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<document id="166D5C44FD54455B594FB93973A5FC1B" ID-CLB-Dataset="8980" ID-DOI="10.11646/zootaxa.4820.3.2" ID-GBIF-Dataset="8f91a52d-de34-47ed-80ac-9dc7c1bf915c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4398015" ID-ZooBank="560370FE-B271-48D2-A720-DB9034CE308E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609211755247" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2020" docId="960C8792160B753956E3FD18B01DBE81" docLanguage="en" docName="zootaxa.4820.3.2.pdf" docOrigin="Zootaxa 4820 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Demoulinia insularis Lugo-Ortiz &amp; McCafferty 1998" docType="treatment" docVersion="8" lastPageNumber="446" masterDocId="6A35FFEA160C75315674FFF5B178BE78" masterDocTitle="Demoulinia Gillies 1990 and two unnamed genera of the plesiomorphon Protopatellata (Ephemeroptera, Baetidae) from South Africa" masterLastPageNumber="464" masterPageNumber="438" pageNumber="445" updateTime="1698848356202" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1D8BB2C87C9781325D3DCB39180C63C4">Demoulinia Gillies 1990 and two unnamed genera of the plesiomorphon Protopatellata (Ephemeroptera, Baetidae) from South Africa</mods:title>
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<treatment id="960C8792160B753956E3FD18B01DBE81" ID-DOI="http://doi.org/10.5281/zenodo.4398133" ID-GBIF-Taxon="176569596" ID-Zenodo-Dep="4398133" LSID="urn:lsid:plazi:treatment:960C8792160B753956E3FD18B01DBE81" httpUri="http://treatment.plazi.org/id/960C8792160B753956E3FD18B01DBE81" lastPageId="8" lastPageNumber="446" pageId="7" pageNumber="445">
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<paragraph id="1E1A3684160B753656E3FD18B251BD7F" blockId="7.[151,809,749,812]" box="[151,809,749,776]" pageId="7" pageNumber="445">
<heading id="455281E8160B753656E3FD18B251BD7F" bold="true" box="[151,809,749,776]" fontSize="11" level="1" pageId="7" pageNumber="445" reason="1">
<taxonomicName id="D9A54D07160B753656E3FD18B251BD7F" ID-CoL="34LVC" authority="Lugo-Ortiz &amp; McCafferty 1998" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1998" box="[151,809,749,776]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="2CD1EA96160B753656E3FD18B251BD7F" bold="true" box="[151,809,749,776]" pageId="7" pageNumber="445">
<emphasis id="2CD1EA96160B753656E3FD18B0E0BD70" bold="true" box="[151,408,749,776]" italics="true" pageId="7" pageNumber="445">Demoulinia insularis</emphasis>
<bibRefCitation id="7A344B75160B753657EBFD18B251BD7F" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[415,809,749,775]" pageId="7" pageNumber="445" pagination="357 - 362" refId="ref11347" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1998) New species of Cloeon and Demoulinia (Ephemeroptera: Baetidae) from Madagascar. Entomological News, 109 (5), 357 - 362." type="journal article" year="1998">Lugo-Ortiz &amp; McCafferty 1998</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="56BF650F160B753656E3FCE4B0A5BD54" box="[151,477,785,812]" pageId="7" pageNumber="445" type="description">
<paragraph id="1E1A3684160B753656E3FCE4B0A5BD54" blockId="7.[151,809,749,812]" box="[151,477,785,812]" pageId="7" pageNumber="445">
(
<figureCitation id="869E2A01160B753656EBFCE4B07CBD53" box="[159,260,785,811]" captionStart="FIGURES 612" captionStartId="6.[151,264,1860,1884]" captionTargetBox="[151,1436,181,1836]" captionTargetId="figure-16@6.[151,1436,181,1836]" captionTargetPageId="6" captionText="FIGURES 612. Labrum of Demoulinia spp., dorsal view. 67, D. insularis (holotype). 812, D. crassi." figureDoi="http://doi.org/10.5281/zenodo.4398019" httpUri="https://zenodo.org/record/4398019/files/figure.png" pageId="7" pageNumber="445">Figs 67</figureCitation>
,
<figureCitation id="869E2A01160B75365764FCE4B020BD54" box="[272,344,785,812]" captionStart="FIGURES 2125" captionStartId="9.[151,264,1790,1814]" captionTargetBox="[151,1436,181,1767]" captionTargetId="figure-32@9.[151,1436,181,1767]" captionTargetPageId="9" captionText="FIGURES 2125. Demoulinia spp. 2123, D. crassi; 2425, D. insularis (holotype); 21, 22, 24, maxilla (21, with focus on ventral side; 22, with focus on dorsal side; d1, d2, d3, dentisetae; p1, p2, p3, segments of maxillary palp); 23, 25, hypopharynx and superlingua." figureDoi="http://doi.org/10.5281/zenodo.4398025" httpUri="https://zenodo.org/record/4398025/files/figure.png" pageId="7" pageNumber="445">2425</figureCitation>
,
<figureCitation id="869E2A01160B75365710FCE4B0FABD53" box="[356,386,785,811]" captionStart="FIGURES 3238" captionStartId="11.[151,264,1921,1945]" captionTargetBox="[151,1436,181,1898]" captionTargetId="figure-32@11.[151,1436,181,1898]" captionTargetPageId="11" captionText="FIGURES 3238. Larval legs of Demoulinia spp. 3237, D. crassi: 32, exuviae of hind leg; 33, inner side of fore tarsus; 34, femoral-tibial of hind leg, focus on anterior side; 3537, claws. 38, D. insularis (holotype), fragment of claw with subimaginal claws inside (arrow shows row of vestigial denticles on inner side)." figureDoi="http://doi.org/10.5281/zenodo.4398029" httpUri="https://zenodo.org/record/4398029/files/figure.png" pageId="7" pageNumber="445">38</figureCitation>
,
<figureCitation id="869E2A01160B753657FAFCE4B0D5BD53" box="[398,429,785,811]" captionStart="FIGURES 3944" captionStartId="12.[151,264,1894,1918]" captionTargetBox="[151,1436,181,1871]" captionTargetId="figure-16@12.[151,1436,181,1871]" captionTargetPageId="12" captionText="FIGURES 3944. Exuviae of larval abdomen of Demoulinia spp. 3943, D. crassi: 39, lateral and posterior area of abdominal tergum VII; 40, anterior-lateral area of abdominal sternum VI; 41, caudalii; 42, fragment of cercus; 43, paraproct. 44, D. insularis (holotype), median margin of paraproct (scale bar refers to Figs 39, 40, 42, 43)." figureDoi="http://doi.org/10.5281/zenodo.4398031" httpUri="https://zenodo.org/record/4398031/files/figure.png" pageId="7" pageNumber="445">44</figureCitation>
,
<figureCitation id="869E2A01160B753657CCFCE4B0ADBD54" box="[440,469,785,812]" captionStart="FIGURES 5761" captionStartId="16.[151,264,1374,1398]" captionTargetBox="[151,1436,310,1351]" captionTargetId="figure-80@16.[151,1436,309,1351]" captionTargetPageId="16" captionText="FIGURES 5761. Male genitalia of Demoulinia spp. 57, D. insularis (paratype of D. assimilis), subimaginal gonostyli under larval cuticle. 5861, D. crassi: 58, subimaginal gonostyli extracted from larva; 59, imaginal genitalia; 6061, apex of imaginal gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4398037" httpUri="https://zenodo.org/record/4398037/files/figure.png" pageId="7" pageNumber="445">57</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="56BF650F160B753656E3FCBAB3BBBDFE" pageId="7" pageNumber="445" type="reference_group">
<paragraph id="1E1A3684160B753656E3FCBAB242BD1F" blockId="7.[151,826,847,902]" box="[151,826,847,871]" pageId="7" pageNumber="445">
<treatmentCitationGroup id="3EB511AA160B753656E3FCBAB242BD1F" box="[151,826,847,871]" pageId="7" pageNumber="445">
<treatmentCitation id="9F041095160B753656E3FCBAB391BD1F" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[151,745,847,871]" page="360" pageId="7" pageNumber="445" year="1998">
<taxonomicName id="D9A54D07160B753656E3FCBAB391BD1F" ID-CoL="34LVC" authority="Lugo-Ortiz &amp; McCafferty 1998: 360" authorityName="Lugo-Ortiz &amp; McCafferty" authorityPageNumber="360" authorityYear="1998" box="[151,745,847,871]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="2CD1EA96160B753656E3FCBAB016BD1F" box="[151,366,847,871]" italics="true" pageId="7" pageNumber="445">Demoulinia insularis</emphasis>
<bibRefCitation id="7A344B75160B75365700FCBAB391BD1F" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[372,745,847,871]" pageId="7" pageNumber="445" pagination="357 - 362" refId="ref11347" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1998) New species of Cloeon and Demoulinia (Ephemeroptera: Baetidae) from Madagascar. Entomological News, 109 (5), 357 - 362." type="journal article" year="1998">Lugo-Ortiz &amp; McCafferty 1998: 360</bibRefCitation>
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(larva).
</treatmentCitationGroup>
</paragraph>
<paragraph id="1E1A3684160B753656E3FC9BB3BBBDFE" blockId="7.[151,826,847,902]" box="[151,707,878,902]" pageId="7" pageNumber="445">
<treatmentCitationGroup id="3EB511AA160B753656E3FC9BB3BBBDFE" box="[151,707,878,902]" pageId="7" pageNumber="445">
<treatmentCitation id="9F041095160B753656E3FC9BB35FBDFE" author="Gattolliat, J. - L." box="[151,551,878,902]" page="2" pageId="7" pageNumber="445" year="2003">
<taxonomicName id="D9A54D07160B753656E3FC9BB35FBDFE" ID-CoL="6CJTT" authority="Gattolliat 2003: 2" authorityName="Gattolliat" authorityPageNumber="2" authorityYear="2003" box="[151,551,878,902]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="2CD1EA96160B753656E3FC9BB016BDFE" box="[151,366,878,902]" italics="true" pageId="7" pageNumber="445">Demoulinia assimilis</emphasis>
<bibRefCitation id="7A344B75160B75365700FC9BB35FBDFE" author="Gattolliat, J. - L." box="[372,551,878,902]" pageId="7" pageNumber="445" pagination="1 - 18" refId="ref10074" refString="Gattolliat, J. - L. (2003) The genera Demoulinia Gillies and Potamocloeon Gillies (Ephemeroptera: Baetidae) in Madagascar. Zootaxa, 184 (1), 1 - 18. https: // doi. org / 10.11646 / zootaxa. 184.1.1" type="journal article" year="2003">Gattolliat 2003: 2</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(larva),
<emphasis id="2CD1EA96160B75365409FC9BB3BBBDFE" bold="true" box="[637,707,878,902]" pageId="7" pageNumber="445">syn. n.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="56BF650F160B753956E3FC58B01DBE81" lastPageId="8" lastPageNumber="446" pageId="7" pageNumber="445" type="materials_examined">
<paragraph id="1E1A3684160B753656E3FC58B0DCBAE4" blockId="7.[151,1437,941,2032]" pageId="7" pageNumber="445">
<emphasis id="2CD1EA96160B753656E3FC58B07EBDBF" bold="true" box="[151,262,941,967]" pageId="7" pageNumber="445">Material.</emphasis>
The
<typeStatus id="C11E8826160B7536574BFC58B0D9BDBF" box="[319,417,941,967]" pageId="7" pageNumber="445" type="holotype">holotype</typeStatus>
of
<taxonomicName id="D9A54D07160B753657B3FC5BB333BDBF" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1998" box="[455,587,941,967]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="2CD1EA96160B753657B3FC5BB333BDBF" box="[455,587,941,967]" italics="true" pageId="7" pageNumber="445">D. insularis</emphasis>
</taxonomicName>
is deposited in Purdue Entomological Research Collection, West Lafayette,
<collectingRegion id="DC61F866160B753656E3FC25B196BD92" box="[151,238,976,1002]" country="United States of America" name="Indiana" pageId="7" pageNumber="445">Indiana</collectingRegion>
,
<collectingCountry id="66B27614160B7536568DFC25B057BD92" box="[249,303,976,1002]" name="United States of America" pageId="7" pageNumber="445">USA</collectingCountry>
(
<bibRefCitation id="7A344B75160B7536574AFC25B3A4BD93" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[318,732,976,1003]" pageId="7" pageNumber="445" pagination="357 - 362" refId="ref11347" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1998) New species of Cloeon and Demoulinia (Ephemeroptera: Baetidae) from Madagascar. Entomological News, 109 (5), 357 - 362." type="journal article" year="1998">Lugo-Ortiz &amp; McCafferty 1998: 357</bibRefCitation>
) and is a mature female larva ready to moult to subimago; certain parts of it are slide-mounted (
<bibRefCitation id="7A344B75160B7536546DFC01B2C7BA77" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[537,959,1012,1039]" pageId="7" pageNumber="445" pagination="357 - 362" refId="ref11347" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1998) New species of Cloeon and Demoulinia (Ephemeroptera: Baetidae) from Madagascar. Entomological News, 109 (5), 357 - 362." type="journal article" year="1998">Lugo-Ortiz &amp; McCafferty 1998: 360</bibRefCitation>
), and the remaining parts are preserved in alcohol; Luke Jacobus kindly reexamined this specimen and made pictures (
<figureCitation id="869E2A01160B7536559CFBE2B535BA49" box="[1000,1101,1047,1073]" captionStart="FIGURES 612" captionStartId="6.[151,264,1860,1884]" captionTargetBox="[151,1436,181,1836]" captionTargetId="figure-16@6.[151,1436,181,1836]" captionTargetPageId="6" captionText="FIGURES 612. Labrum of Demoulinia spp., dorsal view. 67, D. insularis (holotype). 812, D. crassi." figureDoi="http://doi.org/10.5281/zenodo.4398019" httpUri="https://zenodo.org/record/4398019/files/figure.png" pageId="7" pageNumber="445">Figs 67</figureCitation>
,
<figureCitation id="869E2A01160B7536522CFBE2B5D9BA4A" box="[1112,1185,1047,1074]" captionStart="FIGURES 2125" captionStartId="9.[151,264,1790,1814]" captionTargetBox="[151,1436,181,1767]" captionTargetId="figure-32@9.[151,1436,181,1767]" captionTargetPageId="9" captionText="FIGURES 2125. Demoulinia spp. 2123, D. crassi; 2425, D. insularis (holotype); 21, 22, 24, maxilla (21, with focus on ventral side; 22, with focus on dorsal side; d1, d2, d3, dentisetae; p1, p2, p3, segments of maxillary palp); 23, 25, hypopharynx and superlingua." figureDoi="http://doi.org/10.5281/zenodo.4398025" httpUri="https://zenodo.org/record/4398025/files/figure.png" pageId="7" pageNumber="445">2425</figureCitation>
,
<figureCitation id="869E2A01160B753652D8FBE2B5B2BA49" box="[1196,1226,1047,1073]" captionStart="FIGURES 3238" captionStartId="11.[151,264,1921,1945]" captionTargetBox="[151,1436,181,1898]" captionTargetId="figure-32@11.[151,1436,181,1898]" captionTargetPageId="11" captionText="FIGURES 3238. Larval legs of Demoulinia spp. 3237, D. crassi: 32, exuviae of hind leg; 33, inner side of fore tarsus; 34, femoral-tibial of hind leg, focus on anterior side; 3537, claws. 38, D. insularis (holotype), fragment of claw with subimaginal claws inside (arrow shows row of vestigial denticles on inner side)." figureDoi="http://doi.org/10.5281/zenodo.4398029" httpUri="https://zenodo.org/record/4398029/files/figure.png" pageId="7" pageNumber="445">38</figureCitation>
,
<figureCitation id="869E2A01160B753652A1FBE2B58DBA49" box="[1237,1269,1047,1073]" captionStart="FIGURES 3944" captionStartId="12.[151,264,1894,1918]" captionTargetBox="[151,1436,181,1871]" captionTargetId="figure-16@12.[151,1436,181,1871]" captionTargetPageId="12" captionText="FIGURES 3944. Exuviae of larval abdomen of Demoulinia spp. 3943, D. crassi: 39, lateral and posterior area of abdominal tergum VII; 40, anterior-lateral area of abdominal sternum VI; 41, caudalii; 42, fragment of cercus; 43, paraproct. 44, D. insularis (holotype), median margin of paraproct (scale bar refers to Figs 39, 40, 42, 43)." figureDoi="http://doi.org/10.5281/zenodo.4398031" httpUri="https://zenodo.org/record/4398031/files/figure.png" pageId="7" pageNumber="445">44</figureCitation>
). The
<typeStatus id="C11E8826160B7536534EFBE2B4E4BA49" box="[1338,1436,1047,1073]" pageId="7" pageNumber="445" type="holotype">holotype</typeStatus>
and the
<typeStatus id="C11E8826160B7536569BFBCEB022BA2D" box="[239,346,1083,1109]" pageId="7" pageNumber="445" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="D9A54D07160B753657F4FBCEB37CBA2D" authorityName="Gattolliat" authorityYear="2003" box="[384,516,1083,1109]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="2CD1EA96160B753657F4FBCEB37CBA2D" box="[384,516,1083,1109]" italics="true" pageId="7" pageNumber="445">D. assimilis</emphasis>
</taxonomicName>
are deposited in the Museum of Zoology, Lausanne,
<collectingCountry id="66B27614160B75365222FBCEB5A4BA2D" box="[1110,1244,1083,1109]" name="Switzerland" pageId="7" pageNumber="445">Switzerland</collectingCountry>
(
<bibRefCitation id="7A344B75160B7536529EFBCEB1DFBA00" author="Gattolliat, J. - L." pageId="7" pageNumber="445" pagination="1 - 18" refId="ref10074" refString="Gattolliat, J. - L. (2003) The genera Demoulinia Gillies and Potamocloeon Gillies (Ephemeroptera: Baetidae) in Madagascar. Zootaxa, 184 (1), 1 - 18. https: // doi. org / 10.11646 / zootaxa. 184.1.1" type="journal article" year="2003">Gattolliat 2003: 2</bibRefCitation>
); Jean-Luc Gattolliat kindly reexamined certain details, prepared an additional slide of mature male larva and made pictures (
<figureCitation id="869E2A01160B75365737FB77B0EFBAE4" box="[323,407,1153,1180]" captionStart="FIGURES 5761" captionStartId="16.[151,264,1374,1398]" captionTargetBox="[151,1436,310,1351]" captionTargetId="figure-80@16.[151,1436,309,1351]" captionTargetPageId="16" captionText="FIGURES 5761. Male genitalia of Demoulinia spp. 57, D. insularis (paratype of D. assimilis), subimaginal gonostyli under larval cuticle. 5861, D. crassi: 58, subimaginal gonostyli extracted from larva; 59, imaginal genitalia; 6061, apex of imaginal gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4398037" httpUri="https://zenodo.org/record/4398037/files/figure.png" pageId="7" pageNumber="445">Fig. 57</figureCitation>
).
</paragraph>
<paragraph id="1E1A3684160B753656B3FB50B52FBB51" blockId="7.[151,1437,941,2032]" pageId="7" pageNumber="445">
<emphasis id="2CD1EA96160B753656B3FB50B3B1BAC7" bold="true" box="[199,713,1189,1216]" pageId="7" pageNumber="445">
Synonymy of
<taxonomicName id="D9A54D07160B75365704FB50B081BAB8" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1998" box="[368,505,1189,1216]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="2CD1EA96160B75365704FB50B081BAB8" bold="true" box="[368,505,1189,1216]" italics="true" pageId="7" pageNumber="445">D. insularis</emphasis>
</taxonomicName>
and
<taxonomicName id="D9A54D07160B7536544EFB50B3BBBAB8" authorityName="Gattolliat" authorityYear="2003" box="[570,707,1189,1216]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="2CD1EA96160B7536544EFB50B3BBBAB8" bold="true" box="[570,707,1189,1216]" italics="true" pageId="7" pageNumber="445">D. assimilis</emphasis>
</taxonomicName>
.
</emphasis>
The original description of
<taxonomicName id="D9A54D07160B7536527BFB53B5EEBAC7" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1998" box="[1039,1174,1189,1215]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="2CD1EA96160B7536527BFB53B5EEBAC7" box="[1039,1174,1189,1215]" italics="true" pageId="7" pageNumber="445">D. insularis</emphasis>
</taxonomicName>
was based on a single larva. The original description of
<taxonomicName id="D9A54D07160B7536547BFB3CB3EBBA9A" authorityName="Gattolliat" authorityYear="2003" box="[527,659,1224,1250]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="2CD1EA96160B7536547BFB3CB3EBBA9A" box="[527,659,1224,1250]" italics="true" pageId="7" pageNumber="445">D. assimilis</emphasis>
</taxonomicName>
was based on
<specimenCount id="08A3FD0D160B75365543FB3CB2E6BA9A" box="[823,926,1224,1251]" pageId="7" pageNumber="445" type="larva">21 larvae</specimenCount>
(
<bibRefCitation id="7A344B75160B753655D8FB3DB524BA9B" author="Gattolliat, J. - L." box="[940,1116,1224,1251]" pageId="7" pageNumber="445" pagination="1 - 18" refId="ref10074" refString="Gattolliat, J. - L. (2003) The genera Demoulinia Gillies and Potamocloeon Gillies (Ephemeroptera: Baetidae) in Madagascar. Zootaxa, 184 (1), 1 - 18. https: // doi. org / 10.11646 / zootaxa. 184.1.1" type="journal article" year="2003">Gattolliat 2003</bibRefCitation>
). As the differences between
<taxonomicName id="D9A54D07160B753656E3FB18B065BB7E" authorityName="Gattolliat" authorityYear="2003" box="[151,285,1260,1286]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="2CD1EA96160B753656E3FB18B065BB7E" box="[151,285,1260,1286]" italics="true" pageId="7" pageNumber="445">D. assimilis</emphasis>
</taxonomicName>
and
<taxonomicName id="D9A54D07160B75365723FB18B3D3BB7E" authority=", Gattolliat (2003)" authorityName=", Gattolliat" authorityYear="2003" box="[343,683,1260,1287]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="2CD1EA96160B75365723FB18B0A6BB7E" box="[343,478,1260,1286]" italics="true" pageId="7" pageNumber="445">D. insularis</emphasis>
,
<bibRefCitation id="7A344B75160B75365799FB19B3D3BB7E" author="Gattolliat, J. - L." box="[493,683,1260,1286]" pageId="7" pageNumber="445" pagination="1 - 18" refId="ref10074" refString="Gattolliat, J. - L. (2003) The genera Demoulinia Gillies and Potamocloeon Gillies (Ephemeroptera: Baetidae) in Madagascar. Zootaxa, 184 (1), 1 - 18. https: // doi. org / 10.11646 / zootaxa. 184.1.1" type="journal article" year="2003">Gattolliat (2003)</bibRefCitation>
</taxonomicName>
reported three characters: the setation of the dorsal surface of the labrum, the shape of the lingua and the number of the segments of the maxillary palp.
</paragraph>
<paragraph id="1E1A3684160B753656B3FAC6B0F1B85A" blockId="7.[151,1437,941,2032]" pageId="7" pageNumber="445">
Labral setation of
<taxonomicName id="D9A54D07160B753657E2FAC1B363BB35" authorityName="Gattolliat" authorityYear="2003" box="[406,539,1331,1357]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="2CD1EA96160B753657E2FAC1B363BB35" box="[406,539,1331,1357]" italics="true" pageId="7" pageNumber="445">D. assimilis</emphasis>
</taxonomicName>
was described as «subapical row of about 11 long stout setae subparallel to the distal margin, a few long fine setae scattered over surface» (
<bibRefCitation id="7A344B75160B75365531FAA3B569BB09" author="Gattolliat, J. - L." box="[837,1041,1366,1393]" pageId="7" pageNumber="445" pagination="1 - 18" refId="ref10074" refString="Gattolliat, J. - L. (2003) The genera Demoulinia Gillies and Potamocloeon Gillies (Ephemeroptera: Baetidae) in Madagascar. Zootaxa, 184 (1), 1 - 18. https: // doi. org / 10.11646 / zootaxa. 184.1.1" type="journal article" year="2003">Gattolliat 2003: 2</bibRefCitation>
, fig. 1), while labral setation of
<taxonomicName id="D9A54D07160B753653F5FAA2B181BBEC" authorityName=", Gattolliat" authorityYear="2003" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="2CD1EA96160B753653F5FAA2B181BBEC" italics="true" pageId="7" pageNumber="445">D. insularis</emphasis>
</taxonomicName>
was described as «numerous long, fine, simple setae scattered over surface» (
<bibRefCitation id="7A344B75160B75365207FA8FB075BBCF" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." pageId="7" pageNumber="445" pagination="357 - 362" refId="ref11347" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1998) New species of Cloeon and Demoulinia (Ephemeroptera: Baetidae) from Madagascar. Entomological News, 109 (5), 357 - 362." type="journal article" year="1998">Lugo-Ortiz &amp; McCafferty 1998: 360</bibRefCitation>
, fig. 11). The South African species
<taxonomicName id="D9A54D07160B753654C2FA6BB266BBCF" authorityName=": Gillies" authorityYear="1990" box="[694,798,1438,1463]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="crassi">
<emphasis id="2CD1EA96160B753654C2FA6BB266BBCF" box="[694,798,1438,1463]" italics="true" pageId="7" pageNumber="445">D. crassi</emphasis>
</taxonomicName>
demonstrates individual variability of this character: in some individuals long setae are evenly scattered over surface (left side of
<figureCitation id="869E2A01160B75365597FA34B54DBBA3" box="[995,1077,1473,1499]" captionStart="FIGURES 612" captionStartId="6.[151,264,1860,1884]" captionTargetBox="[151,1436,181,1836]" captionTargetId="figure-16@6.[151,1436,181,1836]" captionTargetPageId="6" captionText="FIGURES 612. Labrum of Demoulinia spp., dorsal view. 67, D. insularis (holotype). 812, D. crassi." figureDoi="http://doi.org/10.5281/zenodo.4398019" httpUri="https://zenodo.org/record/4398019/files/figure.png" pageId="7" pageNumber="445">Fig. 11</figureCitation>
), in others they form a more or less distinct subapical row (
<figureCitation id="869E2A01160B753657A1FA11B363BB86" box="[469,539,1508,1534]" captionStart="FIGURES 612" captionStartId="6.[151,264,1860,1884]" captionTargetBox="[151,1436,181,1836]" captionTargetId="figure-16@6.[151,1436,181,1836]" captionTargetPageId="6" captionText="FIGURES 612. Labrum of Demoulinia spp., dorsal view. 67, D. insularis (holotype). 812, D. crassi." figureDoi="http://doi.org/10.5281/zenodo.4398019" httpUri="https://zenodo.org/record/4398019/files/figure.png" pageId="7" pageNumber="445">Fig. 9</figureCitation>
); this fact allows the assumption that the Malagasy species can have the same individual variability.
</paragraph>
<paragraph id="1E1A3684160B753656B3F9DEB407B946" blockId="7.[151,1437,941,2032]" pageId="7" pageNumber="445">
The hypopharynx of
<taxonomicName id="D9A54D07160B753657CDF9D9B338B83D" authorityName="Gattolliat" authorityYear="2003" box="[441,576,1579,1605]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="2CD1EA96160B753657CDF9D9B338B83D" box="[441,576,1579,1605]" italics="true" pageId="7" pageNumber="445">D. assimilis</emphasis>
</taxonomicName>
was described as having «a broad trilobate superlingua, covered only with minute setae; lingua weakly developed laterally, apically with long thin setae» (
<bibRefCitation id="7A344B75160B75365263F9BAB59AB811" author="Gattolliat, J. - L." box="[1047,1250,1615,1641]" pageId="7" pageNumber="445" pagination="1 - 18" refId="ref10074" refString="Gattolliat, J. - L. (2003) The genera Demoulinia Gillies and Potamocloeon Gillies (Ephemeroptera: Baetidae) in Madagascar. Zootaxa, 184 (1), 1 - 18. https: // doi. org / 10.11646 / zootaxa. 184.1.1" type="journal article" year="2003">Gattolliat 2003: 2</bibRefCitation>
, fig. 2); here the term «superlingua» is erroneously applied to hypopharynx (= lingua), and the term «lingua» is erroneously applied to superlingua. The hypopharynx of
<taxonomicName id="D9A54D07160B7536545AF963B3CAB8C8" authorityName=", Gattolliat" authorityYear="2003" box="[558,690,1686,1712]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="2CD1EA96160B7536545AF963B3CAB8C8" box="[558,690,1686,1712]" italics="true" pageId="7" pageNumber="445">D. insularis</emphasis>
</taxonomicName>
was not characterized, and only figured in a resolution which does not show its setation (
<bibRefCitation id="7A344B75160B753657FDF94CB389B8AB" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[393,753,1721,1747]" pageId="7" pageNumber="445" pagination="357 - 362" refId="ref11347" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1998) New species of Cloeon and Demoulinia (Ephemeroptera: Baetidae) from Madagascar. Entomological News, 109 (5), 357 - 362." type="journal article" year="1998">Lugo-Ortiz &amp; McCafferty 1998</bibRefCitation>
: fig. 12). The
<typeStatus id="C11E8826160B753655F8F94CB296B8AB" box="[908,1006,1721,1747]" pageId="7" pageNumber="445" type="holotype">holotype</typeStatus>
of
<taxonomicName id="D9A54D07160B75365267F94FB5EEB8AB" authorityName=", Gattolliat" authorityYear="2003" box="[1043,1174,1721,1747]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="2CD1EA96160B75365267F94FB5EEB8AB" box="[1043,1174,1721,1747]" italics="true" pageId="7" pageNumber="445">D. insularis</emphasis>
</taxonomicName>
is a larva ready to molt to subimago, so its mouth parts are represented by empty cuticle; in such a condition the hypopharynx, which has a very thin cuticle, is crumpled: left lobe of the hypopharynx (= lingua) is compressed, but the right lobe retains its shape (
<figureCitation id="869E2A01160B75365692F8D6B042B946" box="[230,314,1827,1854]" captionStart="FIGURES 2125" captionStartId="9.[151,264,1790,1814]" captionTargetBox="[151,1436,181,1767]" captionTargetId="figure-32@9.[151,1436,181,1767]" captionTargetPageId="9" captionText="FIGURES 2125. Demoulinia spp. 2123, D. crassi; 2425, D. insularis (holotype); 21, 22, 24, maxilla (21, with focus on ventral side; 22, with focus on dorsal side; d1, d2, d3, dentisetae; p1, p2, p3, segments of maxillary palp); 23, 25, hypopharynx and superlingua." figureDoi="http://doi.org/10.5281/zenodo.4398025" httpUri="https://zenodo.org/record/4398025/files/figure.png" pageId="7" pageNumber="445">Fig. 25</figureCitation>
); it shape does not differ from that of
<taxonomicName id="D9A54D07160B75365497F8D1B21FB945" authorityName="Gattolliat" authorityYear="2003" box="[739,871,1827,1853]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="2CD1EA96160B75365497F8D1B21FB945" box="[739,871,1827,1853]" italics="true" pageId="7" pageNumber="445">D. assimilis</emphasis>
</taxonomicName>
(
<bibRefCitation id="7A344B75160B75365501F8D6B55CB946" author="Gattolliat, J. - L." box="[885,1060,1827,1854]" pageId="7" pageNumber="445" pagination="1 - 18" refId="ref10074" refString="Gattolliat, J. - L. (2003) The genera Demoulinia Gillies and Potamocloeon Gillies (Ephemeroptera: Baetidae) in Madagascar. Zootaxa, 184 (1), 1 - 18. https: // doi. org / 10.11646 / zootaxa. 184.1.1" type="journal article" year="2003">Gattolliat 2003</bibRefCitation>
: fig. 2) and
<taxonomicName id="D9A54D07160B753652DDF8D1B477B945" authorityName=": Gillies" authorityYear="1990" box="[1193,1295,1828,1853]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="crassi">
<emphasis id="2CD1EA96160B753652DDF8D1B477B945" box="[1193,1295,1828,1853]" italics="true" pageId="7" pageNumber="445">D. crassi</emphasis>
</taxonomicName>
(
<figureCitation id="869E2A01160B7536536AF8D6B40AB946" box="[1310,1394,1827,1854]" captionStart="FIGURES 2125" captionStartId="9.[151,264,1790,1814]" captionTargetBox="[151,1436,181,1767]" captionTargetId="figure-32@9.[151,1436,181,1767]" captionTargetPageId="9" captionText="FIGURES 2125. Demoulinia spp. 2123, D. crassi; 2425, D. insularis (holotype); 21, 22, 24, maxilla (21, with focus on ventral side; 22, with focus on dorsal side; d1, d2, d3, dentisetae; p1, p2, p3, segments of maxillary palp); 23, 25, hypopharynx and superlingua." figureDoi="http://doi.org/10.5281/zenodo.4398025" httpUri="https://zenodo.org/record/4398025/files/figure.png" pageId="7" pageNumber="445">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="1E1A3684160B753656B3F8B2B2A8B997" blockId="7.[151,1437,941,2032]" pageId="7" pageNumber="445">
The maxillary palp of
<taxonomicName id="D9A54D07160B753657CBF8BDB33AB919" authorityName="Gattolliat" authorityYear="2003" box="[447,578,1863,1889]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="2CD1EA96160B753657CBF8BDB33AB919" box="[447,578,1863,1889]" italics="true" pageId="7" pageNumber="445">D. assimilis</emphasis>
</taxonomicName>
was described as «3-segmented» (
<bibRefCitation id="7A344B75160B753655B6F8B2B5F2B919" author="Gattolliat, J. - L." box="[962,1162,1863,1889]" pageId="7" pageNumber="445" pagination="1 - 18" refId="ref10074" refString="Gattolliat, J. - L. (2003) The genera Demoulinia Gillies and Potamocloeon Gillies (Ephemeroptera: Baetidae) in Madagascar. Zootaxa, 184 (1), 1 - 18. https: // doi. org / 10.11646 / zootaxa. 184.1.1" type="journal article" year="2003">Gattolliat 2003: 4</bibRefCitation>
, fig. 5), while the maxillary palp of
<taxonomicName id="D9A54D07160B75365756F89EB0DFB9FC" authorityName=", Gattolliat" authorityYear="2003" box="[290,423,1898,1924]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="2CD1EA96160B75365756F89EB0DFB9FC" box="[290,423,1898,1924]" italics="true" pageId="7" pageNumber="445">D. insularis</emphasis>
</taxonomicName>
was described as «two segmented» (
<bibRefCitation id="7A344B75160B7536553EF89FB596B9FC" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[842,1262,1898,1925]" pageId="7" pageNumber="445" pagination="357 - 362" refId="ref11347" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1998) New species of Cloeon and Demoulinia (Ephemeroptera: Baetidae) from Madagascar. Entomological News, 109 (5), 357 - 362." type="journal article" year="1998">Lugo-Ortiz &amp; McCafferty 1998: 360</bibRefCitation>
, fig. 15). However, the
<typeStatus id="C11E8826160B75365775F87BB01BB9D0" box="[257,355,1934,1960]" pageId="7" pageNumber="445" type="holotype">holotype</typeStatus>
of
<taxonomicName id="D9A54D07160B753657F8F87AB36AB9D0" authorityName=", Gattolliat" authorityYear="2003" box="[396,530,1934,1960]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="2CD1EA96160B753657F8F87AB36AB9D0" box="[396,530,1934,1960]" italics="true" pageId="7" pageNumber="445">D. insularis</emphasis>
</taxonomicName>
(which is the only specimen of the type series) has one maxillary palp broken; another palp is preserved, mounted on a slide and has a clearly expressed third segment (
<figureCitation id="869E2A01160B7536521AF844B5C7B9B3" box="[1134,1215,1969,1995]" captionStart="FIGURES 2125" captionStartId="9.[151,264,1790,1814]" captionTargetBox="[151,1436,181,1767]" captionTargetId="figure-32@9.[151,1436,181,1767]" captionTargetPageId="9" captionText="FIGURES 2125. Demoulinia spp. 2123, D. crassi; 2425, D. insularis (holotype); 21, 22, 24, maxilla (21, with focus on ventral side; 22, with focus on dorsal side; d1, d2, d3, dentisetae; p1, p2, p3, segments of maxillary palp); 23, 25, hypopharynx and superlingua." figureDoi="http://doi.org/10.5281/zenodo.4398025" httpUri="https://zenodo.org/record/4398025/files/figure.png" pageId="7" pageNumber="445">Fig. 24</figureCitation>
) whose shape is the same as in
<taxonomicName id="D9A54D07160B75365767F823B0EFB997" authorityName="Gattolliat" authorityYear="2003" box="[275,407,2005,2031]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="2CD1EA96160B75365767F823B0EFB997" box="[275,407,2005,2031]" italics="true" pageId="7" pageNumber="445">D. assimilis</emphasis>
</taxonomicName>
(
<bibRefCitation id="7A344B75160B753657D2F820B32CB997" author="Gattolliat, J. - L." box="[422,596,2005,2031]" pageId="7" pageNumber="445" pagination="1 - 18" refId="ref10074" refString="Gattolliat, J. - L. (2003) The genera Demoulinia Gillies and Potamocloeon Gillies (Ephemeroptera: Baetidae) in Madagascar. Zootaxa, 184 (1), 1 - 18. https: // doi. org / 10.11646 / zootaxa. 184.1.1" type="journal article" year="2003">Gattolliat 2003</bibRefCitation>
: fig. 5) and in
<taxonomicName id="D9A54D07160B75365483F823B224B997" authorityName=": Gillies" authorityYear="1990" box="[759,860,2006,2031]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="445" phylum="Arthropoda" rank="species" species="crassi">
<emphasis id="2CD1EA96160B75365483F823B224B997" box="[759,860,2006,2031]" italics="true" pageId="7" pageNumber="445">D. crassi</emphasis>
</taxonomicName>
(
<figureCitation id="869E2A01160B7536551FF820B2BAB997" box="[875,962,2005,2031]" captionStart="FIGURES 2125" captionStartId="9.[151,264,1790,1814]" captionTargetBox="[151,1436,181,1767]" captionTargetId="figure-32@9.[151,1436,181,1767]" captionTargetPageId="9" captionText="FIGURES 2125. Demoulinia spp. 2123, D. crassi; 2425, D. insularis (holotype); 21, 22, 24, maxilla (21, with focus on ventral side; 22, with focus on dorsal side; d1, d2, d3, dentisetae; p1, p2, p3, segments of maxillary palp); 23, 25, hypopharynx and superlingua." figureDoi="http://doi.org/10.5281/zenodo.4398025" httpUri="https://zenodo.org/record/4398025/files/figure.png" pageId="7" pageNumber="445">Figs 21</figureCitation>
).
</paragraph>
<paragraph id="1E1A36841604753956B3FF62B01DBE81" blockId="8.[151,1437,151,249]" pageId="8" pageNumber="446">
Thus, not one of the three reported characters allows
<taxonomicName id="D9A54D07160475395569FF62B2DABEC9" authorityName="Gattolliat" authorityYear="2003" box="[797,930,151,177]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="446" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="2CD1EA96160475395569FF62B2DABEC9" box="[797,930,151,177]" italics="true" pageId="8" pageNumber="446">D. assimilis</emphasis>
</taxonomicName>
to be distinguished from
<taxonomicName id="D9A54D071604753952B6FF62B43FBEC9" authorityName=", Gattolliat" authorityYear="2003" box="[1218,1351,151,177]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="446" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="2CD1EA961604753952B6FF62B43FBEC9" box="[1218,1351,151,177]" italics="true" pageId="8" pageNumber="446">D. insularis</emphasis>
</taxonomicName>
. A new synonymy is proposed here:
<taxonomicName id="D9A54D071604753957AEFF4EB549BEAD" authority="Lugo-Ortiz &amp; McCafferty 1998" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1998" box="[474,1073,187,213]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="446" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="2CD1EA961604753957AEFF4EB3BFBEAD" box="[474,711,187,213]" italics="true" pageId="8" pageNumber="446">Demoulinia insularis</emphasis>
<bibRefCitation id="7A344B751604753954BAFF4EB549BEAD" author="Lugo-Ortiz, C. R. &amp; McCafferty, W. P." box="[718,1073,187,213]" pageId="8" pageNumber="446" pagination="357 - 362" refId="ref11347" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1998) New species of Cloeon and Demoulinia (Ephemeroptera: Baetidae) from Madagascar. Entomological News, 109 (5), 357 - 362." type="journal article" year="1998">Lugo-Ortiz &amp; McCafferty 1998</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="D9A54D0716047539522CFF4EB185BE81" authority="Gattolliat 2003" authorityName="Gattolliat" authorityYear="2003" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="446" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="2CD1EA9616047539522CFF4EB43EBEAD" box="[1112,1350,187,213]" italics="true" pageId="8" pageNumber="446">Demoulinia assimilis</emphasis>
<bibRefCitation id="7A344B75160475395339FF4EB185BE81" author="Gattolliat, J. - L." pageId="8" pageNumber="446" pagination="1 - 18" refId="ref10074" refString="Gattolliat, J. - L. (2003) The genera Demoulinia Gillies and Potamocloeon Gillies (Ephemeroptera: Baetidae) in Madagascar. Zootaxa, 184 (1), 1 - 18. https: // doi. org / 10.11646 / zootaxa. 184.1.1" type="journal article" year="2003">Gattolliat 2003</bibRefCitation>
</taxonomicName>
,
<emphasis id="2CD1EA9616047539577CFF2AB02DBE81" bold="true" box="[264,341,223,249]" pageId="8" pageNumber="446">
<taxonomicNameLabel id="37E257ED16047539577CFF2AB02DBE81" box="[264,341,223,249]" pageId="8" pageNumber="446">syn. n.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>