184 lines
19 KiB
XML
184 lines
19 KiB
XML
<document id="855A71B30E8942CA55DA315CEBF1293F" ID-DOI="10.11646/zootaxa.4088.3.5" ID-GBIF-Dataset="5c560767-5327-4cfd-b788-92ca72baca93" ID-ISSN="1175-5326" ID-Zenodo-Dep="256371" ID-ZooBank="7C9F7AA3-B20E-4F5D-908D-3C9404BC1EBC" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457602836381" checkinUser="plazi" docAuthor="Chamon, Carine C." docDate="2016" docId="B95BCF3A942EB520FF0BF89CFC0CFCFD" docLanguage="en" docName="15579-5110-1-PB.pdf.imf" docOrigin="Zootaxa 4088 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acanthicus Agassiz 1829" docType="treatment" docVersion="10" lastPageNumber="397" masterDocId="4562B742942FB522FF9CFFABFFC9FFA0" masterDocTitle="Redescription of Acanthicus hystrix Agassiz, 1829 (Siluriformes: Loricariidae), with comments on the systematics and distribution of the genus" masterLastPageNumber="408" masterPageNumber="395" pageNumber="396" updateTime="1698177565356" updateUser="plazi">
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<mods:titleInfo id="251A87D4C98CA25B695064CB57CDDF04">
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<mods:title id="B203AF346D3D8B93B1137428C8F5D896">Redescription of Acanthicus hystrix Agassiz, 1829 (Siluriformes: Loricariidae), with comments on the systematics and distribution of the genus</mods:title>
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<mods:namePart id="5FD059383CB50940C4AA3B3D3DBCE4A6">Chamon, Carine C.</mods:namePart>
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<mods:typeOfResource id="AE75AEE026AD01FD7826602C5CEAB40A">text</mods:typeOfResource>
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<mods:title id="2D3873CD0B6402C5C27B375EB7F34840">Zootaxa</mods:title>
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<mods:part id="091969B41D675797895EFC90A64DA8D1">
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<mods:date id="6B9A9CFBA6A26225D8DF38A1223C71E7">2016</mods:date>
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<mods:detail id="84601041D05EB934D239FA0B0DEAD7C9" type="volume">
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<mods:number id="8A3C3B88D5BED172462EE0F16B1B05B2">4088</mods:number>
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<mods:detail id="FA8852BE272840D683D0B929B67792B4" type="issue">
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<mods:number id="F91E4B8510B6F1455AA835B00102EC54">3</mods:number>
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<mods:classification id="23928E0B2D13FDC77761EF689E8F95CA">journal article</mods:classification>
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<mods:identifier id="AEE4CA37EFF15E1B1AB4061B1241087D" type="DOI">10.11646/zootaxa.4088.3.5</mods:identifier>
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<treatment id="B95BCF3A942EB520FF0BF89CFC0CFCFD" ID-DOI="http://doi.org/10.5281/zenodo.5662325" ID-GBIF-Taxon="125783193" ID-Zenodo-Dep="5662325" LSID="urn:lsid:plazi:treatment:B95BCF3A942EB520FF0BF89CFC0CFCFD" httpUri="http://treatment.plazi.org/id/B95BCF3A942EB520FF0BF89CFC0CFCFD" lastPageId="2" lastPageNumber="397" pageId="1" pageNumber="396">
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<subSubSection id="79E82DA7942EB523FF0BF89CFB30F872" pageId="1" pageNumber="396" type="nomenclature">
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<paragraph id="314D7E2C942EB523FF0BF89CFE06F8F2" blockId="1.[151,463,1847,1874]" box="[151,463,1847,1874]" pageId="1" pageNumber="396">
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<heading id="6A05C940942EB523FF0BF89CFE06F8F2" bold="true" box="[151,463,1847,1874]" fontSize="11" level="1" pageId="1" pageNumber="396" reason="1">
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<taxonomicName id="F6F205AF942EB523FF0BF89CFE06F8F2" ID-CoL="LZY" ID-ENA="337731" authority="Agassiz, 1829" authorityName="Agassiz" authorityYear="1829" box="[151,463,1847,1874]" class="Actinopterygii" family="Loricariidae" genus="Acanthicus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="396" phylum="Chordata" rank="genus">
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<emphasis id="0386A23E942EB523FF0BF89CFE06F8F2" bold="true" box="[151,463,1847,1874]" pageId="1" pageNumber="396">
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<emphasis id="0386A23E942EB523FF0BF89CFEE9F8F1" bold="true" box="[151,288,1847,1873]" italics="true" pageId="1" pageNumber="396">Acanthicus</emphasis>
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Agassiz, 1829
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="314D7E2C942EB523FF0BF8D5FD99F813" blockId="1.[151,1437,1918,2002]" pageId="1" pageNumber="396">
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<taxonomicName id="F6F205AF942EB523FF0BF8D5FEADF834" authority="Agassiz" authorityName="Agassiz" box="[151,356,1918,1940]" class="Actinopterygii" family="Loricariidae" genus="Acanthicus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="396" phylum="Chordata" rank="genus">
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<emphasis id="0386A23E942EB523FF0BF8D5FEC1F834" box="[151,264,1918,1940]" italics="true" pageId="1" pageNumber="396">Acanthicus</emphasis>
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Agassiz
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</taxonomicName>
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in Spix & Agassiz, 1829: 2. Type species:
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<taxonomicName id="F6F205AF942EB523FCCFF8D5FB72F834" authority="Agassiz, 1829" authorityName="Agassiz" authorityYear="1829" box="[851,1211,1918,1940]" class="Actinopterygii" family="Loricariidae" genus="Acanthicus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="396" phylum="Chordata" rank="species" species="hystrix">
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<emphasis id="0386A23E942EB523FCCFF8D5FBDCF834" box="[851,1045,1918,1940]" italics="true" pageId="1" pageNumber="396">Acanthicus hystrix</emphasis>
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Agassiz, 1829
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</taxonomicName>
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. Type by monotypy. Authorship follows Kottelat, 1988: 78.
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</paragraph>
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<paragraph id="314D7E2C942EB523FF0BF817FB30F872" blockId="1.[151,1437,1918,2002]" box="[151,1273,1980,2002]" pageId="1" pageNumber="396">
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—Bleeker, 1862 [tribe Acanthicini].—Isbrücker & Nijssen, 1988 [description of
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<taxonomicName id="F6F205AF942EB523FC57F817FBE3F872" box="[971,1066,1980,2002]" class="Actinopterygii" family="Loricariidae" genus="Acanthicus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="396" phylum="Chordata" rank="species" species="adonis">
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<emphasis id="0386A23E942EB523FC57F817FBE3F872" box="[971,1066,1980,2002]" italics="true" pageId="1" pageNumber="396">A. adonis</emphasis>
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</taxonomicName>
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; identification key].
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</paragraph>
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</subSubSection>
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<subSubSection id="79E82DA7942DB520FF0BFF3CFC0CFCFD" pageId="2" pageNumber="397" type="diagnosis">
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<paragraph id="314D7E2C942DB520FF0BFF3CFC0CFCFD" blockId="2.[151,1437,151,861]" pageId="2" pageNumber="397">
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<emphasis id="0386A23E942DB520FF0BFF3CFED8FF10" bold="true" box="[151,273,151,176]" pageId="2" pageNumber="397">Diagnosis.</emphasis>
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<taxonomicName id="F6F205AF942DB520FE87FF3CFE51FF0F" box="[283,408,151,175]" class="Actinopterygii" family="Loricariidae" genus="Acanthicus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="genus">
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<emphasis id="0386A23E942DB520FE87FF3CFE51FF0F" box="[283,408,151,175]" italics="true" pageId="2" pageNumber="397">Acanthicus</emphasis>
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</taxonomicName>
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can be diagnosed from other genera of
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<taxonomicName id="F6F205AF942DB520FCF5FF3CFC3BFF10" box="[873,1010,151,176]" class="Actinopterygii" family="Loricariidae" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="family">Loricariidae</taxonomicName>
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by the combination of the following characters: presence of enlarged compound pterotic, associated to the distention of the swim bladder capsule (
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<figureCitation id="A9C962A9942DB520FAF3FF17FF6CFF58" captionStart="FIGURE 2" captionStartId="5.[151,250,1985,2006]" captionTargetBox="[213,1374,193,1969]" captionTargetId="figure@5.[213,1376,193,1970]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal and ventral view of the cranium in Acanthicus hystrix, MZUSP 34244, 174.6 mm SL." httpUri="https://zenodo.org/record/256373/files/figure.png" pageId="2" pageNumber="397">Fig. 2</figureCitation>
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) (except
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<taxonomicName id="F6F205AF942DB520FE88FF74FE0BFF57" box="[276,450,223,247]" class="Actinopterygii" family="Loricariidae" genus="Megalancistrus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="genus">
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<emphasis id="0386A23E942DB520FE88FF74FE0BFF57" box="[276,450,223,247]" italics="true" pageId="2" pageNumber="397">Megalancistrus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="F6F205AF942DB520FE62FF4BFDABFF57" box="[510,610,224,247]" class="Actinopterygii" family="Loricariidae" genus="Panaque" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="genus">
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<emphasis id="0386A23E942DB520FE62FF4BFDABFF57" box="[510,610,224,247]" italics="true" pageId="2" pageNumber="397">Panaque</emphasis>
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</taxonomicName>
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gr.
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<taxonomicName id="F6F205AF942DB520FD0EFF74FCEFFF57" box="[658,806,223,247]" class="Actinopterygii" family="Loricariidae" genus="Panaque" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="species" species="nigrolineatus">
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<emphasis id="0386A23E942DB520FD0EFF74FCEFFF57" box="[658,806,223,247]" italics="true" pageId="2" pageNumber="397">nigrolineatus</emphasis>
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</taxonomicName>
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); presence of hypertrophied odontodes on opercle and cheek plates; presence of rows of keels, formed by hypertrophied odontodes, along the body.
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<taxonomicName id="F6F205AF942DB520FB50FEAFFA81FEBC" box="[1228,1352,260,284]" class="Actinopterygii" family="Loricariidae" genus="Acanthicus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="genus">
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<emphasis id="0386A23E942DB520FB50FEAFFA81FEBC" box="[1228,1352,260,284]" italics="true" pageId="2" pageNumber="397">Acanthicus</emphasis>
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</taxonomicName>
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can be further diagnosed from
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<taxonomicName id="F6F205AF942DB520FE3EFE8CFD99FE9F" box="[418,592,295,319]" class="Actinopterygii" family="Loricariidae" genus="Megalancistrus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="genus">
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<emphasis id="0386A23E942DB520FE3EFE8CFD99FE9F" box="[418,592,295,319]" italics="true" pageId="2" pageNumber="397">Megalancistrus</emphasis>
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</taxonomicName>
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by the oval shape of the swim bladder capsule surpassing the sixth rib and larger compound pterotic (
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<emphasis id="0386A23E942DB520FE57FEE7FE23FEC3" box="[459,490,332,355]" italics="true" pageId="2" pageNumber="397">vs.</emphasis>
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swimbladder capsule rounded, not surpassing the sixth rib and shorter compound pterotic;
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<figureCitation id="A9C962A9942DB520FF61FEC4FE89FE28" box="[253,320,367,392]" captionStart="FIGURE 3" captionStartId="6.[151,250,1993,2014]" captionTargetBox="[169,1416,193,1969]" captionTargetId="figure@6.[169,1416,193,1970]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal and ventral view of the cranium in Megalancistrus parananus, MZUSP 4434, 249.9 mm CP." httpUri="https://zenodo.org/record/256374/files/figure.png" pageId="2" pageNumber="397">Fig. 3</figureCitation>
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). From
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<taxonomicName id="F6F205AF942DB520FE01FEDBFDC8FE27" box="[413,513,368,391]" class="Actinopterygii" family="Loricariidae" genus="Panaque" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="genus">
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<emphasis id="0386A23E942DB520FE01FEDBFDC8FE27" box="[413,513,368,391]" italics="true" pageId="2" pageNumber="397">Panaque</emphasis>
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</taxonomicName>
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gr.
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<emphasis id="0386A23E942DB520FDB2FEC4FD70FE27" box="[558,697,367,391]" italics="true" pageId="2" pageNumber="397">nigolineatus</emphasis>
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it can be differentiated by the presence of long, narrow, viliform and curved teeth (
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<emphasis id="0386A23E942DB520FEF9FE3FFE4DFE0B" box="[357,388,404,427]" italics="true" pageId="2" pageNumber="397">vs.</emphasis>
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teeth short, wide and spoon shaped).
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<taxonomicName id="F6F205AF942DB520FCA8FE3FFC78FE0C" box="[820,945,404,428]" class="Actinopterygii" family="Loricariidae" genus="Acanthicus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="genus">
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<emphasis id="0386A23E942DB520FCA8FE3FFC78FE0C" box="[820,945,404,428]" italics="true" pageId="2" pageNumber="397">Acanthicus</emphasis>
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</taxonomicName>
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can be further distinguished from all other
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<taxonomicName id="F6F205AF942DB520FF0BFE1CFECEFE70" box="[151,263,439,464]" class="Actinopterygii" family="Loricariidae" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="tribe" tribe="Ancistrini">Ancistrini</taxonomicName>
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(except
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<taxonomicName id="F6F205AF942DB520FEF0FE1CFDC9FE6F" box="[364,512,439,463]" class="Actinopterygii" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="genus">
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<emphasis id="0386A23E942DB520FEF0FE1CFDC9FE6F" box="[364,512,439,463]" italics="true" pageId="2" pageNumber="397">Chaetostoma</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="F6F205AF942DB520FDA7FE13FD28FE6F" box="[571,737,440,463]" class="Actinopterygii" family="Loricariidae" genus="Leptoancistrus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="genus">
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<emphasis id="0386A23E942DB520FDA7FE13FD28FE6F" box="[571,737,440,463]" italics="true" pageId="2" pageNumber="397">Leptoancistrus</emphasis>
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</taxonomicName>
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) by the absence of an adipose fin (
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<emphasis id="0386A23E942DB520FBE0FE13FB5AFE6F" box="[1148,1171,440,463]" italics="true" pageId="2" pageNumber="397">vs</emphasis>
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. adipose fin present in remaining taxa). From
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<taxonomicName id="F6F205AF942DB520FE01FE77FDF8FE54" box="[413,561,476,500]" class="Actinopterygii" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="genus">
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<emphasis id="0386A23E942DB520FE01FE77FDF8FE54" box="[413,561,476,500]" italics="true" pageId="2" pageNumber="397">Chaetostoma</emphasis>
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</taxonomicName>
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it can be diagnosed by the presence of a snout completely covered by plates (
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<emphasis id="0386A23E942DB520FF3CFDABFF7EFDB7" box="[160,183,512,535]" italics="true" pageId="2" pageNumber="397">vs</emphasis>
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. snout with a wide naked area); and from
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<taxonomicName id="F6F205AF942DB520FD3CFDABFC8FFDB7" box="[672,838,512,535]" class="Actinopterygii" family="Loricariidae" genus="Leptoancistrus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="genus">
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<emphasis id="0386A23E942DB520FD3CFDABFC8FFDB7" box="[672,838,512,535]" italics="true" pageId="2" pageNumber="397">Leptoancistrus</emphasis>
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</taxonomicName>
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by the presence of the anal fin (
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<emphasis id="0386A23E942DB520FB21FDABFB1DFDB7" box="[1213,1236,512,535]" italics="true" pageId="2" pageNumber="397">vs</emphasis>
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. anal fin absent). Within the
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<taxonomicName id="F6F205AF942DB520FE85FD8FFE5FFD9C" box="[281,406,548,572]" class="Actinopterygii" family="Loricariidae" genus="Acanthicus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="genus">
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<emphasis id="0386A23E942DB520FE85FD8FFE5FFD9C" box="[281,406,548,572]" italics="true" pageId="2" pageNumber="397">Acanthicus</emphasis>
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</taxonomicName>
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group,
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<taxonomicName id="F6F205AF942DB520FE6DFD8FFDA4FD9C" box="[497,621,548,572]" class="Actinopterygii" family="Loricariidae" genus="Acanthicus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="genus">
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<emphasis id="0386A23E942DB520FE6DFD8FFDA4FD9C" box="[497,621,548,572]" italics="true" pageId="2" pageNumber="397">Acanthicus</emphasis>
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</taxonomicName>
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can be differentiated by the wide premaxillae and dentary (
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<emphasis id="0386A23E942DB520FAB8FD8FFAF2FD9B" box="[1316,1339,548,571]" italics="true" pageId="2" pageNumber="397">vs</emphasis>
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. narrow premaxillae and dentary); snout completely covered by small plates (
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<emphasis id="0386A23E942DB520FC04FDE3FC66FDFF" box="[920,943,584,607]" italics="true" pageId="2" pageNumber="397">vs</emphasis>
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. tip of snout with a naked area); presence of abdominal plates (
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<emphasis id="0386A23E942DB520FEFAFDC7FEB4FD23" box="[358,381,620,643]" italics="true" pageId="2" pageNumber="397">vs</emphasis>
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. abdominal plates absent); pectoral-fin spine, when adpressed, reaching almost the end of the unbranched pelvic-fin ray (
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<emphasis id="0386A23E942DB520FE4CFD3BFE2EFD07" box="[464,487,656,679]" italics="true" pageId="2" pageNumber="397">vs</emphasis>
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. adpressed pectoral-fin spine not surpassing middle of pelvic-fin unbranched ray); upper count of premaxillary teeth (more than 25 in each premaxillae
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<emphasis id="0386A23E942DB520FC3CFD1FFC7EFD6B" box="[928,951,692,715]" italics="true" pageId="2" pageNumber="397">vs</emphasis>
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. less than 25 in each premaxillae); dentary angle oblique (
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<emphasis id="0386A23E942DB520FEDAFD73FE94FD4F" box="[326,349,728,751]" italics="true" pageId="2" pageNumber="397">vs</emphasis>
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. right angle); straight bar shaped opercle (
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<emphasis id="0386A23E942DB520FCC8FD73FCA2FD4F" box="[852,875,728,751]" italics="true" pageId="2" pageNumber="397">vs</emphasis>
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. curved bar shaped opercle in other genera and triangular in
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<taxonomicName id="F6F205AF942DB520FEB6FD57FD86FCB3" box="[298,591,764,788]" class="Actinopterygii" family="Loricariidae" genus="Spectracanthicus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="species" species="murinus">
|
|
<emphasis id="0386A23E942DB520FEB6FD57FD86FCB3" box="[298,591,764,788]" italics="true" pageId="2" pageNumber="397">Spectracanthicus murinus</emphasis>
|
|
</taxonomicName>
|
|
); lateral line surpassing the hypural plate (
|
|
<emphasis id="0386A23E942DB520FBA4FD57FB86FCB3" box="[1080,1103,764,787]" italics="true" pageId="2" pageNumber="397">vs</emphasis>
|
|
. lateral line limited up to the hypural plate; except in
|
|
<taxonomicName id="F6F205AF942DB520FE2FFCB4FDA6FC97" box="[435,623,799,823]" class="Actinopterygii" family="Loricariidae" genus="Leporacanthicus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="genus">
|
|
<emphasis id="0386A23E942DB520FE2FFCB4FDA6FC97" box="[435,623,799,823]" italics="true" pageId="2" pageNumber="397">Leporacanthicus</emphasis>
|
|
</taxonomicName>
|
|
); posterior process of coracoid elongate (
|
|
<emphasis id="0386A23E942DB520FBC9FC8BFBA5FC97" box="[1109,1132,800,823]" italics="true" pageId="2" pageNumber="397">vs</emphasis>
|
|
. short in remaining taxa); presence of caudal-fin filaments (
|
|
<emphasis id="0386A23E942DB520FD93FCEFFDEFFCFB" box="[527,550,836,859]" italics="true" pageId="2" pageNumber="397">vs</emphasis>
|
|
. absent, except in
|
|
<taxonomicName id="F6F205AF942DB520FD6BFCEFFC7CFCFC" box="[759,949,836,860]" class="Actinopterygii" family="Loricariidae" genus="Pseudacanthicus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="397" phylum="Chordata" rank="genus">
|
|
<emphasis id="0386A23E942DB520FD6BFCEFFC7CFCFC" box="[759,949,836,860]" italics="true" pageId="2" pageNumber="397">Pseudacanthicus</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |