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<mods:title id="8150E481DA8932F9E51068B56A815564">Review of the leafhopper tribe Deltocephalini Dallas, 1870 (Hemiptera, Cicadellidae, Deltocephalinae) in Pakistan with description of a new species of Paramesodes</mods:title>
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<mods:namePart id="9619D86892EC635D8502BD1CD1B75BC7">Naveed, Hassan</mods:namePart>
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<mods:affiliation id="432B39A96232BCCCE802994E5F6FE1B0">School of Life Sciences, Jiangsu University, Zhenjiang 212013, China</mods:affiliation>
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<mods:affiliation id="C85911B8B7ADCC58B03A50407FFB5CE9">Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A &amp; F University, 666 Wusu Street, Linan, Hangzhou, Zhejiang 311300, China</mods:affiliation>
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<mods:namePart id="B3BCA36FD504560A15D31419A2AEC0F1">Zhang, Yalin</mods:namePart>
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<mods:affiliation id="6416C00933BBB923CD8434042570B283">Department of Entomology, The University of Agriculture, Peshawar 25100, Pakistan</mods:affiliation>
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<mods:namePart id="90A6B9CC5FD5CC2656230EDE08C43DA1">Chen, Keping</mods:namePart>
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<taxonomicName id="16B5EB00F8EC200ED2E1740AFF1EA651" ID-CoL="8RLVT" LSID="1A88C146-2B57-5AC2-9D08-26A8D3D575AC" authority="(Metcalf)" baseAuthorityName="Metcalf" baseAuthorityYear="1967" class="Insecta" family="Cicadellidae" genus="Maiestas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Maiestas pruthii" order="Hemiptera" pageId="0" pageNumber="207" phylum="Arthropoda" rank="species" species="pruthii">Maiestas pruthii (Metcalf)</taxonomicName>
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<figureCitation id="8C2D1E67742E9B08E311918E0F4B56DE" captionStart="Figures 1229" captionStartId="F3" captionText="Figures 12 - 29. 12 - 14 Deltocephalus vulgaris Dash &amp; Viraktamath: 12 habitus, dorsal view 13 aedeagus and connective, dorsal view 14 aedeagus and connective, lateral view 15 - 17 D. infirmus (Melichar): 15 habitus, dorsal view 16 aedeagus and connective, dorsal view 17 aedeagus and connective, lateral view 18 Maiestas albomaculata (Dash &amp; Viraktamath) habitus, dorsal view 19 M. maculata (Singh-Pruthi) habitus, dorsal view 20 M. indica (Singh-Pruthi) aedeagus and connective, lateral view (after Dash &amp; Viraktamath, 1998) 21, 22 M. pruthii (Metcalf): 21 aedeagus and connective, dorsal view (after Dash &amp; Viraktamath, 1998) 22 apex of aedeagus, lateral view 23 M. trispinosa (Dash &amp; Viraktamath) aedeagus and connective, dorsal view 24 M. subviridis (Metcalf) aedeagus, lateral view (after Khatri &amp; Webb, 2010) 25, 26 M. tareni (Dash &amp; Viraktamath): 25 aedeagus and connective, lateral view 26 subgenital plate, valve and styles, dorsal view 27, 28 M. sinuata Shah &amp; Duan: 27 aedeagus and connective, lateral view 28 subgenital plate, valve and styles, dorsal view 29 M. maculata (Singh-Pruthi) aedeagus, lateral view." figureDoi="10.3897/zookeys.1186.110266.figures12-29" httpUri="https://binary.pensoft.net/fig/952035" pageId="0" pageNumber="207">Figs 21</figureCitation>
<figureCitation id="2CC2C294D2246301443EB967F120365A" captionStart="Figures 1229" captionStartId="F3" captionText="Figures 12 - 29. 12 - 14 Deltocephalus vulgaris Dash &amp; Viraktamath: 12 habitus, dorsal view 13 aedeagus and connective, dorsal view 14 aedeagus and connective, lateral view 15 - 17 D. infirmus (Melichar): 15 habitus, dorsal view 16 aedeagus and connective, dorsal view 17 aedeagus and connective, lateral view 18 Maiestas albomaculata (Dash &amp; Viraktamath) habitus, dorsal view 19 M. maculata (Singh-Pruthi) habitus, dorsal view 20 M. indica (Singh-Pruthi) aedeagus and connective, lateral view (after Dash &amp; Viraktamath, 1998) 21, 22 M. pruthii (Metcalf): 21 aedeagus and connective, dorsal view (after Dash &amp; Viraktamath, 1998) 22 apex of aedeagus, lateral view 23 M. trispinosa (Dash &amp; Viraktamath) aedeagus and connective, dorsal view 24 M. subviridis (Metcalf) aedeagus, lateral view (after Khatri &amp; Webb, 2010) 25, 26 M. tareni (Dash &amp; Viraktamath): 25 aedeagus and connective, lateral view 26 subgenital plate, valve and styles, dorsal view 27, 28 M. sinuata Shah &amp; Duan: 27 aedeagus and connective, lateral view 28 subgenital plate, valve and styles, dorsal view 29 M. maculata (Singh-Pruthi) aedeagus, lateral view." figureDoi="10.3897/zookeys.1186.110266.figures12-29" httpUri="https://binary.pensoft.net/fig/952035" pageId="0" pageNumber="207">, 22</figureCitation>
</paragraph>
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<paragraph id="803186064C66EF40D1D0D8BA8743E13D" pageId="0" pageNumber="207">
<taxonomicName id="B50D2DC7F54D45207809A4B3409D5C1E" family="Cicadellidae" genus="Deltocephlaus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Deltocephlaus notatus" order="Hemiptera" pageId="0" pageNumber="207" rank="species" species="notatus">Deltocephlaus notatus</taxonomicName>
Singh-Pruthi, 1936: 128-129, fig. 139, pl. 9 fig. 10 (Pakistan) (primary homonym:
<taxonomicName id="78D5A6D8845556487CDB332C495F0058" authorityName="Melichar" authorityYear="1896" class="Insecta" family="Cicadellidae" genus="Deltocephalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltocephalus notatus" order="Hemiptera" pageId="0" pageNumber="207" phylum="Arthropoda" rank="species" species="notatus">Deltocephalus notatus</taxonomicName>
Melichar, 1896);
<taxonomicName id="463C9AC18F7AA026727A0A136086EE58" authorityName="Metcalf" authorityYear="1967" class="Insecta" family="Cicadellidae" genus="Deltocephalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltocephalus pruthii" order="Hemiptera" pageId="0" pageNumber="207" phylum="Arthropoda" rank="species" species="pruthii">Deltocephalus pruthii</taxonomicName>
Metcalf, 1967: 1173 (nom. nov. pro
<taxonomicName id="ECE1CF71F6E132A2C6F98B6EAABE3E8F" family="Cicadellidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="D. notatus" order="Hemiptera" pageId="0" pageNumber="207" rank="species" species="notatus">D. notatus</taxonomicName>
Singh-Pruthi, 1936);
<taxonomicName id="31E8BB6FF9A8B8FA8E2B9D27C6F82DD0" authorityName="Metcalf" authorityYear="1967" class="Insecta" family="Cicadellidae" genus="Deltocephalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deltocephalus (Recilia) pruthii" order="Hemiptera" pageId="0" pageNumber="207" phylum="Arthropoda" rank="species" species="pruthii" subGenus="Recilia">Deltocephalus (Recilia) pruthii</taxonomicName>
:
<bibRefCitation id="7021FB3F7A29E03FCF083C1A764C454D" author="Dash, PC" journalOrPublisher="Hexapoda, Insecta Indica" pageId="0" pageNumber="207" pagination="1 - 59" refId="B3" refString="Dash, PC, Viraktamath, CA, 1998. A review of the Indian and Nepalese grass feeding leafhopper genus Deltocephalus (Homoptera, Cicadellidae) with description of new species. Hexapoda, Insecta Indica 10: 1 - 59" title="A review of the Indian and Nepalese grass feeding leafhopper genus Deltocephalus (Homoptera, Cicadellidae) with description of new species." volume="10" year="1998">Dash and Viraktamath 1998</bibRefCitation>
: 22, 23, figs 150-158 (India);
<taxonomicName id="11377AF1A192B51792DF76F48D7D876C" baseAuthorityName="Metcalf" baseAuthorityYear="1967" class="Insecta" family="Cicadellidae" genus="Maiestas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Maiestas pruthii" order="Hemiptera" pageId="0" pageNumber="207" phylum="Arthropoda" rank="species" species="pruthii">Maiestas pruthii</taxonomicName>
:
<bibRefCitation id="2891BB74325BB435E38CDC2BCB536A87" DOI="https://doi.org/10.11646/zootaxa.2163.1.1" author="Webb, MD" journalOrPublisher="Zootaxa" pageId="0" pageNumber="207" pagination="1 - 64" refId="B25" refString="Webb, MD, Viraktamath, CA, 2009. Annotated checklist, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera, Auchenorrhyncha, Cicadellidae). Zootaxa 2163 (1): 1 - 64, DOI: https://doi.org/10.11646/zootaxa.2163.1.1" title="Annotated checklist, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera, Auchenorrhyncha, Cicadellidae)." url="https://doi.org/10.11646/zootaxa.2163.1.1" volume="2163" year="2009">Webb and Viraktamath 2009</bibRefCitation>
: 20, new combination;
<bibRefCitation id="9584497A21996AA2EF4AB96DBA1EB205" DOI="https://doi.org/10.11646/zootaxa.2365.1.1" author="Khatri, I" journalOrPublisher="Zootaxa" pageId="0" pageNumber="207" pagination="1 - 47" refId="B10" refString="Khatri, I, Webb, MD, 2010. The Deltocephalinae leafhoppers of Pakistan (Hemiptera, Cicadellidae). Zootaxa 2365 (1): 1 - 47, DOI: https://doi.org/10.11646/zootaxa.2365.1.1" title="The Deltocephalinae leafhoppers of Pakistan (Hemiptera, Cicadellidae)." url="https://doi.org/10.11646/zootaxa.2365.1.1" volume="2365" year="2010">Khatri and Webb 2010</bibRefCitation>
: 11, pl. 2a fig. 13, misidentification;
<bibRefCitation id="B5C18A1999E5F01936E2EAD9DA070EDE" DOI="https://doi.org/10.35249/rche.45.2.19.16" author="Naveed, H" journalOrPublisher="Revista Chilena de Entomologia" pageId="0" pageNumber="207" pagination="283 - 292" refId="B18" refString="Naveed, H, Sohail, K, Islam, W, Zhang, YL, Bu, WJ, 2019. A review of the leafhopper tribe Deltocephalini (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan. Revista Chilena de Entomologia 45 (2): 283 - 292, DOI: https://doi.org/10.35249/rche.45.2.19.16" title="A review of the leafhopper tribe Deltocephalini (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan." url="https://doi.org/10.35249/rche.45.2.19.16" volume="45" year="2019">Naveed et al. 2019</bibRefCitation>
: 286, fig. 2A-C (incorrectly cited as
<taxonomicName id="39FC3B30D34805917D0C107EECE641A1" family="Cicadellidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="M. subviridis" order="Hemiptera" pageId="0" pageNumber="207" rank="species" species="subviridis">M. subviridis</taxonomicName>
; H. Naveed pers. comm.), Fig.
<figureCitation id="E420C2DEC0185E9207D707F9D7993C5A" captionStart="Figures 18" captionStartId="F1" captionText="Figures 1 - 8. Paramesodes dirensis sp. nov. 1 habitus, dorsal view 2 face 3 male pygofer, lateral view 4 male pygofer, dorsal view 5 subgenital plate, valve and style, dorsal view 6 connective and aedeagus, dorsal view 7 connective and aedeagus, lateral view 8 female abdominal tip and 7 th sternum, dorsal view." figureDoi="10.3897/zookeys.1186.110266.figures1-8" httpUri="https://binary.pensoft.net/fig/952033" pageId="0" pageNumber="207">3i</figureCitation>
, misidentification;
<bibRefCitation id="F923BEDF5283578868664AB9E61C9FDD" DOI="https://doi.org/10.11646/zootaxa.5060.3.6" author="Shah, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="207" pagination="401 - 416" refId="B22" refString="Shah, B, Naveed, H, Webb, M, Duan, YN, 2021. Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records. Zootaxa 5060 (3): 401 - 416, DOI: https://doi.org/10.11646/zootaxa.5060.3.6" title="Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records." url="https://doi.org/10.11646/zootaxa.5060.3.6" volume="5060" year="2021">Shah et al. 2021</bibRefCitation>
: 406, fig. 4F-L (Pakistan).
</paragraph>
</subSubSection>
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<paragraph id="C227505E0447721ABB8D1FAE2D7AD62E" pageId="0" pageNumber="207">Diagnosis.</paragraph>
<paragraph id="547E27A3B0DCB08658A5F228EF787F95" pageId="0" pageNumber="207">
This species has a relatively long, acute head, with a pair of inverted U-shaped markings basally, and forewings with extra cross veins. The identity of this species is based on the figures of
<bibRefCitation id="554EB38F578C6759869617A2EDA60367" author="Dash, PC" journalOrPublisher="Hexapoda, Insecta Indica" pageId="0" pageNumber="207" pagination="1 - 59" refId="B3" refString="Dash, PC, Viraktamath, CA, 1998. A review of the Indian and Nepalese grass feeding leafhopper genus Deltocephalus (Homoptera, Cicadellidae) with description of new species. Hexapoda, Insecta Indica 10: 1 - 59" title="A review of the Indian and Nepalese grass feeding leafhopper genus Deltocephalus (Homoptera, Cicadellidae) with description of new species." volume="10" year="1998">Dash and Viraktamath (1998)</bibRefCitation>
.
</paragraph>
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<subSubSection id="8041F16AFE5EE498EF8C7123C2B42E4C" pageId="0" pageNumber="207" type="distribution">
<paragraph id="CA49CEC71D0AE0341D6362DFE5BCF633" pageId="0" pageNumber="207">Distribution.</paragraph>
<paragraph id="8ADC332E5F7682F4D293094EAC14810C" pageId="0" pageNumber="207">India, Pakistan.</paragraph>
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</treatment>
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