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<mods:title id="3C93988ED9C897A6E937D53BD5DBB4D9">First record of Larsonella pumilus (Teleostei: Gobiidae) from Japan, with phylogenetic placement of the genus Larsonella</mods:title>
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<treatment id="03FEE74FFFC3FFBAFF6BFF2DFBF458E1" ID-DOI="http://doi.org/10.5281/zenodo.5628310" ID-GBIF-Taxon="160238668" ID-Zenodo-Dep="5628310" LSID="urn:lsid:plazi:treatment:03FEE74FFFC3FFBAFF6BFF2DFBF458E1" httpUri="http://treatment.plazi.org/id/03FEE74FFFC3FFBAFF6BFF2DFBF458E1" lastPageId="6" lastPageNumber="373" pageId="3" pageNumber="370">
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<paragraph id="8BE85659FFC3FFBFFF6BFF2DFD4959A5" blockId="3.[151,690,153,251]" box="[151,690,153,180]" pageId="3" pageNumber="370">
<heading id="D0A0E135FFC3FFBFFF6BFF2DFD4959A5" bold="true" box="[151,690,153,180]" fontSize="11" level="1" pageId="3" pageNumber="370" reason="1">
<taxonomicName id="4C572DDAFFC3FFBFFF6BFF2DFD4959A5" ID-CoL="6QPCC" authority="(Larson &amp; Hoese, 1980)" baseAuthorityName="Larson &amp; Hoese" baseAuthorityYear="1980" box="[151,690,153,180]" class="Actinopterygii" family="Gobiidae" genus="Larsonella" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="370" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9238A4BFFC3FFBFFF6BFF2DFD4959A5" bold="true" box="[151,690,153,180]" pageId="3" pageNumber="370">
<emphasis id="B9238A4BFFC3FFBFFF6BFF2DFE7859A2" bold="true" box="[151,387,153,180]" italics="true" pageId="3" pageNumber="370">Larsonella pumilus</emphasis>
(
<bibRefCitation id="EFC62BA8FFC3FFBFFE6EFF2DFD5159A5" author="Larson, H. K. &amp; Hoese, D. F." box="[402,682,153,180]" pageId="3" pageNumber="370" pagination="33 - 43" refId="ref5734" refString="Larson, H. K. &amp; Hoese, D. F. (1980) Fische des Indischen Ozeans. Ergebnisse der ichthyologischen Untersuchungen wahrend der Expedition des Forschungsschiffes &quot; Meteor &quot; in den Indischen Ozean, Oktober 1964 bis Mai 1965. A. Systematischer Teil, XXIII. Gobiidae. Meteor Forschungsergebnisse, Reihe D, Biologie, 32, 33 - 43." type="journal article" year="1980">Larson &amp; Hoese, 1980</bibRefCitation>
)
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</heading>
</paragraph>
<paragraph id="8BE85659FFC3FFBFFF6BFF0AFDE459CE" blockId="3.[151,690,153,251]" box="[151,543,189,216]" pageId="3" pageNumber="370">[New Japanese name: Yuuna-haze]</paragraph>
</subSubSection>
<subSubSection id="C34D05D2FFC3FFBFFF6BFF55FE8F59ED" box="[151,372,225,251]" pageId="3" pageNumber="370" type="description">
<paragraph id="8BE85659FFC3FFBFFF6BFF55FE8F59ED" blockId="3.[151,690,153,251]" box="[151,372,225,251]" pageId="3" pageNumber="370">
(
<figureCitation id="136C4ADCFFC3FFBFFF63FF55FEF759ED" box="[159,268,225,251]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,913,938]" captionStartId-1="3.[151,250,1905,1930]" captionStartId-2="4.[151,250,1419,1444]" captionStartId-3="5.[151,250,1107,1132]" captionTargetBox-0="[156,1431,468,888]" captionTargetBox-1="[202,1388,954,1877]" captionTargetBox-2="[161,1427,194,1391]" captionTargetBox-3="[155,1411,585,1086]" captionTargetId-0="figure@3.[151,1436,463,893]" captionTargetId-1="figure@3.[191,1396,945,1886]" captionTargetId-2="figure@4.[151,1436,184,1400]" captionTargetId-3="figure@5.[151,1419,582,1088]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Larsonella pumilus, OCF-P 3808 (21.6 mm in standard length), in life (A), fresh (B), and after preservation (C)." captionText-1="FIGURE 2. Three-dimensional images of the head of Larsonella pumilus reconstructed from microcomputed tomography data. A, frontal view, showing the outermost teeth. B, mid-sagittal view of the right side (cut along section x-x in A), showing inner teeth without a part of ceratohyal and 5th branchiostegal ray to observe teeth easily. Red, premaxilla and outermost teeth on the premaxilla; blue, dentary and outermost teeth on the dentary. Triangles, vestiges of missing teeth. Scale bars, 1 mm." captionText-2="FIGURE 3. Picture of the caudal peduncle, stained with cyanine-blue solution to show scale arrangements (left) and schematic illustrations of cephalic sensory papillae (right) of Larsonella pumilus (OCF-P 3808, 21.6 mm SL) in dorsal (top), lateral (middle), and ventral (bottom) views. AN and PN, anterior and posterior nares, respectively. Dotted lines, the probable undamaged states of the damaged parts." captionText-3="FIGURE 4. Pigmentations around the eyes of Larsonella pumilus (OCF-P 3808, 21.6 mm SL) in dorsal (left) and lateral (right) views. Photos were taken after preservation." figureDoi-0="http://doi.org/10.5281/zenodo.3534804" figureDoi-1="http://doi.org/10.5281/zenodo.3534806" figureDoi-2="http://doi.org/10.5281/zenodo.3534808" figureDoi-3="http://doi.org/10.5281/zenodo.3534810" httpUri-0="https://zenodo.org/record/3534804/files/figure.png" httpUri-1="https://zenodo.org/record/3534806/files/figure.png" httpUri-2="https://zenodo.org/record/3534808/files/figure.png" httpUri-3="https://zenodo.org/record/3534810/files/figure.png" pageId="3" pageNumber="370">Figs. 14</figureCitation>
;
<tableCitation id="C6D563E2FFC3FFBFFEEBFF55FE9059ED" box="[279,363,225,251]" captionStart="TABLE 2" captionStartId="7.[151,239,153,177]" captionTargetPageId="7" captionText="TABLE 2. Counts and measurements of Larsonella pumilus. Values of the holotype are based on Larson &amp; Hoese (1980). D1, first dorsal fin; D2, second dorsal fin; A, anal fin; C, caudal fin; P1, pectoral fin; P2, pelvic fin; PO, preopercular." httpUri="http://table.plazi.org/id/DF2806D1FFC7FFBBFF6BFF2DFA9D59C3" pageId="3" pageNumber="370" tableUuid="DF2806D1FFC7FFBBFF6BFF2DFA9D59C3">Table 2</tableCitation>
)
</paragraph>
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<paragraph id="8BE85659FFC3FFBFFF6BFE92FC82584A" blockId="3.[151,1328,293,349]" pageId="3" pageNumber="370">
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<taxonomicName id="4C572DDAFFC3FFBFFF6BFE92FD6F5828" ID-CoL="6QPCC" authority="Larson &amp; Hoese, 1980: 41" authorityName="Larson &amp; Hoese" authorityPageNumber="41" authorityYear="1980" box="[151,660,294,318]" class="Actinopterygii" family="Gobiidae" genus="Lubricogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="370" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9238A4BFFC3FFBFFF6BFE92FE7A5828" box="[151,385,294,318]" italics="true" pageId="3" pageNumber="370">Lubricogobius pumilus</emphasis>
<treatmentCitation id="0AF67048FFC3FFBFFE74FE92FD6F5828" author="Larson, H. K. &amp; Hoese, D. F." box="[392,660,294,318]" page="41" pageId="3" pageNumber="370" year="1980">
<bibRefCitation id="EFC62BA8FFC3FFBFFE74FE92FD6F5828" author="Larson, H. K. &amp; Hoese, D. F." box="[392,660,294,318]" pageId="3" pageNumber="370" pagination="33 - 43" refId="ref5734" refString="Larson, H. K. &amp; Hoese, D. F. (1980) Fische des Indischen Ozeans. Ergebnisse der ichthyologischen Untersuchungen wahrend der Expedition des Forschungsschiffes &quot; Meteor &quot; in den Indischen Ozean, Oktober 1964 bis Mai 1965. A. Systematischer Teil, XXIII. Gobiidae. Meteor Forschungsergebnisse, Reihe D, Biologie, 32, 33 - 43." type="journal article" year="1980">Larson &amp; Hoese, 1980: 41</bibRefCitation>
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(
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locality: Indian Ocean,
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<geoCoordinate id="EE63309EFFC3FFBFFBE8FE92FB855828" box="[1044,1150,294,318]" degrees="47" direction="east" minutes="14.8" orientation="longitude" pageId="3" pageNumber="370" precision="92" value="47.246666">47°14.8E</geoCoordinate>
,
<quantity id="4CAFFBBCFFC3FFBFFB74FE92FB19582B" box="[1160,1250,294,318]" metricMagnitude="1" metricUnit="m" metricValue="3.75" metricValueMax="3.8" metricValueMin="3.7" pageId="3" pageNumber="370" unit="m" value="37.5" valueMax="38.0" valueMin="37.0">3738 m</quantity>
depth).
<taxonomicName id="4C572DDAFFC3FFBFFF6BFEF0FDA7584A" ID-CoL="6QPCC" authority="(Larson &amp; Hoese, 1980)" baseAuthorityName="Larson &amp; Hoese" baseAuthorityYear="1980" box="[151,604,324,348]" class="Actinopterygii" family="Gobiidae" genus="Larsonella" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="370" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9238A4BFFC3FFBFFF6BFEF0FEA0584A" box="[151,347,324,348]" italics="true" pageId="3" pageNumber="370">Larsonella pumilus</emphasis>
(
<bibRefCitation id="EFC62BA8FFC3FFBFFE95FEF0FDAF584A" author="Larson, H. K. &amp; Hoese, D. F." box="[361,596,324,348]" pageId="3" pageNumber="370" pagination="33 - 43" refId="ref5734" refString="Larson, H. K. &amp; Hoese, D. F. (1980) Fische des Indischen Ozeans. Ergebnisse der ichthyologischen Untersuchungen wahrend der Expedition des Forschungsschiffes &quot; Meteor &quot; in den Indischen Ozean, Oktober 1964 bis Mai 1965. A. Systematischer Teil, XXIII. Gobiidae. Meteor Forschungsergebnisse, Reihe D, Biologie, 32, 33 - 43." type="journal article" year="1980">Larson &amp; Hoese, 1980</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="0AF67048FFC3FFBFFD99FEF0FC8E584A" author="Randall, J. E. &amp; Senou, H." box="[613,885,324,349]" page="11" pageId="3" pageNumber="370" year="2001">
<bibRefCitation id="EFC62BA8FFC3FFBFFD99FEF0FC8E584A" author="Randall, J. E. &amp; Senou, H." box="[613,885,324,349]" pageId="3" pageNumber="370" pagination="3 - 12" refId="ref6288" refString="Randall, J. E. &amp; Senou, H. (2001) Review of the Indo-Pacific gobiid fish genus Lubricogobius, with description of a new species and a new genus for L. pumilus. Ichthyological Research, 48, 3 - 12. https: // doi. org / 10.1007 / s 10228 - 001 - 8111 - z" type="journal article" year="2001">Randall &amp; Senou 2001: 11</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C34D05D2FFC3FFBFFF6BFE3EFED458DF" pageId="3" pageNumber="370" type="materials_examined">
<paragraph id="8BE85659FFC3FFBFFF6BFE3EFED458DF" blockId="3.[151,1436,394,457]" pageId="3" pageNumber="370">
<emphasis id="B9238A4BFFC3FFBFFF6BFE3EFE7958B3" bold="true" box="[151,386,394,421]" pageId="3" pageNumber="370">Material examined.</emphasis>
OCF-P 3808,
<quantity id="4CAFFBBCFFC3FFBFFDD5FE3FFD7458B3" box="[553,655,395,421]" metricMagnitude="-2" metricUnit="m" metricValue="2.16" pageId="3" pageNumber="370" unit="mm" value="21.6">21.6 mm</quantity>
SL, East
<collectingCountry id="F34016C9FFC3FFBFFCFDFE3FFCBF58B3" box="[769,836,395,421]" name="China" pageId="3" pageNumber="370">China</collectingCountry>
Sea off Seragaki, Onna Village,
<collectingRegion id="499398BBFFC3FFBFFB41FE3FFADE58B3" box="[1213,1317,395,421]" country="Japan" name="Okinawa" pageId="3" pageNumber="370">Okinawa</collectingRegion>
,
<collectingCountry id="F34016C9FFC3FFBFFACEFE3FFA8E58B3" box="[1330,1397,395,421]" name="Japan" pageId="3" pageNumber="370">Japan</collectingCountry>
,
<date id="FFE97099FFC3FFBFFA7DFE3FFED058DF" pageId="3" pageNumber="370" value="2017-08-11">11 August 2017</date>
.
</paragraph>
</subSubSection>
<caption id="DF2806D1FFC3FFBFFF6BFC25FA725ABC" ID-DOI="http://doi.org/10.5281/zenodo.3534804" ID-Zenodo-Dep="3534804" box="[151,1417,913,938]" httpUri="https://zenodo.org/record/3534804/files/figure.png" pageId="3" pageNumber="370" startId="3.[151,250,913,938]" targetBox="[156,1431,468,888]" targetPageId="3">
<paragraph id="8BE85659FFC3FFBFFF6BFC25FA725ABC" blockId="3.[151,1417,913,938]" box="[151,1417,913,938]" pageId="3" pageNumber="370">
<emphasis id="B9238A4BFFC3FFBFFF6BFC25FEE85ABC" bold="true" box="[151,275,913,938]" pageId="3" pageNumber="370">FIGURE 1.</emphasis>
<taxonomicName id="4C572DDAFFC3FFBFFEE5FC25FE255ABF" baseAuthorityName="Larson &amp; Hoese" baseAuthorityYear="1980" box="[281,478,913,937]" class="Actinopterygii" family="Gobiidae" genus="Larsonella" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="370" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9238A4BFFC3FFBFFEE5FC25FE255ABF" box="[281,478,913,937]" italics="true" pageId="3" pageNumber="370">Larsonella pumilus</emphasis>
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, OCF-P 3808 (21.6 mm in standard length), in life (A), fresh (B), and after preservation (C).
</paragraph>
</caption>
<caption id="DF2806D1FFC3FFBFFF6BF8C5FAD65EE0" ID-DOI="http://doi.org/10.5281/zenodo.3534806" ID-Zenodo-Dep="3534806" httpUri="https://zenodo.org/record/3534806/files/figure.png" pageId="3" pageNumber="370" startId="3.[151,250,1905,1930]" targetBox="[202,1388,954,1877]" targetPageId="3">
<paragraph id="8BE85659FFC3FFBFFF6BF8C5FAD65EE0" blockId="3.[151,1437,1905,2038]" pageId="3" pageNumber="370">
<emphasis id="B9238A4BFFC3FFBFFF6BF8C5FEE95E9C" bold="true" box="[151,274,1905,1930]" pageId="3" pageNumber="370">FIGURE 2.</emphasis>
Three-dimensional images of the head of
<taxonomicName id="4C572DDAFFC3FFBFFD45F8C6FC865E9C" baseAuthorityName="Larson &amp; Hoese" baseAuthorityYear="1980" box="[697,893,1906,1930]" class="Actinopterygii" family="Gobiidae" genus="Larsonella" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="370" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9238A4BFFC3FFBFFD45F8C6FC865E9C" box="[697,893,1906,1930]" italics="true" pageId="3" pageNumber="370">Larsonella pumilus</emphasis>
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reconstructed from microcomputed tomography data. A, frontal view, showing the outermost teeth. B, mid-sagittal view of the right side (cut along section x-x in A), showing inner teeth without a part of ceratohyal and 5th branchiostegal ray to observe teeth easily. Red, premaxilla and outermost teeth on the premaxilla; blue, dentary and outermost teeth on the dentary. Triangles, vestiges of missing teeth. Scale bars, 1 mm.
</paragraph>
</caption>
<caption id="DF2806D1FFC4FFB8FF6BFA3FFE4A5F06" ID-DOI="http://doi.org/10.5281/zenodo.3534808" ID-Zenodo-Dep="3534808" httpUri="https://zenodo.org/record/3534808/files/figure.png" pageId="4" pageNumber="371" startId="4.[151,250,1419,1444]" targetBox="[161,1427,194,1391]" targetPageId="4">
<paragraph id="8BE85659FFC4FFB8FF6BFA3FFE4A5F06" blockId="4.[151,1436,1419,1552]" pageId="4" pageNumber="371">
<emphasis id="B9238A4BFFC4FFB8FF6BFA3FFEE85CB2" bold="true" box="[151,275,1419,1444]" pageId="4" pageNumber="371">FIGURE 3.</emphasis>
Picture of the caudal peduncle, stained with cyanine-blue solution to show scale arrangements (left) and schematic illustrations of cephalic sensory papillae (right) of
<taxonomicName id="4C572DDAFFC4FFB8FD6BFA04FCA05CDE" baseAuthorityName="Larson &amp; Hoese" baseAuthorityYear="1980" box="[663,859,1456,1480]" class="Actinopterygii" family="Gobiidae" genus="Larsonella" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9238A4BFFC4FFB8FD6BFA04FCA05CDE" box="[663,859,1456,1480]" italics="true" pageId="4" pageNumber="371">Larsonella pumilus</emphasis>
</taxonomicName>
(OCF-P 3808, 21.6 mm SL) in dorsal (top), lateral (middle), and ventral (bottom) views. AN and PN, anterior and posterior nares, respectively. Dotted lines, the probable undamaged states of the damaged parts.
</paragraph>
</caption>
<subSubSection id="C34D05D2FFC4FFBAFF3BF98AFBF458E1" lastPageId="6" lastPageNumber="373" pageId="4" pageNumber="371" type="description">
<paragraph id="8BE85659FFC4FFB8FF3BF98AFD605E42" blockId="4.[151,1437,1598,2021]" pageId="4" pageNumber="371">
<emphasis id="B9238A4BFFC4FFB8FF3BF98AFEA25F4F" bold="true" box="[199,345,1598,1625]" pageId="4" pageNumber="371">Description.</emphasis>
Counts and measurements are shown in
<tableCitation id="C6D563E2FFC4FFB8FCE0F98AFC8A5F4F" box="[796,881,1598,1625]" captionStart="TABLE 2" captionStartId="7.[151,239,153,177]" captionTargetPageId="7" captionText="TABLE 2. Counts and measurements of Larsonella pumilus. Values of the holotype are based on Larson &amp; Hoese (1980). D1, first dorsal fin; D2, second dorsal fin; A, anal fin; C, caudal fin; P1, pectoral fin; P2, pelvic fin; PO, preopercular." httpUri="http://table.plazi.org/id/DF2806D1FFC7FFBBFF6BFF2DFA9D59C3" pageId="4" pageNumber="371" tableUuid="DF2806D1FFC7FFBBFF6BFF2DFA9D59C3">Table 2</tableCitation>
. Head depressed, trunk nearly cylindrical, and tail compressed. Eyes located dorsolaterally. Mouth strongly oblique with angle to body axis about 60 degrees. Lower jaw protruding beyond upper jaw. Posterior end of upper jaw reaching below middle point between anterior margin of iris and anterior margin of pupil. Canine-like teeth aligned on edges of anterior halves of premaxilla and dentary; four and three teeth on one side of premaxilla and dentary, respectively; posterior teeth larger (
<figureCitation id="136C4ADCFFC4FFB8FB36F97AFADE5FFF" box="[1226,1317,1742,1769]" captionStart="FIGURE 2" captionStartId="3.[151,250,1905,1930]" captionTargetBox="[202,1388,954,1877]" captionTargetId="figure@3.[191,1396,945,1886]" captionTargetPageId="3" captionText="FIGURE 2. Three-dimensional images of the head of Larsonella pumilus reconstructed from microcomputed tomography data. A, frontal view, showing the outermost teeth. B, mid-sagittal view of the right side (cut along section x-x in A), showing inner teeth without a part of ceratohyal and 5th branchiostegal ray to observe teeth easily. Red, premaxilla and outermost teeth on the premaxilla; blue, dentary and outermost teeth on the dentary. Triangles, vestiges of missing teeth. Scale bars, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3534806" httpUri="https://zenodo.org/record/3534806/files/figure.png" pageId="4" pageNumber="371">Fig. 2A</figureCitation>
). An inner row of conical teeth extending from anterior part to more posterior part of dentary than the outer canine-like teeth row (
<figureCitation id="136C4ADCFFC4FFB8FF2EF8A2FED25E27" box="[210,297,1814,1841]" captionStart="FIGURE 2" captionStartId="3.[151,250,1905,1930]" captionTargetBox="[202,1388,954,1877]" captionTargetId="figure@3.[191,1396,945,1886]" captionTargetPageId="3" captionText="FIGURE 2. Three-dimensional images of the head of Larsonella pumilus reconstructed from microcomputed tomography data. A, frontal view, showing the outermost teeth. B, mid-sagittal view of the right side (cut along section x-x in A), showing inner teeth without a part of ceratohyal and 5th branchiostegal ray to observe teeth easily. Red, premaxilla and outermost teeth on the premaxilla; blue, dentary and outermost teeth on the dentary. Triangles, vestiges of missing teeth. Scale bars, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3534806" httpUri="https://zenodo.org/record/3534806/files/figure.png" pageId="4" pageNumber="371">Fig. 2B</figureCitation>
). Additional small conical teeth observed on inner parts of premaxilla and dentary with a stereomicroscope, but no such small teeth with micro-CT.
</paragraph>
<paragraph id="8BE85659FFC4FFB9FF3BF8EAFAB559C1" blockId="4.[151,1437,1598,2021]" lastBlockId="5.[151,1437,153,539]" lastPageId="5" lastPageNumber="372" pageId="4" pageNumber="371">
First dorsal fin with six spines. Second dorsal fin with one spine and nine soft rays. First and second dorsal fins connected by a low membrane behind last spine of the first dorsal fin. Anal fin with one spine and eight soft rays. Caudal fin rounded, with 17 segmented rays. Pectoral fin with 18 soft-rays. Pelvic fin with one spine and five soft rays. Posterior tips of pectoral fins reaching or exceeding position of anus, according to a photograph taken in life (
<figureCitation id="136C4ADCFFC5FFB9FF5CFF2DFF0359A5" box="[160,248,153,179]" captionStart="FIGURE 1" captionStartId="3.[151,250,913,938]" captionTargetBox="[156,1431,468,888]" captionTargetId="figure@3.[151,1436,463,893]" captionTargetPageId="3" captionText="FIGURE 1. Larsonella pumilus, OCF-P 3808 (21.6 mm in standard length), in life (A), fresh (B), and after preservation (C)." figureDoi="http://doi.org/10.5281/zenodo.3534804" httpUri="https://zenodo.org/record/3534804/files/figure.png" pageId="5" pageNumber="372">Fig. 1A</figureCitation>
), although they are broken in the preserved specimen. Pelvic fins without frenum. Vertebrae 10 + 16 = 26; dorsal-fin pterygiophore formula 3-22110; epural 1; anal-fin pterygiophores anterior to first haemal spine 2.
</paragraph>
<paragraph id="8BE85659FFC5FFB9FF3BFF55FAB45871" blockId="5.[151,1437,153,539]" pageId="5" pageNumber="372">
Head and body largely naked except for posterior part of caudal peduncle, involving three rows composed of 710 ctenoid scales along dorsal midline. A row of three ctenoid scales along lateral midline, and three rows composed of 68 ctenoid scales along ventral midline (
<figureCitation id="136C4ADCFFC5FFB9FD32FE9DFCE95855" box="[718,786,297,323]" captionStart="FIGURE 3" captionStartId="4.[151,250,1419,1444]" captionTargetBox="[161,1427,194,1391]" captionTargetId="figure@4.[151,1436,184,1400]" captionTargetPageId="4" captionText="FIGURE 3. Picture of the caudal peduncle, stained with cyanine-blue solution to show scale arrangements (left) and schematic illustrations of cephalic sensory papillae (right) of Larsonella pumilus (OCF-P 3808, 21.6 mm SL) in dorsal (top), lateral (middle), and ventral (bottom) views. AN and PN, anterior and posterior nares, respectively. Dotted lines, the probable undamaged states of the damaged parts." figureDoi="http://doi.org/10.5281/zenodo.3534808" httpUri="https://zenodo.org/record/3534808/files/figure.png" pageId="5" pageNumber="372">Fig. 3</figureCitation>
). No sensory canals or associated pores on head. Cephalic sensory papillae patterns illustrated in
<figureCitation id="136C4ADCFFC5FFB9FDBBFEF9FD715871" box="[583,650,333,359]" captionStart="FIGURE 3" captionStartId="4.[151,250,1419,1444]" captionTargetBox="[161,1427,194,1391]" captionTargetId="figure@4.[151,1436,184,1400]" captionTargetPageId="4" captionText="FIGURE 3. Picture of the caudal peduncle, stained with cyanine-blue solution to show scale arrangements (left) and schematic illustrations of cephalic sensory papillae (right) of Larsonella pumilus (OCF-P 3808, 21.6 mm SL) in dorsal (top), lateral (middle), and ventral (bottom) views. AN and PN, anterior and posterior nares, respectively. Dotted lines, the probable undamaged states of the damaged parts." figureDoi="http://doi.org/10.5281/zenodo.3534808" httpUri="https://zenodo.org/record/3534808/files/figure.png" pageId="5" pageNumber="372">Fig. 3</figureCitation>
. Infraorbital area with five transverse rows of sensory papillae.
</paragraph>
<paragraph id="8BE85659FFC5FFB9FF3BFEC5FB195B0D" blockId="5.[151,1437,153,539]" pageId="5" pageNumber="372">
Color in preservative (
<figureCitation id="136C4ADCFFC5FFB9FE39FEC5FDE6589D" box="[453,541,369,395]" captionStart="FIGURE 1" captionStartId="3.[151,250,913,938]" captionTargetBox="[156,1431,468,888]" captionTargetId="figure@3.[151,1436,463,893]" captionTargetPageId="3" captionText="FIGURE 1. Larsonella pumilus, OCF-P 3808 (21.6 mm in standard length), in life (A), fresh (B), and after preservation (C)." figureDoi="http://doi.org/10.5281/zenodo.3534804" httpUri="https://zenodo.org/record/3534804/files/figure.png" pageId="5" pageNumber="372">Fig. 1C</figureCitation>
): Background of head and tail white, trunk yellowish white. All fin membranes transparent. Melanophores scattered on dorsal half of trunk and on membranes of first and second dorsal, anal, and pelvic fins. Pectoral fin also with a few melanophores. Three rows of tiny melanophores arched between right and left eyes (
<figureCitation id="136C4ADCFFC5FFB9FEFBFE69FE9858E1" box="[263,355,477,503]" captionStart="FIGURE 4" captionStartId="5.[151,250,1107,1132]" captionTargetBox="[155,1411,585,1086]" captionTargetId="figure@5.[151,1419,582,1088]" captionTargetPageId="5" captionText="FIGURE 4. Pigmentations around the eyes of Larsonella pumilus (OCF-P 3808, 21.6 mm SL) in dorsal (left) and lateral (right) views. Photos were taken after preservation." figureDoi="http://doi.org/10.5281/zenodo.3534810" httpUri="https://zenodo.org/record/3534810/files/figure.png" pageId="5" pageNumber="372">Fig. 4A</figureCitation>
). Infraorbital area with two transverse rows of tiny melanophores (
<figureCitation id="136C4ADCFFC5FFB9FBA0FE69FB4F58E1" box="[1116,1204,477,503]" captionStart="FIGURE 4" captionStartId="5.[151,250,1107,1132]" captionTargetBox="[155,1411,585,1086]" captionTargetId="figure@5.[151,1419,582,1088]" captionTargetPageId="5" captionText="FIGURE 4. Pigmentations around the eyes of Larsonella pumilus (OCF-P 3808, 21.6 mm SL) in dorsal (left) and lateral (right) views. Photos were taken after preservation." figureDoi="http://doi.org/10.5281/zenodo.3534810" httpUri="https://zenodo.org/record/3534810/files/figure.png" pageId="5" pageNumber="372">Fig. 4B</figureCitation>
) and another row of tiny melanophores behind eyes (
<figureCitation id="136C4ADCFFC5FFB9FDFEFDB5FDA05B0D" box="[514,603,513,539]" captionStart="FIGURE 4" captionStartId="5.[151,250,1107,1132]" captionTargetBox="[155,1411,585,1086]" captionTargetId="figure@5.[151,1419,582,1088]" captionTargetPageId="5" captionText="FIGURE 4. Pigmentations around the eyes of Larsonella pumilus (OCF-P 3808, 21.6 mm SL) in dorsal (left) and lateral (right) views. Photos were taken after preservation." figureDoi="http://doi.org/10.5281/zenodo.3534810" httpUri="https://zenodo.org/record/3534810/files/figure.png" pageId="5" pageNumber="372">Fig. 4B</figureCitation>
). These melanophore rows arranged radially around eyes.
</paragraph>
<caption id="DF2806D1FFC5FFB9FF6BFBE7FDA35D86" ID-DOI="http://doi.org/10.5281/zenodo.3534810" ID-Zenodo-Dep="3534810" httpUri="https://zenodo.org/record/3534810/files/figure.png" pageId="5" pageNumber="372" startId="5.[151,250,1107,1132]" targetBox="[155,1411,585,1086]" targetPageId="5">
<paragraph id="8BE85659FFC5FFB9FF6BFBE7FDA35D86" blockId="5.[151,1436,1107,1168]" pageId="5" pageNumber="372">
<emphasis id="B9238A4BFFC5FFB9FF6BFBE7FEE95D7A" bold="true" box="[151,274,1107,1132]" pageId="5" pageNumber="372">FIGURE 4.</emphasis>
Pigmentations around the eyes of
<taxonomicName id="4C572DDAFFC5FFB9FD92FBE0FCCA5D7A" baseAuthorityName="Larson &amp; Hoese" baseAuthorityYear="1980" box="[622,817,1108,1132]" class="Actinopterygii" family="Gobiidae" genus="Larsonella" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9238A4BFFC5FFB9FD92FBE0FCCA5D7A" box="[622,817,1108,1132]" italics="true" pageId="5" pageNumber="372">Larsonella pumilus</emphasis>
</taxonomicName>
(OCF-P 3808, 21.6 mm SL) in dorsal (left) and lateral (right) views. Photos were taken after preservation.
</paragraph>
</caption>
<caption id="DF2806D1FFC5FFB9FF6BF9EEFF3D5FC9" ID-DOI="http://doi.org/10.5281/zenodo.3534812" ID-Zenodo-Dep="3534812" httpUri="https://zenodo.org/record/3534812/files/figure.png" pageId="5" pageNumber="372" startId="5.[151,250,1626,1651]" targetBox="[159,1426,1216,1601]" targetPageId="5">
<paragraph id="8BE85659FFC5FFB9FF6BF9EEFF3D5FC9" blockId="5.[151,1436,1626,1759]" pageId="5" pageNumber="372">
<emphasis id="B9238A4BFFC5FFB9FF6BF9EEFEE85F65" bold="true" box="[151,275,1626,1651]" pageId="5" pageNumber="372">FIGURE 5.</emphasis>
Underwater photograph (A) and a dried shell (B and C) of
<taxonomicName id="4C572DDAFFC5FFB9FC89F9EFFBB05F65" baseAuthorityName="Philippi" baseAuthorityYear="1841" box="[885,1099,1627,1651]" class="Gastropoda" family="Xenophoridae" genus="Xenophora" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="372" phylum="Mollusca" rank="species" species="chinensis">
<emphasis id="B9238A4BFFC5FFB9FC89F9EFFBB05F65" box="[885,1099,1627,1651]" italics="true" pageId="5" pageNumber="372">Xenophora chinensis</emphasis>
</taxonomicName>
. A, Collecting the shell in which
<taxonomicName id="4C572DDAFFC5FFB9FF6BF9CBFEA75F81" baseAuthorityName="Larson &amp; Hoese" baseAuthorityYear="1980" box="[151,348,1663,1687]" class="Actinopterygii" family="Gobiidae" genus="Larsonella" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9238A4BFFC5FFB9FF6BF9CBFEA75F81" box="[151,348,1663,1687]" italics="true" pageId="5" pageNumber="372">Larsonella pumilus</emphasis>
</taxonomicName>
may have taken refuge, using a Remotely Operated Vehicle at a depth of 214 m in the East China Sea off Seragaki, Onna Village, Okinawa, Japan. B (dorsal view) and C (ventral view) are likely the same shell as A. Scale bars, 10 mm.
</paragraph>
</caption>
<paragraph id="8BE85659FFC5FFB9FF3BF8B9FD6E5E5A" blockId="5.[151,1436,1805,2012]" pageId="5" pageNumber="372">
Color in life (
<figureCitation id="136C4ADCFFC5FFB9FE9FF8B9FE465E3E" box="[355,445,1805,1832]" captionStart="FIGURE 1" captionStartId="3.[151,250,913,938]" captionTargetBox="[156,1431,468,888]" captionTargetId="figure@3.[151,1436,463,893]" captionTargetPageId="3" captionText="FIGURE 1. Larsonella pumilus, OCF-P 3808 (21.6 mm in standard length), in life (A), fresh (B), and after preservation (C)." figureDoi="http://doi.org/10.5281/zenodo.3534804" httpUri="https://zenodo.org/record/3534804/files/figure.png" pageId="5" pageNumber="372">Fig. 1A</figureCitation>
): Body and all fins yellow or yellowish orange. Arrangement of melanophores same as after preservation, described above (
<figureCitation id="136C4ADCFFC5FFB9FDD2F885FD7C5E5A" box="[558,647,1841,1868]" captionStart="FIGURE 1" captionStartId="3.[151,250,913,938]" captionTargetBox="[156,1431,468,888]" captionTargetId="figure@3.[151,1436,463,893]" captionTargetPageId="3" captionText="FIGURE 1. Larsonella pumilus, OCF-P 3808 (21.6 mm in standard length), in life (A), fresh (B), and after preservation (C)." figureDoi="http://doi.org/10.5281/zenodo.3534804" httpUri="https://zenodo.org/record/3534804/files/figure.png" pageId="5" pageNumber="372">Fig. 1A</figureCitation>
).
</paragraph>
<paragraph id="8BE85659FFC5FFBAFF3BF8E1FAB559ED" blockId="5.[151,1436,1805,2012]" lastBlockId="6.[151,1437,153,503]" lastPageId="6" lastPageNumber="373" pageId="5" pageNumber="372">
<emphasis id="B9238A4BFFC5FFB9FF3BF8E1FED15E66" bold="true" box="[199,298,1877,1904]" pageId="5" pageNumber="372">Habitat.</emphasis>
We collected ten specimens of five invertebrate species from the muddy bottom at a depth of
<quantity id="4CAFFBBCFFC5FFB9FAC6F8E2FF565E82" metricMagnitude="2" metricUnit="m" metricValue="2.145" metricValueMax="2.2" metricValueMin="2.09" pageId="5" pageNumber="372" unit="m" value="214.5" valueMax="220.0" valueMin="209.0">209220 m</quantity>
using the ROV, on
<date id="FFE97099FFC5FFB9FE74F8CEFDC05E82" box="[392,571,1914,1940]" pageId="5" pageNumber="372" value="2017-08-11">11 August 2017</date>
. These included five live comb jellies,
<taxonomicName id="4C572DDAFFC5FFB9FC11F8CEFB265E85" authorityName="Komai" authorityYear="1941" box="[1005,1245,1914,1939]" class="Tentaculata" family="Lyroctenidae" genus="Lyrocteis" kingdom="Animalia" order="Platyctenida" pageId="5" pageNumber="372" phylum="Ctenophora" rank="species" species="imperatoris">
<emphasis id="B9238A4BFFC5FFB9FC11F8CEFB265E85" box="[1005,1245,1914,1939]" italics="true" pageId="5" pageNumber="372">Lyrocteis imperatoris</emphasis>
</taxonomicName>
, a sea cucumber,
<taxonomicName id="4C572DDAFFC5FFB9FF6BF82AFEA95EAE" box="[151,338,1950,1976]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="5" pageNumber="372" phylum="Echinodermata" rank="species" species="dura">
<emphasis id="B9238A4BFFC5FFB9FF6BF82AFEA95EAE" box="[151,338,1950,1976]" italics="true" pageId="5" pageNumber="372">Holothuria dura</emphasis>
</taxonomicName>
, a starfish,
<taxonomicName id="4C572DDAFFC5FFB9FE27F82AFD0F5EA1" authorityName="Imaoka, Irimura, Okutani, Oguro, Oji &amp; Kanazawa, 1" authorityYear="1991" box="[475,756,1950,1976]" class="Asteroidea" family="Asterodiscididae" genus="Asterodiscides" kingdom="Animalia" order="Valvatida" pageId="5" pageNumber="372" phylum="Echinodermata" rank="species" species="japonicus">
<emphasis id="B9238A4BFFC5FFB9FE27F82AFD0F5EA1" box="[475,756,1950,1976]" italics="true" pageId="5" pageNumber="372">Asterodiscides japonicus</emphasis>
</taxonomicName>
, a heart urchin,
<taxonomicName id="4C572DDAFFC5FFB9FC48F82AFBCC5EA1" box="[948,1079,1950,1975]" class="Echinoidea" family="Pericosmidae" genus="Pericosmus" kingdom="Animalia" order="Spatangoida" pageId="5" pageNumber="372" phylum="Echinodermata" rank="genus">
<emphasis id="B9238A4BFFC5FFB9FC48F82AFBCC5EA1" box="[948,1079,1950,1975]" italics="true" pageId="5" pageNumber="372">Pericosmus</emphasis>
</taxonomicName>
sp., and an empty shell of
<taxonomicName id="4C572DDAFFC5FFB9FA89F82AFE9F5ECA" baseAuthorityName="Philippi" baseAuthorityYear="1841" class="Gastropoda" family="Xenophoridae" genus="Xenophora" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="372" phylum="Mollusca" rank="species" species="chinensis">
<emphasis id="B9238A4BFFC5FFB9FA89F82AFE9F5ECA" italics="true" pageId="5" pageNumber="372">Xenophora chinensis</emphasis>
</taxonomicName>
(
<figureCitation id="136C4ADCFFC5FFB9FE8FF875FE4C5ECA" box="[371,439,1985,2012]" captionStart="FIGURE 5" captionStartId="5.[151,250,1626,1651]" captionTargetBox="[159,1426,1216,1601]" captionTargetId="figure@5.[151,1435,1211,1607]" captionTargetPageId="5" captionText="FIGURE 5. Underwater photograph (A) and a dried shell (B and C) of Xenophora chinensis. A, Collecting the shell in which Larsonella pumilus may have taken refuge, using a Remotely Operated Vehicle at a depth of 214 m in the East China Sea off Seragaki, Onna Village, Okinawa, Japan. B (dorsal view) and C (ventral view) are likely the same shell as A. Scale bars, 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3534812" httpUri="https://zenodo.org/record/3534812/files/figure.png" pageId="5" pageNumber="372">Fig. 5</figureCitation>
). After these animals were put into a tank on the boat, the
<taxonomicName id="4C572DDAFFC5FFB9FBC1F876FB485ECA" baseAuthorityName="Larson &amp; Hoese" baseAuthorityYear="1980" box="[1085,1203,1986,2012]" class="Actinopterygii" family="Gobiidae" genus="Larsonella" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9238A4BFFC5FFB9FBC1F876FB485ECA" box="[1085,1203,1986,2012]" italics="true" pageId="5" pageNumber="372">L. pumilus</emphasis>
</taxonomicName>
specimen was found in the tank. Because
<taxonomicName id="4C572DDAFFC6FFBAFE79FF2DFDD359A5" baseAuthorityName="Hoese &amp; Larson" baseAuthorityYear="2010" box="[389,552,153,179]" class="Actinopterygii" family="Gobiidae" genus="Lubricogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="genus">
<emphasis id="B9238A4BFFC6FFBAFE79FF2DFDD359A5" box="[389,552,153,179]" italics="true" pageId="6" pageNumber="373">Lubricogobius</emphasis>
</taxonomicName>
species, close relatives of
<taxonomicName id="4C572DDAFFC6FFBAFCA5FF2DFBCF59A5" baseAuthorityName="Larson &amp; Hoese" baseAuthorityYear="1980" box="[857,1076,153,179]" class="Actinopterygii" family="Gobiidae" genus="Larsonella" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9238A4BFFC6FFBAFCA5FF2DFBCF59A5" box="[857,1076,153,179]" italics="true" pageId="6" pageNumber="373">Larsonella pumilus</emphasis>
</taxonomicName>
, often inhabit empty shells, sea urchin tests, tunicate siphons, bottles, etc. (
<bibRefCitation id="EFC62BA8FFC6FFBAFD86FF09FC8459C1" author="Randall, J. E. &amp; Senou, H." box="[634,895,189,215]" pageId="6" pageNumber="373" pagination="3 - 12" refId="ref6288" refString="Randall, J. E. &amp; Senou, H. (2001) Review of the Indo-Pacific gobiid fish genus Lubricogobius, with description of a new species and a new genus for L. pumilus. Ichthyological Research, 48, 3 - 12. https: // doi. org / 10.1007 / s 10228 - 001 - 8111 - z" type="journal article" year="2001">Randall &amp; Senou 2001</bibRefCitation>
;
<bibRefCitation id="EFC62BA8FFC6FFBAFC76FF09FB6C59CE" author="Allen, G. R. &amp; Erdmann, M. V." box="[906,1175,189,216]" pageId="6" pageNumber="373" pagination="24 - 34" refId="ref4877" refString="Allen, G. R. &amp; Erdmann, M. V. (2016) Lubricogobius tunicatus, a new species of goby (Pisces: Gobiidae) from Papua New Guinea and the first record of L. ornatus from the East Indies. Journal of the Ocean Science Foundation, 24, 24 - 34." type="journal article" year="2016">Allen &amp; Erdmann 2016</bibRefCitation>
), we believe that the
<taxonomicName id="4C572DDAFFC6FFBAFA7AFF0AFF0B59ED" baseAuthorityName="Larson &amp; Hoese" baseAuthorityYear="1980" class="Actinopterygii" family="Gobiidae" genus="Larsonella" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9238A4BFFC6FFBAFA7AFF0AFF0B59ED" italics="true" pageId="6" pageNumber="373">L. pumilus</emphasis>
</taxonomicName>
was inside the empty shell of
<taxonomicName id="4C572DDAFFC6FFBAFDB8FF56FD3059ED" baseAuthorityName="Philippi" baseAuthorityYear="1841" box="[580,715,225,251]" class="Gastropoda" family="Xenophoridae" genus="Xenophora" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="373" phylum="Mollusca" rank="species" species="chinensis">
<emphasis id="B9238A4BFFC6FFBAFDB8FF56FD3059ED" box="[580,715,225,251]" italics="true" pageId="6" pageNumber="373">X. chinensis</emphasis>
</taxonomicName>
(collected at a depth of
<quantity id="4CAFFBBCFFC6FFBAFC26FF55FBD959ED" box="[986,1058,225,251]" metricMagnitude="2" metricUnit="m" metricValue="2.1399999999999997" pageId="6" pageNumber="373" unit="m" value="214.0">214 m</quantity>
) and was collected with it.
</paragraph>
<paragraph id="8BE85659FFC6FFBAFF3BFEB0FBF458E1" blockId="6.[151,1437,153,503]" pageId="6" pageNumber="373">
<emphasis id="B9238A4BFFC6FFBAFF3BFEB0FDDC5809" bold="true" box="[199,551,260,287]" pageId="6" pageNumber="373">Mitochondrial DNA analysis.</emphasis>
We succeeded in assembling the entire mitochondrial genomes of
<taxonomicName id="4C572DDAFFC6FFBAFADFFEB1FF0B5855" baseAuthorityName="Larson &amp; Hoese" baseAuthorityYear="1980" class="Actinopterygii" family="Gobiidae" genus="Larsonella" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9238A4BFFC6FFBAFADFFEB1FF0B5855" italics="true" pageId="6" pageNumber="373">Larsonella pumilus</emphasis>
</taxonomicName>
and 19 related species (
<tableCitation id="C6D563E2FFC6FFBAFE09FE9DFDB35855" box="[501,584,297,323]" captionStart="TABLE 1" captionStartId="2.[151,239,442,466]" captionTargetBox="[166,1404,478,1889]" captionText="TABLE 1. Accession numbers and lengths of mitochondrial genome sequences analyzed in this study. Species Catalog number of Locality Accession number Length of" httpUri="http://table.plazi.org/id/DF2806D1FFC2FFBEFF6BFE0EFB2758C5" pageId="6" pageNumber="373" tableUuid="DF2806D1FFC2FFBEFF6BFE0EFB2758C5">Table 1</tableCitation>
). In the phylogenetic tree, using 15559 bp of aligned mitochondrial genomes (
<figureCitation id="136C4ADCFFC6FFBAFF5CFEF9FF1F587E" box="[160,228,333,360]" captionStart="FIGURE 6" captionStartId="6.[151,250,1849,1874]" captionTargetBox="[151,1428,511,1815]" captionTargetId="figure@6.[151,1436,510,1830]" captionTargetPageId="6" captionText="FIGURE 6. Molecular phylogeny of Larsonella pumilus and 19 related goby species from the Ryukyu Archipelago. Tree topology and branch lengths were obtained from maximum likelihood analysis of aligned mitochondrial genomes (15559 bp) with 100 bootstraps based on the GTR+I+Gamma model. The scale bar indicates 0.1 substitutions per site. Numbers on major nodes represent ML bootstrap support. Bayesian posterior probabilities are summarized as asterisks for values equal to 1." figureDoi="http://doi.org/10.5281/zenodo.3534814" httpUri="https://zenodo.org/record/3534814/files/figure.png" pageId="6" pageNumber="373">Fig. 6</figureCitation>
), most nodes, including
<taxonomicName id="4C572DDAFFC6FFBAFE0AFEFAFD965871" baseAuthorityName="Larson &amp; Hoese" baseAuthorityYear="1980" box="[502,621,333,359]" class="Actinopterygii" family="Gobiidae" genus="Larsonella" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9238A4BFFC6FFBAFE0AFEFAFD965871" box="[502,621,333,359]" italics="true" pageId="6" pageNumber="373">L. pumilus</emphasis>
</taxonomicName>
, were supported by high bootstrap values (100%) and bayesian posterior probabilities (1), indicating that
<taxonomicName id="4C572DDAFFC6FFBAFDF8FEC6FD87589D" baseAuthorityName="Larson &amp; Hoese" baseAuthorityYear="1980" box="[516,636,369,395]" class="Actinopterygii" family="Gobiidae" genus="Larsonella" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9238A4BFFC6FFBAFDF8FEC6FD87589D" box="[516,636,369,395]" italics="true" pageId="6" pageNumber="373">L. pumilus</emphasis>
</taxonomicName>
was placed in a clade including
<taxonomicName id="4C572DDAFFC6FFBAFC12FEC5FBAE589D" box="[1006,1109,369,395]" class="Actinopterygii" family="Gobiidae" genus="Priolepis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="genus">
<emphasis id="B9238A4BFFC6FFBAFC12FEC5FBAE589D" box="[1006,1109,369,395]" italics="true" pageId="6" pageNumber="373">Priolepis</emphasis>
</taxonomicName>
spp. and
<taxonomicName id="4C572DDAFFC6FFBAFB3EFEC6FAE2589D" authorityName="Jordan &amp; Seale" authorityYear="1906" box="[1218,1305,370,395]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="genus">
<emphasis id="B9238A4BFFC6FFBAFB3EFEC6FAE2589D" box="[1218,1305,370,395]" italics="true" pageId="6" pageNumber="373">Trimma</emphasis>
</taxonomicName>
spp., while
<taxonomicName id="4C572DDAFFC6FFBAFF6BFE21FEFC58B9" box="[151,263,405,431]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="genus">
<emphasis id="B9238A4BFFC6FFBAFF6BFE21FEFC58B9" box="[151,263,405,431]" italics="true" pageId="6" pageNumber="373">Gobiodon</emphasis>
</taxonomicName>
spp. was placed in another clade with
<taxonomicName id="4C572DDAFFC6FFBAFD2BFE21FC9B58B9" authorityName="Bleeker" authorityYear="1874" box="[727,864,405,431]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="genus">
<emphasis id="B9238A4BFFC6FFBAFD2BFE21FC9B58B9" box="[727,864,405,431]" italics="true" pageId="6" pageNumber="373">Callogobius</emphasis>
</taxonomicName>
spp.,
<taxonomicName id="4C572DDAFFC6FFBAFC57FE21FBB258B9" authorityName="Smith" authorityYear="1949" box="[939,1097,405,431]" class="Actinopterygii" family="Gobiidae" genus="Vanderhorstia" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="genus">
<emphasis id="B9238A4BFFC6FFBAFC57FE21FBB258B9" box="[939,1097,405,431]" italics="true" pageId="6" pageNumber="373">Vanderhorstia</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="4C572DDAFFC6FFBAFB79FE21FF3858C5" class="Actinopterygii" family="Gobiidae" genus="Asterropteryx" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="species" species="semipunctata">
<emphasis id="B9238A4BFFC6FFBAFB79FE21FF3858C5" italics="true" pageId="6" pageNumber="373">Asterropteryx semipunctata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C572DDAFFC6FFBAFF2DFE0EFEE158C5" authorityName="Jenkins" authorityYear="1903" box="[209,282,442,467]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="genus">
<emphasis id="B9238A4BFFC6FFBAFF2DFE0EFEE158C5" box="[209,282,442,467]" italics="true" pageId="6" pageNumber="373">Eviota</emphasis>
</taxonomicName>
spp.,
<taxonomicName id="4C572DDAFFC6FFBAFEA0FE0DFDA658C5" baseAuthorityName="non Valenciennes" baseAuthorityYear="1837" box="[348,605,441,467]" class="Actinopterygii" family="Gobiidae" genus="Amblygobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="species" species="phalaena">
<emphasis id="B9238A4BFFC6FFBAFEA0FE0DFDA658C5" box="[348,605,441,467]" italics="true" pageId="6" pageNumber="373">Amblygobius phalaena</emphasis>
</taxonomicName>
,
<taxonomicName id="4C572DDAFFC6FFBAFD90FE0DFC7D58C5" baseAuthorityName="Lay &amp; Bennett" baseAuthorityYear="1839" box="[620,902,441,467]" class="Actinopterygii" family="Gobiidae" genus="Valenciennea" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="B9238A4BFFC6FFBAFD90FE0DFC7D58C5" box="[620,902,441,467]" italics="true" pageId="6" pageNumber="373">Valenciennea longipinnis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C572DDAFFC6FFBAFC3FFE0DFBAE58C5" authorityName="Regan" authorityYear="1912" box="[963,1109,441,467]" class="Actinopterygii" family="Microdesmidae" genus="Parioglossus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="genus">
<emphasis id="B9238A4BFFC6FFBAFC3FFE0DFBAE58C5" box="[963,1109,441,467]" italics="true" pageId="6" pageNumber="373">Parioglossus</emphasis>
</taxonomicName>
spp.
<taxonomicName id="4C572DDAFFC6FFBAFB6DFE0DFA9158C5" baseAuthorityName="Larson &amp; Hoese" baseAuthorityYear="1980" box="[1169,1386,441,467]" class="Actinopterygii" family="Gobiidae" genus="Larsonella" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9238A4BFFC6FFBAFB6DFE0DFA9158C5" box="[1169,1386,441,467]" italics="true" pageId="6" pageNumber="373">Larsonella pumilus</emphasis>
</taxonomicName>
was paired with
<taxonomicName id="4C572DDAFFC6FFBAFEE1FE69FE3458E1" baseAuthorityName="Regan" baseAuthorityYear="1908" box="[285,463,477,503]" class="Actinopterygii" family="Gobiidae" genus="Priolepis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="species" species="cincta">
<emphasis id="B9238A4BFFC6FFBAFEE1FE69FE3458E1" box="[285,463,477,503]" italics="true" pageId="6" pageNumber="373">Priolepis cincta</emphasis>
</taxonomicName>
, and they were placed within the
<taxonomicName id="4C572DDAFFC6FFBAFCB5FE69FC4B58E1" box="[841,944,477,503]" class="Actinopterygii" family="Gobiidae" genus="Priolepis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="373" phylum="Chordata" rank="genus">
<emphasis id="B9238A4BFFC6FFBAFCB5FE69FC4B58E1" box="[841,944,477,503]" italics="true" pageId="6" pageNumber="373">Priolepis</emphasis>
</taxonomicName>
lineage.
</paragraph>
</subSubSection>
</treatment>
</document>