooth and clypeo-genal suture. Genae roundedı anterior angles of genae slightly projected. Eyes dorsally smallı dorsal interocular distance 12 times the width of one eye; eyes longer than wideı approximately two times longer than wide.
|
||
</paragraph>
|
||
<caption id="5193228BFFA8124EFF0DEA4EFEE5FA73" ID-DOI="http://doi.org/10.5281/zenodo.3670381" ID-Zenodo-Dep="3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="4" pageNumber="1754" startId="4.[160,225,1336,1359]" targetBox="[172,1150,150,1303]" targetPageId="4">
|
||
<paragraph id="05537203FFA8124EFF0DEA4EFEE5FA73" blockId="4.[160,1155,1336,1506]" pageId="4" pageNumber="1754">
|
||
<emphasis id="3798AE11FFA8124EFF0DEA4EFF06FADE" bold="true" box="[160,251,1336,1359]" pageId="4" pageNumber="1754">Figure 1.</emphasis>
|
||
<taxonomicName id="C2EC0980FFA8124EFEAFEA4EFE5EFADE" authority="Molano-Rendón & Vaz-de- Mello, 2019" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" box="[258,419,1336,1359]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1754" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="3798AE11FFA8124EFEAFEA4EFE5EFADE" box="[258,419,1336,1359]" italics="true" pageId="4" pageNumber="1754">Atlantemolanum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2CAB136AFFA8124EFE04EA4FFDEBFAC1" box="[425,534,1337,1360]" pageId="4" pageNumber="1754" rank="genus">new genus</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="C2EC0980FFA8124EFDE4EA4EFD46FADE" authorityName="Eschscholtz" authorityYear="1822" box="[585,699,1336,1359]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1754" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3798AE11FFA8124EFDE4EA4EFD46FADE" box="[585,699,1336,1359]" italics="true" pageId="4" pageNumber="1754">Deltochilum</emphasis>
|
||
</taxonomicName>
|
||
. (a) male
|
||
<taxonomicName id="C2EC0980FFA8124EFCB2EA4EFB7EFAC1" authority="(Harold 1868)" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[799,1155,1336,1360]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1754" phylum="Arthropoda" rank="species" species="riehli" status="comb. nov.">
|
||
<emphasis id="3798AE11FFA8124EFCB2EA4EFC0BFADE" box="[799,1014,1336,1359]" italics="true" pageId="4" pageNumber="1754">Atlantemolanum riehli</emphasis>
|
||
(
|
||
<bibRefCitation id="617D0FF2FFA8124EFBAFEA4FFB81FAC1" author="Harold E" box="[1026,1148,1337,1360]" pageId="4" pageNumber="1754" pagination="79 - 86" refId="ref6497" refString="Harold E. 1868. Diagnosen Neuer Coprophagen. Coleopterol Hefte. 1: 79 - 86." type="journal article" year="1868">Harold 1868</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2CAB136AFFA8124EFF0DEA20FEACFAFC" box="[160,337,1366,1389]" pageId="4" pageNumber="1754" rank="species">new combination</taxonomicNameLabel>
|
||
, dorsal view; (b) female
|
||
<taxonomicName id="C2EC0980FFA8124EFDE5EA20FCAFFAFC" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" baseAuthorityName="Pereira and d' Andretta" baseAuthorityYear="1955" box="[584,850,1366,1389]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1754" phylum="Arthropoda" rank="species" species="costalimai">
|
||
<emphasis id="3798AE11FFA8124EFDE5EA20FCAFFAFC" box="[584,850,1366,1389]" italics="true" pageId="4" pageNumber="1754">Atlantemolanum costalimai</emphasis>
|
||
</taxonomicName>
|
||
(Pereira and d
|
||
<emphasis id="3798AE11FFA8124EFC44EA20FC13FAFC" box="[1001,1006,1366,1389]" italics="true" pageId="4" pageNumber="1754">’</emphasis>
|
||
Andretta 1955) new combination, dorsal view. (c-e)
|
||
<taxonomicName id="C2EC0980FFA8124EFDA3EA05FDA5FA1B" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[526,600,1395,1418]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1754" phylum="Arthropoda" rank="species" species="riehli">
|
||
<emphasis id="3798AE11FFA8124EFDA3EA05FDA5FA1B" box="[526,600,1395,1418]" italics="true" pageId="4" pageNumber="1754">A. riehli</emphasis>
|
||
</taxonomicName>
|
||
(c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f)
|
||
<taxonomicName id="C2EC0980FFA8124EFCBEEAD8FC0BFA54" baseAuthorityName="Gory" baseAuthorityYear="1831" box="[787,1014,1454,1477]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1754" phylum="Arthropoda" rank="species" species="rubripenne">
|
||
<emphasis id="3798AE11FFA8124EFCBEEAD8FC0BFA54" box="[787,1014,1454,1477]" italics="true" pageId="4" pageNumber="1754">Deltochilum rubripenne</emphasis>
|
||
</taxonomicName>
|
||
, pronotum in dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="05537203FFA8124FFF0DE96BFEE5FEE7" blockId="4.[160,1156,1565,1693]" lastBlockId="5.[160,1155,142,374]" lastPageId="5" lastPageNumber="1755" pageId="4" pageNumber="1754">
|
||
<emphasis id="3798AE11FFA8124EFF0DE96BFDE7F9A7" bold="true" box="[160,538,1565,1590]" italics="true" pageId="4" pageNumber="1754">
|
||
Pronotum (
|
||
<figureCitation id="9DD76E86FFA8124EFE84E96BFE6CF9A7" box="[297,401,1565,1590]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="4" pageNumber="1754">Figures 1</figureCitation>
|
||
(dıe)ı 4(b)).
|
||
</emphasis>
|
||
Globuları two times wider than long; in dorsal viewı margin separating pronotum from hypomeron not visible between medial and posterior angles (
|
||
<figureCitation id="9DD76E86FFA8124EFF57E915FEA4F9EA" box="[250,345,1635,1659]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="4" pageNumber="1754">Figure 1</figureCitation>
|
||
(e) arrow); margin visible in lateral view and interrupted by punctures (
|
||
<figureCitation id="9DD76E86FFA8124EFF05E9F3FEFBF90C" box="[168,262,1669,1693]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="4" pageNumber="1754">Figure 1</figureCitation>
|
||
(d) white arrow). Pronotal anterior angles projectedı slightly directed laterally; margin between anterior and medial angles of pronotum concaveı medial angle rounded; margin between medial and posterior angles of pronotum straight. Pronotal disc with punctures separated by more than one diameterı gradually larger and denser towards pronotal margins. Pronotum in the middle with a callosity near anterior margin (
|
||
<figureCitation id="9DD76E86FFA9124FFF0AEE6FFEFAFEA0" box="[167,263,281,305]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="5" pageNumber="1755">Figure 1</figureCitation>
|
||
(e)). Hypomeron divided by a strong transverse carinaı with deep excavation anteriorly (
|
||
<figureCitation id="9DD76E86FFA9124FFE8CEE4AFE79FEC5" box="[289,388,316,340]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="5" pageNumber="1755">Figure 1</figureCitation>
|
||
(d) black arrow); carina completeı reaching the medial angle of pronotum.
|
||
</paragraph>
|
||
<caption id="5193228BFFA9124FFF0DE91BFC17F92E" ID-DOI="http://doi.org/10.5281/zenodo.3670385" ID-Zenodo-Dep="3670385" httpUri="https://zenodo.org/record/3670385/files/figure.png" pageId="5" pageNumber="1755" startId="5.[160,225,1645,1668]" targetBox="[167,1153,441,1612]" targetPageId="5">
|
||
<paragraph id="05537203FFA9124FFF0DE91BFC17F92E" blockId="5.[160,1156,1645,1727]" pageId="5" pageNumber="1755">
|
||
<emphasis id="3798AE11FFA9124FFF0DE91BFF00F915" bold="true" box="[160,253,1645,1668]" pageId="5" pageNumber="1755">Figure 2.</emphasis>
|
||
Male
|
||
<taxonomicName id="C2EC0980FFA9124FFEECE91BFDE6F915" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[321,539,1645,1668]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1755" phylum="Arthropoda" rank="species" species="riehli">
|
||
<emphasis id="3798AE11FFA9124FFEECE91BFDE6F915" box="[321,539,1645,1668]" italics="true" pageId="5" pageNumber="1755">Atlantemolanum riehli</emphasis>
|
||
</taxonomicName>
|
||
. (a) head, dorsal view; (b) right elytron; (c) elytral apex; (d) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (e) metaesternum; (f) ventrites; (g) mesoleg; (h) metaleg.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="05537203FFAA124CFF0DEFF8FC06FE2D" blockId="6.[160,1156,142,444]" pageId="6" pageNumber="1756">
|
||
<emphasis id="3798AE11FFAA124CFF0DEFF8FDF5FF36" bold="true" box="[160,520,142,167]" italics="true" pageId="6" pageNumber="1756">
|
||
Elytra (
|
||
<figureCitation id="9DD76E86FFAA124CFF5BEFF8FE98FF36" box="[246,357,142,167]" captionStart="Figure 2" captionStartId="5.[160,225,1645,1668]" captionTargetBox="[167,1153,441,1612]" captionTargetId="figure@5.[163,1153,437,1614]" captionTargetPageId="5" captionText="Figure 2. Male Atlantemolanum riehli. (a) head, dorsal view; (b) right elytron; (c) elytral apex; (d) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (e) metaesternum; (f) ventrites; (g) mesoleg; (h) metaleg." figureDoi="http://doi.org/10.5281/zenodo.3670385" httpUri="https://zenodo.org/record/3670385/files/figure.png" pageId="6" pageNumber="1756">Figures 2</figureCitation>
|
||
(b–d)ı 4(c–e)).
|
||
</emphasis>
|
||
With ten striaeı 9th and 10th striae completely separated from the apex to the middle of the elytronı fused from the middle to the base; striae with extended puncturesı punctures almost twice as long as wide (
|
||
<figureCitation id="9DD76E86FFAA124CFC87EFA2FC7DFF7D" box="[810,896,212,236]" captionStart="Figure 2" captionStartId="5.[160,225,1645,1668]" captionTargetBox="[167,1153,441,1612]" captionTargetId="figure@5.[163,1153,437,1614]" captionTargetPageId="5" captionText="Figure 2. Male Atlantemolanum riehli. (a) head, dorsal view; (b) right elytron; (c) elytral apex; (d) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (e) metaesternum; (f) ventrites; (g) mesoleg; (h) metaleg." figureDoi="http://doi.org/10.5281/zenodo.3670385" httpUri="https://zenodo.org/record/3670385/files/figure.png" pageId="6" pageNumber="1756">Figure 2</figureCitation>
|
||
(d)ı 4(e)); strial punctures smallı almost the same width as the strial line (
|
||
<figureCitation id="9DD76E86FFAA124CFD1AEF81FCE1FE9E" box="[695,796,247,271]" captionStart="Figure 2" captionStartId="5.[160,225,1645,1668]" captionTargetBox="[167,1153,441,1612]" captionTargetId="figure@5.[163,1153,437,1614]" captionTargetPageId="5" captionText="Figure 2. Male Atlantemolanum riehli. (a) head, dorsal view; (b) right elytron; (c) elytral apex; (d) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (e) metaesternum; (f) ventrites; (g) mesoleg; (h) metaleg." figureDoi="http://doi.org/10.5281/zenodo.3670385" httpUri="https://zenodo.org/record/3670385/files/figure.png" pageId="6" pageNumber="1756">Figures 2</figureCitation>
|
||
(d)ı 4(e)). 1st
|
||
<emphasis id="3798AE11FFAA124CFC0BEF81FC4EFE9E" box="[934,947,247,271]" italics="true" pageId="6" pageNumber="1756">–</emphasis>
|
||
5th striae widened basallyı twice as wide as at the apex. Interstriae with rounded punctures (
|
||
<figureCitation id="9DD76E86FFAA124CFC6FEE6CFBD8FEA3" box="[962,1061,282,306]" captionStart="Figure 2" captionStartId="5.[160,225,1645,1668]" captionTargetBox="[167,1153,441,1612]" captionTargetId="figure@5.[163,1153,437,1614]" captionTargetPageId="5" captionText="Figure 2. Male Atlantemolanum riehli. (a) head, dorsal view; (b) right elytron; (c) elytral apex; (d) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (e) metaesternum; (f) ventrites; (g) mesoleg; (h) metaleg." figureDoi="http://doi.org/10.5281/zenodo.3670385" httpUri="https://zenodo.org/record/3670385/files/figure.png" pageId="6" pageNumber="1756">Figures 2</figureCitation>
|
||
(d)ı 4(e)). 1stı 3rdı 5th and 7th interstriae with shiny callosities along elytral length; apex of elytra with tubercles on 3rd
|
||
<emphasis id="3798AE11FFAA124CFEFEEE29FE9DFEE6" box="[339,352,351,375]" italics="true" pageId="6" pageNumber="1756">–</emphasis>
|
||
7th intestriae (
|
||
<figureCitation id="9DD76E86FFAA124CFE53EE29FDAAFEE6" box="[510,599,351,375]" captionStart="Figure 4" captionStartId="9.[160,225,1431,1454]" captionTargetBox="[163,1152,142,1396]" captionTargetId="figure@9.[163,1153,142,1399]" captionTargetPageId="9" captionText="Figure 4. Female Atlantemolanum costalimai. (a) head, dorsal view; (b) pronotum, dorsal view; (c) right elytron; (d) elytral apex; (e) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (f) metasternum; (g) ventrites; (h) mesoleg, (i) metaleg." figureDoi="http://doi.org/10.5281/zenodo.3670389" httpUri="https://zenodo.org/record/3670389/files/figure.png" pageId="6" pageNumber="1756">Figure 4</figureCitation>
|
||
(d))ı some times without tubercle on 4th insterstria (
|
||
<figureCitation id="9DD76E86FFAA124CFF0AEEF4FEF8FE0B" box="[167,261,386,410]" captionStart="Figure 2" captionStartId="5.[160,225,1645,1668]" captionTargetBox="[167,1153,441,1612]" captionTargetId="figure@5.[163,1153,437,1614]" captionTargetPageId="5" captionText="Figure 2. Male Atlantemolanum riehli. (a) head, dorsal view; (b) right elytron; (c) elytral apex; (d) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (e) metaesternum; (f) ventrites; (g) mesoleg; (h) metaleg." figureDoi="http://doi.org/10.5281/zenodo.3670385" httpUri="https://zenodo.org/record/3670385/files/figure.png" pageId="6" pageNumber="1756">Figure 2</figureCitation>
|
||
(c)); 7th interstriaı on humeral regionı slightly elevated (
|
||
<figureCitation id="9DD76E86FFAA124CFC25EEF4FC0DFE0B" box="[904,1008,386,410]" captionStart="Figure 2" captionStartId="5.[160,225,1645,1668]" captionTargetBox="[167,1153,441,1612]" captionTargetId="figure@5.[163,1153,437,1614]" captionTargetPageId="5" captionText="Figure 2. Male Atlantemolanum riehli. (a) head, dorsal view; (b) right elytron; (c) elytral apex; (d) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (e) metaesternum; (f) ventrites; (g) mesoleg; (h) metaleg." figureDoi="http://doi.org/10.5281/zenodo.3670385" httpUri="https://zenodo.org/record/3670385/files/figure.png" pageId="6" pageNumber="1756">Figures 2</figureCitation>
|
||
(b)ı 4(c)); 9th interstria carinatedı carina fine and present on the basal fourth of elytral length.
|
||
</paragraph>
|
||
<paragraph id="05537203FFAA124CFF0DEE9FFEF4FDD6" blockId="6.[160,1155,489,583]" pageId="6" pageNumber="1756">
|
||
<emphasis id="3798AE11FFAA124CFF0DEE9FFEEBFD93" bold="true" box="[160,278,489,514]" italics="true" pageId="6" pageNumber="1756">
|
||
<taxonomicName id="C2EC0980FFAA124CFF0DEE9FFEECFD93" box="[160,273,489,514]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="1756" phylum="Chordata" rank="genus">Pygidium</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Triangular and convexı 1.5 times wider than long; with a transverse carina separating it from prepygidium. Pigidial punctures denserı separated by less than one diameter.
|
||
</paragraph>
|
||
<paragraph id="05537203FFAA124CFF0DED02FBC8FCCD" blockId="6.[160,1155,628,860]" pageId="6" pageNumber="1756">
|
||
<emphasis id="3798AE11FFAA124CFF0DED02FD00FD1C" bold="true" box="[160,765,628,653]" italics="true" pageId="6" pageNumber="1756">
|
||
Mesosternum and metasternum (
|
||
<figureCitation id="9DD76E86FFAA124CFD80ED02FD64FD1C" box="[557,665,628,653]" captionStart="Figure 2" captionStartId="5.[160,225,1645,1668]" captionTargetBox="[167,1153,441,1612]" captionTargetId="figure@5.[163,1153,437,1614]" captionTargetPageId="5" captionText="Figure 2. Male Atlantemolanum riehli. (a) head, dorsal view; (b) right elytron; (c) elytral apex; (d) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (e) metaesternum; (f) ventrites; (g) mesoleg; (h) metaleg." figureDoi="http://doi.org/10.5281/zenodo.3670385" httpUri="https://zenodo.org/record/3670385/files/figure.png" pageId="6" pageNumber="1756">Figures 2</figureCitation>
|
||
(e)ı 4(f)).
|
||
</emphasis>
|
||
Mesosternum narrowı in the midline lineı one-third narrower than prosternum. Mesosternum with transversal sulcus anteriorly at mesometasternal suture. Metasternum (
|
||
<figureCitation id="9DD76E86FFAA124CFD5BEDCCFCA3FD43" box="[758,862,698,722]" captionStart="Figure 2" captionStartId="5.[160,225,1645,1668]" captionTargetBox="[167,1153,441,1612]" captionTargetId="figure@5.[163,1153,437,1614]" captionTargetPageId="5" captionText="Figure 2. Male Atlantemolanum riehli. (a) head, dorsal view; (b) right elytron; (c) elytral apex; (d) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (e) metaesternum; (f) ventrites; (g) mesoleg; (h) metaleg." figureDoi="http://doi.org/10.5281/zenodo.3670385" httpUri="https://zenodo.org/record/3670385/files/figure.png" pageId="6" pageNumber="1756">Figures 2</figureCitation>
|
||
(e)ı 4(f)) wideı each mesocoxa separated by approximately three times the width of one mesocoxa. Mesocoxal axis (
|
||
<figureCitation id="9DD76E86FFAA124CFF0AED89FEFFFC86" box="[167,258,767,791]" captionStart="Figure 2" captionStartId="5.[160,225,1645,1668]" captionTargetBox="[167,1153,441,1612]" captionTargetId="figure@5.[163,1153,437,1614]" captionTargetPageId="5" captionText="Figure 2. Male Atlantemolanum riehli. (a) head, dorsal view; (b) right elytron; (c) elytral apex; (d) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (e) metaesternum; (f) ventrites; (g) mesoleg; (h) metaleg." figureDoi="http://doi.org/10.5281/zenodo.3670385" httpUri="https://zenodo.org/record/3670385/files/figure.png" pageId="6" pageNumber="1756">Figure 2</figureCitation>
|
||
(e)) slightly obliqueı almost parallel to the longitudinal body axis. Metasternal disc with longitudinal sulcus in the posterior two-thirds. Metasternum covered by punctures separated by more than one diameterı laterally and posteriorly denser and larger.
|
||
</paragraph>
|
||
<paragraph id="05537203FFAA124CFF0DECFFFD43FB04" blockId="6.[160,1156,905,1173]" pageId="6" pageNumber="1756">
|
||
<emphasis id="3798AE11FFAA124CFF0DECFFFE0EFC33" bold="true" box="[160,499,905,930]" italics="true" pageId="6" pageNumber="1756">
|
||
Abdomen (
|
||
<figureCitation id="9DD76E86FFAA124CFE8FECFFFE70FC33" box="[290,397,905,930]" captionStart="Figure 2" captionStartId="5.[160,225,1645,1668]" captionTargetBox="[167,1153,441,1612]" captionTargetId="figure@5.[163,1153,437,1614]" captionTargetPageId="5" captionText="Figure 2. Male Atlantemolanum riehli. (a) head, dorsal view; (b) right elytron; (c) elytral apex; (d) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (e) metaesternum; (f) ventrites; (g) mesoleg; (h) metaleg." figureDoi="http://doi.org/10.5281/zenodo.3670385" httpUri="https://zenodo.org/record/3670385/files/figure.png" pageId="6" pageNumber="1756">Figures 2</figureCitation>
|
||
(f)ı 4(g)).
|
||
</emphasis>
|
||
With six ventrites; the first ventrite wideı measured along the midlineı as wide as the second and third ventrites combined; 2nd to 6th ventrites almost equalı measured along midlineı in width. Ventrites with puncturesı larger laterallyı gradually smaller to centre; punctures at centre two times smaller than lateral ones. 1st
|
||
<emphasis id="3798AE11FFAA124CFF6CEB62FF33FBBD" box="[193,206,1044,1068]" italics="true" pageId="6" pageNumber="1756">–</emphasis>
|
||
4th ventrites with punctures only at anterior half of each ventrite; 5th ventrite with punctures in the middle line and laterallyı along both anterior and posterior surfaces. Punctures between middle line and lateral ones distributed only at anterior half. Sixth ventrite with completely covered by punctures.
|
||
</paragraph>
|
||
<paragraph id="05537203FFAA124CFF0DEBB7FBB9FA3B" blockId="6.[160,1156,1217,1727]" pageId="6" pageNumber="1756">
|
||
<emphasis id="3798AE11FFAA124CFF0DEBB7FF26FB4B" bold="true" box="[160,219,1217,1242]" italics="true" pageId="6" pageNumber="1756">Legs.</emphasis>
|
||
Protibia with three lateral teethı distributed along the apical two-fifths; with small denticles in the external margin and between the teeth; distance between apical teeth less than distance between second and third teeth; distance between second and third teeth three times the distance between apical teeth. Protibial apex straightly truncatedı with five tarsomeresı 1st and 5th twice wider than longı same length approximately; 2nd
|
||
<emphasis id="3798AE11FFAA124CFB9DEA3AFBC0FAF5" box="[1072,1085,1356,1380]" italics="true" pageId="6" pageNumber="1756">–</emphasis>
|
||
4th as wide as longı same length approximately. Length of 2nd
|
||
<emphasis id="3798AE11FFAA124CFC94EA19FCBBFA16" box="[825,838,1391,1415]" italics="true" pageId="6" pageNumber="1756">–</emphasis>
|
||
4th combined almost same length as 1st. Profemur with dense puncturesı separated by less than one diameter.
|
||
</paragraph>
|
||
<paragraph id="05537203FFAA124CFF6DEAC3FF0EF92E" blockId="6.[160,1156,1217,1727]" pageId="6" pageNumber="1756">
|
||
Mesofemur (
|
||
<figureCitation id="9DD76E86FFAA124CFEF8EAC3FE3EFA5C" box="[341,451,1461,1485]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="6" pageNumber="1756">Figures 1</figureCitation>
|
||
(c)ı 2(g)ı 4(h)) marginated anteriorly and posteriorly on its ventral face. Mesofemur with dense punctures separated by less than one diameter. Mesotibia weakly and gradually widened apicallyı width of the apex approximately oneeighth of the tibial length; mesotibial apex straightly truncated and with one large and curve spur (
|
||
<figureCitation id="9DD76E86FFAA124CFE88E949FE71F9C6" box="[293,396,1599,1623]" captionStart="Figure 1" captionStartId="4.[160,225,1336,1359]" captionTargetBox="[172,1150,150,1303]" captionTargetId="figure@4.[163,1153,142,1305]" captionTargetPageId="4" captionText="Figure 1. Atlantemolanum new genus and Deltochilum. (a) male Atlantemolanum riehli (Harold 1868) new combination, dorsal view; (b) female Atlantemolanum costalimai (Pereira and d’Andretta 1955) new combination, dorsal view. (c-e) A. riehli (c) mesoleg; (d) pronotum in lateral view, white arrow indicates the interrupted carina that separates pronotum from hypomeron, black arrow indicates the anterior hypomeral excavation; (e) pronotum, dorsal view; (f) Deltochilum rubripenne, pronotum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3670381" httpUri="https://zenodo.org/record/3670381/files/figure.png" pageId="6" pageNumber="1756">Figures 1</figureCitation>
|
||
(c)ı 2(g)ı 4(h))ı spur length approximately equal to the length of the first four tarsomeres combined. Mesotibia with five triangular tarsomeresı tarsomeres 1st
|
||
<emphasis id="3798AE11FFAA124CFF6CE9F2FF33F90D" box="[193,206,1668,1692]" italics="true" pageId="6" pageNumber="1756">–</emphasis>
|
||
4th uniform in sizeı and the fifth twice the length of the fourth; mesotarsal claws curved.
|
||
</paragraph>
|
||
<paragraph id="05537203FFAB124DFF6DEFF8FC11FE2D" blockId="7.[160,1156,142,444]" pageId="7" pageNumber="1757">
|
||
Metafemur (
|
||
<figureCitation id="9DD76E86FFAB124DFEE7EFF8FE4DFF37" box="[330,432,142,166]" captionStart="Figure 2" captionStartId="5.[160,225,1645,1668]" captionTargetBox="[167,1153,441,1612]" captionTargetId="figure@5.[163,1153,437,1614]" captionTargetPageId="5" captionText="Figure 2. Male Atlantemolanum riehli. (a) head, dorsal view; (b) right elytron; (c) elytral apex; (d) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (e) metaesternum; (f) ventrites; (g) mesoleg; (h) metaleg." figureDoi="http://doi.org/10.5281/zenodo.3670385" httpUri="https://zenodo.org/record/3670385/files/figure.png" pageId="7" pageNumber="1757">Figures 2</figureCitation>
|
||
(h)ı 4(i)) marginated anteriorly and posteriorly on its ventral face. Metafemur with punctures at anterior half twice as large as the punctures at posterior half; punctures at posterior half of femur denser than those at anterior halfı separated by less than one diameter at posterior half and by more than one diameter at anterior half. Metatibia weakly and gradually widened apicallyı width of the apex approximately oneeleventh of the tibial length; metatibial apex straightly truncated and with one curved spurı spur length approximately equal to the length of the two basal tarsomeres combined; metatibia with five triangular tarsomeresı tarsomeres 1st
|
||
<emphasis id="3798AE11FFAB124DFC6AEEF7FC29FE08" box="[967,980,385,409]" italics="true" pageId="7" pageNumber="1757">–</emphasis>
|
||
4th uniform in sizeı and the fifth twice the length of the fourth; metatarsal claws curved.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4DF62188FFAB124DFF0DEE9EFBBEFDF8" pageId="7" pageNumber="1757" type="etymology">
|
||
<paragraph id="05537203FFAB124DFF0DEE9EFEE0FD90" blockId="7.[160,1155,488,617]" box="[160,285,488,513]" pageId="7" pageNumber="1757">
|
||
<emphasis id="3798AE11FFAB124DFF0DEE9EFEE0FD90" bold="true" box="[160,285,488,513]" italics="true" pageId="7" pageNumber="1757">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="05537203FFAB124DFF0DED7AFBBEFDF8" blockId="7.[160,1155,488,617]" pageId="7" pageNumber="1757">The genus name pays homage to one of the co-authorsı the late Fredy Molano who sadly passed away during the review process of this paper. Associated with its endemicity to the Atlantic rainforestı originates the proposed name. It is male in gender.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4DF62188FFAB124DFF0DEDE0FF1CFD65" pageId="7" pageNumber="1757" type="distribution">
|
||
<paragraph id="05537203FFAB124DFF0DEDE0FE55FD3E" blockId="7.[160,1155,662,756]" box="[160,424,662,687]" pageId="7" pageNumber="1757">
|
||
<emphasis id="3798AE11FFAB124DFF0DEDE0FE55FD3E" bold="true" box="[160,424,662,687]" italics="true" pageId="7" pageNumber="1757">
|
||
Distribution (
|
||
<figureCitation id="9DD76E86FFAB124DFE93EDE0FE5DFD3E" box="[318,416,662,687]" captionStart="Figure 5" captionStartId="11.[160,225,873,896]" captionTargetBox="[163,1152,142,842]" captionTargetId="figure@11.[163,1153,142,842]" captionTargetPageId="11" captionText="Figure 5. Locality map; blue circles, Atlantemolanum riehli (Harold 1868); red star, Atlantemolanum costalimai (Pereira and d’Andretta 1955)." figureDoi="http://doi.org/10.5281/zenodo.3670391" httpUri="https://zenodo.org/record/3670391/files/figure.png" pageId="7" pageNumber="1757">Figure 5</figureCitation>
|
||
)
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="05537203FFAB124DFF0DEDCCFF1CFD65" blockId="7.[160,1155,662,756]" pageId="7" pageNumber="1757">
|
||
This new genus is so far known to be endemic to the central and southern Atlantic Forest of
|
||
<collectingCountry id="7DFB3293FFAB124DFF0DEDAAFF20FD65" box="[160,221,732,756]" name="Brazil" pageId="7" pageNumber="1757">Brazil</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4DF62188FFAB1242FF0DEC57FE74FE9F" lastPageId="8" lastPageNumber="1758" pageId="7" pageNumber="1757" type="discussion">
|
||
<paragraph id="05537203FFAB124DFF0DEC57FEFBFCAB" blockId="7.[160,1157,801,1727]" box="[160,262,801,826]" pageId="7" pageNumber="1757">
|
||
<emphasis id="3798AE11FFAB124DFF0DEC57FEFBFCAB" bold="true" box="[160,262,801,826]" italics="true" pageId="7" pageNumber="1757">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="05537203FFAB124DFF0DEC33FD18FC33" blockId="7.[160,1157,801,1727]" pageId="7" pageNumber="1757">This new genus presents one unique character among all the genera of New World Scarabaeinaeı the presence of only one mesotibial spur; all other described New World Scarabaeinae genera present two mesotibial spurs.</paragraph>
|
||
<paragraph id="05537203FFAB124DFF6DECDBFE05FBE3" blockId="7.[160,1157,801,1727]" pageId="7" pageNumber="1757">
|
||
In the description of the
|
||
<taxonomicName id="C2EC0980FFAB124DFE48ECDBFDC4FC54" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[485,569,941,965]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1757" phylum="Arthropoda" rank="species" species="riehli" status="comb. nov.">
|
||
<emphasis id="3798AE11FFAB124DFE48ECDBFDC4FC54" box="[485,569,941,965]" italics="true" pageId="7" pageNumber="1757">A. riehli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2CAB136AFFAB124DFDEEECDBFD55FC54" box="[579,680,941,965]" pageId="7" pageNumber="1757" rank="species">n. comb.</taxonomicNameLabel>
|
||
<bibRefCitation id="617D0FF2FFAB124DFD1DECDBFCB1FC54" author="Harold E" box="[688,844,941,965]" pageId="7" pageNumber="1757" pagination="79 - 86" refId="ref6497" refString="Harold E. 1868. Diagnosen Neuer Coprophagen. Coleopterol Hefte. 1: 79 - 86." type="journal article" year="1868">Harold (1868)</bibRefCitation>
|
||
did not mention the tibial spurs; for this speciesı
|
||
<bibRefCitation id="617D0FF2FFAB124DFE09ECB9FDBBFC76" author="Paulian R" box="[420,582,975,999]" pageId="7" pageNumber="1757" pagination="1 - 40" refId="ref6929" refString="Paulian R. 1939. Contribution a l ' etude des Canthonides Americains (Coleopt. Lamellic). Ann la Societe Entomol Fr. 108: 1 - 40." type="journal article" year="1939">Paulian (1939)</bibRefCitation>
|
||
described only the protibial and metatibial spurs; and Pereira and d
|
||
<emphasis id="3798AE11FFAB124DFEC4EC84FE92FB9B" box="[361,367,1010,1034]" italics="true" pageId="7" pageNumber="1757">’</emphasis>
|
||
Andretta (1955) stated that this species has two mesotibial spursı one three times longer than the other; howeverı in the figure of the mesotibia (Figure
|
||
<specimenCount id="13EAB98AFFAB124DFBC5EB63FF48FBDE" pageId="7" pageNumber="1757" type="generic">28 p.</specimenCount>
|
||
16) of Pereira and d
|
||
<emphasis id="3798AE11FFAB124DFE3AEB41FE60FBDE" box="[407,413,1079,1103]" italics="true" pageId="7" pageNumber="1757">’</emphasis>
|
||
Andretta (1955) only one spur is evident. The latter was confirmed after examining the
|
||
<typeStatus id="DA57CCA1FFAB124DFE24EB2CFE08FBE3" box="[393,501,1114,1138]" pageId="7" pageNumber="1757" type="lectotype">lectotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="05537203FFAB124DFF6DEB0BFB79FB6C" blockId="7.[160,1157,801,1727]" pageId="7" pageNumber="1757">
|
||
Pereira and d
|
||
<emphasis id="3798AE11FFAB124DFEF9EB0AFEA4FB05" box="[340,345,1148,1172]" italics="true" pageId="7" pageNumber="1757">’</emphasis>
|
||
Andretta (1955)ı in their description of
|
||
<taxonomicName id="C2EC0980FFAB124DFCA4EB0BFC6FFB04" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" baseAuthorityName="Pereira and d' Andretta" baseAuthorityYear="1955" box="[777,914,1149,1173]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1757" phylum="Arthropoda" rank="species" species="costalimai" status="comb. nov.">
|
||
<emphasis id="3798AE11FFAB124DFCA4EB0BFC6FFB04" box="[777,914,1149,1173]" italics="true" pageId="7" pageNumber="1757">A. costalimai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2CAB136AFFAB124DFC34EB0BFC00FB04" box="[921,1021,1149,1173]" pageId="7" pageNumber="1757" rank="species">n. comb.</taxonomicNameLabel>
|
||
ı stated than this species has two mesotibial spursı one larger and curved and the other one smallerı however in the figure of the mesotibia (figure
|
||
<specimenCount id="13EAB98AFFAB124DFD3FEBB4FD3AFB4B" box="[658,711,1218,1242]" pageId="7" pageNumber="1757" type="generic">18 p.</specimenCount>
|
||
13) of Pereira and d
|
||
<emphasis id="3798AE11FFAB124DFC32EBB4FC59FB4B" box="[927,932,1218,1242]" italics="true" pageId="7" pageNumber="1757">’</emphasis>
|
||
Andretta (1955) only one spur is evident; furthermoreı the
|
||
<typeStatus id="DA57CCA1FFAB124DFD94EB93FD61FB6C" box="[569,668,1253,1277]" pageId="7" pageNumber="1757" type="holotype">holotype</typeStatus>
|
||
of this species only has one mesotibial spur.
|
||
</paragraph>
|
||
<paragraph id="05537203FFAB124DFF6DEA71FEC7F9EB" blockId="7.[160,1157,801,1727]" pageId="7" pageNumber="1757">
|
||
One single mesotibial spur is a character shared between
|
||
<taxonomicName id="C2EC0980FFAB124DFCFBEA71FBF7FA8E" authority="Molano-Rendón & Vaz-de- Mello, 2019" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" box="[854,1034,1287,1311]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1754" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="3798AE11FFAB124DFCFBEA71FBF7FA8E" box="[854,1034,1287,1311]" italics="true" pageId="7" pageNumber="1757">Atlantemolanum</emphasis>
|
||
</taxonomicName>
|
||
and some species of the tribes
|
||
<taxonomicName id="C2EC0980FFAB124DFE3FEA5CFDEDFAD3" authorityName="Latreille" authorityYear="1802" box="[402,528,1322,1346]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1757" phylum="Arthropoda" rank="tribe" tribe="Scarabaeini">Scarabaeini</taxonomicName>
|
||
(
|
||
<bibRefCitation id="617D0FF2FFAB124DFDB2EA5CFD33FAD3" author="Janssens A" box="[543,718,1322,1346]" pageId="7" pageNumber="1757" pagination="1 - 81" refId="ref6702" refString="Janssens A. 1940 a. Monographie des Scarabaeus et genres voisins. Memoires du Musee R D ' Histoire Nat Belgique. 2: 1 - 81." type="journal article" year="1940">Janssens 1940a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="617D0FF2FFAB124DFD73EA5CFBFFFAD3" author="Harrison JDGi Philips TK" box="[734,1026,1322,1346]" pageId="7" pageNumber="1757" pagination="47 - 71" refId="ref6555" refString="Harrison JDGi Philips TK. 2003. Phylogeny of Scarabaeus (Pachysoma McLeay) stat. nov. i and related flightless Scarabaeini (Scarabaeidae: Scarabaeinae). Ann Transvaal Museum. 40: 47 - 71." type="journal article" year="2003">Harrison and Philips 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="617D0FF2FFAB124DFBBFEA5CFEC5FAF4" author="Zidek Ji Pokorny S" pageId="7" pageNumber="1757" pagination="1 - 28" refId="ref7243" refString="Zidek Ji Pokorny S. 2008. Illustrated keys to Palearctic Scarabaeus Linne (Scarabaeidae). Animma X. 27: 1 - 28." type="journal article" year="2008">Zídek and Pokorný 2008</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="C2EC0980FFAB124DFED5EA3BFDE1FAF4" authorityName="Lacordaire" authorityYear="1856" box="[376,540,1357,1381]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1757" phylum="Arthropoda" rank="tribe" tribe="Gymnopleurini">Gymnopleurini</taxonomicName>
|
||
(
|
||
<bibRefCitation id="617D0FF2FFAB124DFD84EA3BFD2AFAF4" author="Janssens A" box="[553,727,1357,1381]" pageId="7" pageNumber="1757" pagination="1 - 74" refId="ref6731" refString="Janssens A. 1940 b. Monographie des Gymnopleurides (Coleoptera Lamellicornia). Memoires du Musee R D ' Histoire Nat Belgique. 2: 1 - 74." type="journal article" year="1940">Janssens 1940b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="617D0FF2FFAB124DFD4EEA3BFC0CFAF4" author="Pokorny Si Zidek J" box="[739,1009,1357,1381]" pageId="7" pageNumber="1757" pagination="133 - 149" refId="ref6989" refString="Pokorny Si Zidek J. 2009. Review of the Gymnopleurini (Coleoptera: Scarabaeidae: Scarabaeinae) I. Introduction and the genus Allogymnopleurus. Folia Heyrovskyanai Ser A. 17: 133 - 149." type="journal article" year="2009">Pokorný and Zídek 2009</bibRefCitation>
|
||
<date id="715254C3FFAB124DFC5CEA3BFBC9FAF4" box="[1009,1076,1357,1381]" pageId="7" pageNumber="1757" value="2014-01">
|
||
ı
|
||
<bibRefCitation id="617D0FF2FFAB124DFC53EA3BFBC9FAF4" author="Pokorny Si Zidek J" box="[1022,1076,1357,1381]" pageId="7" pageNumber="1757" pagination="1 - 10" refId="ref7026" refString="Pokorny Si Zidek J. 2014. Review of the Gymnopleurini (Coleoptera: Scarabaeidae: Scarabaeinae). II. The genus Paragymnopleurus Shipp. Insecta Mundi. 395: 1 - 10." type="journal article" year="2014">2014</bibRefCitation>
|
||
</date>
|
||
<date id="715254C3FFAB124DFB99EA3BFB8AFAF4" box="[1076,1143,1357,1381]" pageId="7" pageNumber="1757" value="2016-01">
|
||
ı
|
||
<bibRefCitation id="617D0FF2FFAB124DFBECEA3BFB8AFAF4" author="Pokorny Si Zidek J" box="[1089,1143,1357,1381]" pageId="7" pageNumber="1757" pagination="1 - 8" refId="ref7061" refString="Pokorny Si Zidek J. 2016. Review of the Gymnopleurini (Coleoptera: Scarabaeidae: Scarabaeinae). III. Asian species of Garreta Janssens. Insecta Mundi. 483: 1 - 8." type="journal article" year="2016">2016</bibRefCitation>
|
||
</date>
|
||
). Howeverı this genus can be separated from species of these two tribes by the mesocoxal axis that is strongly obliqueı which contrasts with
|
||
<taxonomicName id="C2EC0980FFAB124DFD7CEAE4FC78FA3B" authority="Molano-Rendón & Vaz-de- Mello, 2019" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" box="[721,901,1426,1450]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1754" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="3798AE11FFAB124DFD7CEAE4FC78FA3B" box="[721,901,1426,1450]" italics="true" pageId="7" pageNumber="1757">Atlantemolanum</emphasis>
|
||
</taxonomicName>
|
||
where the mesocoxae are almost parallel to the longitudinal body axis (
|
||
<figureCitation id="9DD76E86FFAB124DFD12EAC3FCE4FA5C" box="[703,793,1461,1485]" captionStart="Figure 2" captionStartId="5.[160,225,1645,1668]" captionTargetBox="[167,1153,441,1612]" captionTargetId="figure@5.[163,1153,437,1614]" captionTargetPageId="5" captionText="Figure 2. Male Atlantemolanum riehli. (a) head, dorsal view; (b) right elytron; (c) elytral apex; (d) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (e) metaesternum; (f) ventrites; (g) mesoleg; (h) metaleg." figureDoi="http://doi.org/10.5281/zenodo.3670385" httpUri="https://zenodo.org/record/3670385/files/figure.png" pageId="7" pageNumber="1757">Figure 2</figureCitation>
|
||
(e)). Alsoı in the tribe
|
||
<taxonomicName id="C2EC0980FFAB124DFBABEAC3FB79FA5C" authorityName="Latreille" authorityYear="1802" box="[1030,1156,1461,1485]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1757" phylum="Arthropoda" rank="tribe" tribe="Scarabaeini">Scarabaeini</taxonomicName>
|
||
the anterior angles of the genae have a notch-like emarginationı whereas in
|
||
<taxonomicName id="C2EC0980FFAB124DFF0DEA8CFEA9F983" authority="Molano-Rendón & Vaz-de- Mello, 2019" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" box="[160,340,1530,1554]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1754" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="3798AE11FFAB124DFF0DEA8CFEA9F983" box="[160,340,1530,1554]" italics="true" pageId="7" pageNumber="1757">Atlantemolanum</emphasis>
|
||
</taxonomicName>
|
||
the anterior angles of the genae are slightly projected (
|
||
<figureCitation id="9DD76E86FFAB124DFC6CEA8CFBD8F983" box="[961,1061,1530,1554]" captionStart="Figure 2" captionStartId="5.[160,225,1645,1668]" captionTargetBox="[167,1153,441,1612]" captionTargetId="figure@5.[163,1153,437,1614]" captionTargetPageId="5" captionText="Figure 2. Male Atlantemolanum riehli. (a) head, dorsal view; (b) right elytron; (c) elytral apex; (d) elytral sculpture: left, detail of puntctures of the striae and interstria, and right, detail of interstria showing bright callosities; (e) metaesternum; (f) ventrites; (g) mesoleg; (h) metaleg." figureDoi="http://doi.org/10.5281/zenodo.3670385" httpUri="https://zenodo.org/record/3670385/files/figure.png" pageId="7" pageNumber="1757">Figures 2</figureCitation>
|
||
(a)ı 4(a)). The tribe
|
||
<taxonomicName id="C2EC0980FFAB124DFEA7E96BFE53F9A4" authorityName="Lacordaire" authorityYear="1856" box="[266,430,1565,1589]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1757" phylum="Arthropoda" rank="tribe" tribe="Gymnopleurini">Gymnopleurini</taxonomicName>
|
||
is characterised by an elytral emarginationı which exposes part of the pleuron (
|
||
<bibRefCitation id="617D0FF2FFAB124DFE87E949FDCCF9C6" author="Pokorny Si Zidek J" box="[298,561,1599,1623]" pageId="7" pageNumber="1757" pagination="133 - 149" refId="ref6989" refString="Pokorny Si Zidek J. 2009. Review of the Gymnopleurini (Coleoptera: Scarabaeidae: Scarabaeinae) I. Introduction and the genus Allogymnopleurus. Folia Heyrovskyanai Ser A. 17: 133 - 149." type="journal article" year="2009">Pokorný and Zídek 2009</bibRefCitation>
|
||
)ı while the species of
|
||
<taxonomicName id="C2EC0980FFAB124DFCB1E949FC2DF9C6" authority="Molano-Rendón & Vaz-de- Mello, 2019" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" box="[796,976,1599,1623]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1754" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="3798AE11FFAB124DFCB1E949FC2DF9C6" box="[796,976,1599,1623]" italics="true" pageId="7" pageNumber="1757">Atlantemolanum</emphasis>
|
||
</taxonomicName>
|
||
do not have this emargination.
|
||
</paragraph>
|
||
<paragraph id="05537203FFAB1242FF6DE9F3FE74FE9F" blockId="7.[160,1157,801,1727]" lastBlockId="8.[160,1155,142,270]" lastPageId="8" lastPageNumber="1758" pageId="7" pageNumber="1757">
|
||
Additionallyı in a recent studyı
|
||
<bibRefCitation id="617D0FF2FFAB124DFD80E9F3FC88F90C" author="Tarasov Si Dimitrov D" box="[557,885,1669,1693]" pageId="7" pageNumber="1757" pagination="1 - 19" refId="ref7160" refString="Tarasov Si Dimitrov D. 2016. Multigene phylogenetic analysis redefines dung beetles relationships and classification (Coleoptera: Scarabaeidae: Scarabaeinae). BMC Evol Biol. 16: 1 - 19. doi: 10.1186 / s 12862 - 016 - 0822 - x" type="journal article" year="2016">Tarasov and Dimitrov (2016)</bibRefCitation>
|
||
proposed two synapomorphiesı wings with: (1) RP1 posterior sclerite represents small basal appendix of RP1ı and (2) RA4 significantly thinner than RP1ı arcuate and not parallel to RA4; RA4 fused basally with RP1ı for the tribe Deltochiliniı characters which are present in
|
||
<taxonomicName id="C2EC0980FFA41242FF0DEFA2FEAAFF7D" authority="Molano-Rendón & Vaz-de- Mello, 2019" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" box="[160,343,212,236]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1754" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="3798AE11FFA41242FF0DEFA2FEAAFF7D" box="[160,343,212,236]" italics="true" pageId="8" pageNumber="1758">Atlantemolanum</emphasis>
|
||
</taxonomicName>
|
||
. Given thisı
|
||
<taxonomicName id="C2EC0980FFA41242FE58EFA2FD51FF7D" authority="Molano-Rendón & Vaz-de- Mello, 2019" authorityName="Molano-Rendón & Vaz-de- Mello" authorityYear="2019" box="[501,684,212,236]" class="Insecta" family="Scarabaeidae" genus="Atlantemolanum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1754" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="3798AE11FFA41242FE58EFA2FD51FF7D" box="[501,684,212,236]" italics="true" pageId="8" pageNumber="1758">Atlantemolanum</emphasis>
|
||
</taxonomicName>
|
||
is placed in
|
||
<taxonomicName id="C2EC0980FFA41242FCFBEFA2FE78FE9F" authority="sensu Tarasov and Dimitrov (2016)" authorityName="sensu Tarasov and Dimitrov" authorityYear="2016" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1758" phylum="Arthropoda" rank="tribe" tribe="Deltochilini">
|
||
Deltochilini
|
||
<emphasis id="3798AE11FFA41242FC4EEFA2FBE2FF7D" box="[995,1055,212,236]" italics="true" pageId="8" pageNumber="1758">sensu</emphasis>
|
||
<bibRefCitation id="617D0FF2FFA41242FB80EFA2FE78FE9F" author="Tarasov Si Dimitrov D" pageId="8" pageNumber="1758" pagination="1 - 19" refId="ref7160" refString="Tarasov Si Dimitrov D. 2016. Multigene phylogenetic analysis redefines dung beetles relationships and classification (Coleoptera: Scarabaeidae: Scarabaeinae). BMC Evol Biol. 16: 1 - 19. doi: 10.1186 / s 12862 - 016 - 0822 - x" type="journal article" year="2016">Tarasov and Dimitrov (2016)</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |