181 lines
23 KiB
XML
181 lines
23 KiB
XML
<document id="DA693B130DCABC24A681977868614AAB" ID-DOI="10.5281/zenodo.193899" ID-GBIF-Dataset="63d06d60-7dd0-4c4f-93cd-04a9cb970b4b" ID-ISSN="1175-5326" ID-Zenodo-Dep="193899" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460176993965" checkinUser="plazi" docAuthor="Doughty, Paul, Pepper, Mitzy & Keogh, Scott" docDate="2010" docId="87357672412A254CED8B2A82FC586CD0" docLanguage="en" docName="zt02393p045.pdf" docOrigin="Zootaxa 2393" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Diplodactylus savagei Kluge 1963" docType="treatment" docVersion="9" lastPageNumber="38" masterDocId="7B0C0E0A412F2549ED1C2A15FFE76B5A" masterDocTitle="Morphological and molecular assessment of the Diplodactylus savagei species complex in the Pilbara region, Western Australia, with a description of a new species" masterLastPageNumber="45" masterPageNumber="33" pageNumber="38" updateTime="1698236118467" updateUser="plazi">
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<mods:title id="40A917DC7E2B81174FF4ADFA3603ECD8">Morphological and molecular assessment of the Diplodactylus savagei species complex in the Pilbara region, Western Australia, with a description of a new species</mods:title>
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<mods:namePart id="8BFF59A4A53991CB2998B7C48C9228DA">Doughty, Paul</mods:namePart>
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<mods:namePart id="639E7FCDC6833580BB3E775664AB3E39">Pepper, Mitzy</mods:namePart>
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<mods:namePart id="154E20A6A3B408058A2129A537764012">Keogh, Scott</mods:namePart>
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<mods:date id="923A1198807D926D856C7D7AFD6FC8F6">2010</mods:date>
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<treatment id="87357672412A254CED8B2A82FC586CD0" ID-DOI="http://doi.org/10.5281/zenodo.5667374" ID-GBIF-Taxon="119383566" ID-Zenodo-Dep="5667374" LSID="urn:lsid:plazi:treatment:87357672412A254CED8B2A82FC586CD0" httpUri="http://treatment.plazi.org/id/87357672412A254CED8B2A82FC586CD0" lastPageNumber="38" pageId="5" pageNumber="38">
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<subSubSection id="478694EF412A254CED8B2A82FEFF6BA0" pageId="5" pageNumber="38" type="nomenclature">
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<paragraph id="0F23C764412A254CED8B2A82FDDE6BEB" blockId="5.[151,572,151,250]" box="[151,569,151,177]" pageId="5" pageNumber="38">
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<heading id="546B7008412A254CED8B2A82FDDE6BEB" bold="true" box="[151,569,151,177]" fontSize="11" level="1" pageId="5" pageNumber="38" reason="1">
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<taxonomicName id="C89CBCE7412A254CED8B2A82FDDE6BEB" authority="Kluge, 1963" authorityName="Kluge" authorityYear="1963" box="[151,569,151,177]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="savagei">
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<emphasis id="3DE81B76412A254CED8B2A82FDDE6BEB" bold="true" box="[151,569,151,177]" pageId="5" pageNumber="38">
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<emphasis id="3DE81B76412A254CED8B2A82FE796BEB" bold="true" box="[151,414,151,177]" italics="true" pageId="5" pageNumber="38">Diplodactylus savagei</emphasis>
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<bibRefCitation id="6B0DBA95412A254CECB92A82FDDE6BEB" author="Kluge" box="[421,569,151,177]" pageId="5" pageNumber="44" refString="Kluge, A. G. (1963) Three new species of the gekkonid lizard genus Diplodactylus Gray from Australia. Records of the South Australian Museum, 14, 545 - 553." type="journal article" year="1963">Kluge, 1963</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="0F23C764412A254CED8B2AAFFEFF6BA0" blockId="5.[151,572,151,250]" pageId="5" pageNumber="38">
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Southern Pilbara Beak-faced Gecko (
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<figureCitation id="97A7DBE1412A254CED832AF5FF146BA0" box="[159,243,224,250]" captionStart="FIGURE 2" captionStartId="4.[151,255,1886,1910]" captionTargetBox="[152,1433,197,1861]" captionTargetId="figure@4.[152,1435,197,1863]" captionTargetPageId="4" captionText="FIGURE 2. Photographs of live Diplodactylus galaxias sp. nov. (above) from the Burrup Peninsula and D. savagei (below) from near the type locality. Photographs – B. Maryan." httpUri="https://zenodo.org/record/193901/files/figure.png" pageId="5" pageNumber="38">Figs. 2</figureCitation>
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,
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<figureCitation id="97A7DBE1412A254CEDE32AF5FEE96BA0" box="[255,270,224,250]" captionStart="FIGURE 3" captionStartId="7.[151,255,1528,1552]" captionTargetBox="[152,1433,194,1505]" captionTargetId="figure@7.[152,1435,194,1507]" captionTargetPageId="7" captionText="FIGURE 3. Variation of dorsal pattern within Diplodactylus galaxias sp. nov. (top row) and D. savagei (bottom row)." httpUri="https://zenodo.org/record/193902/files/figure.png" pageId="5" pageNumber="38">3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="478694EF412A254CED8B2B3FFE706AD5" pageId="5" pageNumber="38" type="materials_examined">
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<paragraph id="0F23C764412A254CED8B2B3FFE986A30" blockId="5.[151,1437,298,1930]" pageId="5" pageNumber="38">
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<emphasis id="3DE81B76412A254CED8B2B3FFEF56A1E" bold="true" box="[151,274,298,324]" pageId="5" pageNumber="38">
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<typeStatus id="D02779C6412A254CED8B2B3FFEEA6A1E" box="[151,269,298,324]" pageId="5" pageNumber="38" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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R14369, an adult female collected at Marble Bar, Western
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<collectingCountry id="778B87F4412A254CEEC12B3FFBA96A1E" box="[989,1102,298,324]" name="Australia" pageId="5" pageNumber="38">Australia</collectingCountry>
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,
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<collectingCountry id="778B87F4412A254CE9472B3FFB2F6A1E" box="[1115,1224,298,324]" name="Australia" pageId="5" pageNumber="38">Australia</collectingCountry>
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on
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<date id="7B22E1A4412A254CE9EA2B3FFF366A30" pageId="5" pageNumber="38" value="1960-09-22">22 September 1960</date>
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by G.M. Storr.
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</paragraph>
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<paragraph id="0F23C764412A254CEDD82B60FE706AD5" blockId="5.[151,1437,298,1930]" box="[196,407,373,399]" pageId="5" pageNumber="38">
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<emphasis id="3DE81B76412A254CEDD82B60FEAF6AD5" bold="true" box="[196,328,373,399]" pageId="5" pageNumber="38">
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<typeStatus id="D02779C6412A254CEDD82B60FEA46AD5" box="[196,323,373,399]" pageId="5" pageNumber="38" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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None.
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</paragraph>
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</subSubSection>
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<subSubSection id="478694EF412A254CEDD82B8FFB3B697E" pageId="5" pageNumber="38" type="diagnosis">
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<paragraph id="0F23C764412A254CEDD82B8FFB3B697E" blockId="5.[151,1437,298,1930]" pageId="5" pageNumber="38">
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<emphasis id="3DE81B76412A254CEDD82B8FFEAE6AEE" bold="true" box="[196,329,410,436]" pageId="5" pageNumber="38">Diagnosis.</emphasis>
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Differentiated from
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<taxonomicName id="C89CBCE7412A254CEF562B8FFD3E6AE9" box="[586,729,410,435]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="galaxias" status="sp. nov.">
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<emphasis id="3DE81B76412A254CEF562B8FFD3E6AE9" box="[586,729,410,435]" italics="true" pageId="5" pageNumber="38">D. galaxias</emphasis>
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</taxonomicName>
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<emphasis id="3DE81B76412A254CEFFD2B8FFCA36AEE" bold="true" box="[737,836,410,436]" pageId="5" pageNumber="38">
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<taxonomicNameLabel id="26DBA60D412A254CEFFD2B8FFCA36AEE" box="[737,836,410,436]" pageId="5" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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in having slightly larger body size, more gabled dorsal scales, slightly shorter and wider tail, heavy yellowish spots that form transverse rows (not scattered fine spots) on darker reddish-brown background colour, pale dorsal border to dark loreal stripe rarely present and dorsal-ventral colouration transition abrupt or marked by spots or stippling (not gradual).
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</paragraph>
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</subSubSection>
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<subSubSection id="478694EF412A254CEDD8283AFF026D95" pageId="5" pageNumber="38" type="description">
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<paragraph id="0F23C764412A254CEDD8283AFCA36815" blockId="5.[151,1437,298,1930]" pageId="5" pageNumber="38">
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<emphasis id="3DE81B76412A254CEDD8283AFEB86913" bold="true" box="[196,351,559,585]" pageId="5" pageNumber="38">Description.</emphasis>
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Small-bodied
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<taxonomicName id="C89CBCE7412A254CEF0C2825FD5F6913" box="[528,696,560,585]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="38" phylum="Chordata" rank="genus">
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<emphasis id="3DE81B76412A254CEF0C2825FD5F6913" box="[528,696,560,585]" italics="true" pageId="5" pageNumber="38">Diplodactylus</emphasis>
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</taxonomicName>
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with cylindrical body shape and a small, depressed head; tail cylindrical, slightly increasing in width towards tip until about four-fifths of length, then tapering at roughly a 60° angle to tip. Eyes small with no spiny ridges on upper eyelid; snout relatively long and pointed with blunt tip. Scales contacting nostril: supranasal and 4–6 postnasals; rostral roughly hexagonal without crease, bordered by enlarged supralabials, supranasals and internarials; other than first, enlarged supralabials absent with scales along upper jaw similar in size to adjoining rows of scales; mental wider than long with adjacent enlarged gular scales in 2–3 rows; no enlarged infralabials scales with typical-sized scales along lower jaw; ear aperature small and located near retroarticular process.
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</paragraph>
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<paragraph id="0F23C764412A254CEDD8294FFD2B68E5" blockId="5.[151,1437,298,1930]" pageId="5" pageNumber="38">Limbs slender and of moderate length; undersurfaces of digits terminating with claw between moderately enlarged apical plates and 6–10 rows of enlarged paired lamellae along length of digit until decreasing to the size of tubercles on palmar and plantar surfaces.</paragraph>
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<paragraph id="0F23C764412A254CEDD829DFFCBB6F75" blockId="5.[151,1437,298,1930]" pageId="5" pageNumber="38">Scales on body small; dorsal and ventral scales approximately the same size. Dorsal scales slightly gabled, with the apex towards the posterior edge of scale. Caudal scales on original tail enlarged and flattened, tending to form transverse rows; scattered enlarged scales on tail tip.</paragraph>
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<paragraph id="0F23C764412A254CEDD82E2FFCAE6E6E" blockId="5.[151,1437,298,1930]" pageId="5" pageNumber="38">
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<emphasis id="3DE81B76412A254CEDD82E2FFE816F0E" bold="true" box="[196,358,1082,1108]" pageId="5" pageNumber="38">Colouration.</emphasis>
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In life, dorsum a rich reddish brown with small (1–5 scales) yellowish spots that usually align and/or abut to clearly form 8–12 transverse rows between nape and base of tail; spots rarely extending onto limbs but, if present, also tending to form rows; dark loreal-temporal streak through eye, weakly connecting nape; dorsal border loreal streak rarely with pale edge; top of head slightly pale compared to rest of dorsal colouration. Transition from dorsal to ventral colouration abrupt: demarcated by a sharp transition or stippling. Ventral surfaces pale cream. In preservative, ground colour faded, pale cap on head more prominent and spots creamy white (almost no trace of yellow colour).
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</paragraph>
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<paragraph id="0F23C764412A254CEDD82F2AFDFE6E00" blockId="5.[151,1437,298,1930]" box="[196,537,1343,1370]" pageId="5" pageNumber="38">
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<emphasis id="3DE81B76412A254CEDD82F2AFE646E03" bold="true" box="[196,387,1343,1369]" pageId="5" pageNumber="38">Measurements.</emphasis>
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See
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<tableCitation id="421EF2DF412A254CECA72F55FDF46E00" box="[443,531,1344,1370]" captionStart="TABLE 1" captionStartId="6.[151,239,151,175]" captionTargetBox="[151,1437,515,1997]" captionTargetPageId="6" captionText="TABLE 1. Summaries of characters and ratios measured for Diplodactylus galaxias sp. nov. and D. savagei. Mean ± S. D. (range). Sample sizes are listed in column headings, unless noted for individual characters below and are based on all specimens examined in the type series and Appendix 1. Abbreviations: snout-vent length—SVL; trunk length (from axilla to groin) — TrunkL; head length (from tip of snout to retroarticular process) — HeadL; head width (at widest point) — HeadW; head depth (at deepest point): HeadD; ratio of length / width of mental scale—MenL / W; internarial distance (measured at medial edge of nares) — INar; interorbital width (at anterior centre of eyes) — IO; number of interial scales— IntNar; number of postnasal scales—PostNas; foreleg length (from elbow to base of palm) — ArmL; tibia length (from upper surface of knee to heel) – LegL; number of rows of lamellae on fourth finger— 4 FLam; number of rows of lamellae on fourth toe—TLam; tail length (original tails only) — TL; tail width (at widest point on original tails) — TailW; tail length as a percentage of SVL – Tail % SVL; number of subcaudal scales (from fracture plane to tip) — NoSC; number of enlarged cloacal spurs—CSpurs." httpUri="http://table.plazi.org/id/5BE397EC4129254FED8B2A82FE136AB5" pageId="5" pageNumber="38" tableUuid="5BE397EC4129254FED8B2A82FE136AB5">Table 1</tableCitation>
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.
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</paragraph>
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<paragraph id="0F23C764412A254CEDD82F70FADE6D05" blockId="5.[151,1437,298,1930]" pageId="5" pageNumber="38">
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<emphasis id="3DE81B76412A254CEDD82F70FED86E25" bold="true" box="[196,319,1381,1407]" pageId="5" pageNumber="38">Variation.</emphasis>
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Males possess an average of 7 enlarged pointed cloacal spurs on either side of cloaca. Females attain larger body sizes and trunk lengths than males (
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<tableCitation id="421EF2DF412A254CEE1B2F9FFCB96EFE" box="[775,862,1418,1444]" captionStart="TABLE 1" captionStartId="6.[151,239,151,175]" captionTargetBox="[151,1437,515,1997]" captionTargetPageId="6" captionText="TABLE 1. Summaries of characters and ratios measured for Diplodactylus galaxias sp. nov. and D. savagei. Mean ± S. D. (range). Sample sizes are listed in column headings, unless noted for individual characters below and are based on all specimens examined in the type series and Appendix 1. Abbreviations: snout-vent length—SVL; trunk length (from axilla to groin) — TrunkL; head length (from tip of snout to retroarticular process) — HeadL; head width (at widest point) — HeadW; head depth (at deepest point): HeadD; ratio of length / width of mental scale—MenL / W; internarial distance (measured at medial edge of nares) — INar; interorbital width (at anterior centre of eyes) — IO; number of interial scales— IntNar; number of postnasal scales—PostNas; foreleg length (from elbow to base of palm) — ArmL; tibia length (from upper surface of knee to heel) – LegL; number of rows of lamellae on fourth finger— 4 FLam; number of rows of lamellae on fourth toe—TLam; tail length (original tails only) — TL; tail width (at widest point on original tails) — TailW; tail length as a percentage of SVL – Tail % SVL; number of subcaudal scales (from fracture plane to tip) — NoSC; number of enlarged cloacal spurs—CSpurs." httpUri="http://table.plazi.org/id/5BE397EC4129254FED8B2A82FE136AB5" pageId="5" pageNumber="38" tableUuid="5BE397EC4129254FED8B2A82FE136AB5">Table 1</tableCitation>
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). Ground colour shows some limited variation in hue, but the pattern of spotting is more variable, and even more variable than in
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<taxonomicName id="C89CBCE7412A254CE9552FA5FB376E93" box="[1097,1232,1456,1481]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="galaxias" status="sp. nov.">
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<emphasis id="3DE81B76412A254CE9552FA5FB376E93" box="[1097,1232,1456,1481]" italics="true" pageId="5" pageNumber="38">D. galaxias</emphasis>
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</taxonomicName>
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<emphasis id="3DE81B76412A254CE9C42FBAFAD16E93" bold="true" box="[1240,1334,1455,1481]" pageId="5" pageNumber="38">
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<taxonomicNameLabel id="26DBA60D412A254CE9C42FBAFAD16E93" box="[1240,1334,1455,1481]" pageId="5" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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In some individuals the spots align to form solid transverse bars across the dorsum, although the individual spots comprising the bars are still evident (e.g., R162852, R
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<date id="7B22E1A4412A254CEE3A2FEFFC676D4E" box="[806,896,1530,1556]" pageId="5" pageNumber="38" value="11-02-17">170211</date>
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, R
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<date id="7B22E1A4412A254CEEBE2FEFFC196D4E" box="[930,1022,1530,1556]" pageId="5" pageNumber="38" value="96-01-17">170196</date>
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, R
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<date id="7B22E1A4412A254CE9032FEFFB9C6D4E" box="[1055,1147,1530,1556]" pageId="5" pageNumber="38" value="75-02-17">170275</date>
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;
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<figureCitation id="97A7DBE1412A254CE9942FEFFB366D4E" box="[1160,1233,1530,1556]" captionStart="FIGURE 3" captionStartId="7.[151,255,1528,1552]" captionTargetBox="[152,1433,194,1505]" captionTargetId="figure@7.[152,1435,194,1507]" captionTargetPageId="7" captionText="FIGURE 3. Variation of dorsal pattern within Diplodactylus galaxias sp. nov. (top row) and D. savagei (bottom row)." httpUri="https://zenodo.org/record/193902/files/figure.png" pageId="5" pageNumber="38">Fig. 3</figureCitation>
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). The size of the spots also varied, from very small (<5; e.g., R158144) to large (~10 scales; e.g., R
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<date id="7B22E1A4412A254CE9702C35FB2F6D60" box="[1132,1224,1568,1594]" pageId="5" pageNumber="38" value="27-01-17">170127</date>
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). R
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<date id="7B22E1A4412A254CE9F32C35FAA16D60" box="[1263,1350,1568,1594]" pageId="5" pageNumber="38" value="12-01-16">160112</date>
|
||
had an unusual pattern of strong transverse rows that alternated with a weak transverse row of spots (
|
||
<figureCitation id="97A7DBE1412A254CE9FE2C50FACD6D05" box="[1250,1322,1605,1631]" captionStart="FIGURE 3" captionStartId="7.[151,255,1528,1552]" captionTargetBox="[152,1433,194,1505]" captionTargetId="figure@7.[152,1435,194,1507]" captionTargetPageId="7" captionText="FIGURE 3. Variation of dorsal pattern within Diplodactylus galaxias sp. nov. (top row) and D. savagei (bottom row)." httpUri="https://zenodo.org/record/193902/files/figure.png" pageId="5" pageNumber="38">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="0F23C764412A254CEDD82C7FFF026D95" blockId="5.[151,1437,298,1930]" pageId="5" pageNumber="38">
|
||
<emphasis id="3DE81B76412A254CEDD82C7FFEC96DDE" bold="true" box="[196,302,1642,1668]" pageId="5" pageNumber="38">Habitat.</emphasis>
|
||
Collection records of
|
||
<taxonomicName id="C89CBCE7412A254CEF5D2C7EFD246DDE" box="[577,707,1643,1668]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="savagei">
|
||
<emphasis id="3DE81B76412A254CEF5D2C7EFD246DDE" box="[577,707,1643,1668]" italics="true" pageId="5" pageNumber="38">D. savagei</emphasis>
|
||
</taxonomicName>
|
||
indicate an association with stony hills with spinifex, scree slopes,
|
||
<taxonomicName id="C89CBCE7412A254CEDF32C85FE966DF3" box="[239,369,1680,1705]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="38" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="3DE81B76412A254CEDF32C85FE966DF3" box="[239,369,1680,1705]" italics="true" pageId="5" pageNumber="38">Eucalyptus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C89CBCE7412A254CECB02C85FE1A6DF3" box="[428,509,1680,1705]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="38" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="3DE81B76412A254CECB02C85FE1A6DF3" box="[428,509,1680,1705]" italics="true" pageId="5" pageNumber="38">Acacia</emphasis>
|
||
</taxonomicName>
|
||
spp, but also occasionally recorded from termitaria, cracking clays and loamy plains.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="478694EF412A254CEDD82CCFFA9D6C3E" pageId="5" pageNumber="38" type="distribution">
|
||
<paragraph id="0F23C764412A254CEDD82CCFFA9D6C3E" blockId="5.[151,1437,298,1930]" pageId="5" pageNumber="38">
|
||
<emphasis id="3DE81B76412A254CEDD82CCFFE8F6DAE" bold="true" box="[196,360,1754,1780]" pageId="5" pageNumber="38">Distribution.</emphasis>
|
||
Pilbara and Ashburton regions (
|
||
<figureCitation id="97A7DBE1412A254CEFEF2CCFFCDC6DAE" box="[755,827,1754,1780]" captionStart="FIGURE 1" captionStartId="2.[151,259,1709,1733]" captionTargetBox="[152,1426,383,1708]" captionTargetId="figure@2.[151,1436,372,1708]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of Diplodactylus savagei populations in the Pilbara and Ashburton regions of Western Australia. Key: blue circles— D. savagei; red circles— D. galaxias sp. nov.; stars—genotyped individuals." httpUri="https://zenodo.org/record/193900/files/figure.png" pageId="5" pageNumber="38">Fig. 1</figureCitation>
|
||
): eastern Pilbara from Wodgina to the edge of the Great Sandy Desert, then extending south and west along the Hamersley Range, not occurring north of the Fortescue Marsh. Two widely-separated southeastern locality records are from the Barlee Ranges and Waldburg Station; however, the intervening area is poorly surveyed and the species may be common there.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="478694EF412A254CEDD82D7AFC586CD0" box="[196,959,1903,1930]" pageId="5" pageNumber="38" type="etymology">
|
||
<paragraph id="0F23C764412A254CEDD82D7AFC586CD0" blockId="5.[151,1437,298,1930]" box="[196,959,1903,1930]" pageId="5" pageNumber="38">
|
||
<emphasis id="3DE81B76412A254CEDD82D7AFEB66CD3" bold="true" box="[196,337,1903,1929]" pageId="5" pageNumber="38">Etymology.</emphasis>
|
||
Named after American herpetologist Jay M. Savage.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |