treatments-xml/data/87/5F/1C/875F1C5EFFDDFFDCFC23FEC3FAC3FB08.xml

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<mods:title id="161B8B0A9FD5B10FA70299DDEF4D2141">New cave species of Pectenoniscus Andersson, 1960 (Isopoda: Oniscidea: Styloniscidae) and an identification key for the genus</mods:title>
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<mods:namePart id="C38AE2B675154ECBDEBA468266E7821B">Ferreira, Giovanna Monticelli Cardoso Rafaela Bastos-Pereira Leila Aparecida Souza Rodrigo Lopes</mods:namePart>
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<heading id="54011A24FFDDFFC0FC23FEC3FAECFEE9" bold="true" box="[933,1376,266,295]" centered="true" fontSize="12" level="2" pageId="12" pageNumber="13" reason="2">
<emphasis id="3D82715AFFDDFFC0FC23FEC3FAECFEE9" bold="true" box="[933,1376,266,295]" pageId="12" pageNumber="13">
<taxonomicName id="C8F6D6CBFFDDFFC0FC23FEC3FA9AFEE9" ID-CoL="84C4H" authorityName="Ferreira" authorityYear="2020" box="[933,1302,266,295]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="carinhanhensis" status="sp. nov.">
<emphasis id="3D82715AFFDDFFC0FC23FEC3FA9AFEE9" bold="true" box="[933,1302,266,295]" italics="true" pageId="12" pageNumber="13">Pectenoniscus carinhanhensis</emphasis>
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<taxonomicNameLabel id="26B1CC21FFDDFFC0FA9BFEC3FAECFEE9" box="[1309,1376,267,295]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="0F49AD48FFDDFFC0FC7BFEFAFA84FE81" blockId="12.[864,1445,266,413]" box="[1021,1288,306,335]" pageId="12" pageNumber="13">
(
<figureCitation id="97CDB1CDFFDDFFC0FB8FFEFAFB1FFE81" box="[1033,1171,306,335]" captionStart-0="Figure 11" captionStart-1="Figure 12" captionStart-2="Figure 13" captionStartId-0="13.[181,248,1403,1426]" captionStartId-1="14.[181,250,1898,1921]" captionStartId-2="15.[181,249,1528,1551]" captionTargetBox-0="[198,1457,156,1367]" captionTargetBox-1="[200,1440,696,1866]" captionTargetBox-2="[181,1466,151,1498]" captionTargetId-0="figure-26@13.[181,1466,151,1379]" captionTargetId-1="figure-232@14.[193,1452,695,1874]" captionTargetId-2="figure-27@15.[181,1466,151,1498]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="Figure 11.Pectenoniscus carinhanhensis n.sp.Maleparatype:A, habitus, dorsalview; B, cephalon,frontal view; C, pleonite 5,pleotelson and uropod, dorsal view; D, antennula; E, antenna; F, left mandible; G, right mandible; H, maxillula; I, maxilla; J, maxilliped. Scale bars: A: 0,5 mm; B–J: 0.2 mm." captionText-1="Figure 12. Pectenoniscus carinhanhensis n. sp. Male paratype: A, uropod; B, pereopod 1; C, pereopod 6; D, pereopod 7; E, genital papilla and pleopod 1; F, pleopod 2; G, pleopod 3 exopod; H, pleopod 4 exopod; I, pleopod 5 exopod. Scale bars: 0.2 mm." captionText-2="Figure 13. Pectenoniscus carinhanhensisn. sp.A, Águas Claras karst system (extensionof caves in red, circles mark species occurrence; B, karst area of Serra do Ramalho; C, main conduit of Águas Claras cave; D, accumulated organic material in the conduit of Águas Claras cave; E, main conduit of Peixes II cave; F, specimen of Pectenoniscus carinhanhensis n. sp., approximately 3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14668050" figureDoi-1="http://doi.org/10.5281/zenodo.14668052" figureDoi-2="http://doi.org/10.5281/zenodo.14668054" httpUri-0="https://zenodo.org/record/14668050/files/figure.png" httpUri-1="https://zenodo.org/record/14668052/files/figure.png" httpUri-2="https://zenodo.org/record/14668054/files/figure.png" pageId="12" pageNumber="13">Figs. 11–13</figureCitation>
, 
<figureCitation id="97CDB1CDFFDDFFC0FB18FEFAFB31FE81" box="[1182,1213,306,335]" captionStart="Figure 21" captionStartId="24.[181,250,1235,1258]" captionTargetBox="[181,1466,174,1180]" captionTargetId="figure-27@24.[181,1466,166,1181]" captionTargetPageId="24" captionText="Figure 21. Map of South America highlighting Brazil and the states of Bahia (BA), Minas Gerais (MG), and Santa Catarina (SC), where the species of Pectenoniscus are recorded. Brown and green colors indicate the elevation of the terrain, with brown for higher levels, green for lower levels; gray indicates karstic areas." figureDoi="http://doi.org/10.5281/zenodo.14668070" httpUri="https://zenodo.org/record/14668070/files/figure.png" pageId="12" pageNumber="13">21</figureCitation>
, 
<figureCitation id="97CDB1CDFFDDFFC0FB41FEFAFB77FE81" box="[1223,1275,306,335]" captionStart="Figure 22" captionStartId="26.[184,255,1379,1402]" captionTargetBox="[181,1469,151,1342]" captionTargetId="figure-26@26.[181,1469,151,1342]" captionTargetPageId="26" captionText="Figure 22. Satellite images of localities: A, Abrigo da Ema (Montalvânia municipality); B, Gruta do Tabuleirinho (Juvenilia municipality); C, Baixa da Fortuna cave (Iuiu municipality); D, Gruna da Água Clara Cave, Gruna dos Índios cave and Gruna dos Peixes II, from top to bottom (Carinhanha municipality); E, Gruta do Padre cave (Santana municipality); F, Gruta dos Brejões (Morro do Chapéu municipality)." figureDoi="http://doi.org/10.5281/zenodo.14668072" httpUri="https://zenodo.org/record/14668072/files/figure.png" pageId="12" pageNumber="13">22D</figureCitation>
)
</paragraph>
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Zoobank: 
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urn:lsid:zoobank.org:act: 
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<emphasis id="3D82715AFFDDFFC0FCF7FE06FB97FE24" box="[881,1051,461,490]" italics="true" pageId="12" pageNumber="13">Type material</emphasis>
: 
<materialsCitation id="BF9EA715FFDDFFC0FBABFE05FB5AFD91" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Brazil" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
<typeStatus id="D04D13EAFFDDFFC0FBABFE05FB26FE24" box="[1069,1194,461,490]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
, male (ISLA 50648), 
<collectingCountry id="77E1EDD8FFDDFFC0FCCBFE3CFC1AFDDF" box="[845,918,500,529]" name="Brazil" pageId="12" pageNumber="13">Brazil</collectingCountry>
, 
<collectingRegion id="CD3263AAFFDDFFC0FC18FE3CFC69FDDF" box="[926,997,500,529]" country="Brazil" name="Bahia" pageId="12" pageNumber="13">Bahia</collectingRegion>
, 
<collectingCounty id="E628D5C4FFDDFFC0FC69FE3CFB4BFDDF" box="[1007,1223,500,529]" pageId="12" pageNumber="13">Serra do Ramalho</collectingCounty>
, 
<collectingMunicipality id="EF2D3732FFDDFFC0FB56FE3CFAE8FDDF" box="[1232,1380,500,529]" pageId="12" pageNumber="13">Carinhanha</collectingMunicipality>
, 
<location id="0A29FB93FFDDFFC0FAE8FE3CFBCDFDF6" LSID="urn:lsid:plazi:treatment:875F1C5EFFDDFFDCFC23FEC3FAC3FB08:0A29FB93FFDDFFC0FAE8FE3CFBCDFDF6" country="Brazil" county="Serra do Ramalho" municipality="Carinhanha" name="Gruna da Agua Clara Cave" pageId="12" pageNumber="13" stateProvince="Bahia">Gruna da Água Clara Cave</location>
(-13,801182º -43,951733º), 
<date id="7B488B88FFDDFFC0FA1AFDD3FC79FD91" pageId="12" pageNumber="13" value="2017-10-11">
<collectingDate id="6B0C7260FFDDFFC0FA1AFDD3FC79FD91" pageId="12" pageNumber="13" value="2017-10-11">11 October 2017</collectingDate>
</date>
, leg. 
<collectionCode id="69E7358DFFDDFFC0FBB4FD8BFBC6FD91" box="[1074,1098,579,607]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="13">R</collectionCode>
. 
<collectorName id="A203C89EFFDDFFC0FBC8FD8BFB5AFD91" box="[1102,1238,578,607]" pageId="12" pageNumber="13">L. Ferreira</collectorName>
</materialsCitation>
. 
<materialsCitation id="BF9EA715FFDDFFC0FB66FD8BFB72FD48" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" stateProvince="Bahia" typeStatus="paratype">
<typeStatus id="D04D13EAFFDDFFC0FB66FD8BFAD2FD91" box="[1248,1374,579,607]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="19F066C1FFDDFFC0FAECFD8AFA30FD91" box="[1386,1468,578,607]" count="1" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FCD2FDA1FC14FD48" box="[852,920,617,646]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50614) same data as holotype
</materialsCitation>
; 
<materialsCitation id="BF9EA715FFDDFFC0FA81FDA1FB41FD63" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FA81FDA1FAE7FD48" box="[1287,1387,617,646]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FAFEFDA1FA30FD48" box="[1400,1468,617,646]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50615), same data as holotype
</materialsCitation>
; 
<materialsCitation id="BF9EA715FFDDFFC0FB5CFD58FA8CFD1A" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FB5CFD58FAB6FD63" box="[1242,1338,656,685]" count="2" pageId="12" pageNumber="13" type="male">2 males</specimenCount>
<specimenCount id="19F066C1FFDDFFC0FAC4FD58FA34FD63" box="[1346,1464,656,685]" count="3" pageId="12" pageNumber="13" type="female">3 females</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FCD2FD7FFC1BFD1A" box="[852,919,695,724]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50620), same data as holotype
</materialsCitation>
; 
<materialsCitation id="BF9EA715FFDDFFC0FA8FFD7FFB45FD35" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FA8FFD7FFAE0FD1A" box="[1289,1388,695,724]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FAFFFD7FFA31FD1A" box="[1401,1469,695,724]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50633), same data as holotype
</materialsCitation>
; 
<specimenCount id="19F066C1FFDDFFC0FB50FD16FAA5FD35" box="[1238,1321,734,763]" count="1" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(in slide) 
<materialsCitation id="BF9EA715FFDDFFC0FA2CFD16FA11FCEC" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="2" specimenCount-female="2" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FA2CFD16FC22FCEC" count="2" pageId="12" pageNumber="13" type="female">2 females</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FC47FCCDFB80FCEC" box="[961,1036,773,802]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50635), same data as holotype
</materialsCitation>
; 
<materialsCitation id="BF9EA715FFDDFFC0FA2AFCCEFAC3FC87" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FA2AFCCEFC16FC87" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FC20FCE5FC65FC84" box="[934,1001,813,842]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50643), same data as holotype
</materialsCitation>
; 
<materialsCitation id="BF9EA715FFDDFFC0FADEFCE5FB72FCBE" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FADEFCE5FA37FC87" box="[1368,1467,812,842]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FCD7FC9CFC18FCBF" box="[849,916,852,881]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50644), same data as holotype
</materialsCitation>
; 
<materialsCitation id="BF9EA715FFDDFFC0FA81FC9CFB44FC59" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FA81FC9CFAE5FCBE" box="[1287,1385,851,881]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FAF0FC9CFA35FCBF" box="[1398,1465,852,881]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50645), same data as holotype
</materialsCitation>
; 
<materialsCitation id="BF9EA715FFDDFFC0FB52FCB3FA83FC70" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="16" specimenCount-female="13" specimenCount-male="3" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FB52FCB3FAB9FC59" box="[1236,1333,890,920]" count="3" pageId="12" pageNumber="13" type="male">3 males</specimenCount>
, 
<specimenCount id="19F066C1FFDDFFC0FAC7FCB3FA35FC59" box="[1345,1465,890,920]" count="13" pageId="12" pageNumber="13" type="female">13 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FCD2FC6AFC15FC71" box="[852,921,930,959]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50653), same data as holotype
</materialsCitation>
; 
<materialsCitation id="BF9EA715FFDDFFC0FA9FFC6AFB29FC28" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FA9FFC6AFAE5FC70" box="[1305,1385,929,959]" count="1" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FAF1FC6AFA30FC71" box="[1399,1468,930,959]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50655), same data as holotype
</materialsCitation>
; 
<materialsCitation id="BF9EA715FFDDFFC0FB2BFC01FBE1FBC3" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FB2BFC01FA82FC28" box="[1197,1294,969,998]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FA9FFC01FAD7FC28" box="[1305,1371,969,998]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50656), same data as holotype
</materialsCitation>
; 
<materialsCitation id="BF9EA715FFDDFFC0FBFDFC38FBD3FBFA" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="2" specimenCount-female="2" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FBFDFC38FB79FBC3" box="[1147,1269,1008,1037]" count="2" pageId="12" pageNumber="13" type="female">2 females</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FA81FC38FAC3FBC3" box="[1287,1359,1008,1037]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50674), same data as holotype
</materialsCitation>
; 
<materialsCitation id="BF9EA715FFDDFFC0FBEDFBDFFBBBFB95" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FBEDFBDFFB31FBFA" box="[1131,1213,1047,1076]" count="1" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FB4AFBDFFA9DFBFA" box="[1228,1297,1047,1076]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50649), same location as holotype
</materialsCitation>
, 
<date id="7B488B88FFDDFFC0FBC6FBF6FA8EFB95" box="[1088,1282,1086,1115]" pageId="12" pageNumber="13" value="2017-10-10">10October 2017</date>
; 
<materialsCitation id="BF9EA715FFDDFFC0FA8DFBF6FB6EFB4C" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FA8DFBF6FAE0FB95" box="[1291,1388,1086,1115]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FAFFFBF6FA30FB95" box="[1401,1468,1086,1115]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50646), same location as holotype
</materialsCitation>
, 
<date id="7B488B88FFDDFFC0FB6DFBADFA39FB4C" box="[1259,1461,1125,1154]" pageId="12" pageNumber="13" value="2017-10-12">12 October 2017</date>
; 
<materialsCitation id="BF9EA715FFDDFFC0FCCCFB44FA1EFB67" box="[842,1426,1164,1193]" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FCCCFB44FC20FB67" box="[842,940,1164,1193]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FC3FFB44FC71FB67" box="[953,1021,1164,1193]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50650), same location as holotype
</materialsCitation>
, 
<date id="7B488B88FFDDFFC0FA1DFB44FC7FFB1E" pageId="12" pageNumber="13" value="2017-10-12">12 October 2017</date>
; 
<materialsCitation id="BF9EA715FFDDFFC0FC78FB7BFBBAFB39" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FC78FB7BFBC3FB1E" box="[1022,1103,1203,1232]" count="1" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
<specimenCount id="19F066C1FFDDFFC0FBD0FB7BFB31FB1E" box="[1110,1213,1203,1232]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FB4AFB7BFA9DFB1E" box="[1228,1297,1203,1232]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50673), same location as holotype
</materialsCitation>
, 
<date id="7B488B88FFDDFFC0FBB9FB12FA8EFB39" box="[1087,1282,1242,1271]" pageId="12" pageNumber="13" value="2017-10-12">12 October 2017</date>
; 
<materialsCitation id="BF9EA715FFDDFFC0FA8DFB12FB6EFAD0" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FA8DFB12FAE0FB39" box="[1291,1388,1242,1271]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FAFEFB12FA37FB39" box="[1400,1467,1242,1271]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50630), same location as holotype
</materialsCitation>
, 
<date id="7B488B88FFDDFFC0FB6DFAC9FA38FAD0" box="[1259,1460,1281,1310]" pageId="12" pageNumber="13" value="2017-10-16">16 October 2017</date>
; 
<materialsCitation id="BF9EA715FFDDFFC0FCCCFAE0FA18FA8B" box="[842,1428,1320,1349]" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FCCCFAE0FC20FA8B" box="[842,940,1320,1349]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FC3FFAE0FC71FA8B" box="[953,1021,1320,1349]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50647), same location as holotype
</materialsCitation>
, 
<date id="7B488B88FFDDFFC0FA1AFAE0FC62FAA2" pageId="12" pageNumber="13" value="2017-10-16">16 October 2017</date>
; 
<materialsCitation id="BF9EA715FFDDFFC0FC71FA87FC5FFA5D" collectingDate="2017-10-11" collectionCode="R" collectorName="L. Ferreira" country="Iceland" county="Serra do Ramalho" location="Gruna da Agua Clara Cave" municipality="Carinhanha" pageId="12" pageNumber="13" specimenCount="2" specimenCount-female="2" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FC71FA87FBE8FAA2" box="[1015,1124,1359,1388]" count="2" pageId="12" pageNumber="13" type="female">2 females</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FBF4FA87FB34FAA2" box="[1138,1208,1359,1388]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50658), same location as holotype
</materialsCitation>
, 
<date id="7B488B88FFDDFFC0FC5AFABEFB2DFA5D" box="[988,1185,1398,1427]" pageId="12" pageNumber="13" value="2017-10-17">17 October 2017</date>
; 
<materialsCitation id="BF9EA715FFDDFFC0FB2CFABEFA3BFA2F" collectingDate="2017-10-14" collectorName="Carinhanha &amp; Gruna dos Peixes II" country="Iceland" location="Bahia" municipality="Serra do Ramalho" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FB2CFABEFA80FA5D" box="[1194,1292,1398,1427]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FA9FFABEFAD2FA5D" box="[1305,1374,1398,1427]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50632), 
<location id="0A29FB93FFDDFFC0FCCFFA55FC1FFA74" LSID="urn:lsid:plazi:treatment:875F1C5EFFDDFFDCFC23FEC3FAC3FB08:0A29FB93FFDDFFC0FCCFFA55FC1FFA74" box="[841,915,1437,1466]" country="Iceland" municipality="Serra do Ramalho" name="Bahia" pageId="12" pageNumber="13">Bahia</location>
, 
<collectingMunicipality id="EF2D3732FFDDFFC0FC19FA55FB0FFA74" box="[927,1155,1437,1466]" pageId="12" pageNumber="13">Serra do Ramalho</collectingMunicipality>
, 
<collectorName id="A203C89EFFDDFFC0FB16FA55FAA5FA74" box="[1168,1321,1437,1466]" pageId="12" pageNumber="13">Carinhanha</collectorName>
, 
<collectorName id="A203C89EFFDDFFC0FAB3FA55FC3CFA2F" pageId="12" pageNumber="13">Gruna dos Peixes II</collectorName>
(-13.822815º -43.956776º), 
<date id="7B488B88FFDDFFC0FB73FA0CFA3BFA2F" box="[1269,1463,1476,1505]" pageId="12" pageNumber="13" value="2017-10-14">
<collectingDate id="6B0C7260FFDDFFC0FB73FA0CFA3BFA2F" box="[1269,1463,1476,1505]" pageId="12" pageNumber="13" value="2017-10-14">14 October 2017</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="BF9EA715FFDDFFC0FCCFFA23FB73F9C6" box="[841,1279,1515,1544]" collectingDate="2017-10-14" collectorName="Carinhanha &amp; Gruna dos Peixes II" country="Iceland" location="Bahia" municipality="Serra do Ramalho" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FCCFFA23FC20F9C6" box="[841,940,1515,1544]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FC31FA23FC76F9C6" box="[951,1018,1515,1544]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50651), same location
</materialsCitation>
; 
<materialsCitation id="BF9EA715FFDDFFC0FA8EFA23FBECF9E1" collectingDate="2017-10-14" collectorName="Carinhanha &amp; Gruna dos Peixes II" country="Iceland" location="Bahia" municipality="Serra do Ramalho" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FA8EFA23FAE6F9C6" box="[1288,1386,1515,1544]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FAF0FA23FA35F9C6" box="[1398,1465,1515,1544]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50659), same location
</materialsCitation>
, 
<date id="7B488B88FFDDFFC0FBEBF9DAFACEF9E1" box="[1133,1346,1554,1583]" pageId="12" pageNumber="13" value="2017-10-10">10 October 2017</date>
; 
<materialsCitation id="BF9EA715FFDDFFC0FAC8F9DAFB05F998" collectingDate="2017-10-14" collectorName="Carinhanha &amp; Gruna dos Peixes II" country="Iceland" location="Bahia" municipality="Serra do Ramalho" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FAC8F9DAFA35F9E1" box="[1358,1465,1554,1583]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FCD3F9F2FC17F999" box="[853,923,1594,1623]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50661), same data
</materialsCitation>
; 
<materialsCitation id="BF9EA715FFDDFFC0FB10F9F2FC04F902" collectingDate="2017-10-14" collectorName="Carinhanha &amp; Gruna" country="Iceland" county="Indios cave" location="Bahia" municipality="Serra do Ramalho" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="19F066C1FFDDFFC0FB10F9F2FA8CF998" box="[1174,1280,1593,1623]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectingCountry id="77E1EDD8FFDDFFC0FA89F9F2FAD9F999" box="[1295,1365,1594,1623]" name="Iceland" pageId="12" pageNumber="13">ISLA</collectingCountry>
50641), 
<location id="0A29FB93FFDDFFC0FCCAF9A8FC1AF9B3" LSID="urn:lsid:plazi:treatment:875F1C5EFFDDFFDCFC23FEC3FAC3FB08:0A29FB93FFDDFFC0FCCAF9A8FC1AF9B3" box="[844,918,1632,1661]" country="Iceland" county="Indios cave" municipality="Serra do Ramalho" name="Bahia" pageId="12" pageNumber="13">Bahia</location>
, 
<collectingMunicipality id="EF2D3732FFDDFFC0FC24F9A9FB0AF9B3" box="[930,1158,1632,1662]" pageId="12" pageNumber="13">Serra do Ramalho</collectingMunicipality>
, 
<collectorName id="A203C89EFFDDFFC0FB15F9A8FAA0F9B3" box="[1171,1324,1632,1661]" pageId="12" pageNumber="13">Carinhanha</collectorName>
, 
<collectorName id="A203C89EFFDDFFC0FABEF9A9FA05F9B0" box="[1336,1417,1633,1662]" pageId="12" pageNumber="13">Gruna</collectorName>
dos 
<collectingCounty id="E628D5C4FFDDFFC0FCCAF942FC5BF96A" box="[844,983,1672,1706]" pageId="12" pageNumber="13">Índios cave</collectingCounty>
(-13,801308º -43,951374º), 
<date id="7B488B88FFDDFFC0FAB7F940FC04F902" pageId="12" pageNumber="13" value="2017-10-14">
<collectingDate id="6B0C7260FFDDFFC0FAB7F940FC04F902" pageId="12" pageNumber="13" value="2017-10-14">14 October 2017</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="0F49AD48FFDDFFC0FCF7F934FA87F87A" blockId="12.[843,1468,1787,1972]" pageId="12" pageNumber="13">
<emphasis id="3D82715AFFDDFFC0FCF7F934FC63F8D6" box="[881,1007,1788,1816]" italics="true" pageId="12" pageNumber="13">Diagnosis.</emphasis>
Antennula with eleven aesthetascs; male pleopod 1 exopod triangular, with round apex and sinuous lateral margin; male pleopod 2 exopod subtriangular, and pleopod 2 endopod apex twisted with pointed apex, directed outward.
</paragraph>
<caption id="5B89FDC0FFDCFFC1FF33FAB3FE6BFA1C" ID-DOI="http://doi.org/10.5281/zenodo.14668050" ID-Zenodo-Dep="14668050" httpUri="https://zenodo.org/record/14668050/files/figure.png" pageId="13" pageNumber="14" startId="13.[181,248,1403,1426]" targetBox="[198,1457,156,1367]" targetPageId="13" targetType="figure">
<paragraph id="0F49AD48FFDCFFC1FF33FAB3FE6BFA1C" blockId="13.[180,1467,1402,1490]" pageId="13" pageNumber="14">
<emphasis id="3D82715AFFDCFFC1FF33FAB3FE90FA5C" bold="true" box="[181,284,1403,1426]" pageId="13" pageNumber="14">Figure 11.</emphasis>
<taxonomicName id="C8F6D6CBFFDCFFC1FE99FAB3FDA6FA5C" authorityName="Ferreira" authorityYear="2020" box="[287,554,1402,1426]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="carinhanhensis" status="sp. nov.">
<emphasis id="3D82715AFFDCFFC1FE99FAB3FDA6FA5C" box="[287,554,1402,1426]" italics="true" pageId="13" pageNumber="14">Pectenoniscus carinhanhensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26B1CC21FFDCFFC1FDA8FAB3FDD3FA5C" box="[558,607,1403,1426]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
Male paratype: 
<emphasis id="3D82715AFFDCFFC1FD71FAB2FC8BFA5C" bold="true" box="[759,775,1402,1426]" pageId="13" pageNumber="14">A</emphasis>
, habitus, dorsal view; 
<emphasis id="3D82715AFFDCFFC1FC5FFAB2FC65FA5C" bold="true" box="[985,1001,1402,1426]" pageId="13" pageNumber="14">B</emphasis>
, cephalon, frontal view; 
<emphasis id="3D82715AFFDCFFC1FB54FAB2FB6FFA5C" bold="true" box="[1234,1251,1402,1426]" pageId="13" pageNumber="14">C</emphasis>
, pleonite 5, pleotelson and uropod, dorsal view; 
<emphasis id="3D82715AFFDCFFC1FE28FA52FE4DFA7C" bold="true" box="[430,449,1434,1458]" pageId="13" pageNumber="14">D</emphasis>
, antennula; 
<emphasis id="3D82715AFFDCFFC1FDBCFA52FDC6FA7C" bold="true" box="[570,586,1434,1458]" pageId="13" pageNumber="14">E</emphasis>
, antenna; 
<emphasis id="3D82715AFFDCFFC1FD29FA52FD32FA7C" bold="true" box="[687,702,1434,1458]" pageId="13" pageNumber="14">F</emphasis>
, left mandible; 
<emphasis id="3D82715AFFDCFFC1FCD0FA52FCE5FA7C" bold="true" box="[854,873,1434,1458]" pageId="13" pageNumber="14">G</emphasis>
, right mandible; 
<emphasis id="3D82715AFFDCFFC1FB97FA52FBA9FA7C" bold="true" box="[1041,1061,1434,1458]" pageId="13" pageNumber="14">H</emphasis>
, maxillula; 
<emphasis id="3D82715AFFDCFFC1FB10FA52FB2CFA7C" bold="true" box="[1174,1184,1434,1458]" pageId="13" pageNumber="14">I</emphasis>
, maxilla; 
<emphasis id="3D82715AFFDCFFC1FB78FA52FA84FA7C" bold="true" box="[1278,1288,1434,1458]" pageId="13" pageNumber="14">J</emphasis>
, maxilliped. Scale bars: 
<emphasis id="3D82715AFFDCFFC1FF6DFA72FF70FA1C" bold="true" box="[235,252,1466,1490]" pageId="13" pageNumber="14">A</emphasis>
: 0,5 mm; 
<emphasis id="3D82715AFFDCFFC1FEE6FA72FE04FA1C" bold="true" box="[352,392,1466,1490]" pageId="13" pageNumber="14">B–J</emphasis>
: 0.2 mm.
</paragraph>
</caption>
<paragraph id="0F49AD48FFDCFFC2FF5DF9C3FD8FFDD6" blockId="13.[179,807,1546,1964]" lastBlockId="14.[180,806,153,653]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="3D82715AFFDCFFC1FF5DF9C3FEFCF9E9" box="[219,368,1547,1575]" italics="true" pageId="13" pageNumber="14">Description.</emphasis>
Maximum length: male, 
<quantity id="C80E00ADFFDCFFC1FD3CF9C2FCADF9E8" box="[698,801,1546,1575]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="14" unit="mm" value="3.5">3.5 mm</quantity>
. Colorless, eyes absent (
<figureCitation id="97CDB1CDFFDCFFC1FE77F9F9FDE2F980" box="[497,622,1585,1614]" captionStart="Figure 11" captionStartId="13.[181,248,1403,1426]" captionTargetBox="[198,1457,156,1367]" captionTargetId="figure-26@13.[181,1466,151,1379]" captionTargetPageId="13" captionText="Figure 11.Pectenoniscus carinhanhensis n.sp.Maleparatype:A, habitus, dorsalview; B, cephalon,frontal view; C, pleonite 5,pleotelson and uropod, dorsal view; D, antennula; E, antenna; F, left mandible; G, right mandible; H, maxillula; I, maxilla; J, maxilliped. Scale bars: A: 0,5 mm; B–J: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668050" httpUri="https://zenodo.org/record/14668050/files/figure.png" pageId="13" pageNumber="14">Figs. 11B</figureCitation>
, 
<figureCitation id="97CDB1CDFFDCFFC1FDF9F9F9FD23F980" box="[639,687,1585,1614]" captionStart="Figure 13" captionStartId="15.[181,249,1528,1551]" captionTargetBox="[181,1466,151,1498]" captionTargetId="figure-27@15.[181,1466,151,1498]" captionTargetPageId="15" captionText="Figure 13. Pectenoniscus carinhanhensisn. sp.A, Águas Claras karst system (extensionof caves in red, circles mark species occurrence; B, karst area of Serra do Ramalho; C, main conduit of Águas Claras cave; D, accumulated organic material in the conduit of Águas Claras cave; E, main conduit of Peixes II cave; F, specimen of Pectenoniscus carinhanhensis n. sp., approximately 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14668054" httpUri="https://zenodo.org/record/14668054/files/figure.png" pageId="13" pageNumber="14">13F</figureCitation>
). Dorsal granulations in three rows on pereonite 1, two on pereonites2 to7,smooth pleon (
<figureCitation id="97CDB1CDFFDCFFC1FDA4F9B6FD11F952" box="[546,669,1662,1692]" captionStart="Figure 11" captionStartId="13.[181,248,1403,1426]" captionTargetBox="[198,1457,156,1367]" captionTargetId="figure-26@13.[181,1466,151,1379]" captionTargetPageId="13" captionText="Figure 11.Pectenoniscus carinhanhensis n.sp.Maleparatype:A, habitus, dorsalview; B, cephalon,frontal view; C, pleonite 5,pleotelson and uropod, dorsal view; D, antennula; E, antenna; F, left mandible; G, right mandible; H, maxillula; I, maxilla; J, maxilliped. Scale bars: A: 0,5 mm; B–J: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668050" httpUri="https://zenodo.org/record/14668050/files/figure.png" pageId="13" pageNumber="14">Fig.11A,C</figureCitation>
).Cephalon with round antennal lobes; supra-antennal line bent in middle (
<figureCitation id="97CDB1CDFFDCFFC1FEC5F904FE41F927" box="[323,461,1740,1770]" captionStart="Figure 11" captionStartId="13.[181,248,1403,1426]" captionTargetBox="[198,1457,156,1367]" captionTargetId="figure-26@13.[181,1466,151,1379]" captionTargetPageId="13" captionText="Figure 11.Pectenoniscus carinhanhensis n.sp.Maleparatype:A, habitus, dorsalview; B, cephalon,frontal view; C, pleonite 5,pleotelson and uropod, dorsal view; D, antennula; E, antenna; F, left mandible; G, right mandible; H, maxillula; I, maxilla; J, maxilliped. Scale bars: A: 0,5 mm; B–J: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668050" httpUri="https://zenodo.org/record/14668050/files/figure.png" pageId="13" pageNumber="14">Fig. 11A, B</figureCitation>
). Pereonite 1 with anterior margin not surpassing median portion of cephalon; pereonites 3–7 posterior margin progressively more concave; pleonites 3–5 epimera posterior point slightly developed; pleon narrower than pereon (
<figureCitation id="97CDB1CDFFDCFFC1FD33F8A0FF46F862" captionStart="Figure 11" captionStartId="13.[181,248,1403,1426]" captionTargetBox="[198,1457,156,1367]" captionTargetId="figure-26@13.[181,1466,151,1379]" captionTargetPageId="13" captionText="Figure 11.Pectenoniscus carinhanhensis n.sp.Maleparatype:A, habitus, dorsalview; B, cephalon,frontal view; C, pleonite 5,pleotelson and uropod, dorsal view; D, antennula; E, antenna; F, left mandible; G, right mandible; H, maxillula; I, maxilla; J, maxilliped. Scale bars: A: 0,5 mm; B–J: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668050" httpUri="https://zenodo.org/record/14668050/files/figure.png" pageId="13" pageNumber="14">Fig. 11A, C</figureCitation>
). Pleotelson with concave margin and round apex (
<figureCitation id="97CDB1CDFFDCFFC1FCD0F9C2FC39F9E9" box="[854,949,1546,1575]" captionStart="Figure 11" captionStartId="13.[181,248,1403,1426]" captionTargetBox="[198,1457,156,1367]" captionTargetId="figure-26@13.[181,1466,151,1379]" captionTargetPageId="13" captionText="Figure 11.Pectenoniscus carinhanhensis n.sp.Maleparatype:A, habitus, dorsalview; B, cephalon,frontal view; C, pleonite 5,pleotelson and uropod, dorsal view; D, antennula; E, antenna; F, left mandible; G, right mandible; H, maxillula; I, maxilla; J, maxilliped. Scale bars: A: 0,5 mm; B–J: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668050" httpUri="https://zenodo.org/record/14668050/files/figure.png" pageId="13" pageNumber="14">Fig.11C</figureCitation>
). Antennula with three articles, distal article with eleven aesthetascs (
<figureCitation id="97CDB1CDFFDCFFC1FBF2F9F9FB55F980" box="[1140,1241,1585,1614]" captionStart="Figure 11" captionStartId="13.[181,248,1403,1426]" captionTargetBox="[198,1457,156,1367]" captionTargetId="figure-26@13.[181,1466,151,1379]" captionTargetPageId="13" captionText="Figure 11.Pectenoniscus carinhanhensis n.sp.Maleparatype:A, habitus, dorsalview; B, cephalon,frontal view; C, pleonite 5,pleotelson and uropod, dorsal view; D, antennula; E, antenna; F, left mandible; G, right mandible; H, maxillula; I, maxilla; J, maxilliped. Scale bars: A: 0,5 mm; B–J: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668050" httpUri="https://zenodo.org/record/14668050/files/figure.png" pageId="13" pageNumber="14">Fig. 11D</figureCitation>
). Antenna reaches distal margin of pereonite 2 when extended backward, fifth article of peduncle shorter than f lagellum, with one seta longer than first f lagellum article; f lagellum with three articles (
<figureCitation id="97CDB1CDFFDCFFC1FBB7F904FB02F924" box="[1073,1166,1740,1770]" captionStart="Figure 11" captionStartId="13.[181,248,1403,1426]" captionTargetBox="[198,1457,156,1367]" captionTargetId="figure-26@13.[181,1466,151,1379]" captionTargetPageId="13" captionText="Figure 11.Pectenoniscus carinhanhensis n.sp.Maleparatype:A, habitus, dorsalview; B, cephalon,frontal view; C, pleonite 5,pleotelson and uropod, dorsal view; D, antennula; E, antenna; F, left mandible; G, right mandible; H, maxillula; I, maxilla; J, maxilliped. Scale bars: A: 0,5 mm; B–J: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668050" httpUri="https://zenodo.org/record/14668050/files/figure.png" pageId="13" pageNumber="14">Fig.11E</figureCitation>
). Left mandible with two penicils (
<figureCitation id="97CDB1CDFFDCFFC1FC3DF93BFB97F8DE" box="[955,1051,1779,1808]" captionStart="Figure 11" captionStartId="13.[181,248,1403,1426]" captionTargetBox="[198,1457,156,1367]" captionTargetId="figure-26@13.[181,1466,151,1379]" captionTargetPageId="13" captionText="Figure 11.Pectenoniscus carinhanhensis n.sp.Maleparatype:A, habitus, dorsalview; B, cephalon,frontal view; C, pleonite 5,pleotelson and uropod, dorsal view; D, antennula; E, antenna; F, left mandible; G, right mandible; H, maxillula; I, maxilla; J, maxilliped. Scale bars: A: 0,5 mm; B–J: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668050" httpUri="https://zenodo.org/record/14668050/files/figure.png" pageId="13" pageNumber="14">Fig. 11F</figureCitation>
); right mandible with one penicil, 
<emphasis id="3D82715AFFDCFFC1FCCDF8D2FB8DF8F9" box="[843,1025,1818,1847]" italics="true" pageId="13" pageNumber="14">lacinia mobilis</emphasis>
leaf-shaped (
<figureCitation id="97CDB1CDFFDCFFC1FB35F8D2FAADF8F9" box="[1203,1313,1818,1847]" captionStart="Figure 11" captionStartId="13.[181,248,1403,1426]" captionTargetBox="[198,1457,156,1367]" captionTargetId="figure-26@13.[181,1466,151,1379]" captionTargetPageId="13" captionText="Figure 11.Pectenoniscus carinhanhensis n.sp.Maleparatype:A, habitus, dorsalview; B, cephalon,frontal view; C, pleonite 5,pleotelson and uropod, dorsal view; D, antennula; E, antenna; F, left mandible; G, right mandible; H, maxillula; I, maxilla; J, maxilliped. Scale bars: A: 0,5 mm; B–J: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668050" httpUri="https://zenodo.org/record/14668050/files/figure.png" pageId="13" pageNumber="14">Fig. 11G</figureCitation>
). Maxillula outer branch with 4+4 teeth, apically entire, and one plumose seta; inner branch with three penicils (
<figureCitation id="97CDB1CDFFDCFFC1FA0BF8A0FCF3F862" captionStart="Figure 11" captionStartId="13.[181,248,1403,1426]" captionTargetBox="[198,1457,156,1367]" captionTargetId="figure-26@13.[181,1466,151,1379]" captionTargetPageId="13" captionText="Figure 11.Pectenoniscus carinhanhensis n.sp.Maleparatype:A, habitus, dorsalview; B, cephalon,frontal view; C, pleonite 5,pleotelson and uropod, dorsal view; D, antennula; E, antenna; F, left mandible; G, right mandible; H, maxillula; I, maxilla; J, maxilliped. Scale bars: A: 0,5 mm; B–J: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668050" httpUri="https://zenodo.org/record/14668050/files/figure.png" pageId="13" pageNumber="14">Fig. 11H</figureCitation>
). Maxilla with bilobate apex, inner lobe wider than outer lobe with several setae on distal margin (
<figureCitation id="97CDB1CDFFDFFFC2FF47FF08FE9AFF13" box="[193,278,192,221]" captionStart="Figure 11" captionStartId="13.[181,248,1403,1426]" captionTargetBox="[198,1457,156,1367]" captionTargetId="figure-26@13.[181,1466,151,1379]" captionTargetPageId="13" captionText="Figure 11.Pectenoniscus carinhanhensis n.sp.Maleparatype:A, habitus, dorsalview; B, cephalon,frontal view; C, pleonite 5,pleotelson and uropod, dorsal view; D, antennula; E, antenna; F, left mandible; G, right mandible; H, maxillula; I, maxilla; J, maxilliped. Scale bars: A: 0,5 mm; B–J: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668050" httpUri="https://zenodo.org/record/14668050/files/figure.png" pageId="14" pageNumber="15">Fig.11I</figureCitation>
). Maxilliped basis enlarged on distal portion; palp with two setae; endite triangular, apex with one triangular penicil and two lateral spines (
<figureCitation id="97CDB1CDFFDFFFC2FD31FEC7FC9EFEE2" box="[695,786,271,300]" captionStart="Figure 11" captionStartId="13.[181,248,1403,1426]" captionTargetBox="[198,1457,156,1367]" captionTargetId="figure-26@13.[181,1466,151,1379]" captionTargetPageId="13" captionText="Figure 11.Pectenoniscus carinhanhensis n.sp.Maleparatype:A, habitus, dorsalview; B, cephalon,frontal view; C, pleonite 5,pleotelson and uropod, dorsal view; D, antennula; E, antenna; F, left mandible; G, right mandible; H, maxillula; I, maxilla; J, maxilliped. Scale bars: A: 0,5 mm; B–J: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668050" httpUri="https://zenodo.org/record/14668050/files/figure.png" pageId="14" pageNumber="15">Fig. 11J</figureCitation>
). Pereopod 1 antennal grooming brush composed of serrated scale setae on propodus and on sternal margin of carpus (
<figureCitation id="97CDB1CDFFDFFFC2FEB9FE4DFE2AFE6C" box="[319,422,389,418]" captionStart="Figure 12" captionStartId="14.[181,250,1898,1921]" captionTargetBox="[200,1440,696,1866]" captionTargetId="figure-232@14.[193,1452,695,1874]" captionTargetPageId="14" captionText="Figure 12. Pectenoniscus carinhanhensis n. sp. Male paratype: A, uropod; B, pereopod 1; C, pereopod 6; D, pereopod 7; E, genital papilla and pleopod 1; F, pleopod 2; G, pleopod 3 exopod; H, pleopod 4 exopod; I, pleopod 5 exopod. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668052" httpUri="https://zenodo.org/record/14668052/files/figure.png" pageId="14" pageNumber="15">Fig. 12B</figureCitation>
); dactylar seta bifid with thin setule. Uropod protopod triangular, as long as distal margin of pleotelson; exopod longer than endopod, inserted distally (
<figureCitation id="97CDB1CDFFDFFFC2FE0DFE32FE7CFDD6" box="[395,496,506,536]" captionStart="Figure 12" captionStartId="14.[181,250,1898,1921]" captionTargetBox="[200,1440,696,1866]" captionTargetId="figure-232@14.[193,1452,695,1874]" captionTargetPageId="14" captionText="Figure 12. Pectenoniscus carinhanhensis n. sp. Male paratype: A, uropod; B, pereopod 1; C, pereopod 6; D, pereopod 7; E, genital papilla and pleopod 1; F, pleopod 2; G, pleopod 3 exopod; H, pleopod 4 exopod; I, pleopod 5 exopod. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668052" httpUri="https://zenodo.org/record/14668052/files/figure.png" pageId="14" pageNumber="15">Fig. 12A</figureCitation>
).
</paragraph>
<paragraph id="0F49AD48FFDFFFC2FF5DFDEAFBA6FD43" blockId="14.[180,806,153,653]" lastBlockId="14.[841,1468,153,653]" pageId="14" pageNumber="15">
<emphasis id="3D82715AFFDFFFC2FF5DFDEAFE93FDF1" box="[219,287,546,575]" italics="true" pageId="14" pageNumber="15">Male:</emphasis>
Pereopods 6 and 7 (
<figureCitation id="97CDB1CDFFDFFFC2FD90FDEAFD12FDF0" box="[534,670,546,575]" captionStart="Figure 12" captionStartId="14.[181,250,1898,1921]" captionTargetBox="[200,1440,696,1866]" captionTargetId="figure-232@14.[193,1452,695,1874]" captionTargetPageId="14" captionText="Figure 12. Pectenoniscus carinhanhensis n. sp. Male paratype: A, uropod; B, pereopod 1; C, pereopod 6; D, pereopod 7; E, genital papilla and pleopod 1; F, pleopod 2; G, pleopod 3 exopod; H, pleopod 4 exopod; I, pleopod 5 exopod. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668052" httpUri="https://zenodo.org/record/14668052/files/figure.png" pageId="14" pageNumber="15">Fig. 12C, D</figureCitation>
) propodus with tufts of setae on tergal margin; with water conducting system; pereopod 7 ischium with straight sternal margin (
<figureCitation id="97CDB1CDFFDFFFC2FBA4FF51FB1FFF78" box="[1058,1171,153,182]" captionStart="Figure 12" captionStartId="14.[181,250,1898,1921]" captionTargetBox="[200,1440,696,1866]" captionTargetId="figure-232@14.[193,1452,695,1874]" captionTargetPageId="14" captionText="Figure 12. Pectenoniscus carinhanhensis n. sp. Male paratype: A, uropod; B, pereopod 1; C, pereopod 6; D, pereopod 7; E, genital papilla and pleopod 1; F, pleopod 2; G, pleopod 3 exopod; H, pleopod 4 exopod; I, pleopod 5 exopod. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668052" httpUri="https://zenodo.org/record/14668052/files/figure.png" pageId="14" pageNumber="15">Fig. 12D</figureCitation>
). Genital papilla (
<figureCitation id="97CDB1CDFFDFFFC2FA0CFF51FCF6FF2F" captionStart="Figure 12" captionStartId="14.[181,250,1898,1921]" captionTargetBox="[200,1440,696,1866]" captionTargetId="figure-232@14.[193,1452,695,1874]" captionTargetPageId="14" captionText="Figure 12. Pectenoniscus carinhanhensis n. sp. Male paratype: A, uropod; B, pereopod 1; C, pereopod 6; D, pereopod 7; E, genital papilla and pleopod 1; F, pleopod 2; G, pleopod 3 exopod; H, pleopod 4 exopod; I, pleopod 5 exopod. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668052" httpUri="https://zenodo.org/record/14668052/files/figure.png" pageId="14" pageNumber="15">Fig. 12E</figureCitation>
) medial portion slightly enlarged with narrow apex. Pleopod 1 (
<figureCitation id="97CDB1CDFFDFFFC2FBAEFF27FB1CFEC2" box="[1064,1168,239,268]" captionStart="Figure 12" captionStartId="14.[181,250,1898,1921]" captionTargetBox="[200,1440,696,1866]" captionTargetId="figure-232@14.[193,1452,695,1874]" captionTargetPageId="14" captionText="Figure 12. Pectenoniscus carinhanhensis n. sp. Male paratype: A, uropod; B, pereopod 1; C, pereopod 6; D, pereopod 7; E, genital papilla and pleopod 1; F, pleopod 2; G, pleopod 3 exopod; H, pleopod 4 exopod; I, pleopod 5 exopod. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668052" httpUri="https://zenodo.org/record/14668052/files/figure.png" pageId="14" pageNumber="15">Fig. 12E</figureCitation>
) protopod rectangular, two times wider than long; exopod triangular, with round apex and sinuous lateral margin; endopod longer than exopod. Pleopod 2 (
<figureCitation id="97CDB1CDFFDFFFC2FB55FEA7FAB9FE42" box="[1235,1333,367,396]" captionStart="Figure 12" captionStartId="14.[181,250,1898,1921]" captionTargetBox="[200,1440,696,1866]" captionTargetId="figure-232@14.[193,1452,695,1874]" captionTargetPageId="14" captionText="Figure 12. Pectenoniscus carinhanhensis n. sp. Male paratype: A, uropod; B, pereopod 1; C, pereopod 6; D, pereopod 7; E, genital papilla and pleopod 1; F, pleopod 2; G, pleopod 3 exopod; H, pleopod 4 exopod; I, pleopod 5 exopod. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668052" httpUri="https://zenodo.org/record/14668052/files/figure.png" pageId="14" pageNumber="15">Fig. 12F</figureCitation>
) protopod rectangular; exopod subtriangular, twice wider than long; endopod with distal article fourfold longer than proximal article, apex twisted with pointed apex, directed outward. Pleopod 3–5 exopods (
<figureCitation id="97CDB1CDFFDFFFC2FAACFDD3FA22FDF6" box="[1322,1454,539,568]" captionStart="Figure 12" captionStartId="14.[181,250,1898,1921]" captionTargetBox="[200,1440,696,1866]" captionTargetId="figure-232@14.[193,1452,695,1874]" captionTargetPageId="14" captionText="Figure 12. Pectenoniscus carinhanhensis n. sp. Male paratype: A, uropod; B, pereopod 1; C, pereopod 6; D, pereopod 7; E, genital papilla and pleopod 1; F, pleopod 2; G, pleopod 3 exopod; H, pleopod 4 exopod; I, pleopod 5 exopod. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14668052" httpUri="https://zenodo.org/record/14668052/files/figure.png" pageId="14" pageNumber="15">Fig. 12F–H</figureCitation>
) trapezoidal, bearing setae on outer margin; exopod 5 with dorsal lobe.
</paragraph>
<caption id="5B89FDC0FFDFFFC2FF33F8A2FAFFF86F" ID-DOI="http://doi.org/10.5281/zenodo.14668052" ID-Zenodo-Dep="14668052" httpUri="https://zenodo.org/record/14668052/files/figure.png" pageId="14" pageNumber="15" startId="14.[181,250,1898,1921]" targetBox="[200,1440,696,1866]" targetPageId="14" targetType="figure">
<paragraph id="0F49AD48FFDFFFC2FF33F8A2FAFFF86F" blockId="14.[181,1467,1897,1953]" pageId="14" pageNumber="15">
<emphasis id="3D82715AFFDFFFC2FF33F8A2FEADF84F" bold="true" box="[181,289,1898,1921]" pageId="14" pageNumber="15">Figure 12.</emphasis>
<taxonomicName id="C8F6D6CBFFDFFFC2FEACF8A2FDB7F84F" authorityName="Ferreira" authorityYear="2020" box="[298,571,1897,1921]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carinhanhensis" status="sp. nov.">
<emphasis id="3D82715AFFDFFFC2FEACF8A2FDB7F84F" box="[298,571,1897,1921]" italics="true" pageId="14" pageNumber="15">Pectenoniscus carinhanhensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26B1CC21FFDFFFC2FDC6F8A2FDF8F84F" box="[576,628,1898,1921]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
Male paratype: 
<emphasis id="3D82715AFFDFFFC2FC93F8A1FCAAF84F" bold="true" box="[789,806,1897,1921]" pageId="14" pageNumber="15">A</emphasis>
, uropod; 
<emphasis id="3D82715AFFDFFFC2FC0EF8A1FC15F84F" bold="true" box="[904,921,1897,1921]" pageId="14" pageNumber="15">B</emphasis>
, pereopod 1; 
<emphasis id="3D82715AFFDFFFC2FBA5F8A1FBB9F84F" bold="true" box="[1059,1077,1897,1921]" pageId="14" pageNumber="15">C</emphasis>
, pereopod 6; 
<emphasis id="3D82715AFFDFFFC2FB39F8A1FB5EF84F" bold="true" box="[1215,1234,1897,1921]" pageId="14" pageNumber="15">D</emphasis>
, pereopod 7; 
<emphasis id="3D82715AFFDFFFC2FADBF8A1FAE1F84F" bold="true" box="[1373,1389,1897,1921]" pageId="14" pageNumber="15">E</emphasis>
, genital papilla and pleopod 1; 
<emphasis id="3D82715AFFDFFFC2FE1EF841FE2BF86F" bold="true" box="[408,423,1929,1953]" pageId="14" pageNumber="15">F</emphasis>
, pleopod 2; 
<emphasis id="3D82715AFFDFFFC2FDA4F841FDB9F86F" bold="true" box="[546,565,1929,1953]" pageId="14" pageNumber="15">G</emphasis>
, pleopod 3 exopod; 
<emphasis id="3D82715AFFDFFFC2FC87F841FC99F86F" bold="true" box="[769,789,1929,1953]" pageId="14" pageNumber="15">H</emphasis>
, pleopod 4 exopod; 
<emphasis id="3D82715AFFDFFFC2FC66F841FC66F86F" bold="true" box="[992,1002,1929,1953]" pageId="14" pageNumber="15">I</emphasis>
, pleopod 5 exopod. Scale bars: 0.2 mm.
</paragraph>
</caption>
<caption id="5B89FDC0FFDEFFC3FF33FA30FAB1F980" ID-DOI="http://doi.org/10.5281/zenodo.14668054" ID-Zenodo-Dep="14668054" httpUri="https://zenodo.org/record/14668054/files/figure.png" pageId="15" pageNumber="16" startId="15.[181,249,1528,1551]" targetBox="[181,1466,151,1498]" targetPageId="15" targetType="figure">
<paragraph id="0F49AD48FFDEFFC3FF33FA30FAB1F980" blockId="15.[181,1467,1527,1615]" pageId="15" pageNumber="16">
<emphasis id="3D82715AFFDEFFC3FF33FA30FE92F9C1" bold="true" box="[181,286,1528,1551]" pageId="15" pageNumber="16">Figure 13.</emphasis>
<taxonomicName id="C8F6D6CBFFDEFFC3FEA4FA30FDBEF9C1" authorityName="Ferreira" authorityYear="2020" box="[290,562,1527,1551]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="carinhanhensis" status="sp. nov.">
<emphasis id="3D82715AFFDEFFC3FEA4FA30FDBEF9C1" box="[290,562,1527,1551]" italics="true" pageId="15" pageNumber="16">Pectenoniscus carinhanhensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26B1CC21FFDEFFC3FDB3FA3FFDEBF9C0" box="[565,615,1527,1550]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="3D82715AFFDEFFC3FDECFA3FFDF7F9C1" bold="true" box="[618,635,1527,1551]" pageId="15" pageNumber="16">A</emphasis>
, Águas Claras karst system (extension of caves in red, circles mark species occurrence; 
<emphasis id="3D82715AFFDEFFC3FF33F9DFFF4AF9E1" bold="true" box="[181,198,1559,1583]" pageId="15" pageNumber="16">B</emphasis>
, karst area of Serra do Ramalho; 
<emphasis id="3D82715AFFDEFFC3FD88F9DFFDACF9E1" bold="true" box="[526,544,1559,1583]" pageId="15" pageNumber="16">C</emphasis>
, main conduit of Águas Claras cave; 
<emphasis id="3D82715AFFDEFFC3FC0AF9DFFC13F9E1" bold="true" box="[908,927,1559,1583]" pageId="15" pageNumber="16">D</emphasis>
, accumulated organic material in the conduit of Águas Claras cave; 
<emphasis id="3D82715AFFDEFFC3FEB7F9FFFECDF981" bold="true" box="[305,321,1591,1615]" pageId="15" pageNumber="16">E</emphasis>
, main conduit of Peixes II cave; 
<emphasis id="3D82715AFFDEFFC3FD06F9FFFD03F981" bold="true" box="[640,655,1591,1615]" pageId="15" pageNumber="16">F</emphasis>
, specimen of 
<taxonomicName id="C8F6D6CBFFDEFFC3FC91F9F0FBA4F981" authorityName="Ferreira" authorityYear="2020" box="[791,1064,1591,1615]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="carinhanhensis" status="sp. nov.">
<emphasis id="3D82715AFFDEFFC3FC91F9F0FBA4F981" box="[791,1064,1591,1615]" italics="true" pageId="15" pageNumber="16">Pectenoniscus carinhanhensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26B1CC21FFDEFFC3FBABF9FFFBEDF980" box="[1069,1121,1591,1614]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
, approximately 3 mm.
</paragraph>
</caption>
<paragraph id="0F49AD48FFDEFFC3FF5DF969FD43F8C2" blockId="15.[181,805,1697,1804]" pageId="15" pageNumber="16">
<emphasis id="3D82715AFFDEFFC3FF5DF969FED2F970" box="[219,350,1697,1726]" italics="true" pageId="15" pageNumber="16">Etymology.</emphasis>
The specific epithet 
<taxonomicName id="C8F6D6CBFFDEFFC3FDE7F969FC95F970" authorityName="Ferreira" authorityYear="2020" box="[609,793,1697,1726]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="carinhanhensis">
“ 
<emphasis id="3D82715AFFDEFFC3FDECF969FC95F970" box="[618,793,1697,1726]" italics="true" pageId="15" pageNumber="16">carinhanhensis</emphasis>
</taxonomicName>
” refers to the municipality of Carinhanha (state of 
<collectingRegion id="CD3263AAFFDEFFC3FF31F927FE8DF8C2" box="[183,257,1775,1804]" country="Brazil" name="Bahia" pageId="15" pageNumber="16">Bahia</collectingRegion>
), where the new species was collected.
</paragraph>
<paragraph id="0F49AD48FFDEFFDCFF5DF8F3FE6AFE51" blockId="15.[181,805,1851,1959]" lastBlockId="16.[180,807,151,415]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="3D82715AFFDEFFC3FF5DF8F3FD5EF896" box="[219,722,1851,1880]" italics="true" pageId="15" pageNumber="16">
Remarks. 
<taxonomicName id="C8F6D6CBFFDEFFC3FED8F8F4FD5EF896" authorityName="Ferreira" authorityYear="2020" box="[350,722,1851,1880]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="carinhanhensis" status="sp. nov.">Pectenoniscus carinhanhensis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="26B1CC21FFDEFFC3FD5BF8F4FCA9F896" box="[733,805,1852,1880]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
differs from 
<taxonomicName id="C8F6D6CBFFDEFFC3FECBF8ABFE50F8B1" authorityName="Andersson" authorityYear="1960" box="[333,476,1890,1919]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="3D82715AFFDEFFC3FECBF8ABFE50F8B1" box="[333,476,1890,1919]" italics="true" pageId="15" pageNumber="16">P. angulatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="C8F6D6CBFFDEFFC3FD91F8ABFDE0F8B1" authorityName="Campo-Filho, Bichuette and Taiti" authorityYear="2019" box="[535,620,1890,1919]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="liliae">
<emphasis id="3D82715AFFDEFFC3FD91F8ABFDE0F8B1" box="[535,620,1890,1919]" italics="true" pageId="15" pageNumber="16">P. liliae</emphasis>
</taxonomicName>
by the number of aesthetascs on antennula (11 
<emphasis id="3D82715AFFDEFFC3FDA3F842FDE5F868" box="[549,617,1930,1958]" italics="true" pageId="15" pageNumber="16">versus</emphasis>
<quantity id="C80E00ADFFDEFFC3FDEAF842FD14F868" box="[620,664,1929,1959]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="15" pageNumber="16" unit="in" value="9.0">9 in</quantity>
<taxonomicName id="C8F6D6CBFFDEFFC3FD1AF842FCA9F868" authorityName="Andersson" authorityYear="1960" box="[668,805,1929,1958]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="3D82715AFFDEFFC3FD1AF842FCA9F868" box="[668,805,1929,1958]" italics="true" pageId="15" pageNumber="16">P.angulatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="C8F6D6CBFFDEFFC3FCFAF96AFC45F970" authorityName="Campo-Filho, Bichuette and Taiti" authorityYear="2019" box="[892,969,1697,1726]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="liliae">
<emphasis id="3D82715AFFDEFFC3FCFAF96AFC45F970" box="[892,969,1697,1726]" italics="true" pageId="15" pageNumber="16">P. liliae</emphasis>
</taxonomicName>
), by the sternal margin of male pereopod 7 ischium (straight 
<emphasis id="3D82715AFFDEFFC3FB91F901FBD7F92B" box="[1047,1115,1737,1765]" italics="true" pageId="15" pageNumber="16">versus</emphasis>
with a triangular projection in 
<taxonomicName id="C8F6D6CBFFDEFFC3FCCDF938FC5EF8C2" authorityName="Andersson" authorityYear="1960" box="[843,978,1775,1804]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="3D82715AFFDEFFC3FCCDF938FC5EF8C2" box="[843,978,1775,1804]" italics="true" pageId="15" pageNumber="16">P.angulatus</emphasis>
</taxonomicName>
and convex margin in 
<taxonomicName id="C8F6D6CBFFDEFFC3FB56F938FA90F8C2" authorityName="Campo-Filho, Bichuette and Taiti" authorityYear="2019" box="[1232,1308,1775,1804]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="liliae">
<emphasis id="3D82715AFFDEFFC3FB56F938FA90F8C2" box="[1232,1308,1775,1804]" italics="true" pageId="15" pageNumber="16">P.liliae</emphasis>
</taxonomicName>
), the shape of male pleopod 1 exopod (sinuous lateral margin 
<emphasis id="3D82715AFFDEFFC3FAF2F8DFFA35F8FD" box="[1396,1465,1815,1843]" italics="true" pageId="15" pageNumber="16">versus</emphasis>
straight in 
<taxonomicName id="C8F6D6CBFFDEFFC3FC4EF8F6FBDFF894" authorityName="Andersson" authorityYear="1960" box="[968,1107,1853,1882]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="3D82715AFFDEFFC3FC4EF8F6FBDFF894" box="[968,1107,1853,1882]" italics="true" pageId="15" pageNumber="16">P.angulatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="C8F6D6CBFFDEFFC3FB0FF8F6FB54F894" authorityName="Campo-Filho, Bichuette and Taiti" authorityYear="2019" box="[1161,1240,1853,1882]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="liliae">
<emphasis id="3D82715AFFDEFFC3FB0FF8F6FB54F894" box="[1161,1240,1853,1882]" italics="true" pageId="15" pageNumber="16">P. liliae</emphasis>
</taxonomicName>
), the shape of male pleopod 2 exopod (subtriangular 
<emphasis id="3D82715AFFDEFFC3FB48F8ACFA9EF84E" box="[1230,1298,1892,1920]" italics="true" pageId="15" pageNumber="16">versus</emphasis>
rectangular in 
<taxonomicName id="C8F6D6CBFFDEFFC3FCCDF843FC5BF869" authorityName="Andersson" authorityYear="1960" box="[843,983,1930,1959]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="3D82715AFFDEFFC3FCCDF843FC5BF869" box="[843,983,1930,1959]" italics="true" pageId="15" pageNumber="16">P.angulatus</emphasis>
</taxonomicName>
and ovoid in 
<taxonomicName id="C8F6D6CBFFDEFFC3FBF2F843FB48F869" authorityName="Campo-Filho, Bichuette and Taiti" authorityYear="2019" box="[1140,1220,1930,1959]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="liliae">
<emphasis id="3D82715AFFDEFFC3FBF2F843FB48F869" box="[1140,1220,1930,1959]" italics="true" pageId="15" pageNumber="16">P. liliae</emphasis>
</taxonomicName>
) and the orientation of endopod (twisted 
<emphasis id="3D82715AFFC1FFDCFE21FF50FE67FF7A" box="[423,491,152,180]" italics="true" pageId="16" pageNumber="17">versus</emphasis>
straight in 
<taxonomicName id="C8F6D6CBFFC1FFDCFDEDFF50FD79FF7A" authorityName="Andersson" authorityYear="1960" box="[619,757,151,180]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="3D82715AFFC1FFDCFDEDFF50FD79FF7A" box="[619,757,151,180]" italics="true" pageId="16" pageNumber="17">P.angulatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="C8F6D6CBFFC1FFDCFF32FF77FE8AFF15" authorityName="Campo-Filho, Bichuette and Taiti" authorityYear="2019" box="[180,262,190,219]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="liliae">
<emphasis id="3D82715AFFC1FFDCFF32FF77FE8AFF15" box="[180,262,190,219]" italics="true" pageId="16" pageNumber="17">P. liliae</emphasis>
</taxonomicName>
). 
<taxonomicName id="C8F6D6CBFFC1FFDCFE98FF77FDE3FF15" authorityName="Ferreira" authorityYear="2020" box="[286,623,190,219]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="carinhanhensis" status="sp. nov.">
<emphasis id="3D82715AFFC1FFDCFE98FF77FDE3FF15" box="[286,623,190,219]" italics="true" pageId="16" pageNumber="17">Pectenoniscus carinhanhensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26B1CC21FFC1FFDCFDF2FF77FD3EFF15" box="[628,690,191,219]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
is similar to 
<taxonomicName id="C8F6D6CBFFC1FFDCFF50FF2EFED5FECC" authorityName="Ferreira" authorityYear="2020" box="[214,345,230,258]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="iuiuensis" status="n. sp.">
<emphasis id="3D82715AFFC1FFDCFF50FF2EFED5FECC" box="[214,345,230,258]" italics="true" pageId="16" pageNumber="17">P. iuiuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26B1CC21FFC1FFDCFEE7FF2EFE2DFECC" box="[353,417,230,258]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
in the triangular shape of male pleopod 1 exopod, with round apex and sinuous lateral margin, however in 
<taxonomicName id="C8F6D6CBFFC1FFDCFE1EFEFCFD9EFE9E" authorityName="Ferreira" authorityYear="2020" box="[408,530,308,336]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="iuiuensis" status="n. sp.">
<emphasis id="3D82715AFFC1FFDCFE1EFEFCFD9EFE9E" box="[408,530,308,336]" italics="true" pageId="16" pageNumber="17">P. iuiuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26B1CC21FFC1FFDCFD9EFEFCFDDFFE9E" box="[536,595,308,336]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
the exopod seems to be directed backward while in 
<taxonomicName id="C8F6D6CBFFC1FFDCFDC6FE94FC85FEB6" authorityName="Ferreira" authorityYear="2020" box="[576,777,347,376]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="carinhanhensis" status="n. sp.">
<emphasis id="3D82715AFFC1FFDCFDC6FE94FC85FEB6" box="[576,777,347,376]" italics="true" pageId="16" pageNumber="17">P. carinhanhensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26B1CC21FFC1FFDCFC88FE93FF5BFE50" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
it is directed outward.
</paragraph>
<paragraph id="0F49AD48FFC1FFDCFF5DFE07FAC3FB08" blockId="16.[180,807,463,1900]" lastBlockId="16.[841,1468,151,1222]" pageId="16" pageNumber="17">
<emphasis id="3D82715AFFC1FFDCFF5DFE07FDD5FE22" box="[219,601,463,492]" italics="true" pageId="16" pageNumber="17">Habitat and ecological remarks.</emphasis>
Specimens of 
<taxonomicName id="C8F6D6CBFFC1FFDCFC89FE18FEE1FDDD" authorityName="Ferreira" authorityYear="2020" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="carinhanhensis" status="n. sp.">
<emphasis id="3D82715AFFC1FFDCFC89FE18FEE1FDDD" italics="true" pageId="16" pageNumber="17">P. carinhanhensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26B1CC21FFC1FFDCFEF5FE3FFE3AFDDD" box="[371,438,503,531]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
were found in three caves of the Água Clara Cave System (ACCS) (
<figureCitation id="97CDB1CDFFC1FFDCFD3FFDD5FCACFDF4" box="[697,800,541,570]" captionStart="Figure 21" captionStartId="24.[181,250,1235,1258]" captionTargetBox="[181,1466,174,1180]" captionTargetId="figure-27@24.[181,1466,166,1181]" captionTargetPageId="24" captionText="Figure 21. Map of South America highlighting Brazil and the states of Bahia (BA), Minas Gerais (MG), and Santa Catarina (SC), where the species of Pectenoniscus are recorded. Brown and green colors indicate the elevation of the terrain, with brown for higher levels, green for lower levels; gray indicates karstic areas." figureDoi="http://doi.org/10.5281/zenodo.14668070" httpUri="https://zenodo.org/record/14668070/files/figure.png" pageId="16" pageNumber="17">Figs. 21</figureCitation>
, 
<figureCitation id="97CDB1CDFFC1FFDCFF33FD8CFF65FDAF" box="[181,233,580,609]" captionStart="Figure 22" captionStartId="26.[184,255,1379,1402]" captionTargetBox="[181,1469,151,1342]" captionTargetId="figure-26@26.[181,1469,151,1342]" captionTargetPageId="26" captionText="Figure 22. Satellite images of localities: A, Abrigo da Ema (Montalvânia municipality); B, Gruta do Tabuleirinho (Juvenilia municipality); C, Baixa da Fortuna cave (Iuiu municipality); D, Gruna da Água Clara Cave, Gruna dos Índios cave and Gruna dos Peixes II, from top to bottom (Carinhanha municipality); E, Gruta do Padre cave (Santana municipality); F, Gruta dos Brejões (Morro do Chapéu municipality)." figureDoi="http://doi.org/10.5281/zenodo.14668072" httpUri="https://zenodo.org/record/14668072/files/figure.png" pageId="16" pageNumber="17">22D</figureCitation>
). This system is formed by four limestone caves interconnected by an intermittent drainage (Gruna da Água Clara, Gruna dos Índios, Lapa dos Peixes I, and Lapa dos Peixes II – 
<figureCitation id="97CDB1CDFFC1FFDCFD93FD71FDF3FD19" box="[533,639,697,727]" captionStart="Figure 13" captionStartId="15.[181,249,1528,1551]" captionTargetBox="[181,1466,151,1498]" captionTargetId="figure-27@15.[181,1466,151,1498]" captionTargetPageId="15" captionText="Figure 13. Pectenoniscus carinhanhensisn. sp.A, Águas Claras karst system (extensionof caves in red, circles mark species occurrence; B, karst area of Serra do Ramalho; C, main conduit of Águas Claras cave; D, accumulated organic material in the conduit of Águas Claras cave; E, main conduit of Peixes II cave; F, specimen of Pectenoniscus carinhanhensis n. sp., approximately 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14668054" httpUri="https://zenodo.org/record/14668054/files/figure.png" pageId="16" pageNumber="17">Fig. 13A</figureCitation>
). Two of the caves that compose the system are the tenth and the fourteenth biggest caves in 
<collectingCountry id="77E1EDD8FFC1FFDCFE72FCC0FDB1FCEB" box="[500,573,776,805]" name="Brazil" pageId="16" pageNumber="17">Brazil</collectingCountry>
, respectively (Água Clara Cave – 13,880 meters and Lapa dos Peixes cave – 9,320 meters long), while the other two are smaller (Gruna dos Índios cave – 570 meters and Lapa dos Peixes II – 2,100 meters). Together these four caves account for 
<quantity id="C80E00ADFFC1FFDCFECCFC03FE4CFC26" box="[330,448,971,1000]" metricMagnitude="4" metricUnit="m" metricValue="2.585" pageId="16" pageNumber="17" unit="km" value="25.85">25.85 km</quantity>
of subterranean passages. In general, the caves present wide vadose conduits (
<figureCitation id="97CDB1CDFFC1FFDCFD7EFC3AFE89FBF8" captionStart="Figure 13" captionStartId="15.[181,249,1528,1551]" captionTargetBox="[181,1466,151,1498]" captionTargetId="figure-27@15.[181,1466,151,1498]" captionTargetPageId="15" captionText="Figure 13. Pectenoniscus carinhanhensisn. sp.A, Águas Claras karst system (extensionof caves in red, circles mark species occurrence; B, karst area of Serra do Ramalho; C, main conduit of Águas Claras cave; D, accumulated organic material in the conduit of Águas Claras cave; E, main conduit of Peixes II cave; F, specimen of Pectenoniscus carinhanhensis n. sp., approximately 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14668054" httpUri="https://zenodo.org/record/14668054/files/figure.png" pageId="16" pageNumber="17">Fig. 13C, E</figureCitation>
) through which a huge amount of water f lows during the rainy periods, noticeable by the massive plant debris (such as tree trunks) observed at several points within the caves (
<figureCitation id="97CDB1CDFFC1FFDCFE75FB47FDD3FB62" box="[499,607,1167,1196]" captionStart="Figure 13" captionStartId="15.[181,249,1528,1551]" captionTargetBox="[181,1466,151,1498]" captionTargetId="figure-27@15.[181,1466,151,1498]" captionTargetPageId="15" captionText="Figure 13. Pectenoniscus carinhanhensisn. sp.A, Águas Claras karst system (extensionof caves in red, circles mark species occurrence; B, karst area of Serra do Ramalho; C, main conduit of Águas Claras cave; D, accumulated organic material in the conduit of Águas Claras cave; E, main conduit of Peixes II cave; F, specimen of Pectenoniscus carinhanhensis n. sp., approximately 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14668054" httpUri="https://zenodo.org/record/14668054/files/figure.png" pageId="16" pageNumber="17">Fig. 13D</figureCitation>
). Although the water f lows along the caves only in the rainy periods, flooded chambers occur in some areas even in the dry periods, as in the Lapa dos Peixes II cave (
<figureCitation id="97CDB1CDFFC1FFDCFD37FACCFC9EFAEC" box="[689,786,1284,1314]" captionStart="Figure 13" captionStartId="15.[181,249,1528,1551]" captionTargetBox="[181,1466,151,1498]" captionTargetId="figure-27@15.[181,1466,151,1498]" captionTargetPageId="15" captionText="Figure 13. Pectenoniscus carinhanhensisn. sp.A, Águas Claras karst system (extensionof caves in red, circles mark species occurrence; B, karst area of Serra do Ramalho; C, main conduit of Águas Claras cave; D, accumulated organic material in the conduit of Águas Claras cave; E, main conduit of Peixes II cave; F, specimen of Pectenoniscus carinhanhensis n. sp., approximately 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14668054" httpUri="https://zenodo.org/record/14668054/files/figure.png" pageId="16" pageNumber="17">Fig. 13E</figureCitation>
). Individuals of 
<taxonomicName id="C8F6D6CBFFC1FFDCFEE2FAE4FDA1FA86" authorityName="Ferreira" authorityYear="2020" box="[356,557,1323,1352]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="carinhanhensis" status="n. sp.">
<emphasis id="3D82715AFFC1FFDCFEE2FAE4FDA1FA86" box="[356,557,1323,1352]" italics="true" pageId="16" pageNumber="17">P. carinhanhensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26B1CC21FFC1FFDCFDB5FAE4FDFDFA86" box="[563,625,1324,1352]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
were observed in several areas along the caves, always associated with moist substrates (
<figureCitation id="97CDB1CDFFC1FFDCFE01FAB2FE6AFA59" box="[391,486,1402,1431]" captionStart="Figure 13" captionStartId="15.[181,249,1528,1551]" captionTargetBox="[181,1466,151,1498]" captionTargetId="figure-27@15.[181,1466,151,1498]" captionTargetPageId="15" captionText="Figure 13. Pectenoniscus carinhanhensisn. sp.A, Águas Claras karst system (extensionof caves in red, circles mark species occurrence; B, karst area of Serra do Ramalho; C, main conduit of Águas Claras cave; D, accumulated organic material in the conduit of Águas Claras cave; E, main conduit of Peixes II cave; F, specimen of Pectenoniscus carinhanhensis n. sp., approximately 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14668054" httpUri="https://zenodo.org/record/14668054/files/figure.png" pageId="16" pageNumber="17">Fig. 13F</figureCitation>
). Most specimens (around 90% of the observed specimens in the ACCS) were found in Água Clara cave (
<figureCitation id="97CDB1CDFFC1FFDCFE6AFA00FDC2FA2B" box="[492,590,1480,1509]" captionStart="Figure 13" captionStartId="15.[181,249,1528,1551]" captionTargetBox="[181,1466,151,1498]" captionTargetId="figure-27@15.[181,1466,151,1498]" captionTargetPageId="15" captionText="Figure 13. Pectenoniscus carinhanhensisn. sp.A, Águas Claras karst system (extensionof caves in red, circles mark species occurrence; B, karst area of Serra do Ramalho; C, main conduit of Águas Claras cave; D, accumulated organic material in the conduit of Águas Claras cave; E, main conduit of Peixes II cave; F, specimen of Pectenoniscus carinhanhensis n. sp., approximately 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14668054" httpUri="https://zenodo.org/record/14668054/files/figure.png" pageId="16" pageNumber="17">Fig. 13C</figureCitation>
), which is the first (upstream) and biggest cave of the ACCS. The large number of specimens observed in this cave (dozens of individuals) can be related to its high humidity throughout the whole year, due to its dimensions. Few specimens (less than 10) were observed in Lapa do Índio cave, where they were restricted to a few moistened areas (a considerable extension of the cave was dry when the sampling was carried out). Finally, specimens were also collected in Lapa dos Peixes II cave, the last (downstream) cave of the ACCS. Although this species was not found in Lapa dos Peixes cave I, the possibility of its occurrence in this cave is not excluded, since only part of this cave was inventoried. In this sense it is plausible to assume that this species occurs in the whole ACCS. Specimens of 
<taxonomicName id="C8F6D6CBFFC1FFDCFCCCFEA8FB9CFEB2" authorityName="Ferreira" authorityYear="2020" box="[842,1040,351,380]" class="Malacostraca" family="Styloniscidae" genus="Pectenoniscus" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="carinhanhensis" status="n. sp.">
<emphasis id="3D82715AFFC1FFDCFCCCFEA8FB9CFEB2" box="[842,1040,351,380]" italics="true" pageId="16" pageNumber="17">P. carinhanhensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26B1CC21FFC1FFDCFB90FEA8FBDFFEB2" box="[1046,1107,352,380]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
were observed in very distinct conditions regarding the presence of visible organic matter. In some areas they were encountered close to vegetable debris, while in the deeper portions of the caves (especially in Água Clara cave) organisms were found in areas completely devoid of visible organic resources. The ACCS also shelters six other troglobitic species: the isopod 
<taxonomicName id="C8F6D6CBFFC1FFDCFBA1FDB1FBE8FD73" authority="Campos-Filho, Araujo and Taiti, 2014" authorityName="Campos-Filho, Araujo and Taiti" authorityYear="2014" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aganju">
<emphasis id="3D82715AFFC1FFDCFBA1FDB1FA8FFD5B" box="[1063,1283,633,661]" italics="true" pageId="16" pageNumber="17">Xangoniscus aganju</emphasis>
<bibRefCitation id="6B67D0B9FFC1FFDCFA8FFDB0FBE8FD73" author="Campos-Filho, I. S. &amp; Araujo, P. B. &amp; Bichuette, M. E. &amp; Trajano, E. &amp; Taiti, S." pageId="16" pageNumber="17" pagination="360 - 425" refId="ref12859" refString="Campos-Filho, I. S.; Araujo, P. B.; Bichuette, M. E.; Trajano, E. and Taiti, S. 2014. Terrestrial isopods (Crustacea: Isopoda: Oniscidea) from Brazilian caves. Zoological Journal of the Linnean Society, 172: 360 - 425." type="journal article" year="2014">Campos-Filho, Araujo and Taiti, 2014</bibRefCitation>
</taxonomicName>
, the harvestmen 
<taxonomicName id="C8F6D6CBFFC1FFDCFABDFD68FB3DFD2B" authority="Perez and Kury, 2002" authorityName="Perez and Kury" authorityYear="2002" class="Arachnida" family="Gonyleptidae" genus="Giupponia" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="chagasi">
<emphasis id="3D82715AFFC1FFDCFABDFD68FC2CFD2B" italics="true" pageId="16" pageNumber="17">Giupponia chagasi</emphasis>
Pérez and 
<bibRefCitation id="6B67D0B9FFC1FFDCFBA0FD01FB3DFD2B" author="Kury, A. B. &amp; Gonzalez, A. P." box="[1062,1201,712,741]" pageId="16" pageNumber="17" pagination="43 - 50" refId="ref13557" refString="Kury, A. B. and Gonzalez, A. P. 2002. A new remarkable troglomorphic gonyleptid from Brazil (Arachnida, Opiliones, Laniatores). Revista Iberica de aracnologia, 5: 43 - 50." type="journal article" year="2002">Kury, 2002</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="6B67D0B9FFC1FFDCFB47FD00FC02FCC3" author="Kury, A. B. &amp; Gonzalez, A. P." pageId="16" pageNumber="17" pagination="43 - 50" refId="ref13557" refString="Kury, A. B. and Gonzalez, A. P. 2002. A new remarkable troglomorphic gonyleptid from Brazil (Arachnida, Opiliones, Laniatores). Revista Iberica de aracnologia, 5: 43 - 50." type="journal article" year="2002">Kury and González, 2002</bibRefCitation>
), the whip spider 
<taxonomicName id="C8F6D6CBFFC1FFDCFBEEFD38FBCEFCFB" authority="Baptista and Giupponi, 2002" authorityName="Baptista and Giupponi" authorityYear="2002" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="troglobius">
<emphasis id="3D82715AFFC1FFDCFBEEFD38FADCFCC3" box="[1128,1360,752,781]" italics="true" pageId="16" pageNumber="17">Charinus troglobius</emphasis>
<bibRefCitation id="6B67D0B9FFC1FFDCFAD0FD38FBCEFCFB" author="Baptista, R. L. C. &amp; Giupponi, A. D. L." pageId="16" pageNumber="17" pagination="105 - 110" refId="ref12638" refString="Baptista, R. L. C. and Giupponi, A. D. L. 2002. A new troglomorphic Charinus from brazil (Arachnida: Amblypygi: Charinidae). Revista Iberica de Aracnologia, 6: 105 - 110." type="journal article" year="2002">Baptista and Giupponi, 2002</bibRefCitation>
</taxonomicName>
, the earwig 
<taxonomicName id="C8F6D6CBFFC1FFDCFB56FCD0FB36FC93" authority="Kamimura and Ferreira, 2018" authorityName="Kamimura and Ferreira" authorityYear="2018" class="Insecta" family="Diplatyidae" genus="Mesodiplatys" kingdom="Animalia" order="Dermaptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="falcifer">
<emphasis id="3D82715AFFC1FFDCFB56FCD0FA37FCFB" box="[1232,1467,792,821]" italics="true" pageId="16" pageNumber="17">Mesodiplatys falcifer</emphasis>
<bibRefCitation id="6B67D0B9FFC1FFDCFCCDFC88FB36FC93" author="Kamimura, Y. &amp; Ferreira, R. L." box="[843,1210,832,861]" pageId="16" pageNumber="17" pagination="87 - 100" refId="ref13451" refString="Kamimura, Y. and Ferreira, R. L. 2018. Description of a second South American species in the Malagasy earwig genus Mesodiplatys from a cave habitat, with notes on the definition of Haplodiplatyidae (Insecta, Dermaptera). ZooKeys, 790: 87 - 100." type="journal article" year="2018">Kamimura and Ferreira, 2018</bibRefCitation>
</taxonomicName>
, the snail 
<taxonomicName id="C8F6D6CBFFC1FFDCFAB0FC88FBE8FC4B" authority="Simone, 2012" authorityName="Simone" authorityYear="2012" class="Gastropoda" family="Cochliopidae" genus="Spiripockia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="punctata">
<emphasis id="3D82715AFFC1FFDCFAB0FC88FC38FC4B" italics="true" pageId="16" pageNumber="17">Spiripockia punctata</emphasis>
<bibRefCitation id="6B67D0B9FFC1FFDCFC3DFCA0FBE8FC4B" author="Simone, L. R. L." box="[955,1124,872,901]" pageId="16" pageNumber="17" pagination="515 - 524" refId="ref14041" refString="Simone, L. R. L. 2012. A new genus and species of cavernicolous Pomatiopsidae (Mollusca, Caenogastropoda) in Bahia, Brazil. Papeis Avulsos de Zoologia, 52: 515 - 524." type="journal article" year="2012">Simone, 2012</bibRefCitation>
</taxonomicName>
and the fish 
<taxonomicName id="C8F6D6CBFFC1FFDCFA83FCA0FAADFC60" authority="Bichuette and Rizatto, 2012" authorityName="Bichuette and Rizatto" authorityYear="2012" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="rubiolli">
<emphasis id="3D82715AFFC1FFDCFA83FCA0FC29FC63" italics="true" pageId="16" pageNumber="17">Trichomycterus rubiolli</emphasis>
Bichuette and Rizatto, 2012
</taxonomicName>
. The native habitat surrounding the caves consists of the Caatinga formation, which is severely altered, especially due to the removal of the original vegetation for charcoal production, as well as for the establishment of pastures and monocultures (
<figureCitation id="97CDB1CDFFC1FFDCFBBAFB91FB12FBB8" box="[1084,1182,1113,1142]" captionStart="Figure 13" captionStartId="15.[181,249,1528,1551]" captionTargetBox="[181,1466,151,1498]" captionTargetId="figure-27@15.[181,1466,151,1498]" captionTargetPageId="15" captionText="Figure 13. Pectenoniscus carinhanhensisn. sp.A, Águas Claras karst system (extensionof caves in red, circles mark species occurrence; B, karst area of Serra do Ramalho; C, main conduit of Águas Claras cave; D, accumulated organic material in the conduit of Águas Claras cave; E, main conduit of Peixes II cave; F, specimen of Pectenoniscus carinhanhensis n. sp., approximately 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14668054" httpUri="https://zenodo.org/record/14668054/files/figure.png" pageId="16" pageNumber="17">Fig. 13B</figureCitation>
). The caves seem to be only visited by speleologists, so that the impacts of trampling over the cave fauna is negligible.
</paragraph>
</subSubSection>
</treatment>
</document>