278 lines
28 KiB
XML
278 lines
28 KiB
XML
<document id="A330F8063D085FC4C793BCCA12C4C0D9" ID-DOI="10.5252/cryptogamie-bryologie2023v44a1" ID-ISSN="1776-0992" ID-Zenodo-Dep="7822577" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1681312727720" checkinUser="felipe" docAuthor="Thouvenot, Louis" docDate="2023" docId="CD138796FFC8FFB1FE9F2133FB03FA19" docLanguage="en" docName="CryptogamieBryologie.44.1.1-60.pdf" docOrigin="Cryptogamie, Bryologie 20 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.4878.3.2" docStyle="DocumentStyle:3F4A000F5B047ABA0EAB2943AA9373AE.3:CryptogamieBryologie.2019-.journal_article" docStyleId="3F4A000F5B047ABA0EAB2943AA9373AE" docStyleName="CryptogamieBryologie.2019-.journal_article" docStyleVersion="3" docTitle="Heteroscyphus etesseanus Thouvenot 2023, comb. nov." docType="treatment" docVersion="8" lastPageNumber="26" masterDocId="312AFFEEFFD3FFAAFFDE2540FF9AFFE2" masterDocTitle="A taxonomic revision of the Lophocoleaceae Vanden Berghen (Marchantiophyta) of New Caledonia" masterLastPageNumber="60" masterPageNumber="1" pageNumber="26" updateTime="1707318641535" updateUser="guilherme" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="C691CA15B3540F326222DCBE1A4973E6">
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<mods:title id="CA98883CB650AC558BF7EC4D15A4AF0C">A taxonomic revision of the Lophocoleaceae Vanden Berghen (Marchantiophyta) of New Caledonia</mods:title>
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<mods:name id="0F3AAD8158CF61F491A02FD2BDADF631" type="personal">
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<mods:namePart id="FA4F51ECFCD6EB3275DECBA03982C044">Thouvenot, Louis</mods:namePart>
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<mods:affiliation id="5427241DC294D41588E50AF24344FB3F">Saint Léon, 66000 Perpignan (France) thouvenot. louis @ orange. fr</mods:affiliation>
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<mods:nameIdentifier id="F3569F3EEB9837EA806B3F81C80C66F7" type="email">louis@orange.fr</mods:nameIdentifier>
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<mods:title id="C0D46C0100D50452B16B775C4269CC0F">Cryptogamie, Bryologie</mods:title>
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</mods:titleInfo>
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<mods:part id="114B4FF582A4C25BBEBF25604D2663DA">
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<mods:date id="82202F5560561FA024E0955593CDB972">2023</mods:date>
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<mods:number id="198489EF9E3147B6A21CBEDDF7B25B27">2023-01-16</mods:number>
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<mods:url id="DD0925D8591DB7885F644B026290FF6F">http://dx.doi.org/10.11646/zootaxa.4878.3.2</mods:url>
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<mods:classification id="93B1FE8757DFA1945E793DE53061DDE6">journal article</mods:classification>
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<mods:identifier id="E5D50FF81942876532CCA68821545364" type="DOI">10.5252/cryptogamie-bryologie2023v44a1</mods:identifier>
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<mods:identifier id="EC225BABA9ABD23084BC7E7FC76699D1" type="ISSN">1776-0992</mods:identifier>
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<treatment id="CD138796FFC8FFB1FE9F2133FB03FA19" LSID="urn:lsid:plazi:treatment:CD138796FFC8FFB1FE9F2133FB03FA19" httpUri="http://treatment.plazi.org/id/CD138796FFC8FFB1FE9F2133FB03FA19" lastPageNumber="26" pageId="27" pageNumber="26">
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<subSubSection id="0DA0650BFFC8FFB1FE9F2133FDECFB4C" pageId="27" pageNumber="26" type="nomenclature">
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<paragraph id="45053680FFC8FFB1FE9F2133FDECFB4C" blockId="27.[278,630,1139,1199]" pageId="27" pageNumber="26">
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<heading id="1E4D81ECFFC8FFB1FE9F2133FDECFB4C" centered="true" fontSize="11" level="2" pageId="27" pageNumber="26" reason="4">
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<taxonomicName id="82BA4D03FFC8FFB1FE9F2133FE6AFB4C" ID-CoL="BLRGV" authority="(Steph.) Thouvenot" authorityName="Thouvenot" authorityYear="2023" baseAuthorityName="Thouvenot" baseAuthorityYear="2023" class="Jungermanniopsida" family="Lophocoleaceae" genus="Heteroscyphus" kingdom="Plantae" order="Jungermanniales" pageId="27" pageNumber="26" phylum="Marchantiophyta" rank="species" species="etesseanus" status="comb. nov.">
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<emphasis id="77CEEA92FFC8FFB1FE9F2133FDD0FB6C" bold="true" box="[321,586,1139,1166]" italics="true" pageId="27" pageNumber="26">Heteroscyphus etesseanus</emphasis>
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||
(Steph.) Thouvenot
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||
</taxonomicName>
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||
,
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||
<taxonomicNameLabel id="6CFD57E9FFC8FFB1FE2521D4FDECFB4C" box="[507,630,1172,1198]" pageId="27" pageNumber="26" rank="species">comb. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0DA0650BFFC8FFB1FE4821F4FE6FFB2C" box="[406,501,1204,1230]" pageId="27" pageNumber="26" type="description">
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<paragraph id="45053680FFC8FFB1FE4821F4FE6FFB2C" blockId="27.[406,501,1204,1230]" box="[406,501,1204,1230]" pageId="27" pageNumber="26">
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<heading id="1E4D81ECFFC8FFB1FE4821F4FE6FFB2C" box="[406,501,1204,1230]" centered="true" fontSize="11" level="3" pageId="27" pageNumber="26" reason="3">
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||
(
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<figureCitation id="DD812A05FFC8FFB1FE4121F4FE76FB2C" box="[415,492,1204,1230]" captionStart="FIG" captionStartId="28.[132,143,1604,1622]" captionTargetBox="[133,1453,215,1558]" captionTargetId="figure-168@28.[132,1455,215,1559]" captionTargetPageId="28" captionText="FIG. 13. — Heteroscyphus etesseanus (Steph.) Thouvenot, comb.nov.: A, shoot portion,ventral view; B-D, leaves; E-I, underleaves;J, median leaf cells; K, tooth at leaf apex; L, androecium; M, female bracteole; N, perianth tearing showing the inserted calyptra; O, female bract; P, perianth. A, L, M-P, drawn from the lectotype of Chiloscyphus etesseanus Steph. (G00061024); B-D, K, from Thouvenot NC2752; E-J, from Thouvenot NC2753. Scale bars: A, L, 1 mm; B-D, M-P, 500 µm; E-I, 200 µm; J, K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.7822620" httpUri="https://zenodo.org/record/7822620/files/figure.png" pageId="27" pageNumber="26">Fig. 13</figureCitation>
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)
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0DA0650BFFC8FFB1FF4521ADFF5FFAFD" pageId="27" pageNumber="26" type="reference_group">
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||
<paragraph id="45053680FFC8FFB1FF4521ADFF5FFAFD" blockId="27.[131,775,1261,1364]" pageId="27" pageNumber="26">
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||
<treatmentCitation id="C41B1091FFC8FFB1FF4521ADFF5BFAFD" author="Steph." journal="Species Hepaticarum" page="217" pageId="27" pageNumber="26" score="0.0033280253576670055" volume="3" year="1906">
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||
<taxonomicName id="82BA4D03FFC8FFB1FF4521ADFE30FAE7" ID-CoL="5XSDS" authority="Steph." authorityName="Steph." box="[155,426,1261,1285]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Chiloscyphus" kingdom="Plantae" order="Jungermanniales" pageId="27" pageNumber="26" phylum="Marchantiophyta" rank="species" species="etesseanus">
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||
<emphasis id="77CEEA92FFC8FFB1FF4521ADFEFDFAE6" box="[155,359,1261,1285]" italics="true" pageId="27" pageNumber="26">Chiloscyphus etesseanus</emphasis>
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Steph.
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</taxonomicName>
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||
,
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<bibCitation id="90903EECFFC8FFB1FE7121ADFF5BFAFD" journal="Species Hepaticarum" page="217" pageId="27" pageNumber="26" score="0.0033280253576670055" volume="3" year="1906">
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||
<emphasis id="77CEEA92FFC8FFB1FE7121ADFDFDFAE6" box="[431,615,1261,1284]" italics="true" pageId="27" pageNumber="26">Species Hepaticarum</emphasis>
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||
3: 217 (
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||
<bibRefCitation id="212B4B71FFC8FFB1FD6A21ADFF2DFAFD" author="STEPHANI F." pageId="27" pageNumber="26" pagination="935 - 966" refId="ref42244" refString="STEPHANI F. 1906. - Species hepaticarum 3. Bulletin de l'Herbier Boissier, serie 2, 6 (11): 935 - 966." type="journal article" year="1906">Stephani 1906</bibRefCitation>
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||
)
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||
</bibCitation>
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||
</treatmentCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0DA0650BFFC8FFB1FF142048FF79FAFD" box="[202,227,1288,1311]" pageId="27" pageNumber="26" type="multiple">
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||
<paragraph id="45053680FFC8FFB1FF142048FF79FAFD" blockId="27.[131,775,1261,1364]" box="[202,227,1288,1311]" pageId="27" pageNumber="26">—</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0DA0650BFFC8FFB1FF392048FC9CFA5B" pageId="27" pageNumber="26" type="materials_examined">
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||
<paragraph id="45053680FFC8FFB1FF392048FE59FAB6" blockId="27.[131,775,1261,1364]" pageId="27" pageNumber="26">
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||
<materialsCitation id="F5D23CDDFFC8FFB1FF392048FE59FAB6" collectionCode="G" collectorName="Etesse" country="New Caledonia" location="Noumeana" pageId="27" pageNumber="26" specimenCode="G00061024" specimenCount="1" typeStatus="lectotype">
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||
Type:
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||
<emphasis id="77CEEA92FFC8FFB1FEC12047FE2DFAC3" bold="true" box="[287,439,1287,1313]" pageId="27" pageNumber="26">
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||
<collectingCountry id="3DAD7610FFC8FFB1FEC12047FE2DFAC3" box="[287,439,1287,1313]" name="New Caledonia" pageId="27" pageNumber="26">New Caledonia</collectingCountry>
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||
</emphasis>
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||
.
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<emphasis id="77CEEA92FFC8FFB1FE1B2048FD84FAFD" box="[453,542,1288,1311]" italics="true" pageId="27" pageNumber="26">
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||
<collectorName id="E84F5356FFC8FFB1FE1B2048FE63FAFD" box="[453,505,1288,1311]" pageId="27" pageNumber="26">Etesse</collectorName>
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||
s.n.
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||
</emphasis>
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||
(
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||
<typeStatus id="9A018822FFC8FFB1FDF72047FDF9FAFD" box="[553,611,1287,1311]" pageId="27" pageNumber="26">lecto-</typeStatus>
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||
, here designated,
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||
<collectionCode id="23ABAE45FFC8FFB1FF5D2062FF0FFAD8" box="[131,149,1314,1338]" pageId="27" pageNumber="26">G</collectionCode>
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||
[“ex Herb. E. G. Paris, Nova Caledonia, in ditione
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||
<location id="4065605BFFC8FFB1FD4D2062FC98FADB" LSID="urn:lsid:plazi:treatment:CD138796FFC8FFB1FE9F2133FB03FA19:4065605BFFC8FFB1FD4D2062FC98FADB" box="[659,770,1314,1337]" country="New Caledonia" name="Noumeana" pageId="27" pageNumber="26">Noumeana</location>
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||
, 1904”,
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<emphasis id="77CEEA92FFC8FFB1FF10207DFEBBFAB6" box="[206,289,1340,1364]" italics="true" pageId="27" pageNumber="26">Etesse 12</emphasis>
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||
,
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||
<specimenCode id="151C9EFBFFC8FFB1FEF0207CFE3DFAB6" box="[302,423,1340,1364]" pageId="27" pageNumber="26">G00061024</specimenCode>
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||
]!).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="45053680FFC8FFB1FF5A202CFC9CFA5B" blockId="27.[131,776,1388,1465]" pageId="27" pageNumber="26">
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||
FURTHER SPECIMEN EXAMINED. —
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||
<materialsCitation id="F5D23CDDFFC8FFB1FE12202CFC9CFA5B" collectedFrom="on rocks in wet mesophilous forest" collectingDate="2019-10-13" collectionCode="PC" collectorName="Thouvenot" country="New Caledonia" elevation="163" location="Amoa valley" municipality="Poindimie" pageId="27" pageNumber="26" specimenCode="PC0779851" specimenCount="1" stateProvince="North Province">
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||
<emphasis id="77CEEA92FFC8FFB1FE12202CFDFBFA67" bold="true" box="[460,609,1388,1413]" pageId="27" pageNumber="26">
|
||
<collectingCountry id="3DAD7610FFC8FFB1FE12202CFDFBFA67" box="[460,609,1388,1413]" name="New Caledonia" pageId="27" pageNumber="26">New Caledonia</collectingCountry>
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||
</emphasis>
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||
.
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||
<collectingRegion id="877EF862FFC8FFB1FDB0202CFC9EFA66" box="[622,772,1388,1412]" country="New Caledonia" name="Province Nord" pageId="27" pageNumber="26">North Province</collectingRegion>
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||
,
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<collectingMunicipality id="A561ACFAFFC8FFB1FF5D20C6FF70FA7C" box="[131,234,1414,1438]" pageId="27" pageNumber="26">Poindimié</collectingMunicipality>
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||
,
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||
<location id="4065605BFFC8FFB1FF2C20C6FEF2FA7C" LSID="urn:lsid:plazi:treatment:CD138796FFC8FFB1FE9F2133FB03FA19:4065605BFFC8FFB1FF2C20C6FEF2FA7C" box="[242,360,1414,1438]" country="New Caledonia" municipality="Poindimie" name="Amoa valley" pageId="27" pageNumber="26" stateProvince="North Province">Amoa valley</location>
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||
,
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||
<location id="4065605BFFC8FFB1FEB120C6FE74FA7C" LSID="urn:lsid:plazi:treatment:CD138796FFC8FFB1FE9F2133FB03FA19:4065605BFFC8FFB1FEB120C6FE74FA7C" box="[367,494,1414,1438]" country="New Caledonia" municipality="Poindimie" name="Tipwadabwe" pageId="27" pageNumber="26" stateProvince="North Province">Tipwadabwé</location>
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||
,
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||
<collectedFrom id="F6A6757DFFC8FFB1FE2820C7FF26FA5B" pageId="27" pageNumber="26">on rocks in wet mesophilous forest</collectedFrom>
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||
,
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||
<quantity id="82429B65FFC8FFB1FF1A20E1FE9CFA5A" box="[196,262,1441,1465]" metricMagnitude="2" metricUnit="m" metricValue="1.63" pageId="27" pageNumber="26" unit="m" value="163.0">
|
||
<elevation id="CE97D1B3FFC8FFB1FF1A20E1FE9CFA5A" box="[196,262,1441,1465]" metricMagnitude="2" metricUnit="m" metricValue="1.63" pageId="27" pageNumber="26" unit="m" value="163.0">163 m</elevation>
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||
</quantity>
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||
,
|
||
<date id="31041040FFC8FFB1FECE20E1FEE0FA5B" box="[272,378,1441,1465]" pageId="27" pageNumber="26" value="2019-10-13">
|
||
<collectingDate id="2140E9A8FFC8FFB1FECE20E1FEE0FA5B" box="[272,378,1441,1465]" pageId="27" pageNumber="26" value="2019-10-13">13.X.2019</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="77CEEA92FFC8FFB1FE5B20E1FDD8FA5B" box="[389,578,1441,1465]" italics="true" pageId="27" pageNumber="26">
|
||
<collectorName id="E84F5356FFC8FFB1FE5B20E1FE7DFA5B" box="[389,487,1441,1465]" pageId="27" pageNumber="26">Thouvenot</collectorName>
|
||
NC2752
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="23ABAE45FFC8FFB1FD8E20E1FDE8FA5B" box="[592,626,1441,1465]" pageId="27" pageNumber="26">PC</collectionCode>
|
||
[
|
||
<specimenCode id="151C9EFBFFC8FFB1FDA620E0FD6BFA5A" box="[632,753,1440,1464]" pageId="27" pageNumber="26">PC0779851</specimenCode>
|
||
]).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DA0650BFFC8FFB1FF5A2091FE08F8E4" pageId="27" pageNumber="26" type="distribution">
|
||
<paragraph id="45053680FFC8FFB1FF5A2091FECCF95E" blockId="27.[131,777,1488,1724]" pageId="27" pageNumber="26">
|
||
DISTRIBUTION IN
|
||
<collectingCountry id="3DAD7610FFC8FFB1FE9F2091FE6EFA05" box="[321,500,1488,1512]" name="New Caledonia" pageId="27" pageNumber="26">NEW CALEDONIA</collectingCountry>
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||
. — Hitherto only known from
|
||
<collectingRegion id="877EF862FFC8FFB1FF1E20ABFEF9F9E1" box="[192,355,1515,1539]" country="New Caledonia" name="Province Nord" pageId="27" pageNumber="26">North Province</collectingRegion>
|
||
, rarely collected on rocks at low altitude (
|
||
<quantity id="82429B65FFC8FFB1FF532345FF65F9FF" box="[141,255,1541,1565]" metricMagnitude="2" metricUnit="m" metricValue="2.05" metricValueMax="2.5" metricValueMin="1.6" pageId="27" pageNumber="26" unit="m" value="205.0" valueMax="250.0" valueMin="160.0">160-250 m</quantity>
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||
) in wet forest with tall trees. The mention “Nouméa” on the label of the
|
||
<typeStatus id="9A018822FFC8FFB1FE902360FEE3F9DA" box="[334,377,1568,1592]" pageId="27" pageNumber="26">type</typeStatus>
|
||
specimen is not reliable since Etesse’s specimens were not precisely located and were commonly labelled as having originated from the capital of New Caledonia. In fact, his bryophyte gatherings were made along the Tipindjé river in
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||
<collectingRegion id="877EF862FFC8FFB1FF5823C9FEBAF943" box="[134,288,1673,1697]" country="New Caledonia" name="Province Nord" pageId="27" pageNumber="26">North Province</collectingRegion>
|
||
(
|
||
<bibRefCitation id="212B4B71FFC8FFB1FEF023CAFE3DF940" author="MORAT P." box="[302,423,1674,1698]" pageId="27" pageNumber="26" pagination="159 - 216" refId="ref41005" refString="MORAT P. 2010. - Les botanistes recolteurs en Nouvelle-Caledonie de 1774 a 2005. Adansonia 32 (2): 159 - 216. https: // doi. org / 10.5252 / a 2010 n 2 a 1" type="journal article" year="2010">Morat 2010</bibRefCitation>
|
||
), about
|
||
<quantity id="82429B65FFC8FFB1FE2323CAFDA4F943" box="[509,574,1673,1698]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="26" unit="km" value="30.0">30 km</quantity>
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||
from the locality of the recent collection.
|
||
</paragraph>
|
||
<paragraph id="45053680FFC8FFB1FF5A2394FE08F8E4" blockId="27.[131,775,1747,1798]" pageId="27" pageNumber="26">
|
||
TOTAL RANGE. — Endemic to
|
||
<collectingRegion id="877EF862FFC8FFB1FE782393FDA7F909" box="[422,573,1747,1771]" country="New Caledonia" name="Province Nord" pageId="27" pageNumber="26">North Province</collectingRegion>
|
||
, on the north-eastern versant of the central range.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DA0650BFFC8FFB1FF5A2272FC74FC1B" pageId="27" pageNumber="26" type="description">
|
||
<paragraph id="45053680FFC8FFB1FF5A2272FE8FF8A8" blockId="27.[132,277,1842,1868]" box="[132,277,1842,1868]" pageId="27" pageNumber="26">
|
||
<heading id="1E4D81ECFFC8FFB1FF5A2272FE8FF8A8" box="[132,277,1842,1868]" fontSize="9" level="3" pageId="27" pageNumber="26" reason="5">DESCRIPTION</heading>
|
||
</paragraph>
|
||
<paragraph id="45053680FFC8FFB1FF5A2211FF63F889" blockId="27.[132,249,1873,1899]" box="[132,249,1873,1899]" pageId="27" pageNumber="26">Dioecious.</paragraph>
|
||
<paragraph id="45053680FFC8FFB1FF5A22D1FF58F849" blockId="27.[132,194,1937,1963]" box="[132,194,1937,1963]" pageId="27" pageNumber="26">
|
||
<heading id="1E4D81ECFFC8FFB1FF5A22D1FF58F849" box="[132,194,1937,1963]" fontSize="11" level="3" pageId="27" pageNumber="26" reason="9">
|
||
<emphasis id="77CEEA92FFC8FFB1FF5A22D1FF58F849" box="[132,194,1937,1963]" italics="true" pageId="27" pageNumber="26">Habit</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="45053680FFC8FFB1FF5A22F1FC93F809" blockId="27.[132,777,1969,2028]" pageId="27" pageNumber="26">
|
||
Plants small, wet shoots
|
||
<quantity id="82429B65FFC8FFB1FE4C22F1FE62F82E" box="[402,504,1969,1996]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="27" pageNumber="26" unit="mm" value="1.6">1.60 mm</quantity>
|
||
wide, stems rigid, leaves horizontally spreading, slightly convex, contiguous, alternate.
|
||
</paragraph>
|
||
<paragraph id="45053680FFC8FFB1FCF32598FCF5FF13" blockId="27.[813,879,216,241]" box="[813,879,216,241]" pageId="27" pageNumber="26">
|
||
<heading id="1E4D81ECFFC8FFB1FCF32598FCF5FF13" box="[813,879,216,241]" fontSize="11" level="3" pageId="27" pageNumber="26" reason="9">
|
||
<emphasis id="77CEEA92FFC8FFB1FCF32598FCF5FF13" box="[813,879,216,241]" italics="true" pageId="27" pageNumber="26">Leaves</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="45053680FFC8FFB1FCF325B8FB71FE70" blockId="27.[810,1458,248,402]" pageId="27" pageNumber="26">
|
||
Rectangular,
|
||
<quantity id="82429B65FFC8FFB1FC6725B8FB86FEF1" box="[953,1052,248,275]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="27" pageNumber="26" unit="mm" value="0.6">0.60 mm</quantity>
|
||
long,
|
||
<quantity id="82429B65FFC8FFB1FB8125B8FB58FEF1" box="[1119,1218,248,275]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="26" unit="mm" value="0.5">0.50 mm</quantity>
|
||
wide, apices truncate, concave, both distal angles triangular acute, usually with a short linear tooth three cells long, some leaves pseudo-trilobate, sometimes with a further tooth, apical or subapical on the ventral margin, margins otherwise entire.
|
||
</paragraph>
|
||
<paragraph id="45053680FFC8FFB1FCF324F9FCC4FE31" blockId="27.[813,862,441,467]" box="[813,862,441,467]" pageId="27" pageNumber="26">
|
||
<heading id="1E4D81ECFFC8FFB1FCF324F9FCC4FE31" box="[813,862,441,467]" fontSize="11" level="3" pageId="27" pageNumber="26" reason="9">
|
||
<emphasis id="77CEEA92FFC8FFB1FCF324F9FCC4FE31" box="[813,862,441,467]" italics="true" pageId="27" pageNumber="26">Cells</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="45053680FFC8FFB1FCF3249BFC19FDF7" blockId="27.[813,1455,475,533]" pageId="27" pageNumber="26">Leaf cells hexagonal, 24-40 µm wide, firm walled without trigone.</paragraph>
|
||
<paragraph id="45053680FFC8FFB1FCF3277CFC33FDB4" blockId="27.[813,937,572,598]" box="[813,937,572,598]" pageId="27" pageNumber="26">
|
||
<heading id="1E4D81ECFFC8FFB1FCF3277CFC33FDB4" box="[813,937,572,598]" fontSize="11" level="3" pageId="27" pageNumber="26" reason="9">
|
||
<emphasis id="77CEEA92FFC8FFB1FCF3277CFC33FDB4" box="[813,937,572,598]" italics="true" pageId="27" pageNumber="26">Underleaves</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="45053680FFC8FFB1FCF3271DFC56FD15" blockId="27.[812,1457,605,759]" pageId="27" pageNumber="26">Small, hardly wider than the stem, obconical, free on both sides, deeply bifid, the discs sometimes strongly reduced, lobes divergent, setaceous, sinus variable, V-like or lunate or nearly flat, both lateral margins with a single tooth at most, small to linear.</paragraph>
|
||
<paragraph id="45053680FFC8FFB1FCF3265EFC36FCDA" blockId="27.[813,940,798,824]" box="[813,940,798,824]" pageId="27" pageNumber="26">
|
||
<heading id="1E4D81ECFFC8FFB1FCF3265EFC36FCDA" box="[813,940,798,824]" fontSize="11" level="3" pageId="27" pageNumber="26" reason="9">
|
||
<emphasis id="77CEEA92FFC8FFB1FCF3265EFC36FCDA" box="[813,940,798,824]" italics="true" pageId="27" pageNumber="26">Gametangia</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="45053680FFC8FFB1FCF32600FC74FC1B" blockId="27.[812,1455,831,1017]" pageId="27" pageNumber="26">
|
||
On short leafless, lateral-intercalary branches; gynoecia with laciniate bracts, bifid bracteoles with a single small tooth on both lateral margins; perianths about
|
||
<quantity id="82429B65FFC8FFB1FB10263FFABCFC78" box="[1230,1318,895,922]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="26" unit="mm" value="1.5">1.5 mm</quantity>
|
||
hight, campanulate, deeply trilobate, lobes laciniate-toothed, calyptrae very shorter than the perianths; androecia in spikes of up to six pairs of bracts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DA0650BFFC8FFB1FCF32160FB03FA19" pageId="27" pageNumber="26" type="discussion">
|
||
<paragraph id="45053680FFC8FFB1FCF32160FC36FBDB" blockId="27.[813,940,1056,1082]" box="[813,940,1056,1082]" pageId="27" pageNumber="26">
|
||
<heading id="1E4D81ECFFC8FFB1FCF32160FC36FBDB" box="[813,940,1056,1082]" fontSize="9" level="3" pageId="27" pageNumber="26" reason="5">COMMENTS</heading>
|
||
</paragraph>
|
||
<paragraph id="45053680FFC8FFB1FCF32102FB03FA19" blockId="27.[810,1456,1089,1532]" pageId="27" pageNumber="26">
|
||
Some vegetative features of the species such as the alternate leaves, the leaf cells without trigones and the small, free underleaves are suggestive of
|
||
<taxonomicName id="82BA4D03FFC8FFB1FC3921C1FBF2FB79" box="[999,1128,1153,1179]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Chiloscyphus" kingdom="Plantae" order="Jungermanniales" pageId="27" pageNumber="26" phylum="Marchantiophyta" rank="genus">
|
||
<emphasis id="77CEEA92FFC8FFB1FC3921C1FBF2FB79" box="[999,1128,1153,1179]" italics="true" pageId="27" pageNumber="26">Chiloscyphus</emphasis>
|
||
</taxonomicName>
|
||
. But the spicate androecia on abbreviated ventral branches are typical of the genus
|
||
<taxonomicName id="82BA4D03FFC8FFB1FA9721E2FCC0FB39" class="Jungermanniopsida" family="Lophocoleaceae" genus="Heteroscyphus" kingdom="Plantae" order="Jungermanniales" pageId="27" pageNumber="26" phylum="Marchantiophyta" rank="genus">
|
||
<emphasis id="77CEEA92FFC8FFB1FA9721E2FCC0FB39" italics="true" pageId="27" pageNumber="26">Heteroscyphus</emphasis>
|
||
</taxonomicName>
|
||
. Recent studies on
|
||
<taxonomicName id="82BA4D03FFC8FFB1FBC42181FB39FB39" box="[1050,1187,1217,1243]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Heteroscyphus" kingdom="Plantae" order="Jungermanniales" pageId="27" pageNumber="26" phylum="Marchantiophyta" rank="genus">
|
||
<emphasis id="77CEEA92FFC8FFB1FBC42181FB39FB39" box="[1050,1187,1217,1243]" italics="true" pageId="27" pageNumber="26">Heteroscyphus</emphasis>
|
||
</taxonomicName>
|
||
in Australasia treat species with similar vegetative features and highlight the importance of androecia and perianth
|
||
<typeStatus id="9A018822FFC8FFB1FBFF2041FBCDFAF9" box="[1057,1111,1281,1307]" pageId="27" pageNumber="26">types</typeStatus>
|
||
to discriminating these two genera (e.g.
|
||
<bibRefCitation id="212B4B71FFC8FFB1FC582061FBDEFAD9" author="ENGEL J. J. & HE X." box="[902,1092,1313,1339]" pageId="27" pageNumber="26" pagination="149 - 163" refId="ref39378" refString="ENGEL J. J. & HE X. 2010. - Studies on Lophocoleaceae. XIX. The systematic identity of Cyanolophocolea R. M. Schust., an intriguing liverwort from New Zealand and Australia, based on morphological and molecular evidence. The Bryologist 113 (1): 149 - 163. https: // doi. org / 10.1639 / 0007 - 2745 - 113.1.149" type="journal article" year="2010">Engel & He 2010</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="212B4B71FFC8FFB1FB8C2061FB50FAD9" author="ENGEL J. J." box="[1106,1226,1313,1339]" pageId="27" pageNumber="26" pagination="95 - 106" refId="ref39098" refString="ENGEL J. J. 2013. - Studies on Lophocoleaceae XXII. New taxa and combinations in New Zealand Heteroscyphus Schiffn. Polish Botanical Journal 58 (1): 95 - 106. https: // doi. org / 10.2478 / pbj- 2013 - 0011" type="journal article" year="2013">Engel 2013</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="212B4B71FFC8FFB1FB072060FA95FAD9" author="ENGEL J. J." box="[1241,1295,1312,1339]" pageId="27" pageNumber="26" pagination="553 - 582" refId="ref39200" refString="ENGEL J. J. 2015. - Studies on Lophocoleaceae. XXV. A conspectus of Heteroscyphus Schiffn. in temperate Australia together with nomenclatural changes in Chiloscyphus Corda and Leptoscyphus Mitt., refinements and a range extension of Clasmatocolea Spruce, and a range extension in Stolonivector J. J. Engel. Nova Hedwigia 100 (3 - 4): 553 - 582. https: // doi. org / 10.1127 / nova _ hedwigia / 2015 / 0260" type="journal article" year="2015">2015</bibRefCitation>
|
||
). These studies warrant the transfer of
|
||
<taxonomicName id="82BA4D03FFC8FFB1FBC62000FB67FAB8" box="[1048,1277,1344,1370]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Chiloscyphus" kingdom="Plantae" order="Jungermanniales" pageId="27" pageNumber="26" phylum="Marchantiophyta" rank="species" species="etesseanus">
|
||
<emphasis id="77CEEA92FFC8FFB1FBC62000FB67FAB8" box="[1048,1277,1344,1370]" italics="true" pageId="27" pageNumber="26">Chiloscyphus etesseanus</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="82BA4D03FFC8FFB1FAC02000FA32FAB8" box="[1310,1448,1344,1370]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Heteroscyphus" kingdom="Plantae" order="Jungermanniales" pageId="27" pageNumber="26" phylum="Marchantiophyta" rank="genus">
|
||
<emphasis id="77CEEA92FFC8FFB1FAC02000FA32FAB8" box="[1310,1448,1344,1370]" italics="true" pageId="27" pageNumber="26">Heteroscyphus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="82BA4D03FFC8FFB1FC9D2022FA82FA9E" authority="(Steph.) Thouvenot" authorityName="Thouvenot" authorityYear="2023" baseAuthorityName="Thouvenot" baseAuthorityYear="2023" box="[835,1304,1378,1405]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Heteroscyphus" kingdom="Plantae" order="Jungermanniales" pageId="27" pageNumber="26" phylum="Marchantiophyta" rank="species" species="etesseanus" status="comb. nov.">
|
||
<emphasis id="77CEEA92FFC8FFB1FC9D2022FBA1FA9E" box="[835,1083,1378,1404]" italics="true" pageId="27" pageNumber="26">Heteroscyphus etesseanus</emphasis>
|
||
(Steph.) Thouvenot
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="6CFD57E9FFC8FFB1FAFC2022FA03FA9E" box="[1314,1433,1378,1404]" pageId="27" pageNumber="26" rank="species">comb. nov.</taxonomicNameLabel>
|
||
is superficially similar to
|
||
<taxonomicName id="82BA4D03FFC8FFB1FBFB20C3FB0EFA7E" box="[1061,1172,1411,1436]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Heteroscyphus" kingdom="Plantae" order="Jungermanniales" pageId="27" pageNumber="26" phylum="Marchantiophyta" rank="species" species="argutus">
|
||
<emphasis id="77CEEA92FFC8FFB1FBFB20C3FB0EFA7E" box="[1061,1172,1411,1436]" italics="true" pageId="27" pageNumber="26">H. argutus</emphasis>
|
||
</taxonomicName>
|
||
, but the latter species can be easily distinguished from
|
||
<taxonomicName id="82BA4D03FFC8FFB1FB8F20E3FA33FA5E" authority="(Steph.) Thouvenot" authorityName="Thouvenot" authorityYear="2023" baseAuthorityName="Thouvenot" baseAuthorityYear="2023" box="[1105,1449,1442,1469]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Heteroscyphus" kingdom="Plantae" order="Jungermanniales" pageId="27" pageNumber="26" phylum="Marchantiophyta" rank="species" species="etesseanus" status="comb. nov.">
|
||
<emphasis id="77CEEA92FFC8FFB1FB8F20E3FB4EFA5E" box="[1105,1236,1443,1468]" italics="true" pageId="27" pageNumber="26">H. etesseanus</emphasis>
|
||
(Steph.) Thouvenot
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="6CFD57E9FFC8FFB1FCF52082FC3BFA3E" box="[811,929,1474,1500]" pageId="27" pageNumber="26" rank="species">comb. nov.</taxonomicNameLabel>
|
||
by its larger size and the rounded instead of retuse leaf apices with at least four teeth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |