347 lines
56 KiB
XML
347 lines
56 KiB
XML
<document id="A0342672086C797D29EB201CF20E4DE2" ID-DOI="10.11646/zootaxa.4756.1.1" ID-GBIF-Dataset="4d48d52e-6505-4f62-909f-5da5c2cfb489" ID-ISSN="1175-5334" ID-Zenodo-Dep="3736895" ID-ZooBank="urn:lsid:zoobank.org:pub:9FF62616-A7FA-4331-AC51-0F534400631D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1585767638433" checkinUser="carolina" docAuthor="Kluge, Nikita J." docDate="2020" docId="039787A6FFC9843E8CFBFF7FDA93FAAD" docLanguage="en" docName="zootaxa.4756.1.1.pdf.imf" docOrigin="Zootaxa 4756 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Thraulodes flavus Kluge, 2020, sp. n." docType="treatment" docVersion="7" lastPageNumber="91" masterDocId="FFAEFFDEFF9D84648C6CFFE6DB3EFFA2" masterDocTitle="Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species" masterLastPageNumber="142" masterPageNumber="1" pageNumber="85" updateTime="1698734461607" updateUser="plazi">
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<mods:title id="2F1A790A2225F063E4BAF7228A9F1B8F">Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species</mods:title>
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<mods:name id="FF6DFC16350408EE49A1A88AD04270EB" type="personal">
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<mods:namePart id="1E428FBB175955AD4352F3785801A446">Kluge, Nikita J.</mods:namePart>
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<mods:title id="44E8C0D5DDE20165FD30196E82A3A5BB">Zootaxa</mods:title>
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<mods:part id="DDC73A0F67677672A4177084E4D8147F">
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<mods:date id="D7123547FCA84873BA0BA8B0CE1E8F89">2020</mods:date>
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<mods:number id="53ABC48CE7B1AEF639D2D612FBCACEAE">2020-03-26</mods:number>
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<mods:number id="4BBEB5525061F9A5EF5F2C10D02C86EC">4756</mods:number>
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<mods:classification id="D7170E3E64C05E04DFF0D34443B09CEB">journal article</mods:classification>
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<mods:identifier id="5EAAA9D4C6CC5A400EED075AEA97C7AC" type="DOI">10.11646/zootaxa.4756.1.1</mods:identifier>
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<mods:identifier id="F35290450EFDADC5C64C32217E9FD5EC" type="GBIF-Dataset">4d48d52e-6505-4f62-909f-5da5c2cfb489</mods:identifier>
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<mods:identifier id="D17393C96DB37E33E3A3A94814489E8E" type="ISSN">1175-5334</mods:identifier>
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<treatment id="039787A6FFC9843E8CFBFF7FDA93FAAD" ID-DOI="http://doi.org/10.5281/zenodo.3811781" ID-GBIF-Taxon="163420782" ID-Zenodo-Dep="3811781" LSID="urn:lsid:plazi:treatment:039787A6FFC9843E8CFBFF7FDA93FAAD" httpUri="http://treatment.plazi.org/id/039787A6FFC9843E8CFBFF7FDA93FAAD" lastPageId="90" lastPageNumber="91" pageId="84" pageNumber="85">
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<subSubSection id="C324653BFFC984308CFBFF7FDAD8FF16" box="[151,486,153,180]" pageId="84" pageNumber="85" type="nomenclature">
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<paragraph id="8B8136B0FFC984308CFBFF7FDAD8FF16" blockId="84.[151,486,153,216]" box="[151,486,153,180]" pageId="84" pageNumber="85">
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<heading id="D0C981DCFFC984308CFBFF7FDAD8FF16" bold="true" box="[151,486,153,180]" fontSize="10" level="2" pageId="84" pageNumber="85" reason="2">
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<emphasis id="B94AEAA2FFC984308CFBFF7FDAD8FF16" bold="true" box="[151,486,153,180]" pageId="84" pageNumber="85">
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11.
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<taxonomicName id="4C3E4D33FFC984308CAEFF7FDAA2FF16" box="[194,412,153,180]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="flavus" status="sp. nov.">
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<emphasis id="B94AEAA2FFC984308CAEFF7FDAA2FF16" bold="true" box="[194,412,153,180]" italics="true" pageId="84" pageNumber="85">Thraulodes flavus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27957D9FFC984308DC8FF7CDAD8FF16" box="[420,486,154,180]" pageId="84" pageNumber="85" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C324653BFFC984308CFBFF5BDA7DFF7A" box="[151,323,189,216]" pageId="84" pageNumber="85" type="description">
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<paragraph id="8B8136B0FFC984308CFBFF5BDA7DFF7A" blockId="84.[151,486,153,216]" box="[151,323,189,216]" pageId="84" pageNumber="85">
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(
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<figureCitation id="13052A35FFC984308CF3FF5BDA02FF7A" box="[159,316,189,216]" captionStart-0="FIGURES 350–360" captionStart-1="FIGURES 361–364" captionStart-2="FIGURES 365–370" captionStart-3="FIGURES 371–376" captionStart-4="FIGURES 377–378" captionStartId-0="85.[151,264,1890,1915]" captionStartId-1="86.[151,264,1839,1864]" captionStartId-2="87.[151,264,1781,1806]" captionStartId-3="88.[151,264,1781,1806]" captionStartId-4="89.[151,264,1998,2023]" captionTargetBox-0="[169,1419,166,1871]" captionTargetBox-1="[151,1430,166,1820]" captionTargetBox-2="[151,1436,166,1762]" captionTargetBox-3="[157,1425,171,1752]" captionTargetBox-4="[151,1436,181,1972]" captionTargetId-0="figure@85.[169,1419,166,1871]" captionTargetId-1="figure@86.[151,1436,166,1820]" captionTargetId-2="figure@87.[151,1436,166,1762]" captionTargetId-3="figure@88.[151,1436,166,1761]" captionTargetId-4="figure@89.[151,1436,181,1972]" captionTargetPageId-0="85" captionTargetPageId-1="86" captionTargetPageId-2="87" captionTargetPageId-3="88" captionTargetPageId-4="89" captionText-0="FIGURES 350–360. Thraulodes flavus sp. n. 350–356, larval exuviae at the same magnification: 350, abdominal terga; 351– 353, parts of head with labrum and mandibles; 354, left half of pronotum and mesonotum; 355–356, fore and hind legs; 357, hind tibia with focus on outer-posterior row of stout setae; 358, the same with focus on outer-anterior and inner-anterior rows of stout setae; 359, anterior side of fore femur; 360, anterior side of fore tibia (350–356, 359–360, holotype)." captionText-1="FIGURES 361–364. Thraulodes flavus sp. n. (holotype). 361, tergalii; 362–363, distal segment of maxillary palp in ventral and dorsal view; 364, hind femur in anterior view." captionText-2="FIGURES 365–370. Thraulodes flavus sp. n. 365–366, male imago in lateral and dorsal view; 367, female imago; 368, left half of subimaginal exuviae of mesonotum; 369, subimaginal exuviae of fore leg of male; 370, genitalia of male imago (368–370, holotype)." captionText-3="FIGURES 371–376. Thraulodes flavus sp. n. 371, genitalia of male imago in lateral view; 372, apex of penis in dorsal view; 373, genitalia of male imago in ventral view; 374, their subimaginal exuviae; 375, larval protogonostyli and protopenis; 376, hind wing (372–376, holotype)." captionText-4="FIGURES 377–378. Thraulodes flavus sp. n., eggs." figureDoi-0="http://doi.org/10.5281/zenodo.3736993" figureDoi-1="http://doi.org/10.5281/zenodo.3736995" figureDoi-2="http://doi.org/10.5281/zenodo.3736997" figureDoi-3="http://doi.org/10.5281/zenodo.3736999" figureDoi-4="http://doi.org/10.5281/zenodo.3737001" httpUri-0="https://zenodo.org/record/3736993/files/figure.png" httpUri-1="https://zenodo.org/record/3736995/files/figure.png" httpUri-2="https://zenodo.org/record/3736997/files/figure.png" httpUri-3="https://zenodo.org/record/3736999/files/figure.png" httpUri-4="https://zenodo.org/record/3737001/files/figure.png" pageId="84" pageNumber="85">Figs 350–378</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C324653BFFC984308CFBFEE0D999FEBD" box="[151,679,261,288]" pageId="84" pageNumber="85" type="etymology">
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<paragraph id="8B8136B0FFC984308CFBFEE0D999FEBD" blockId="84.[151,1437,261,2016]" box="[151,679,261,288]" pageId="84" pageNumber="85">
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<emphasis id="B94AEAA2FFC984308CFBFEE0DA20FE82" bold="true" box="[151,286,262,288]" pageId="84" pageNumber="85">Etymology.</emphasis>
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Allusion to yellow color of imago.
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</paragraph>
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</subSubSection>
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<subSubSection id="C324653BFFC984308CABFECFDE49FE29" pageId="84" pageNumber="85" type="materials_examined">
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<paragraph id="8B8136B0FFC984308CABFECFDE49FE29" blockId="84.[151,1437,261,2016]" pageId="84" pageNumber="85">
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<emphasis id="B94AEAA2FFC984308CABFECFDA8DFEE6" bold="true" box="[199,435,297,324]" pageId="84" pageNumber="85">Material examined.</emphasis>
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<materialsCitation id="3B563CEDFFC984308DD7FECCDBCDFE29" ID-GBIF-Occurrence="2594509916" collectingDate="2018-01-27" collectionCode="XII" collectorName="N. Kluge & L. Sheyko." country="Panama" county="Bosque" latitude="8.793612" location="Altos del Valle" longLatPrecision="21" longitude="-82.193054" municipality="Palo Seco" pageId="84" pageNumber="85" specimenCount="1" specimenCount-male="1" stateProvince="Provincia de Bocas del Toro" typeStatus="holotype">
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||
<emphasis id="B94AEAA2FFC984308DD7FECCD90CFEE6" bold="true" box="[443,562,298,324]" pageId="84" pageNumber="85">
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||
<typeStatus id="54858812FFC984308DD7FECCD912FEE6" box="[443,556,298,324]" pageId="84" pageNumber="85" type="holotype">Holotype</typeStatus>
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||
:
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</emphasis>
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L-S-I
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<specimenCount id="9D38FD39FFC984308E16FECFD9B3FEE1" box="[634,653,297,323]" pageId="84" pageNumber="85" type="male">♂</specimenCount>
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{specimen [XII](10)2018},
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||
<collectingCountry id="F3297620FFC984308FA6FECCDF7AFEE6" box="[970,1092,298,324]" name="Panama" pageId="84" pageNumber="85">PANAMA</collectingCountry>
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||
,
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||
<collectingRegion id="49FAF852FFC98430883DFECFDEA6FEE6" box="[1105,1432,297,324]" country="Panama" name="Bocas del Toro" pageId="84" pageNumber="85">Provincia de Bocas del Toro</collectingRegion>
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||
,
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||
<collectingCounty id="62E04E3CFFC984308CFBFEA8DBD5FECA" box="[151,235,334,360]" pageId="84" pageNumber="85">Bosque</collectingCounty>
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||
protector
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||
<collectingMunicipality id="6BE5ACCAFFC984308D33FEABDAEFFEC5" box="[351,465,333,359]" pageId="84" pageNumber="85">Palo Seco</collectingMunicipality>
|
||
,
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||
<location id="8EE1606BFFC984308DB0FEABD9BBFEC5" LSID="urn:lsid:plazi:treatment:039787A6FFC9843E8CFBFF7FDA93FAAD:8EE1606BFFC984308DB0FEABD9BBFEC5" box="[476,645,333,359]" country="Panama" county="Bosque" latitude="8.793612" longLatPrecision="21" longitude="-82.193054" municipality="Palo Seco" name="Altos del Valle" pageId="84" pageNumber="85" stateProvince="Provincia de Bocas del Toro">Altos del Valle</location>
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||
,
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||
<location id="8EE1606BFFC984308EFDFEABD9C0FEC5" LSID="urn:lsid:plazi:treatment:039787A6FFC9843E8CFBFF7FDA93FAAD:8EE1606BFFC984308EFDFEABD9C0FEC5" box="[657,766,333,359]" country="Panama" county="Bosque" latitude="8.793612" longLatPrecision="21" longitude="-82.193054" municipality="Palo Seco" name="Rio Buris" pageId="84" pageNumber="85" stateProvince="Provincia de Bocas del Toro">Rio Buris</location>
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||
(
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||
<geoCoordinate id="EE0A5077FFC984308F62FEABD8B1FEC5" box="[782,911,333,359]" degrees="8" direction="north" minutes="47" orientation="latitude" pageId="84" pageNumber="85" precision="15" seconds="37" value="8.793612">8°47’37’’N</geoCoordinate>
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,
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||
<geoCoordinate id="EE0A5077FFC984308FF0FEABDF0FFECA" box="[924,1073,333,360]" degrees="82" direction="west" minutes="11" orientation="longitude" pageId="84" pageNumber="85" precision="15" seconds="35" value="-82.193054">82°11’35’’W</geoCoordinate>
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),
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<date id="FF801070FFC98430882AFEABDF8AFEC5" box="[1094,1204,333,359]" pageId="84" pageNumber="85" value="2018-01-27">
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||
<collectingDate id="EFC4E998FFC98430882AFEABDF8AFEC5" box="[1094,1204,333,359]" pageId="84" pageNumber="85" value="2018-01-27">27.I.2018</collectingDate>
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</date>
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, coll.
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<collectorName id="26CB5366FFC984308894FEA8DE61FEC5" box="[1272,1375,333,360]" pageId="84" pageNumber="85">N. Kluge</collectorName>
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&
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<collectorName id="26CB5366FFC9843089E8FEA8DBCDFE29" pageId="84" pageNumber="85">L. Sheyko.</collectorName>
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</materialsCitation>
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||
<materialsCitation id="3B563CEDFFC984308C95FE94DE49FE29" ID-GBIF-Occurrence="2594509932" box="[249,1399,369,396]" collectingDate="2012-01" collectingDateMax="2018-01-28" collectingDateMin="2012-01" collectionCode="XII" collectorName="N. Kluge & L. Sheyko." country="Panama" county="Bosque" latitude="8.793612" location="Altos del Valle" longLatPrecision="21" longitude="-82.193054" municipality="Palo Seco" pageId="84" pageNumber="85" specimenCount="8" specimenCount-female="2" specimenCount-larva="5" specimenCount-male="1" stateProvince="Provincia de Bocas del Toro" typeStatus="paratype">
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||
<emphasis id="B94AEAA2FFC984308C95FE94DA44FE2E" bold="true" box="[249,378,370,396]" pageId="84" pageNumber="85">
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||
<typeStatus id="54858812FFC984308C95FE94DA4BFE2E" box="[249,373,370,396]" pageId="84" pageNumber="85" type="paratype">Paratypes</typeStatus>
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||
:
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||
</emphasis>
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||
the same locality and collectors,
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||
<date id="FF801070FFC984308E9CFE97D8B7FE29" box="[752,905,369,395]" pageId="84" pageNumber="85" value="2018-01-24" valueMax="2018-01-28" valueMin="2018-01-24">
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||
<collectingDate id="EFC4E998FFC984308E9CFE97D8B7FE29" box="[752,905,369,395]" pageId="84" pageNumber="85" value="2018-01-24" valueMax="2018-01-28" valueMin="2018-01-24">24–28.I.2018</collectingDate>
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||
</date>
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||
:
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||
<date id="FF801070FFC984308FF8FE97D88DFE2E" box="[916,947,369,396]" pageId="84" pageNumber="85">
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||
<collectingDate id="EFC4E998FFC984308FF8FE97D88DFE2E" box="[916,947,369,396]" pageId="84" pageNumber="85">8 I</collectingDate>
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||
</date>
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||
<specimenCount id="9D38FD39FFC984308FDFFE94D8F6FE2E" box="[947,968,370,396]" pageId="84" pageNumber="85" type="male">♂</specimenCount>
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||
and
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||
<date id="FF801070FFC984308F92FE97DF13FE2E" box="[1022,1069,369,396]" pageId="84" pageNumber="85" value="2012-01">
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||
<collectingDate id="EFC4E998FFC984308F92FE97DF13FE2E" box="[1022,1069,369,396]" pageId="84" pageNumber="85" value="2012-01">12 I</collectingDate>
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||
</date>
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||
<specimenCount id="9D38FD39FFC984308842FE94DF7EFE2E" box="[1070,1088,370,396]" pageId="84" pageNumber="85" type="female">♀</specimenCount>
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||
(at light),
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||
<specimenCount id="9D38FD39FFC9843088DAFE97DFDEFE29" box="[1206,1248,369,395]" pageId="84" pageNumber="85" type="larva">1 L</specimenCount>
|
||
/S
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<specimenCount id="9D38FD39FFC984308897FE97DE35FE29" box="[1275,1291,369,395]" pageId="84" pageNumber="85" type="female">♀</specimenCount>
|
||
,
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||
<specimenCount id="9D38FD39FFC98430897AFE97DE4DFE29" box="[1302,1395,369,395]" pageId="84" pageNumber="85" type="larva">4 larvae</specimenCount>
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||
.
|
||
</materialsCitation>
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324653BFFC9843E8CABFE73DF3DFB95" lastPageId="90" lastPageNumber="91" pageId="84" pageNumber="85" type="description">
|
||
<paragraph id="8B8136B0FFC984308CABFE73DA5AFE12" blockId="84.[151,1437,261,2016]" box="[199,356,405,432]" pageId="84" pageNumber="85">
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||
<emphasis id="B94AEAA2FFC984308CABFE73DA5AFE12" bold="true" box="[199,356,405,432]" pageId="84" pageNumber="85">Descriptions.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC984308CABFE5CD814FDC1" blockId="84.[151,1437,261,2016]" pageId="84" pageNumber="85">
|
||
<emphasis id="B94AEAA2FFC984308CABFE5CDA2CFE76" bold="true" box="[199,274,442,468]" italics="true" pageId="84" pageNumber="85">Larva.</emphasis>
|
||
CUTICULAR COLORATION (
|
||
<figureCitation id="13052A35FFC984308EEDFE5FD81FFE76" box="[641,801,441,468]" captionStart="FIGURES 350–360" captionStartId="85.[151,264,1890,1915]" captionTargetBox="[169,1419,166,1871]" captionTargetId="figure@85.[169,1419,166,1871]" captionTargetPageId="85" captionText="FIGURES 350–360. Thraulodes flavus sp. n. 350–356, larval exuviae at the same magnification: 350, abdominal terga; 351– 353, parts of head with labrum and mandibles; 354, left half of pronotum and mesonotum; 355–356, fore and hind legs; 357, hind tibia with focus on outer-posterior row of stout setae; 358, the same with focus on outer-anterior and inner-anterior rows of stout setae; 359, anterior side of fore femur; 360, anterior side of fore tibia (350–356, 359–360, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736993" httpUri="https://zenodo.org/record/3736993/files/figure.png" pageId="84" pageNumber="85">Figs 350–356</figureCitation>
|
||
). Dorsal side of head, thorax and abdomen mostly brown with few blanks: pronotum bordered by light laterally, with or without other blanks; abdominal terga with blanks on lateral parts, area mediad of tergalii attachments nearly unicolor brown, terga VIII–IX with diffusive blank medi- ally. Fore protoptera with indistinct darkenings corresponding to dark maculae on imaginal wings. Femora mostly brown, with blanks; tibiae and tarsi uniformly light brown.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC984308CABFD8BDA79FD0E" blockId="84.[151,1437,261,2016]" pageId="84" pageNumber="85">
|
||
HYPODERMAL COLORATION. Abdomen without blackish hypodermal markings. Tergalii yellowish with gray (
|
||
<figureCitation id="13052A35FFC984308CB4FD77DA05FD0E" box="[216,315,657,684]" captionStart="FIGURES 361–364" captionStartId="86.[151,264,1839,1864]" captionTargetBox="[151,1430,166,1820]" captionTargetId="figure@86.[151,1436,166,1820]" captionTargetPageId="86" captionText="FIGURES 361–364. Thraulodes flavus sp. n. (holotype). 361, tergalii; 362–363, distal segment of maxillary palp in ventral and dorsal view; 364, hind femur in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3736995" httpUri="https://zenodo.org/record/3736995/files/figure.png" pageId="84" pageNumber="85">Fig. 361</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC984308CABFD53D90FFC9E" blockId="84.[151,1437,261,2016]" pageId="84" pageNumber="85">
|
||
SHAPE AND SETATION. Clypeus widened distally; labrum 1.3–1.45 times wider than clypeus (
|
||
<figureCitation id="13052A35FFC984308947FD53DEAEFD72" box="[1323,1424,693,720]" captionStart="FIGURES 350–360" captionStartId="85.[151,264,1890,1915]" captionTargetBox="[169,1419,166,1871]" captionTargetId="figure@85.[169,1419,166,1871]" captionTargetPageId="85" captionText="FIGURES 350–360. Thraulodes flavus sp. n. 350–356, larval exuviae at the same magnification: 350, abdominal terga; 351– 353, parts of head with labrum and mandibles; 354, left half of pronotum and mesonotum; 355–356, fore and hind legs; 357, hind tibia with focus on outer-posterior row of stout setae; 358, the same with focus on outer-anterior and inner-anterior rows of stout setae; 359, anterior side of fore femur; 360, anterior side of fore tibia (350–356, 359–360, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736993" httpUri="https://zenodo.org/record/3736993/files/figure.png" pageId="84" pageNumber="85">Fig. 351</figureCitation>
|
||
). Labrum widest at 0.4 length from base; initial fore margin (turned ventrally) without median emargination, with all 5 denticles wide; anterior transverse setal row regular (as in
|
||
<figureCitation id="13052A35FFC984308F54FD1BD8B4FCB5" box="[824,906,765,791]" captionStart="FIGURES 86–93" captionStartId="37.[151,264,188,213]" captionTargetId="figure@36.[151,1436,342,2018]" captionTargetPageId="36" captionText="FIGURES 86–93. Thraulodes telegraphicus. 86, median longitudinal section of labrum in usual condition; 87, the same, in spread condition; 88, labrum in spread condition in dorsal view (setal rows shown as dotted lines); 89, transverse section of hind tibia; 90–93, tergalii I, III, VI and VII. Abbreviations: anter.r, anterior transverse row of setae; dent., five denticles separated by six sensilla; poster.r, posterior transverse row of setae; i.f, inner field of stout pointed setae; i-a.h, inner-anterior recurved hairs; i-a.r, inner-anterior row of stout setae; o-a.r, outer-anterior row of stout setae; o.h, outer hairs; o-p.r, outer-posterior row of stout setae; p.r, row of posterior pointed pectinate setae." figureDoi="http://doi.org/10.5281/zenodo.3736921" httpUri="https://zenodo.org/record/3736921/files/figure.png" pageId="84" pageNumber="85">Fig. 88</figureCitation>
|
||
), as wide as all 5 denticles. Maxilla with 16–17 pectinate setae in apical-ventral row.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC984308CABFCA0D967FC26" blockId="84.[151,1437,261,2016]" pageId="84" pageNumber="85">
|
||
Femora: Stout setae on anterior surface narrowed distally and blunt (
|
||
<figureCitation id="13052A35FFC984308FAEFCA3DF1AFCC2" box="[962,1060,837,864]" captionStart="FIGURES 350–360" captionStartId="85.[151,264,1890,1915]" captionTargetBox="[169,1419,166,1871]" captionTargetId="figure@85.[169,1419,166,1871]" captionTargetPageId="85" captionText="FIGURES 350–360. Thraulodes flavus sp. n. 350–356, larval exuviae at the same magnification: 350, abdominal terga; 351– 353, parts of head with labrum and mandibles; 354, left half of pronotum and mesonotum; 355–356, fore and hind legs; 357, hind tibia with focus on outer-posterior row of stout setae; 358, the same with focus on outer-anterior and inner-anterior rows of stout setae; 359, anterior side of fore femur; 360, anterior side of fore tibia (350–356, 359–360, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736993" httpUri="https://zenodo.org/record/3736993/files/figure.png" pageId="84" pageNumber="85">Fig. 359</figureCitation>
|
||
). Irregular row of hairs near inner margin present on all femora (
|
||
<figureCitation id="13052A35FFC984308D85FC8FD972FC26" box="[489,588,873,900]" captionStart="FIGURES 361–364" captionStartId="86.[151,264,1839,1864]" captionTargetBox="[151,1430,166,1820]" captionTargetId="figure@86.[151,1436,166,1820]" captionTargetPageId="86" captionText="FIGURES 361–364. Thraulodes flavus sp. n. (holotype). 361, tergalii; 362–363, distal segment of maxillary palp in ventral and dorsal view; 364, hind femur in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3736995" httpUri="https://zenodo.org/record/3736995/files/figure.png" pageId="84" pageNumber="85">Fig. 364</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC984308CABFC68DF36FBB1" blockId="84.[151,1437,261,2016]" pageId="84" pageNumber="85">
|
||
Fore tibia (
|
||
<figureCitation id="13052A35FFC984308D2AFC6BDA95FC0A" box="[326,427,909,936]" captionStart="FIGURES 350–360" captionStartId="85.[151,264,1890,1915]" captionTargetBox="[169,1419,166,1871]" captionTargetId="figure@85.[169,1419,166,1871]" captionTargetPageId="85" captionText="FIGURES 350–360. Thraulodes flavus sp. n. 350–356, larval exuviae at the same magnification: 350, abdominal terga; 351– 353, parts of head with labrum and mandibles; 354, left half of pronotum and mesonotum; 355–356, fore and hind legs; 357, hind tibia with focus on outer-posterior row of stout setae; 358, the same with focus on outer-anterior and inner-anterior rows of stout setae; 359, anterior side of fore femur; 360, anterior side of fore tibia (350–356, 359–360, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736993" httpUri="https://zenodo.org/record/3736993/files/figure.png" pageId="84" pageNumber="85">Fig. 360</figureCitation>
|
||
): outer hairs form two irregular rows; inner-anterior row of recurved hairs absent; inneranterior row of stout setae represented by few (4–7) stout setae in proximal part of tibia; inner field of stout pointed setae sparse, denser on middle 1/3 of tibia, consists of pointed setae of subequal length, among which bipectinate setae form 1 row, and non-pectinate setae form 2 irregular rows posteriad of it.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC984308CABFBFBD97FFB01" blockId="84.[151,1437,261,2016]" pageId="84" pageNumber="85">
|
||
Hind tibia (
|
||
<figureCitation id="13052A35FFC984308D27FBFBDAD7FB9A" box="[331,489,1053,1080]" captionStart="FIGURES 350–360" captionStartId="85.[151,264,1890,1915]" captionTargetBox="[169,1419,166,1871]" captionTargetId="figure@85.[169,1419,166,1871]" captionTargetPageId="85" captionText="FIGURES 350–360. Thraulodes flavus sp. n. 350–356, larval exuviae at the same magnification: 350, abdominal terga; 351– 353, parts of head with labrum and mandibles; 354, left half of pronotum and mesonotum; 355–356, fore and hind legs; 357, hind tibia with focus on outer-posterior row of stout setae; 358, the same with focus on outer-anterior and inner-anterior rows of stout setae; 359, anterior side of fore femur; 360, anterior side of fore tibia (350–356, 359–360, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736993" httpUri="https://zenodo.org/record/3736993/files/figure.png" pageId="84" pageNumber="85">Fig. 357–358</figureCitation>
|
||
): both outer-anterior and outer-posterior rows of stout setae consist of more or less pointed stout setae of various lengths; hairs located between these rows, numerous and form more than one row (besides row of hairs posteriad of outer-posterior row of stout setae); stout setae of inner-anterior row moderately long, narrowed distally and truncated.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC984308CABFB4BDE23FB65" blockId="84.[151,1437,261,2016]" box="[199,1309,1197,1224]" pageId="84" pageNumber="85">Claws with 6–7 denticles on rigid portion, with or without minute denticles on articulatory portion.</paragraph>
|
||
<paragraph id="8B8136B0FFC984308CABFB37DF0FFB49" blockId="84.[151,1437,261,2016]" box="[199,1073,1233,1260]" pageId="84" pageNumber="85">
|
||
Tergalii (
|
||
<figureCitation id="13052A35FFC984308D40FB37DAACFB4E" box="[300,402,1233,1260]" captionStart="FIGURES 350–360" captionStartId="85.[151,264,1890,1915]" captionTargetBox="[169,1419,166,1871]" captionTargetId="figure@85.[169,1419,166,1871]" captionTargetPageId="85" captionText="FIGURES 350–360. Thraulodes flavus sp. n. 350–356, larval exuviae at the same magnification: 350, abdominal terga; 351– 353, parts of head with labrum and mandibles; 354, left half of pronotum and mesonotum; 355–356, fore and hind legs; 357, hind tibia with focus on outer-posterior row of stout setae; 358, the same with focus on outer-anterior and inner-anterior rows of stout setae; 359, anterior side of fore femur; 360, anterior side of fore tibia (350–356, 359–360, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736993" httpUri="https://zenodo.org/record/3736993/files/figure.png" pageId="84" pageNumber="85">Figs 350</figureCitation>
|
||
,
|
||
<figureCitation id="13052A35FFC984308DF1FB37DAF5FB4E" box="[413,459,1233,1260]" captionStart="FIGURES 361–364" captionStartId="86.[151,264,1839,1864]" captionTargetBox="[151,1430,166,1820]" captionTargetId="figure@86.[151,1436,166,1820]" captionTargetPageId="86" captionText="FIGURES 361–364. Thraulodes flavus sp. n. (holotype). 361, tergalii; 362–363, distal segment of maxillary palp in ventral and dorsal view; 364, hind femur in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3736995" httpUri="https://zenodo.org/record/3736995/files/figure.png" pageId="84" pageNumber="85">361</figureCitation>
|
||
): each lamella slender, tracheae without side branches.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC984308CABFB13DE5EFA91" blockId="84.[151,1437,261,2016]" pageId="84" pageNumber="85">
|
||
Male genitalia in last larval instar (
|
||
<figureCitation id="13052A35FFC984308E20FB13D991FAB2" box="[588,687,1269,1296]" captionStart="FIGURES 371–376" captionStartId="88.[151,264,1781,1806]" captionTargetBox="[157,1425,171,1752]" captionTargetId="figure@88.[151,1436,166,1761]" captionTargetPageId="88" captionText="FIGURES 371–376. Thraulodes flavus sp. n. 371, genitalia of male imago in lateral view; 372, apex of penis in dorsal view; 373, genitalia of male imago in ventral view; 374, their subimaginal exuviae; 375, larval protogonostyli and protopenis; 376, hind wing (372–376, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736999" httpUri="https://zenodo.org/record/3736999/files/figure.png" pageId="84" pageNumber="85">Fig. 375</figureCitation>
|
||
): protogonostyli separated one from another by deep emargination. Each protopenis lobe with gonopore-bearing process located near median margin; gonopore opened caudally.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC984308CABFADBDA4CF98D" blockId="84.[151,1437,261,2016]" pageId="84" pageNumber="85">
|
||
<emphasis id="B94AEAA2FFC984308CABFADBDA02FAFA" bold="true" box="[199,316,1341,1368]" italics="true" pageId="84" pageNumber="85">Subimago</emphasis>
|
||
(
|
||
<figureCitation id="13052A35FFC984308D26FADBDAD8FAF5" box="[330,486,1341,1367]" captionStart="FIGURES 365–370" captionStartId="87.[151,264,1781,1806]" captionTargetBox="[151,1436,166,1762]" captionTargetId="figure@87.[151,1436,166,1762]" captionTargetPageId="87" captionText="FIGURES 365–370. Thraulodes flavus sp. n. 365–366, male imago in lateral and dorsal view; 367, female imago; 368, left half of subimaginal exuviae of mesonotum; 369, subimaginal exuviae of fore leg of male; 370, genitalia of male imago (368–370, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736997" httpUri="https://zenodo.org/record/3736997/files/figure.png" pageId="84" pageNumber="85">Figs 368, 369</figureCitation>
|
||
,
|
||
<figureCitation id="13052A35FFC984308D9CFADBD920FAF5" box="[496,542,1341,1367]" captionStart="FIGURES 371–376" captionStartId="88.[151,264,1781,1806]" captionTargetBox="[157,1425,171,1752]" captionTargetId="figure@88.[151,1436,166,1761]" captionTargetPageId="88" captionText="FIGURES 371–376. Thraulodes flavus sp. n. 371, genitalia of male imago in lateral view; 372, apex of penis in dorsal view; 373, genitalia of male imago in ventral view; 374, their subimaginal exuviae; 375, larval protogonostyli and protopenis; 376, hind wing (372–376, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736999" httpUri="https://zenodo.org/record/3736999/files/figure.png" pageId="84" pageNumber="85">374</figureCitation>
|
||
). CUTICULAR COLORATION. Cuticle mostly colorless. Flagellum of antenna brownish. Pronotum tinged with light brownish laterally. Mesonotum with antelateroparapsidal suture contrastingly colored with dark brown, some other sutures shed with brownish; chromozones as colorless as achromozones (
|
||
<figureCitation id="13052A35FFC984308903FA63DBFBFA66" captionStart="FIGURES 365–370" captionStartId="87.[151,264,1781,1806]" captionTargetBox="[151,1436,166,1762]" captionTargetId="figure@87.[151,1436,166,1762]" captionTargetPageId="87" captionText="FIGURES 365–370. Thraulodes flavus sp. n. 365–366, male imago in lateral and dorsal view; 367, female imago; 368, left half of subimaginal exuviae of mesonotum; 369, subimaginal exuviae of fore leg of male; 370, genitalia of male imago (368–370, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736997" httpUri="https://zenodo.org/record/3736997/files/figure.png" pageId="84" pageNumber="85">Fig. 368</figureCitation>
|
||
). On fore leg femur colored with dark brown on outer side and close to apex, tibia brown at very base and light brownish apically, tarsal segments 2–4 light brownish (
|
||
<figureCitation id="13052A35FFC984308E92FA2BD85EFA4A" box="[766,864,1485,1512]" captionStart="FIGURES 365–370" captionStartId="87.[151,264,1781,1806]" captionTargetBox="[151,1436,166,1762]" captionTargetId="figure@87.[151,1436,166,1762]" captionTargetPageId="87" captionText="FIGURES 365–370. Thraulodes flavus sp. n. 365–366, male imago in lateral and dorsal view; 367, female imago; 368, left half of subimaginal exuviae of mesonotum; 369, subimaginal exuviae of fore leg of male; 370, genitalia of male imago (368–370, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736997" httpUri="https://zenodo.org/record/3736997/files/figure.png" pageId="84" pageNumber="85">Fig. 369</figureCitation>
|
||
). Middle and hind legs with tibiae and tarsi slightly colored with ocher. Cuticle of wings colorless, microtrichia light brownish. Other cuticle of body, legs, caudalii and gonostyli colorless.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC984308CABF9DCDA1CF939" blockId="84.[151,1437,261,2016]" pageId="84" pageNumber="85">
|
||
TEXTURE. On tarsi of all legs, 1st tarsomere with microtrichia (as tibia), 2nd–5th tarsomeres coved by blunt microlepides; pointed microlepides present near apical margins of 2nd–4th tarsomeres of middle and hind legs (as in
|
||
<figureCitation id="13052A35FFC984308CD8F967DA2BF939" box="[180,277,1665,1691]" captionStart="FIGURES 464–473" captionStartId="106.[151,264,188,213]" captionTargetId="figure@105.[151,1436,181,1987]" captionTargetPageId="105" captionText="FIGURES 464–473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus." figureDoi="http://doi.org/10.5281/zenodo.3737024" httpUri="https://zenodo.org/record/3737024/files/figure.png" pageId="84" pageNumber="85">Fig. 473</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC984308CABF943DFD8F91D" blockId="84.[151,1437,261,2016]" box="[199,1254,1701,1728]" pageId="84" pageNumber="85">
|
||
<emphasis id="B94AEAA2FFC984308CABF943DA6FF91D" bold="true" box="[199,337,1701,1728]" italics="true" pageId="84" pageNumber="85">Male imago</emphasis>
|
||
(
|
||
<figureCitation id="13052A35FFC984308D0CF943DAC1F962" box="[352,511,1701,1728]" captionStart="FIGURES 365–370" captionStartId="87.[151,264,1781,1806]" captionTargetBox="[151,1436,166,1762]" captionTargetId="figure@87.[151,1436,166,1762]" captionTargetPageId="87" captionText="FIGURES 365–370. Thraulodes flavus sp. n. 365–366, male imago in lateral and dorsal view; 367, female imago; 368, left half of subimaginal exuviae of mesonotum; 369, subimaginal exuviae of fore leg of male; 370, genitalia of male imago (368–370, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736997" httpUri="https://zenodo.org/record/3736997/files/figure.png" pageId="84" pageNumber="85">Figs 365–366</figureCitation>
|
||
). Coloration yellow with orange, with contrasting black markings.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC984308CABF92FDAD7F889" blockId="84.[151,1437,261,2016]" pageId="84" pageNumber="85">Head yellow with black, antennae entirely yellow. Dorsal eyes contiguous medially, dull orange. Pronotum yellow with pair of large black maculae laterally and narrower black line medially. Mesothorax and metathorax yellow, with black maculae on pleura.</paragraph>
|
||
<paragraph id="8B8136B0FFC9843E8CABF8D3DA66FEE1" blockId="84.[151,1437,261,2016]" lastBlockId="90.[151,1437,153,1295]" lastPageId="90" lastPageNumber="91" pageId="84" pageNumber="85">
|
||
Cuticle of legs mostly colorless; on fore leg, apex of femur and base of tibia with cuticle light brown. Legs with following hypodermal pigmentation on ocher background: Fore femur orange with dark brown apex; posterior side with longitudinal blackish-brown macula at middle; inner side with blackish-brown macula near base and blackish-brown stripe on distal 2/3. Fore tibia orange with black apex. Fore tarsus with middle areas of 2nd-4th segments brown. Middle and hind femora with 2 wide, diffusive, orange bands, one near base and another at apex; proximal band with longitudinal blackish-brown macula on posterior side near middle of femur; apical band with longitudinal blackish-brown stripe on posterior-inner margin. Middle and hind tibiae with small brown apical spot; tarsus with orange spots on 2nd–5th segments. In
|
||
<typeStatus id="54858812FFC7843E8E8EFF07D879FF59" box="[738,839,225,251]" pageId="90" pageNumber="91" type="holotype">holotype</typeStatus>
|
||
, relation of fore femur to fore wing length 85:355; proportions femur/tibia/tarsomeres on fore leg 85:120:3:38:30:20:10; on middle leg 82:108:3:6:5:4:9; on hind leg 95:115:3:6:5:4:9.
|
||
</paragraph>
|
||
<caption id="DF416638FFC884318CFBF884DFF5F845" ID-DOI="http://doi.org/10.5281/zenodo.3736993" ID-Zenodo-Dep="3736993" httpUri="https://zenodo.org/record/3736993/files/figure.png" pageId="85" pageNumber="86" startId="85.[151,264,1890,1915]" targetBox="[169,1419,166,1871]" targetPageId="85">
|
||
<paragraph id="8B8136B0FFC884318CFBF884DFF5F845" blockId="85.[151,1437,1890,2023]" pageId="85" pageNumber="86">
|
||
<emphasis id="B94AEAA2FFC884318CFBF884DA53F8D9" bold="true" box="[151,365,1890,1915]" pageId="85" pageNumber="86">FIGURES 350–360.</emphasis>
|
||
<taxonomicName id="4C3E4D33FFC884318D19F885D912F8D9" box="[373,556,1891,1915]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="flavus" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFC884318D19F885D912F8D9" box="[373,556,1891,1915]" italics="true" pageId="85" pageNumber="86">Thraulodes flavus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFC884318E5FF885D952F8D9" bold="true" box="[563,620,1891,1915]" pageId="85" pageNumber="86">
|
||
<taxonomicNameLabel id="A27957D9FFC884318E5FF885D952F8D9" box="[563,620,1891,1915]" pageId="85" pageNumber="86" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
350–356, larval exuviae at the same magnification: 350, abdominal terga; 351– 353, parts of head with labrum and mandibles; 354, left half of pronotum and mesonotum; 355–356, fore and hind legs; 357, hind tibia with focus on outer-posterior row of stout setae; 358, the same with focus on outer-anterior and inner-anterior rows of stout setae; 359, anterior side of fore femur; 360, anterior side of fore tibia (350–356, 359–360, holotype).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF416638FFCB84328CFBF8C9D957F8CE" ID-DOI="http://doi.org/10.5281/zenodo.3736995" ID-Zenodo-Dep="3736995" httpUri="https://zenodo.org/record/3736995/files/figure.png" pageId="86" pageNumber="87" startId="86.[151,264,1839,1864]" targetBox="[151,1430,166,1820]" targetPageId="86">
|
||
<paragraph id="8B8136B0FFCB84328CFBF8C9D957F8CE" blockId="86.[151,1437,1839,1900]" pageId="86" pageNumber="87">
|
||
<emphasis id="B94AEAA2FFCB84328CFBF8C9DA53F8EA" bold="true" box="[151,365,1839,1864]" pageId="86" pageNumber="87">FIGURES 361–364.</emphasis>
|
||
<taxonomicName id="4C3E4D33FFCB84328D1EF8D6D916F8EA" box="[370,552,1840,1864]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="flavus" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFCB84328D1EF8D6D916F8EA" box="[370,552,1840,1864]" italics="true" pageId="86" pageNumber="87">Thraulodes flavus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFCB84328E41F8D6D95BF8EA" bold="true" box="[557,613,1840,1864]" pageId="86" pageNumber="87">
|
||
<taxonomicNameLabel id="A27957D9FFCB84328E41F8D6D95BF8EA" box="[557,613,1840,1864]" pageId="86" pageNumber="87" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype). 361, tergalii; 362–363, distal segment of maxillary palp in ventral and dorsal view; 364, hind femur in anterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF416638FFCA84338CFBF913DA3EF8F4" ID-DOI="http://doi.org/10.5281/zenodo.3736997" ID-Zenodo-Dep="3736997" httpUri="https://zenodo.org/record/3736997/files/figure.png" pageId="87" pageNumber="88" startId="87.[151,264,1781,1806]" targetBox="[151,1436,166,1762]" targetPageId="87">
|
||
<paragraph id="8B8136B0FFCA84338CFBF913DA3EF8F4" blockId="87.[151,1436,1781,1878]" pageId="87" pageNumber="88">
|
||
<emphasis id="B94AEAA2FFCA84338CFBF913DA52F8AC" bold="true" box="[151,364,1781,1806]" pageId="87" pageNumber="88">FIGURES 365–370.</emphasis>
|
||
<taxonomicName id="4C3E4D33FFCA84338D1EF910D916F8AC" box="[370,552,1782,1806]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="flavus" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFCA84338D1EF910D916F8AC" box="[370,552,1782,1806]" italics="true" pageId="87" pageNumber="88">Thraulodes flavus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFCA84338E41F910D95BF8AC" bold="true" box="[557,613,1782,1806]" pageId="87" pageNumber="88">
|
||
<taxonomicNameLabel id="A27957D9FFCA84338E41F910D95BF8AC" box="[557,613,1782,1806]" pageId="87" pageNumber="88" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
365–366, male imago in lateral and dorsal view; 367, female imago; 368, left half of subimaginal exuviae of mesonotum; 369, subimaginal exuviae of fore leg of male; 370, genitalia of male imago (368–370, holotype).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF416638FFC5843C8CFBF913DAE4F8F4" ID-DOI="http://doi.org/10.5281/zenodo.3736999" ID-Zenodo-Dep="3736999" httpUri="https://zenodo.org/record/3736999/files/figure.png" pageId="88" pageNumber="89" startId="88.[151,264,1781,1806]" targetBox="[157,1425,171,1752]" targetPageId="88">
|
||
<paragraph id="8B8136B0FFC5843C8CFBF913DAE4F8F4" blockId="88.[151,1436,1781,1878]" pageId="88" pageNumber="89">
|
||
<emphasis id="B94AEAA2FFC5843C8CFBF913DA50F8AC" bold="true" box="[151,366,1781,1806]" pageId="88" pageNumber="89">FIGURES 371–376.</emphasis>
|
||
<taxonomicName id="4C3E4D33FFC5843C8D19F913D912F8AF" box="[373,556,1781,1805]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="flavus" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFC5843C8D19F913D912F8AF" box="[373,556,1781,1805]" italics="true" pageId="88" pageNumber="89">Thraulodes flavus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFC5843C8E5FF913D953F8AF" bold="true" box="[563,621,1781,1805]" pageId="88" pageNumber="89">
|
||
<taxonomicNameLabel id="A27957D9FFC5843C8E5FF913D953F8AF" box="[563,621,1781,1805]" pageId="88" pageNumber="89" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
371, genitalia of male imago in lateral view; 372, apex of penis in dorsal view; 373, genitalia of male imago in ventral view; 374, their subimaginal exuviae; 375, larval protogonostyli and protopenis; 376, hind wing (372–376, holotype).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF416638FFC4843D8CFBF828D995F845" ID-DOI="http://doi.org/10.5281/zenodo.3737001" ID-Zenodo-Dep="3737001" box="[151,683,1998,2023]" httpUri="https://zenodo.org/record/3737001/files/figure.png" pageId="89" pageNumber="90" startId="89.[151,264,1998,2023]" targetBox="[151,1436,181,1972]" targetPageId="89">
|
||
<paragraph id="8B8136B0FFC4843D8CFBF828D995F845" blockId="89.[151,683,1998,2023]" box="[151,683,1998,2023]" pageId="89" pageNumber="90">
|
||
<emphasis id="B94AEAA2FFC4843D8CFBF828DA53F845" bold="true" box="[151,365,1998,2023]" pageId="89" pageNumber="90">FIGURES 377–378.</emphasis>
|
||
<taxonomicName id="4C3E4D33FFC4843D8D1FF828D914F844" box="[371,554,1998,2022]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="flavus" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFC4843D8D1FF828D914F844" box="[371,554,1998,2022]" italics="true" pageId="89" pageNumber="90">Thraulodes flavus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFC4843D8E5CF829D957F845" bold="true" box="[560,617,1999,2023]" pageId="89" pageNumber="90">
|
||
<taxonomicNameLabel id="A27957D9FFC4843D8E5CF829D957F845" box="[560,617,1999,2023]" pageId="89" pageNumber="90" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, eggs.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8136B0FFC7843E8CABFEABDBDBFD9D" blockId="90.[151,1437,153,1295]" pageId="90" pageNumber="91">
|
||
Fore wing with oblique cross veins in costal field, both proximad and distad of bulla; pterostigmatic cross veins dense and partly anastomosed. Costal field yellow, with cross veins proximad and posteriad of bulla bordered by brown. Subcostal field orange with apex brown. Orange or brownish transverse stripe posteriad of costal brace. Cross vein near bulla and furcation of MA bordered by brown. Longitudinal veins C, Sc and RA orange, distal part of RA brown; other longitudinal veins ocher. Cross veins posteriad of RA brown. Hind wing without coloration (
|
||
<figureCitation id="13052A35FFC7843E8903FE3BDBFBFDBE" captionStart="FIGURES 371–376" captionStartId="88.[151,264,1781,1806]" captionTargetBox="[157,1425,171,1752]" captionTargetId="figure@88.[151,1436,166,1761]" captionTargetPageId="88" captionText="FIGURES 371–376. Thraulodes flavus sp. n. 371, genitalia of male imago in lateral view; 372, apex of penis in dorsal view; 373, genitalia of male imago in ventral view; 374, their subimaginal exuviae; 375, larval protogonostyli and protopenis; 376, hind wing (372–376, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736999" httpUri="https://zenodo.org/record/3736999/files/figure.png" pageId="90" pageNumber="91">Fig. 376</figureCitation>
|
||
). Coloration of fore wing partly appears during larval development, being in subimago less contrasting than in imago.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC7843E8CABFDAFDEAAFD09" blockId="90.[151,1437,153,1295]" pageId="90" pageNumber="91">Abdomen whitish-yellowish, with diffusive narrow grayish bands on posterior margins of terga I–X, with oblique lateral orange streaks on terga II–VI; terga VII and IX–X mostly orange; stigmatic spots absent. Caudalii ocher, each segment proximally dark (brown or orange); segments with longer and shorter dark portions alternate.</paragraph>
|
||
<paragraph id="8B8136B0FFC7843E8CABFD53DA81FCFD" blockId="90.[151,1437,153,1295]" pageId="90" pageNumber="91">
|
||
Genitalia (
|
||
<figureCitation id="13052A35FFC7843E8D2DFD53DADBFD6D" box="[321,485,693,719]" captionStart-0="FIGURES 365–370" captionStart-1="FIGURES 371–376" captionStartId-0="87.[151,264,1781,1806]" captionStartId-1="88.[151,264,1781,1806]" captionTargetBox-0="[151,1436,166,1762]" captionTargetBox-1="[157,1425,171,1752]" captionTargetId-0="figure@87.[151,1436,166,1762]" captionTargetId-1="figure@88.[151,1436,166,1761]" captionTargetPageId-0="87" captionTargetPageId-1="88" captionText-0="FIGURES 365–370. Thraulodes flavus sp. n. 365–366, male imago in lateral and dorsal view; 367, female imago; 368, left half of subimaginal exuviae of mesonotum; 369, subimaginal exuviae of fore leg of male; 370, genitalia of male imago (368–370, holotype)." captionText-1="FIGURES 371–376. Thraulodes flavus sp. n. 371, genitalia of male imago in lateral view; 372, apex of penis in dorsal view; 373, genitalia of male imago in ventral view; 374, their subimaginal exuviae; 375, larval protogonostyli and protopenis; 376, hind wing (372–376, holotype)." figureDoi-0="http://doi.org/10.5281/zenodo.3736997" figureDoi-1="http://doi.org/10.5281/zenodo.3736999" httpUri-0="https://zenodo.org/record/3736997/files/figure.png" httpUri-1="https://zenodo.org/record/3736999/files/figure.png" pageId="90" pageNumber="91">Figs 370–374</figureCitation>
|
||
): Styliger, gonostyli and penis light ocher. Dorsal extension of styliger small and prominent. Penis lobes widely divergent; each lobe gradually widened apically, without lateral pouch. Apico-lateral area forming well-expressed ear; ventro-apical plate moderately developed, hiding base of spear from ventral view. Telopenes in form of spear-like rolls, directed medially-caudally-ventrally, nearly straight, with groove opened dorsal-laterally (
|
||
<figureCitation id="13052A35FFC7843E8D3CFCA3DA8CFCFD" box="[336,434,837,863]" captionStart="FIGURES 371–376" captionStartId="88.[151,264,1781,1806]" captionTargetBox="[157,1425,171,1752]" captionTargetId="figure@88.[151,1436,166,1761]" captionTargetPageId="88" captionText="FIGURES 371–376. Thraulodes flavus sp. n. 371, genitalia of male imago in lateral view; 372, apex of penis in dorsal view; 373, genitalia of male imago in ventral view; 374, their subimaginal exuviae; 375, larval protogonostyli and protopenis; 376, hind wing (372–376, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736999" httpUri="https://zenodo.org/record/3736999/files/figure.png" pageId="90" pageNumber="91">Fig. 372</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC7843E8CABFC8EDF5EFC69" blockId="90.[151,1437,153,1295]" pageId="90" pageNumber="91">
|
||
<emphasis id="B94AEAA2FFC7843E8CABFC8EDA50FC21" bold="true" box="[199,366,872,899]" italics="true" pageId="90" pageNumber="91">Female imago</emphasis>
|
||
(
|
||
<figureCitation id="13052A35FFC7843E8D11FC8FDADEFC26" box="[381,480,873,900]" captionStart="FIGURES 365–370" captionStartId="87.[151,264,1781,1806]" captionTargetBox="[151,1436,166,1762]" captionTargetId="figure@87.[151,1436,166,1762]" captionTargetPageId="87" captionText="FIGURES 365–370. Thraulodes flavus sp. n. 365–366, male imago in lateral and dorsal view; 367, female imago; 368, left half of subimaginal exuviae of mesonotum; 369, subimaginal exuviae of fore leg of male; 370, genitalia of male imago (368–370, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736997" httpUri="https://zenodo.org/record/3736997/files/figure.png" pageId="90" pageNumber="91">Fig. 367</figureCitation>
|
||
). Coloration of head, thorax, wings, legs and caudalii as in male. Dorsal side of head varies from entirely yellow to yellow with wide black transverse band connecting eyes and all three ocelli. Coloration of abdomen similar to that of male, but with tergum VII not colored with orange.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC7843E8CABFC33DF66FBB1" blockId="90.[151,1437,153,1295]" pageId="90" pageNumber="91">
|
||
<emphasis id="B94AEAA2FFC7843E8CABFC33DA3FFC4D" bold="true" box="[199,257,981,1007]" italics="true" pageId="90" pageNumber="91">Eggs</emphasis>
|
||
(
|
||
<figureCitation id="13052A35FFC7843E8D63FC33DA91FC4D" box="[271,431,981,1007]" captionStart="FIGURES 377–378" captionStartId="89.[151,264,1998,2023]" captionTargetBox="[151,1436,181,1972]" captionTargetId="figure@89.[151,1436,181,1972]" captionTargetPageId="89" captionText="FIGURES 377–378. Thraulodes flavus sp. n., eggs." figureDoi="http://doi.org/10.5281/zenodo.3737001" httpUri="https://zenodo.org/record/3737001/files/figure.png" pageId="90" pageNumber="91">Figs 377–378</figureCitation>
|
||
). Mostly barrel-shaped. Each KTC surrounded by ring-like cover. Chorion between KTCs with rugose ridges forming hexagonal or pentagonal cells, with one KTC in each cell.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFC7843E8CABFBFADF3DFB95" blockId="90.[151,1437,153,1295]" box="[199,1027,1052,1080]" pageId="90" pageNumber="91">
|
||
<emphasis id="B94AEAA2FFC7843E8CABFBFADA70FB95" bold="true" box="[199,334,1052,1079]" pageId="90" pageNumber="91">Dimension.</emphasis>
|
||
Fore wing length (and approximate body length)
|
||
<quantity id="4CC69B55FFC7843E8F10FBFAD8C1FB95" box="[892,1023,1052,1079]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="10.0" metricValueMin="8.5" pageId="90" pageNumber="91" unit="mm" value="9.25" valueMax="10.0" valueMin="8.5">8.5–10 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324653BFFC7843E8CABFBA6DA93FAAD" pageId="90" pageNumber="91" type="discussion">
|
||
<paragraph id="8B8136B0FFC7843E8CABFBA6DA93FAAD" blockId="90.[151,1437,153,1295]" pageId="90" pageNumber="91">
|
||
<emphasis id="B94AEAA2FFC7843E8CABFBA6DA5CFBF9" bold="true" box="[199,354,1088,1115]" pageId="90" pageNumber="91">Comparison.</emphasis>
|
||
Eggs with net-like relief, besides
|
||
<taxonomicName id="4C3E4D33FFC7843E8E88FBA7D86DFBF9" box="[740,851,1089,1115]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="flavus" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFC7843E8E88FBA7D86DFBF9" box="[740,851,1089,1115]" italics="true" pageId="90" pageNumber="91">Th. flavus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFC7843E8F37FBA7D8A5FBF9" bold="true" box="[859,923,1089,1115]" pageId="90" pageNumber="91">
|
||
<taxonomicNameLabel id="A27957D9FFC7843E8F37FBA7D8A5FBF9" box="[859,923,1089,1115]" pageId="90" pageNumber="91" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, were reported only for
|
||
<taxonomicName id="4C3E4D33FFC7843E88C2FBA7DA7EFBDD" authority="Traver & Edmunds 1967" authorityName="Traver & Edmunds" authorityYear="1967" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="packeri">
|
||
<emphasis id="B94AEAA2FFC7843E88C2FBA7DE10FBF9" box="[1198,1326,1089,1115]" italics="true" pageId="90" pageNumber="91">Th. packeri</emphasis>
|
||
<bibRefCitation id="EFAF4B41FFC7843E8959FBA7DA7EFBDD" author="Traver, J. R. & Edmunds, G. F. Jr." pageId="90" pageNumber="91" pagination="349 - 395" refId="ref72596" refString="Traver, J. R. & Edmunds, G. F. Jr. (1967) A revision of the genus Thraulodes (Ephemeroptera: Leptophlebiidae). Miscellaneous Publications of the Entomological Society of America, 5 (8), 349 - 395." type="journal article" year="1967">Traver & Edmunds 1967</bibRefCitation>
|
||
</taxonomicName>
|
||
and some unnamed species from
|
||
<collectingCountry id="F3297620FFC7843E8ED5FB83D83EFBDD" box="[697,768,1125,1151]" name="Brazil" pageId="90" pageNumber="91">Brazil</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3297620FFC7843E8F67FB83D876FBDD" box="[779,840,1125,1151]" name="Chile" pageId="90" pageNumber="91">Chile</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3297620FFC7843E8F12FB83D8C9FBDD" box="[894,1015,1125,1151]" name="Costa Rica" pageId="90" pageNumber="91">Costa Rica</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFAF4B41FFC7843E886AFB83DE1FFB22" author="Traver, J. R. & Edmunds, G. F. Jr." box="[1030,1313,1125,1152]" pageId="90" pageNumber="91" pagination="349 - 395" refId="ref72596" refString="Traver, J. R. & Edmunds, G. F. Jr. (1967) A revision of the genus Thraulodes (Ephemeroptera: Leptophlebiidae). Miscellaneous Publications of the Entomological Society of America, 5 (8), 349 - 395." type="journal article" year="1967">Traver & Edmunds 1967</bibRefCitation>
|
||
). The male imago of
|
||
<taxonomicName id="4C3E4D33FFC7843E8D68FB6FDA4AFB01" box="[260,372,1161,1187]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="flavus" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFC7843E8D68FB6FDA4AFB01" box="[260,372,1161,1187]" italics="true" pageId="90" pageNumber="91">Th. flavus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFC7843E8D17FB6FDA82FB01" bold="true" box="[379,444,1161,1187]" pageId="90" pageNumber="91">
|
||
<taxonomicNameLabel id="A27957D9FFC7843E8D17FB6FDA82FB01" box="[379,444,1161,1187]" pageId="90" pageNumber="91" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C3E4D33FFC7843E8E3EFB6FD9ECFB01" authorityName="Traver & Edmunds" authorityYear="1967" box="[594,722,1161,1187]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="packeri">
|
||
<emphasis id="B94AEAA2FFC7843E8E3EFB6FD9ECFB01" box="[594,722,1161,1187]" italics="true" pageId="90" pageNumber="91">Th. packeri</emphasis>
|
||
</taxonomicName>
|
||
by the entirely light yellow color of mesonotum (in contrast to pale reddish brown with dark and black in
|
||
<taxonomicName id="4C3E4D33FFC7843E8E14FB4BD9C6FB65" authorityName="Traver & Edmunds" authorityYear="1967" box="[632,760,1197,1223]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="packeri">
|
||
<emphasis id="B94AEAA2FFC7843E8E14FB4BD9C6FB65" box="[632,760,1197,1223]" italics="true" pageId="90" pageNumber="91">Th. packeri</emphasis>
|
||
</taxonomicName>
|
||
), the yellow coloration of costal and subcostal fields of fore wing, absence of black or dark brown markings on abdomen, wider dorsal extension of styliger and absence of lateral pouches on penis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |