288 lines
36 KiB
XML
288 lines
36 KiB
XML
<document id="56851B898D21915FE38690F891DA8D68" ID-DOI="10.1111/zoj.12362" ID-ISSN="0024-4082" ID-Zenodo-Dep="10109299" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630235092894" checkinUser="marcus" docAuthor="Dellapé, Pablo M., Melo, María C. & Henry, Thomas J." docDate="2016" docId="69250127E966500EFC3C373AFAAEE876" docLanguage="en" docName="zoj12362.pdf" docOrigin="Zoological Journal of the Linnean Society 177 (1)" docSource="http://dx.doi.org/10.1111/zoj.12362" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Heraeus nicaraguensis Dellapé & Melo & Henry 2016, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="99" masterDocId="951C795FE9235048FF9E3541FFCCEE5A" masterDocTitle="A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species" masterLastPageNumber="134" masterPageNumber="29" pageNumber="98" updateTime="1700019240479" updateUser="ExternalLinkService">
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<mods:title id="B34FCCC938868E36BB7B43FC86024135">A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species</mods:title>
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<mods:namePart id="71EF7616C9E4FE27439F008E970CA480">Dellapé, Pablo M.</mods:namePart>
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<mods:namePart id="F0841A20540D258D21E1B88733B103D2">Melo, María C.</mods:namePart>
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<mods:namePart id="311500B9FF9DC52FD903B8A0DD94A406">Henry, Thomas J.</mods:namePart>
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<mods:title id="2EB18DC2FD252C65510CD3EEC8B4F6B2">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="05841D25D30248A7F2177DE5A06D3497">2016</mods:date>
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<treatment id="69250127E966500EFC3C373AFAAEE876" LSID="urn:lsid:plazi:treatment:69250127E966500EFC3C373AFAAEE876" httpUri="http://treatment.plazi.org/id/69250127E966500EFC3C373AFAAEE876" lastPageId="70" lastPageNumber="99" pageId="69" pageNumber="98">
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<subSubSection id="A996E3BAE966500DFC3C373AFAF5ECD5" box="[930,1337,634,656]" pageId="69" pageNumber="98" type="nomenclature">
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<paragraph id="E133B031E966500DFC3C373AFAF5ECD5" blockId="69.[930,1337,634,656]" box="[930,1337,634,656]" pageId="69" pageNumber="98">
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<heading id="BA7B075DE966500DFC3C373AFAF5ECD5" box="[930,1337,634,656]" centered="true" fontSize="9" level="2" pageId="69" pageNumber="98" reason="4">
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<taxonomicName id="268CCBB2E966500DFC3C373AFB1AECD5" ID-CoL="3KXML" authority="Dellapé & Melo & Henry, 2016" authorityName="Dellapé & Melo & Henry" authorityYear="2016" box="[930,1238,635,656]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="69" pageNumber="100" phylum="Arthropoda" rank="species" species="nicaraguensis" status="sp. nov.">HERAEUS NICARAGUENSIS</taxonomicName>
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<emphasis id="D3F86C23E966500DFB40373FFAF5ECD5" bold="true" box="[1246,1337,634,655]" pageId="69" pageNumber="98">
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<taxonomicNameLabel id="C8CBD158E966500DFB40373FFAF5ECD5" box="[1246,1337,634,655]" pageId="69" pageNumber="98" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="A996E3BAE966500EFC5537E0FAAEE876" lastPageId="70" lastPageNumber="99" pageId="69" pageNumber="98" type="description">
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<paragraph id="E133B031E966500DFC5537E0FADCECED" blockId="69.[971,1296,673,695]" box="[971,1296,673,695]" pageId="69" pageNumber="98">
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(
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<figureCitation id="79B7ACB4E966500DFC4D37E0FB8EECED" box="[979,1090,673,695]" captionStart="Figure 36" captionStartId="65.[164,244,1794,1813]" captionTargetBox="[164,1444,194,1763]" captionTargetId="figure-0@65.[164,1444,193,1764]" captionTargetPageId="65" captionText="Figure 36. Habitus dorsal: A, Heraeus mesoamericanus sp. nov.; B, Heraeus mexicanus sp. nov.; C, Heraeus morganae sp. nov.; D, Heraeus nicaraguensis sp. nov.; E, Heraeus pacificus Barber, 1925; F, Heraeus pallidinervis sp. nov.; G, Heraeus penai sp. nov.; H, Heraeus plebejus Stål, 1874; I, Heraeus pulchellus Barber, 1954." figureDoi="http://doi.org/10.5281/zenodo.10128651" httpUri="https://zenodo.org/record/10128651/files/figure.png" pageId="69" pageNumber="98">FIGS 36D</figureCitation>
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,
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<figureCitation id="79B7ACB4E966500DFBD137E0FBA1ECEC" box="[1103,1133,673,694]" captionStart="Figure 38" captionStartId="68.[144,222,1166,1185]" captionTargetBox="[144,1424,195,1136]" captionTargetId="figure-238@68.[144,1424,195,1136]" captionTargetPageId="68" captionText="Figure 38. Distributional map of Heraeus mexicanus sp. nov., Heraeus nicaraguensis sp. nov., and Heraeus pacificus Barber, 1925." figureDoi="http://doi.org/10.5281/zenodo.10128655" httpUri="https://zenodo.org/record/10128655/files/figure.png" pageId="69" pageNumber="98">38</figureCitation>
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,
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<figureCitation id="79B7ACB4E966500DFBE437E0FB67ECEC" box="[1146,1195,673,694]" captionStart="Figure 39" captionStartId="70.[144,220,1059,1078]" captionTargetBox="[305,1261,196,1028]" captionTargetId="figure-296@70.[303,1263,194,1030]" captionTargetPageId="70" captionText="Figure 39. Legs: A, Heraeus nicaraguensis sp. nov.; B, Heraeus pacificus Barber, 1925; C, Heraeus pallidinervis sp. nov.; D, Heraeus penai sp. nov.; E, Heraeus plebejus Stål, 1874; F, Heraeus pulchellus Barber, 1954." figureDoi="http://doi.org/10.5281/zenodo.10128659" httpUri="https://zenodo.org/record/10128659/files/figure.png" pageId="69" pageNumber="98">39A</figureCitation>
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,
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<figureCitation id="79B7ACB4E966500DFB2637E0FAC5ECED" box="[1208,1289,673,695]" captionStart="Figure 40" captionStartId="71.[164,243,1714,1733]" captionTargetBox="[254,1349,195,1677]" captionTargetId="figure-0@71.[250,1356,192,1684]" captionTargetPageId="71" captionText="Figure 40. Male genitalia. Heraeus nicaraguensis sp. nov.: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus pacificus Barber, 1925: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus pallidinervis sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus penai sp. nov.: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." figureDoi="http://doi.org/10.5281/zenodo.10128661" httpUri="https://zenodo.org/record/10128661/files/figure.png" pageId="69" pageNumber="98">40A–D</figureCitation>
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)
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</paragraph>
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<paragraph id="E133B031E966500DFCA43789FC66EC87" blockId="69.[826,1442,712,1101]" box="[826,938,712,733]" pageId="69" pageNumber="98">
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<emphasis id="D3F86C23E966500DFCA43789FC66EC87" box="[826,938,712,733]" italics="true" pageId="69" pageNumber="98">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="E133B031E966500DFCA437A7FAE2ED02" blockId="69.[826,1442,712,1101]" pageId="69" pageNumber="98">Labium extending to mesocoxae. Distiflagellomere with a pale band. Membrane with an apical pale spot. Metafemur with a subapical dark band. Apex of pygophore pointed dorsally in lateral view.</paragraph>
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<paragraph id="E133B031E966500DFCCD3621FAC3EA17" blockId="69.[826,1442,712,1101]" pageId="69" pageNumber="98">
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The diagnostic characters relate
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<taxonomicName id="268CCBB2E966500DFA1A3620FC18EDCF" authority="Dellapé & Melo & Henry, 2016" authorityName="Dellapé & Melo & Henry" authorityYear="2016" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="69" pageNumber="98" phylum="Arthropoda" rank="species" species="nicaraguensis" status="sp. nov.">
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<emphasis id="D3F86C23E966500DFA1A3620FC18EDCF" italics="true" pageId="69" pageNumber="98">H. nicaraguensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C8CBD158E966500DFC4436C1FBE1EDCF" box="[986,1069,896,917]" pageId="69" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
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to
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<taxonomicName id="268CCBB2E966500DFBCF36C1FB17EDCE" authority="Dellapé & Melo & Henry, 2016" authorityName="Dellapé & Melo & Henry" authorityYear="2016" box="[1105,1243,895,917]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="69" pageNumber="98" phylum="Arthropoda" rank="species" species="bahiensis" status="sp. nov.">
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<emphasis id="D3F86C23E966500DFBCF36C1FB17EDCE" box="[1105,1243,895,917]" italics="true" pageId="69" pageNumber="98">H. bahiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C8CBD158E966500DFB7F36C1FAF8EDCF" box="[1249,1332,896,917]" pageId="69" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
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They can be distinguished by the darker general colouration of
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<taxonomicName id="268CCBB2E966500DFCA436FCFC36ED88" authority="Dellapé & Melo & Henry, 2016" authorityName="Dellapé & Melo & Henry" authorityYear="2016" box="[826,1018,957,978]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="69" pageNumber="98" phylum="Arthropoda" rank="species" species="nicaraguensis" status="sp. nov.">
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<emphasis id="D3F86C23E966500DFCA436FCFC36ED88" box="[826,1018,957,978]" italics="true" pageId="69" pageNumber="98">H. nicaraguensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C8CBD158E966500DFB9F36FCFB94ED88" box="[1025,1112,957,978]" pageId="69" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
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, with a blackish head and a dark posterior pronotal lobe with four longitudinal pale stripes, whereas in
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<taxonomicName id="268CCBB2E966500DFBBD36BBFB7DEA54" authority="Dellapé & Melo & Henry, 2016" authorityName="Dellapé & Melo & Henry" authorityYear="2016" box="[1059,1201,1017,1039]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="69" pageNumber="98" phylum="Arthropoda" rank="species" species="bahiensis" status="sp. nov.">
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<emphasis id="D3F86C23E966500DFBBD36BBFB7DEA54" box="[1059,1201,1017,1039]" italics="true" pageId="69" pageNumber="98">H. bahiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C8CBD158E966500DFB2636BBFAC3EA55" box="[1208,1295,1018,1039]" pageId="69" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
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, the head is dark reddish brown and the posterior pronotal lobe is brown, with diffuse longitudinal stripes.
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</paragraph>
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<paragraph id="E133B031E966500DFCA4313BFBFBEAD4" blockId="69.[826,1442,1145,1902]" box="[826,1079,1145,1167]" pageId="69" pageNumber="98">
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<emphasis id="D3F86C23E966500DFCA4313BFBFBEAD4" box="[826,1079,1145,1167]" italics="true" pageId="69" pageNumber="98">
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Description (
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<figureCitation id="79B7ACB4E966500DFC53313BFBE3EAD4" box="[973,1071,1145,1167]" captionStart="Figure 36" captionStartId="65.[164,244,1794,1813]" captionTargetBox="[164,1444,194,1763]" captionTargetId="figure-0@65.[164,1444,193,1764]" captionTargetPageId="65" captionText="Figure 36. Habitus dorsal: A, Heraeus mesoamericanus sp. nov.; B, Heraeus mexicanus sp. nov.; C, Heraeus morganae sp. nov.; D, Heraeus nicaraguensis sp. nov.; E, Heraeus pacificus Barber, 1925; F, Heraeus pallidinervis sp. nov.; G, Heraeus penai sp. nov.; H, Heraeus plebejus Stål, 1874; I, Heraeus pulchellus Barber, 1954." figureDoi="http://doi.org/10.5281/zenodo.10128651" httpUri="https://zenodo.org/record/10128651/files/figure.png" pageId="69" pageNumber="98">Fig. 36D</figureCitation>
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)
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</emphasis>
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</paragraph>
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<paragraph id="E133B031E966500DFCA431D9FB32EAF4" blockId="69.[826,1442,1145,1902]" box="[826,1278,1176,1198]" pageId="69" pageNumber="98">Total length 5.32. Dark brown species.</paragraph>
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<paragraph id="E133B031E966500DFCCD31F6FB35E822" blockId="69.[826,1442,1145,1902]" pageId="69" pageNumber="98">
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<emphasis id="D3F86C23E966500DFCCD31F6FC5AEA96" box="[851,918,1207,1228]" italics="true" pageId="69" pageNumber="98">Head:</emphasis>
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Dark brown, shiny, coriaceous, with short recumbent and long erect setae dorsally. Head length 1.10, width 0.83, postocular length 0.43. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli at level of an imaginary line passing the posterior border of eyes. Interocular width 0.40, interocellar width 0.24. Labium pale brown, with erect setae; extending to mesocoxae. Labial segment lengths: I 0.70, II 0.75, III 0.48, and IV 0.29. Antennae brown, basal half of basiflagellomere and base of distiflagellomere darkened, distiflagellomere with a wide sub-basal yellowish band; setae abundant, short, and recumbent. Antennal lengths: scape 0.42, pedicel 0.99, basiflagellomere 0.80, and distiflagellomere 1.04. Length of pale band on distiflagellomere 0.40.
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</paragraph>
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<paragraph id="E133B031E966500EFCCD33C0FF33EBAF" blockId="69.[826,1442,1145,1902]" lastBlockId="70.[144,759,1166,1800]" lastPageId="70" lastPageNumber="99" pageId="69" pageNumber="98">
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<emphasis id="D3F86C23E966500DFCCD33C0FC6BE8CC" box="[851,935,1665,1686]" italics="true" pageId="69" pageNumber="98">Thorax:</emphasis>
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Pronotum dark brown, posterior pronotal lobe slightly paler, with four longitudinal pale bands and a pale spot on each humeral angle. Anterior and posterior lobes with short recumbent setae and long erect setae, longer on anterior lobe. Collar length 0.08, anterior lobe length 0.54, posterior lobe length 0.46; anterior lobe width 0.86, posterior lobe width 1.28. Scutellum dark brown, apex paler, pruinose, punctate, with short recumbent and erect setae, as on anterior lobe. Pleurae dark brown, punctate, with short recumbent setae. Evaporative area extended. Hemelytra dark brown, with a pale inner corial spot and a subapical corial spot; costal margin pale on proximal three-quarters; membrane brown, with veins paler and an elongate apical pale spot (
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<figureCitation id="79B7ACB4E965500EFE943007FEA3EB06" box="[266,367,1350,1372]" captionStart="Figure 36" captionStartId="65.[164,244,1794,1813]" captionTargetBox="[164,1444,194,1763]" captionTargetId="figure-0@65.[164,1444,193,1764]" captionTargetPageId="65" captionText="Figure 36. Habitus dorsal: A, Heraeus mesoamericanus sp. nov.; B, Heraeus mexicanus sp. nov.; C, Heraeus morganae sp. nov.; D, Heraeus nicaraguensis sp. nov.; E, Heraeus pacificus Barber, 1925; F, Heraeus pallidinervis sp. nov.; G, Heraeus penai sp. nov.; H, Heraeus plebejus Stål, 1874; I, Heraeus pulchellus Barber, 1954." figureDoi="http://doi.org/10.5281/zenodo.10128651" httpUri="https://zenodo.org/record/10128651/files/figure.png" pageId="70" pageNumber="99">Fig. 36D</figureCitation>
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). Legs: Coxae, protrochanter, and profemur dark brown, at most with apices paler, meso- and metafemur with a subapical dark band; remain- der of legs pale brown, apex of tibiae, tarsi, and pretarsi darker (
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<figureCitation id="79B7ACB4E965500EFF6C3081FE94EB8C" box="[242,344,1472,1494]" captionStart="Figure 39" captionStartId="70.[144,220,1059,1078]" captionTargetBox="[305,1261,196,1028]" captionTargetId="figure-296@70.[303,1263,194,1030]" captionTargetPageId="70" captionText="Figure 39. Legs: A, Heraeus nicaraguensis sp. nov.; B, Heraeus pacificus Barber, 1925; C, Heraeus pallidinervis sp. nov.; D, Heraeus penai sp. nov.; E, Heraeus plebejus Stål, 1874; F, Heraeus pulchellus Barber, 1954." figureDoi="http://doi.org/10.5281/zenodo.10128659" httpUri="https://zenodo.org/record/10128659/files/figure.png" pageId="70" pageNumber="99">Fig. 39A</figureCitation>
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); setae long and erect, longest on profemur.
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</paragraph>
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<paragraph id="E133B031E965500EFF0E3162FB51EA0E" blockId="70.[144,1421,1059,1108]" pageId="70" pageNumber="99">
|
||
<emphasis id="D3F86C23E965500EFF0E3162FECCEA6C" bold="true" box="[144,256,1059,1078]" pageId="70" pageNumber="99">Figure 39.</emphasis>
|
||
Legs: A,
|
||
<emphasis id="D3F86C23E965500EFEFE3165FD7AEA6D" bold="true" box="[352,694,1059,1079]" pageId="70" pageNumber="99">
|
||
<taxonomicName id="268CCBB2E965500EFEFE3165FDADEA6C" authority="Dellapé & Melo & Henry, 2016" authorityName="Dellapé & Melo & Henry" authorityYear="2016" box="[352,609,1059,1079]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="70" pageNumber="99" phylum="Arthropoda" rank="species" species="nicaraguensis" status="sp. nov.">
|
||
<emphasis id="D3F86C23E965500EFEFE3165FDADEA6C" bold="true" box="[352,609,1059,1079]" italics="true" pageId="70" pageNumber="99">Heraeus nicaraguensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C8CBD158E965500EFDF83165FD7AEA6D" box="[614,694,1060,1079]" pageId="70" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; B,
|
||
<taxonomicName id="268CCBB2E965500EFD7E3165FBDBEA6C" authority="Barber, 1925" authorityName="Barber" authorityYear="1925" box="[736,1047,1059,1079]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="70" pageNumber="99" phylum="Arthropoda" rank="species" species="pacificus">
|
||
<emphasis id="D3F86C23E965500EFD7E3165FC43EA6D" box="[736,911,1060,1079]" italics="true" pageId="70" pageNumber="99">Heraeus pacificus</emphasis>
|
||
<bibRefCitation id="851DCDC0E965500EFC0B3162FBDBEA6C" author="Barber HG" box="[917,1047,1059,1078]" pageId="70" pageNumber="99" pagination="241 - 254" refId="ref75170" refString="Barber HG. 1925. Hemiptera-Heteroptera from the Williams Galapagos expeditions. Zoologica 5: 241 - 254." type="journal article" year="1925">Barber, 1925</bibRefCitation>
|
||
</taxonomicName>
|
||
; C,
|
||
<emphasis id="D3F86C23E965500EFBA03165FA4BEA6D" bold="true" box="[1086,1415,1059,1079]" pageId="70" pageNumber="99">
|
||
<taxonomicName id="268CCBB2E965500EFBA03165FAFDEA6C" authority="Dellapé & Melo & Henry, 2016" authorityName="Dellapé & Melo & Henry" authorityYear="2016" box="[1086,1329,1059,1079]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="70" pageNumber="99" phylum="Arthropoda" rank="species" species="pallidinervis" status="sp. nov.">
|
||
<emphasis id="D3F86C23E965500EFBA03165FAFDEA6C" bold="true" box="[1086,1329,1059,1079]" italics="true" pageId="70" pageNumber="99">Heraeus pallidinervis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C8CBD158E965500EFAA83165FA4BEA6D" box="[1334,1415,1060,1079]" pageId="70" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; D,
|
||
<emphasis id="D3F86C23E965500EFF2F3100FE74EA0E" bold="true" box="[177,440,1089,1108]" pageId="70" pageNumber="99">
|
||
<taxonomicName id="268CCBB2E965500EFF2F3100FE90EA0E" authority="Dellapé & Melo & Henry, 2016" authorityName="Dellapé & Melo & Henry" authorityYear="2016" box="[177,348,1089,1108]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="70" pageNumber="99" phylum="Arthropoda" rank="species" species="penai" status="sp. nov.">
|
||
<emphasis id="D3F86C23E965500EFF2F3100FE90EA0E" bold="true" box="[177,348,1089,1108]" italics="true" pageId="70" pageNumber="99">Heraeus penai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C8CBD158E965500EFEFC3100FE74EA0E" box="[354,440,1089,1108]" pageId="70" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; E,
|
||
<taxonomicName id="268CCBB2E965500EFE793100FCDEEA0E" authority="Stal, 1874" authorityName="Stal" authorityYear="1874" box="[487,786,1088,1108]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="70" pageNumber="99" phylum="Arthropoda" rank="species" species="plebejus">
|
||
<emphasis id="D3F86C23E965500EFE793100FD56EA0E" box="[487,666,1089,1108]" italics="true" pageId="70" pageNumber="99">Heraeus plebejus</emphasis>
|
||
<bibRefCitation id="851DCDC0E965500EFD3D3101FCDEEA0E" author="Stal C" box="[675,786,1088,1108]" pageId="70" pageNumber="99" pagination="1 - 186" refId="ref78329" refString="Stal C. 1874. Enumeratio Lygaeidarum Extraeuropaearum. Svenska Vetenskaps Akademiens Handlingar 12: 1 - 186. Pp. 98 - 170. In Enumeratio Hemipterorum pt. 4 Kongl." type="book chapter" year="1874">Stål, 1874</bibRefCitation>
|
||
</taxonomicName>
|
||
; F,
|
||
<taxonomicName id="268CCBB2E965500EFCA43100FB55EA0E" authority="Barber, 1954" authorityName="Barber" authorityYear="1954" box="[826,1177,1089,1108]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="70" pageNumber="99" phylum="Arthropoda" rank="species" species="pulchellus">
|
||
<emphasis id="D3F86C23E965500EFCA43100FBC8EA0E" box="[826,1028,1089,1108]" italics="true" pageId="70" pageNumber="99">Heraeus pulchellus</emphasis>
|
||
Barber, 1954
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E133B031E965500EFF3630BCFF25E952" blockId="70.[144,759,1166,1800]" pageId="70" pageNumber="99">
|
||
<emphasis id="D3F86C23E965500EFF3630BCFED4E848" box="[168,280,1533,1554]" italics="true" pageId="70" pageNumber="99">Abdomen:</emphasis>
|
||
Dark brown, with abundant short recumbent setae. Male genitalia: Pygophore (
|
||
<figureCitation id="79B7ACB4E965500EFDF6335DFD22E868" box="[616,750,1564,1586]" captionStart="Figure 40" captionStartId="71.[164,243,1714,1733]" captionTargetBox="[254,1349,195,1677]" captionTargetId="figure-0@71.[250,1356,192,1684]" captionTargetPageId="71" captionText="Figure 40. Male genitalia. Heraeus nicaraguensis sp. nov.: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus pacificus Barber, 1925: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus pallidinervis sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus penai sp. nov.: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." figureDoi="http://doi.org/10.5281/zenodo.10128661" httpUri="https://zenodo.org/record/10128661/files/figure.png" pageId="70" pageNumber="99">Fig. 40A, B</figureCitation>
|
||
) rounded, apex with a dorsally directed protuberance, anterior margin of dorsal aperture slightly rounded. Parameres:
|
||
<figureCitation id="79B7ACB4E965500EFE833339FE41E8D7" box="[285,397,1656,1678]" captionStart="Figure 40" captionStartId="71.[164,243,1714,1733]" captionTargetBox="[254,1349,195,1677]" captionTargetId="figure-0@71.[250,1356,192,1684]" captionTargetPageId="71" captionText="Figure 40. Male genitalia. Heraeus nicaraguensis sp. nov.: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus pacificus Barber, 1925: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus pallidinervis sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus penai sp. nov.: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." figureDoi="http://doi.org/10.5281/zenodo.10128661" httpUri="https://zenodo.org/record/10128661/files/figure.png" pageId="70" pageNumber="99">Figure 40</figureCitation>
|
||
(C, D). Aedeagus: spinose; conjunctiva with spines laterally beneath the ejaculatory reservoir; vesica with spines laterally and sclerotized lobes with spines distally;
|
||
<emphasis id="D3F86C23E965500EFE253395FD41E8B3" box="[443,653,1748,1769]" italics="true" pageId="70" pageNumber="99">processus gonopori</emphasis>
|
||
long and slender.
|
||
</paragraph>
|
||
<paragraph id="E133B031E965500EFF0E3278FED7E914" blockId="70.[144,562,1849,1902]" box="[144,283,1849,1870]" pageId="70" pageNumber="99">
|
||
<emphasis id="D3F86C23E965500EFF0E3278FED7E914" box="[144,283,1849,1870]" italics="true" pageId="70" pageNumber="99">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E133B031E965500EFF0E3219FC51EAF9" blockId="70.[144,562,1849,1902]" box="[144,925,1166,1902]" lastBlockId="70.[806,1421,1166,1280]" pageId="70" pageNumber="99">
|
||
<collectingCountry id="999BF0A1E965500EFF0E3219FEC0E934" box="[144,268,1880,1902]" name="Costa Rica" pageId="70" pageNumber="99">Costa Rica</collectingCountry>
|
||
and
|
||
<collectingCountry id="999BF0A1E965500EFED53219FE0EE934" box="[331,450,1880,1902]" name="Nicaragua" pageId="70" pageNumber="99">Nicaragua</collectingCountry>
|
||
(
|
||
<figureCitation id="79B7ACB4E965500EFE4A3219FDE9E934" box="[468,549,1880,1902]" captionStart="Figure 38" captionStartId="68.[144,222,1166,1185]" captionTargetBox="[144,1424,195,1136]" captionTargetId="figure-238@68.[144,1424,195,1136]" captionTargetPageId="68" captionText="Figure 38. Distributional map of Heraeus mexicanus sp. nov., Heraeus nicaraguensis sp. nov., and Heraeus pacificus Barber, 1925." figureDoi="http://doi.org/10.5281/zenodo.10128655" httpUri="https://zenodo.org/record/10128655/files/figure.png" pageId="70" pageNumber="99">Fig. 38</figureCitation>
|
||
).
|
||
<emphasis id="D3F86C23E965500EFCB831CFFC51EAF9" box="[806,925,1166,1187]" italics="true" pageId="70" pageNumber="99">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E133B031E965500EFCB831ECFB63EB5A" blockId="70.[806,1421,1166,1280]" pageId="70" pageNumber="99">
|
||
<materialsCitation id="51E4BA6CE965500EFCB831ECFB67EB5A" country="Nicaragua" location="The" municipality="Central American" pageId="70" pageNumber="99" specimenCount="1" typeStatus="holotype">
|
||
<location id="E453E6EAE965500EFCB831ECFC9EEA99" LSID="urn:lsid:plazi:treatment:69250127E966500EFC3C373AFAAEE876:E453E6EAE965500EFCB831ECFC9EEA99" box="[806,850,1197,1219]" country="Nicaragua" municipality="Central American" name="The" pageId="70" pageNumber="99">The</location>
|
||
specific epithet ‘
|
||
<taxonomicName id="268CCBB2E965500EFB8A31EFFB7EEA99" authorityName="Dellapé & Melo & Henry" authorityYear="2016" box="[1044,1202,1198,1219]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="70" pageNumber="99" phylum="Arthropoda" rank="species" species="nicaraguensis">
|
||
<emphasis id="D3F86C23E965500EFB8A31EFFB7EEA99" box="[1044,1202,1198,1219]" italics="true" pageId="70" pageNumber="99">nicaraguensis</emphasis>
|
||
</taxonomicName>
|
||
’ is an adjective referring to
|
||
<collectingCountry id="999BF0A1E965500EFC3F318AFBD7EABB" box="[929,1051,1227,1249]" name="Nicaragua" pageId="70" pageNumber="99">Nicaragua</collectingCountry>
|
||
, the
|
||
<collectingMunicipality id="01572A4BE965500EFBC5318AFAE6EABB" box="[1115,1322,1227,1249]" pageId="70" pageNumber="99">Central American</collectingMunicipality>
|
||
country where the
|
||
<typeStatus id="3E370E93E965500EFC3A31ABFBC9EB5A" box="[932,1029,1258,1280]" pageId="70" pageNumber="99" type="holotype">holotype</typeStatus>
|
||
was collected
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E133B031E965500EFCB83009FC0FEB06" blockId="70.[806,1420,1351,1435]" box="[806,963,1351,1373]" pageId="70" pageNumber="99">
|
||
<emphasis id="D3F86C23E965500EFCB83009FC0FEB06" box="[806,963,1351,1373]" italics="true" pageId="70" pageNumber="99">Type material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E133B031E965500EFCB83027FB56EBC1" blockId="70.[806,1420,1351,1435]" pageId="70" pageNumber="99">
|
||
<materialsCitation id="51E4BA6CE965500EFCB83027FB56EBC1" collectingDate="1971-07" collectionCode="USNM" collectorName="J. Maldonado" country="Nicaragua" location="Puerto Cabezas" pageId="70" pageNumber="99" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="D3F86C23E965500EFCB83027FC58EB21" box="[806,916,1382,1403]" italics="true" pageId="70" pageNumber="99">
|
||
<typeStatus id="3E370E93E965500EFCB83027FC42EB21" box="[806,910,1382,1403]" pageId="70" pageNumber="99" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="F78A7BB8E965500EFC3F3024FC76EB21" box="[929,954,1381,1403]" pageId="70" pageNumber="99" type="male">♂</specimenCount>
|
||
,
|
||
<emphasis id="D3F86C23E965500EFC533027FBB5EB26" bold="true" box="[973,1145,1382,1404]" pageId="70" pageNumber="99">
|
||
<collectingCountry id="999BF0A1E965500EFC533027FBB5EB26" box="[973,1145,1382,1404]" name="Nicaragua" pageId="70" pageNumber="99">NICARAGUA</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<location id="E453E6EAE965500EFB133027FA80EB26" LSID="urn:lsid:plazi:treatment:69250127E966500EFC3C373AFAAEE876:E453E6EAE965500EFB133027FA80EB26" box="[1165,1356,1382,1404]" country="Nicaragua" name="Puerto Cabezas" pageId="70" pageNumber="99">Puerto Cabezas</location>
|
||
,
|
||
<date id="953296F1E965500EFAC33027FC93EBC0" pageId="70" pageNumber="99" value="1971-07">
|
||
<collectingDate id="85766F19E965500EFAC33027FC93EBC0" pageId="70" pageNumber="99" value="1971-07">VII- 1971</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="4C79D5E7E965500EFCF030C4FBCBEBC0" box="[878,1031,1412,1434]" pageId="70" pageNumber="99">J. Maldonado</collectorName>
|
||
C. (
|
||
<collectionCode id="879D28F4E965500EFBA530C4FB43EBC1" box="[1083,1167,1413,1435]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="70" pageNumber="99" type="Museum">USNM</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="E133B031E965500EFCB830FAFAAEE876" blockId="70.[806,1421,1466,1580]" pageId="70" pageNumber="99">
|
||
<materialsCitation id="51E4BA6CE965500EFCB830FAFA45EB8B" box="[806,1417,1466,1489]" collectingDate="1971-07" collectionCode="USNM" collectorName="J. Maldonado" country="Costa Rica" location="Puerto Cabezas" pageId="70" pageNumber="99" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||
<emphasis id="D3F86C23E965500EFCB830FAFC56EB8A" box="[806,922,1467,1488]" italics="true" pageId="70" pageNumber="99">
|
||
<typeStatus id="3E370E93E965500EFCB830FAFC58EB8A" box="[806,916,1467,1488]" pageId="70" pageNumber="99" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="F78A7BB8E965500EFC3F30FAFC0AEB8A" box="[929,966,1466,1488]" pageId="70" pageNumber="99" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="F78A7BB8E965500EFC4B30FAFC3AEB8A" box="[981,1014,1466,1488]" pageId="70" pageNumber="99" type="female">1♀</specimenCount>
|
||
, same data as for holotype (
|
||
<collectionCode id="879D28F4E965500EFAAF30FAFA4EEB8B" box="[1329,1410,1467,1489]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="70" pageNumber="99" type="Museum">USNM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="51E4BA6CE965500EFCA13098FA94E854" collectingDate="1988-05-23" collectionCode="UCMS" collectorName="La Selva & Station & J. O'Donnell" country="Costa Rica" location="Heredia" pageId="70" pageNumber="99" specimenCount="1" specimenCount-female="1" stateProvince="Heredia" typeStatus="paratype">
|
||
<emphasis id="D3F86C23E965500EFCA13098FC20EBB5" bold="true" box="[831,1004,1497,1519]" pageId="70" pageNumber="99">
|
||
<collectingCountry id="999BF0A1E965500EFCA13098FC2AEBB5" box="[831,998,1497,1519]" name="Costa Rica" pageId="70" pageNumber="99">COSTA RICA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<emphasis id="D3F86C23E965500EFC6D3098FB81EBB4" box="[1011,1101,1497,1518]" italics="true" pageId="70" pageNumber="99">
|
||
<collectingRegion id="23487ED3E965500EFC6D3098FB81EBB4" box="[1011,1101,1497,1518]" country="Costa Rica" name="Heredia" pageId="70" pageNumber="99">Heredia</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="F78A7BB8E965500EFBC23098FBB2EBB5" box="[1116,1150,1497,1519]" pageId="70" pageNumber="99" type="female">1♀</specimenCount>
|
||
,
|
||
<collectorName id="4C79D5E7E965500EFB133098FB3DEBB5" box="[1165,1265,1497,1519]" pageId="70" pageNumber="99">La Selva</collectorName>
|
||
OTS
|
||
<collectorName id="4C79D5E7E965500EFAAA3098FA45EBB5" box="[1332,1417,1497,1519]" pageId="70" pageNumber="99">Station</collectorName>
|
||
, 22/
|
||
<date id="953296F1E965500EFCFD30B9FC1AE857" box="[867,982,1528,1549]" pageId="70" pageNumber="99" value="1988-05-23">
|
||
<collectingDate id="85766F19E965500EFCFD30B9FC1AE857" box="[867,982,1528,1549]" pageId="70" pageNumber="99" value="1988-05-23">23-V-1988</collectingDate>
|
||
</date>
|
||
, blacklight,
|
||
<collectorName id="4C79D5E7E965500EFBFD30B9FB3CE854" box="[1123,1264,1528,1550]" pageId="70" pageNumber="99">J. O’Donnell</collectorName>
|
||
(
|
||
<collectionCode id="879D28F4E965500EFA9E30B9FA9DE854" box="[1280,1361,1528,1550]" country="USA" httpUri="http://grbio.org/cool/gg3r-fyna" name="University of Connecticut Biodiversity Research Collections" pageId="70" pageNumber="99" type="University or college">UCMS</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="51E4BA6CE965500EFAFA30B9FA92E876" collectingDate="1988-05-25" collectionCode="UCMS" collectorName="J. O'Donnell" country="Costa Rica" pageId="70" pageNumber="99" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="F78A7BB8E965500EFAFA30B9FA4BE857" box="[1380,1415,1527,1549]" pageId="70" pageNumber="99" type="female">1♀</specimenCount>
|
||
, 24/
|
||
<date id="953296F1E965500EFCFD3357FC1AE876" box="[867,982,1558,1580]" pageId="70" pageNumber="99" value="1988-05-25">
|
||
<collectingDate id="85766F19E965500EFCFD3357FC1AE876" box="[867,982,1558,1580]" pageId="70" pageNumber="99" value="1988-05-25">25-V-1988</collectingDate>
|
||
</date>
|
||
, blacklight,
|
||
<collectorName id="4C79D5E7E965500EFBF83356FB38E876" box="[1126,1268,1558,1580]" pageId="70" pageNumber="99">J. O’Donnell</collectorName>
|
||
(
|
||
<collectionCode id="879D28F4E965500EFA9B3357FA9AE876" box="[1285,1366,1558,1580]" country="USA" httpUri="http://grbio.org/cool/gg3r-fyna" name="University of Connecticut Biodiversity Research Collections" pageId="70" pageNumber="99" type="University or college">UCMS</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |