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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1073.71843" ID-GBIF-Dataset="5161ac99-ec0f-4443-ac73-a9d016a9383d" ID-PMC="PMC8648685" ID-Pensoft-Pub="1313-2970-1073-55" ID-Pensoft-UUID="F401EBF007E0519AB4B8EC43D0F5EFE4" ID-PubMed="34949951" ID-ZooBank="18D5AF2786E54D21BCC527D09FB384DA" ModsDocID="1313-2970-1073-55" checkinTime="1638245621972" checkinUser="pensoft" docAuthor="Barr, Cheryl B. &amp; Shepard, William D." docDate="2021" docId="2672ACF7B63C5F11A342AD0AEDC2B95A" docLanguage="en" docName="ZooKeys 1073: 55-117" docOrigin="ZooKeys 1073" docPubDate="2021-11-29" docSource="http://dx.doi.org/10.3897/zookeys.1073.71843" docTitle="Stetholus longipennis Barr &amp; Shepard 2021, sp. nov." docType="treatment" docUuid="4A0A3280-EACD-42E2-9851-7BB81665144D" docUuidSource="ZooBank" docVersion="4" id="F401EBF007E0519AB4B8EC43D0F5EFE4" lastPageNumber="55" masterDocId="F401EBF007E0519AB4B8EC43D0F5EFE4" masterDocTitle="A review of the Larainae of Australia with description of seven new species and the new genus Australara (Coleoptera, Byrrhoidea, Elmidae)" masterLastPageNumber="117" masterPageNumber="55" pageNumber="55" updateTime="1668151180298" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of the Larainae of Australia with description of seven new species and the new genus Australara (Coleoptera, Byrrhoidea, Elmidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Barr, Cheryl B.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6707-4301</mods:nameIdentifier>
<mods:affiliation>Essig Museum of Entomology, 1101 Life Sciences Bldg. # 4780, University of California, Berkeley, CA 94720 USA</mods:affiliation>
<mods:nameIdentifier type="email">cbarr@berkeley.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shepard, William D.</mods:namePart>
<mods:affiliation>Essig Museum of Entomology, 1101 Life Sciences Bldg. # 4780, University of California, Berkeley, CA 94720 USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-11-29</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1073</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>55</mods:start>
<mods:end>117</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1073.71843</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1073.71843</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1073-55</mods:identifier>
<mods:identifier type="ZooBank">18D5AF2786E54D21BCC527D09FB384DA</mods:identifier>
<mods:identifier type="Pensoft-UUID">F401EBF007E0519AB4B8EC43D0F5EFE4</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="190874587" LSID="urn:lsid:zoobank.org:act:4A0A3280-EACD-42E2-9851-7BB81665144D" httpUri="http://treatment.plazi.org/id/2672ACF7B63C5F11A342AD0AEDC2B95A" lastPageNumber="55" pageId="0" pageNumber="55">
<subSubSection pageId="0" pageNumber="55" type="nomenclature">
<paragraph pageId="0" pageNumber="55">
<taxonomicName LSID="http://zoobank.org/4A0A3280-EACD-42E2-9851-7BB81665144D" authority="Barr &amp; Shepard, 2021" authorityName="Barr &amp; Shepard" authorityYear="2021" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus longipennis" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="longipennis" status="sp. nov.">Stetholus longipennis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="55">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="description">
<paragraph pageId="0" pageNumber="55">
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. Geographical distribution of species records 1 Australara glaisteri gen. nov., sp. nov. 2 Hydora laticeps 3 Ovolara australis 4 Ovolara lawrencei sp. nov. 5 Ovolara leai 6 Ovolara monteithi sp. nov. 7 Potamophilinus papuanus 8 Stetholus carinatus sp. nov. 9 Stetholus elongatus 10 Stetholus longipennis sp. nov. 11 Stetholus metatibialis sp. nov. 12 Stetholus woronora sp. nov." figureDoi="10.3897/zookeys.1073.71843.figures1-12" httpUri="https://binary.pensoft.net/fig/615728" pageId="0" pageNumber="55">Figs 10</figureCitation>
<figureCitation captionStart="Figure 27" captionStartId="F10" captionText="Figure 27. Ovolara lawrencei sp. nov., type locality, and Stetholus longipennis sp. nov., collection locality: Emerald Creek, east of Mareeba, QLD, Australia (photograph courtesy of David Rentz, James Cook University, Smithfield, QLD)." figureDoi="10.3897/zookeys.1073.71843.figure27" httpUri="https://binary.pensoft.net/fig/615737" pageId="0" pageNumber="55">, 27</figureCitation>
<figureCitation captionStart="Figures 38, 39" captionStartId="F16" captionText="Figures 38, 39. Stetholus longipennis sp. nov., male 38 habitus, 4.2 mm long A dorsal B ventral 39 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures38-39" httpUri="https://binary.pensoft.net/fig/615743" pageId="0" pageNumber="55">, 38</figureCitation>
<figureCitation captionStart="Figures 38, 39" captionStartId="F16" captionText="Figures 38, 39. Stetholus longipennis sp. nov., male 38 habitus, 4.2 mm long A dorsal B ventral 39 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures38-39" httpUri="https://binary.pensoft.net/fig/615743" pageId="0" pageNumber="55">, 39</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="type locality">
<paragraph pageId="0" pageNumber="55">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="55">
Hunters Creek north of Mount Molloy;
<geoCoordinate degrees="16.6324" direction="south" orientation="latitude" precision="5" value="-16.6324">16.6324° S</geoCoordinate>
,
<geoCoordinate degrees="145. 3254" direction="east" orientation="longitude" precision="5" value="145.3254">145. 3254° E</geoCoordinate>
; north Queensland, Australia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="type material">
<paragraph pageId="0" pageNumber="55">Type material.</paragraph>
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">
<emphasis italics="true" pageId="0" pageNumber="55">Holotype</emphasis>
male.
</emphasis>
&quot;AUSTRALIA: Queensland / 5 km N Mount Molloy / 17 I 2001 / Hunters Creek / S
<geoCoordinate degrees="16" direction="east" minutes="38" orientation="longitude" precision="15" seconds="00" value="16.633333">
16°38
<normalizedToken originalValue="00”">'00&quot;</normalizedToken>
E
</geoCoordinate>
145°19
<normalizedToken originalValue="27”">'27&quot;</normalizedToken>
(WDS-A-1368 on reverse) // William D. / Shepard, leg. // HOLOTYPE /
<taxonomicName authorityName="Carter &amp; Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">Stetholus</taxonomicName>
/
<taxonomicName authorityName="Barr &amp; Shepard" authorityYear="2021" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus longipennis" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="longipennis">Stetholus longipennis</taxonomicName>
/ Barr &amp; Shepard&quot; [red label, handwritten]. Dry pinned. Deposited in the Australian National Insect Collection, Canberra; ANIC Database Number 25-077642.
<emphasis bold="true" pageId="0" pageNumber="55">
<emphasis italics="true" pageId="0" pageNumber="55">Paratypes</emphasis>
(108).
</emphasis>
Same data as for holotype (1&gt;AM, 1 ANIC, 13 EMEC, 1 QM); AUSTRALIA: no. QLD / Hunters Creek at Hwy. 44 / 5 rd. km. N Mount Molloy /
<geoCoordinate degrees="16" direction="south" minutes="38" orientation="latitude" precision="15" seconds="00" value="-16.633333">
16°38
<normalizedToken originalValue="00”">'00&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate degrees="145" direction="east" minutes="19" orientation="longitude" precision="15" seconds="27" value="145.32417">
145°19
<normalizedToken originalValue="27”">'27&quot;</normalizedToken>
E
</geoCoordinate>
/ 17-I-2001, coll. C. B. Barr (5 EMEC, 1 QM); AUSTRALIA: no. QLD / Bushy Creek at Hwy. 44 / just W of Julatten /
<geoCoordinate degrees="16" direction="south" minutes="36" orientation="latitude" precision="15" seconds="40" value="-16.61111">
16°36
<normalizedToken originalValue="40”">'40&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate degrees="45" direction="east" minutes="20" orientation="longitude" precision="15" seconds="10" value="45.33611">
45°20
<normalizedToken originalValue="10”">'10&quot;</normalizedToken>
E
</geoCoordinate>
/ 17-I-2001, coll. C.B.Barr (1 EMEC); AUSTRALIA: no. QLD / Emerald Creek at Hwy. 1 / E of Mareeba /
<geoCoordinate degrees="16" direction="south" minutes="59" orientation="latitude" precision="15" seconds="12" value="-16.986668">
16°59
<normalizedToken originalValue="12”">'12&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate degrees="145" direction="east" minutes="28" orientation="longitude" precision="15" seconds="21" value="145.47249">
145°28
<normalizedToken originalValue="21”">'21&quot;</normalizedToken>
E
</geoCoordinate>
/ 17-I-2001, coll. C.B. Barr (6 EMEC); AUSTRALIA: Queensland / Emerald Creek Store / 17 I 2001 / Emerald Creek / S
<geoCoordinate degrees="16" direction="east" minutes="59" orientation="longitude" precision="15" seconds="12" value="16.986668">16°59'12&quot; E</geoCoordinate>
145°28'21&quot; (WDS-A-1369 on reverse) // William D. / Shepard, leg. (1&gt;AM, 9 EMEC); AUSTRALIA: no. QLD / Freshwater, Freshwater / Cr. at Ryan Weare Park /
<geoCoordinate degrees="16" direction="south" minutes="53" orientation="latitude" precision="15" seconds="13" value="-16.886944">
16°53
<normalizedToken originalValue="13”">'13&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate degrees="145" direction="east" minutes="42" orientation="longitude" precision="15" seconds="05" value="145.70139">
145°42
<normalizedToken originalValue="05”">'05&quot;</normalizedToken>
E
</geoCoordinate>
/ 18-I-2001, coll. C.B. Barr (6 EMEC); AUSTRALIA: Queensland / Freshwater / 18 I 2001 / Freshwater Creek (WDS-A-1370 on reverse) // William D. / Shepard, leg. (2 EMEC); AUSTRALIA: no. QLD / Mulgrave River at Hwy. 1 / 1 rd. km. S of Gordonvale /
<geoCoordinate degrees="17" direction="south" minutes="06" orientation="latitude" precision="15" seconds="10" value="-17.102777">
17°06
<normalizedToken originalValue="10”">'10&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate degrees="145" direction="east" minutes="47" orientation="longitude" precision="15" seconds="15" value="145.7875">
145°47
<normalizedToken originalValue="15”">'15&quot;</normalizedToken>
E
</geoCoordinate>
/ 18-I-2001, coll. C. B. Barr (5 EMEC, 1 QM); AUSTRALIA: Queensland / 1 km S Gordonvale, 18 I 2001 94 ft / Mulgrave River / (WDS-A-1371 on reverse) // William D. / Shepard, leg. (1&gt;AM, 1 ANIC, 2 EMEC, 1 QM); AUSTRALIA: no. QLD / Fishery Creek at / Hwy. 1, Fishery Falls /
<geoCoordinate degrees="17" direction="south" minutes="11" orientation="latitude" precision="15" seconds="10" value="-17.18611">
17°11
<normalizedToken originalValue="10”">'10&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate degrees="145" direction="east" minutes="53" orientation="longitude" precision="15" seconds="11" value="145.88638">
145°53
<normalizedToken originalValue="11”">'11&quot;</normalizedToken>
E
</geoCoordinate>
/ 18-I-2001, C. B. Barr (6 EMEC); AUSTRALIA: Queensland / Fishery Falls / 18 I 2001 / Fishery Creek (WDS-A-1372 on reverse) // William D. / Shepard, leg. (1&gt;AM, 1 ANIC, 2 EMEC); QLD. Babinda / Apr. 1946 / J.G.Brooks (3 ANIC); 16.03S to 16.05S / 145.28E Cape / Tribulation area / QLD 21-28Mar.1984 / A.Calder &amp; T.Weir // on rocks / in stream (1 ANIC); 16.03S to 16.05S / 145.28E QLD, Cape / Tribulation area / 1-11 May 1992 / J.F.Lawrence // on rocks / in stream (1 ANIC); same data as for preceding // Genitalia prep. / HS-243 ♂ / A.Calder 1997 (1 ANIC); same data as for preceding // Genitalia prep. / HS-274 ♀ / A.Calder 1997 (1 ANIC); QLD. Cardstone / 23 Jan. 1965 / J.G.Brooks / at light (4 ANIC); same data as for preceding // Genitalia prep. / HS-307 ♀ / A.Calder 1999 (1 ANIC); Cardstone, N.Q. / 23.i.65. J.G. &amp; / J.A.G.Brooks // Genitalia prep. / HS-241 ♂ / A.Calder 1997 (1 ANIC); Crystal Cascades / Via Cairns, N.Qld. / 22.xii.1964. / G. Monteith // EX UQIC / DONATED / 2011 (7 QM); same locality / H.A.Rose. (1 QM); Crystal Cascades / Cairns, N.Qld. / 30.xii.1963. / G. Monteith // EX UQIC / DONATED / 2011 (1 QM); QLD. Gordonvale / Apr. 1946 / J.G.Brooks // J. G. Brooks / Bequest, 1976 (4 ANIC); Little Mulgrave R. / Gordonvale, N.Q. / Apr. 1946 / J. G. Brooks // Australian Museum / K 579979 (2&gt;AM); Mossman, Q. / 25 Mar 1967 / M.S. Upton (1 ANIC); Mossman Gorge / Via Mossman, N.Qld. / 25-26.xii.1964. / G. Monteith // EX UQIC / DONATED / 2011 (2 QM); Mt. Molloy, Q. / Station Creek, at light / 30.xii.69 J.G.Brooks (1 ANIC); 32km S [N?] of Ravenshoe, Q. / (17.38S, 145.29E) /
<taxonomicName genus="K." lsidName="K." pageId="0" pageNumber="55" rank="genus">K.</taxonomicName>
Hyde // 12.ii.1966, Genitalia prep. / HS-239 ♂ / A.Calder 1997 (1 ANIC); same data as for preceding // Genitalia prep. / HS-240 ♂ / A.Calder 1997 (1 ANIC); Upper Daintree R. / Via Daintree, / 27.xii.1964. N.Qld. / G. Monteith // EX UQIC / DONATED / 2011 (3 QM); Upper Finch Hatton / Ck., Via Finch / Hatton, N.Qld. / 3.i.1965. / G. Monteith // EX UQIC / DONATED / 2011 (1 QM); Upper Mulgrave River, / 30.iv.1970, N.Qld. / G. B. Monteith // EX UQIC / DONATED / 2011 (2 QM). Paratypes all with the following label: PARATYPE /
<taxonomicName authorityName="Carter &amp; Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Stetholus</emphasis>
</taxonomicName>
/
<taxonomicName authorityName="Barr &amp; Shepard" authorityYear="2021" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus longipennis" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="longipennis">
<emphasis italics="true" pageId="0" pageNumber="55">Stetholus longipennis</emphasis>
</taxonomicName>
/ Barr &amp; Shepard [yellow label, printed].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="materials_examined">
<paragraph pageId="0" pageNumber="55">Other material examined</paragraph>
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">(17).</emphasis>
<materialsCitation collectingDate="1964-03-14" collectingDateMax="1981-11-09" collectingDateMin="1964-03-14" collectorName="Archers Ck., Q., J. G. Brooks, Bellenden Ker Range, NQ, A. N., Voucher No., Cardstone, N. Q., J. A. G. Brooks, Cardstone, Hyde, Henrietta Ck., Palmerston Nat., Pk., N. Qld., G. Monteith, Millstream, Archers, Ck. N. Q. Mt. Garnet, Rd., Genitalia, A. Calder, Mossman Gorge, Via Mossman, N., Mossman, Q., I. F. B. Common, M. S. Upton, Ravenshoe, Q., Spring Ck. N. Q." county="Heberton Rd." determinerName="T. A. Weir" elevation="100" latitude="-17.38" location="Mt. Garnet Rd." longLatPrecision="767" longitude="145.29" municipality="Mt. Garnet" specimenCount="614" specimenCount-female="614">
<collectorName>Archers Ck., Q.</collectorName>
/
<location LSID="urn:lsid:plazi:treatment:2672ACF7B63C5F11A342AD0AEDC2B95A:0A6842E3921E8CD0C92071851E5E0B3E" county="Heberton Rd." latitude="-17.38" longLatPrecision="767" longitude="145.29" municipality="Mt. Garnet" name="Mt. Garnet Rd.">Mt. Garnet Rd.</location>
, /
<collectingDate value="1964-12-28">28.xii.1964</collectingDate>
/
<collectorName>J.G.Brooks</collectorName>
(1 ANIC);
<collectorName>Bellenden Ker Range, NQ</collectorName>
/
<location LSID="urn:lsid:plazi:treatment:2672ACF7B63C5F11A342AD0AEDC2B95A:019428A9D2FE9D9AEDA3336840E3DF16" county="Heberton Rd." latitude="-17.38" longLatPrecision="767" longitude="145.29" municipality="Mt. Garnet" name="Cableway Base Stn">Cableway Base Stn</location>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100m</elevation>
</quantity>
/
<collectingDate value="1981-10-17" valueMax="1981-11-09" valueMin="1981-10-17">17 Oct.-9 Nov. 1981</collectingDate>
/ EARTHWATCH/QLD. MUSEUM / MV light, rainforest //
<collectorName name="AF Newton">A.N.</collectorName>
I.C. /
<taxonomicName authorityName="Carter &amp; Zeck" authorityYear="1929" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="55" rank="order">COLEOPTERA</taxonomicName>
/
<collectorName>Voucher No.</collectorName>
/ 83-0610 [green label] //
<taxonomicName authorityName="Carter &amp; Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">Stetholus</taxonomicName>
/ sp. 1 / det.
<determinerName>T. A. Weir</determinerName>
1983 (1 QM);
<collectorName>Cardstone, N.Q.</collectorName>
/ 23.i.65. J.G. &amp; /
<collectorName>J.A.G.Brooks</collectorName>
(4 ANIC);
<collectorName>Cardstone</collectorName>
QLD /
<collectingDate value="1966-03-10" valueMax="1966-03-13" valueMin="1966-03-10">10-13.iii.1966</collectingDate>
/
<taxonomicName genus="K." lsidName="K." pageId="0" pageNumber="55" rank="genus">K.</taxonomicName>
<collectorName>Hyde</collectorName>
(1 ANIC);
<collectorName>Henrietta Ck.</collectorName>
, /
<collectorName>Palmerston Nat.</collectorName>
/
<collectorName>Pk., N.Qld.</collectorName>
/
<collectingDate value="1964-12-29">29.xii.1964</collectingDate>
. /
<collectorName>G. Monteith</collectorName>
// EX UQIC / DONATED / 2011 (1 QM);
<collectorName>Millstream</collectorName>
at
<collectorName>Archers</collectorName>
/
<collectorName>
Ck. N. Q.
<collectingMunicipality>Mt. Garnet</collectingMunicipality>
</collectorName>
/
<collectorName>Rd.</collectorName>
28.xii.64 /
<collectorName>J.G. Brooks</collectorName>
(1 ANIC); same data as for preceding //
<collectorName>Genitalia</collectorName>
prep. / HS-
<specimenCount type="female">306 ♀</specimenCount>
/
<collectorName>A.Calder</collectorName>
1999 (1 ANIC); same data as for preceding //
<collectorName>Genitalia</collectorName>
prep. / HS-
<specimenCount type="female">308 ♀</specimenCount>
/
<collectorName>A.Calder</collectorName>
1999 (1 ANIC);
<collectorName>Mossman Gorge</collectorName>
/
<collectorName>Via Mossman, N.</collectorName>
Qld. /
<collectingDate value="1964-12-25" valueMax="1964-12-26" valueMin="1964-12-25">25-26.xii.1964</collectingDate>
. /
<collectorName>G. Monteith</collectorName>
// EX UQIC / DONATED / 2011 (1 QM); 3 mls.W. of /
<collectorName>Mossman, Q.</collectorName>
/
<collectingDate value="1964-03-14">14 Mar. 1964</collectingDate>
/
<collectorName>I.F.B.Common</collectorName>
/ &amp;
<collectorName>M.S.Upton</collectorName>
(1 ANIC);
<quantity metricMagnitude="4" metricUnit="m" metricValue="3.2" unit="km" value="32.0">32km</quantity>
S [N?] of
<collectorName>Ravenshoe, Q.</collectorName>
/ (
<geoCoordinate degrees="17.38" direction="south" orientation="latitude" precision="555" value="-17.38">17.38S</geoCoordinate>
,
<geoCoordinate degrees="145.29" direction="east" orientation="longitude" precision="555" value="145.29">145.29E</geoCoordinate>
) /
<taxonomicName genus="K." lsidName="K." pageId="0" pageNumber="55" rank="genus">K.</taxonomicName>
Hyde //
<collectingDate value="1966-02-16">16.ii.1966</collectingDate>
(3 ANIC);
<collectorName>Spring Ck. N. Q.</collectorName>
/
<collectingCounty>Heberton Rd.</collectingCounty>
/ 30.xii.64 /
<collectorName>J.G. Brooks</collectorName>
(1 ANIC)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="differential diagnosis">
<paragraph pageId="0" pageNumber="55">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="55">
<taxonomicName authorityName="Barr &amp; Shepard" authorityYear="2021" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus longipennis" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="longipennis">
<emphasis italics="true" pageId="0" pageNumber="55">Stetholus longipennis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figures 38, 39" captionStartId="F16" captionText="Figures 38, 39. Stetholus longipennis sp. nov., male 38 habitus, 4.2 mm long A dorsal B ventral 39 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures38-39" httpUri="https://binary.pensoft.net/fig/615743" pageId="0" pageNumber="55">38</figureCitation>
,
<figureCitation captionStart="Figures 38, 39" captionStartId="F16" captionText="Figures 38, 39. Stetholus longipennis sp. nov., male 38 habitus, 4.2 mm long A dorsal B ventral 39 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures38-39" httpUri="https://binary.pensoft.net/fig/615743" pageId="0" pageNumber="55">39</figureCitation>
) can be distinguished from other species of
<taxonomicName authorityName="Carter &amp; Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Stetholus</emphasis>
</taxonomicName>
(Figs
<figureCitation captionStart="Figures 34, 35" captionStartId="F14" captionText="Figures 34, 35. Stetholus carinatus sp. nov., holotype male 34 habitus, 3.7 mm long A dorsal B ventral 35 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures34-35" httpUri="https://binary.pensoft.net/fig/615741" pageId="0" pageNumber="55">34</figureCitation>
-
<figureCitation captionStart="Figures 36, 37" captionStartId="F15" captionText="Figures 36, 37. Stetholus elongatus 36 female habitus, 5.2 mm long A dorsal B ventral 37 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures36-37" httpUri="https://binary.pensoft.net/fig/615742" pageId="0" pageNumber="55">37</figureCitation>
,
<figureCitation captionStart="Figure 40" captionStartId="F17" captionText="Figure 40. Stetholus metatibialis sp. nov., holotype female, 3.9 mm long A dorsal habitus B ventral habitus C head, frontal view D head, ventral view." figureDoi="10.3897/zookeys.1073.71843.figure40" httpUri="https://binary.pensoft.net/fig/615744" pageId="0" pageNumber="55">40</figureCitation>
-
<figureCitation captionStart="Figures 41, 42" captionStartId="F18" captionText="Figures 41, 42. Stetholus woronora sp. nov., holotype male 41 habitus, 5.2 mm long A dorsal B ventral 42 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures41-42" httpUri="https://binary.pensoft.net/fig/615745" pageId="0" pageNumber="55">42</figureCitation>
) by a combination of the following characters: Length usually shorter than or equal to 4.6 mm; antennae slender, almost moniliform; pronotum smooth or lightly sculptured, sublateral carinae absent; metatibiae entirely setose; male genitalia with penis very slender and much longer than parameres. Conversely,
<taxonomicName lsidName="S. metatibialis" pageId="0" pageNumber="55" rank="species" species="metatibialis">
<emphasis italics="true" pageId="0" pageNumber="55">S. metatibialis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 40" captionStartId="F17" captionText="Figure 40. Stetholus metatibialis sp. nov., holotype female, 3.9 mm long A dorsal habitus B ventral habitus C head, frontal view D head, ventral view." figureDoi="10.3897/zookeys.1073.71843.figure40" httpUri="https://binary.pensoft.net/fig/615744" pageId="0" pageNumber="55">40</figureCitation>
) and
<taxonomicName lsidName="S. carinatus" pageId="0" pageNumber="55" rank="species" species="carinatus">
<emphasis italics="true" pageId="0" pageNumber="55">S. carinatus</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figures 34, 35" captionStartId="F14" captionText="Figures 34, 35. Stetholus carinatus sp. nov., holotype male 34 habitus, 3.7 mm long A dorsal B ventral 35 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures34-35" httpUri="https://binary.pensoft.net/fig/615741" pageId="0" pageNumber="55">34</figureCitation>
) both have long, distinct sublateral pronotal carinae;
<taxonomicName lsidName="S. woronora" pageId="0" pageNumber="55" rank="species" species="woronora">
<emphasis italics="true" pageId="0" pageNumber="55">S. woronora</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figures 41, 42" captionStartId="F18" captionText="Figures 41, 42. Stetholus woronora sp. nov., holotype male 41 habitus, 5.2 mm long A dorsal B ventral 42 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures41-42" httpUri="https://binary.pensoft.net/fig/615745" pageId="0" pageNumber="55">41</figureCitation>
) has very short carinae; and all three have distinctly clavate antennae.
<taxonomicName authorityName="Carter &amp; Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus elongatus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis italics="true" pageId="0" pageNumber="55">Stetholus elongatus</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figures 36, 37" captionStartId="F15" captionText="Figures 36, 37. Stetholus elongatus 36 female habitus, 5.2 mm long A dorsal B ventral 37 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures36-37" httpUri="https://binary.pensoft.net/fig/615742" pageId="0" pageNumber="55">36</figureCitation>
) also lacks pronotal carinae, but is usually longer than 4.6 mm and the antennae are clavate. The male genitalia of
<taxonomicName lsidName="S. longipennis" pageId="0" pageNumber="55" rank="species" species="longipennis">
<emphasis italics="true" pageId="0" pageNumber="55">S. longipennis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figures 38, 39" captionStartId="F16" captionText="Figures 38, 39. Stetholus longipennis sp. nov., male 38 habitus, 4.2 mm long A dorsal B ventral 39 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures38-39" httpUri="https://binary.pensoft.net/fig/615743" pageId="0" pageNumber="55">39</figureCitation>
) are strikingly different from those of other
<taxonomicName authorityName="Carter &amp; Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Stetholus</emphasis>
</taxonomicName>
(Figs
<figureCitation captionStart="Figures 34, 35" captionStartId="F14" captionText="Figures 34, 35. Stetholus carinatus sp. nov., holotype male 34 habitus, 3.7 mm long A dorsal B ventral 35 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures34-35" httpUri="https://binary.pensoft.net/fig/615741" pageId="0" pageNumber="55">35</figureCitation>
,
<figureCitation captionStart="Figures 36, 37" captionStartId="F15" captionText="Figures 36, 37. Stetholus elongatus 36 female habitus, 5.2 mm long A dorsal B ventral 37 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures36-37" httpUri="https://binary.pensoft.net/fig/615742" pageId="0" pageNumber="55">37</figureCitation>
,
<figureCitation captionStart="Figures 41, 42" captionStartId="F18" captionText="Figures 41, 42. Stetholus woronora sp. nov., holotype male 41 habitus, 5.2 mm long A dorsal B ventral 42 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures41-42" httpUri="https://binary.pensoft.net/fig/615745" pageId="0" pageNumber="55">42</figureCitation>
) excluding
<taxonomicName lsidName="S. metatibialis" pageId="0" pageNumber="55" rank="species" species="metatibialis">
<emphasis italics="true" pageId="0" pageNumber="55">S. metatibialis</emphasis>
</taxonomicName>
for which males are currently unknown.
</paragraph>
<caption doi="10.3897/zookeys.1073.71843.figure40" httpUri="https://binary.pensoft.net/fig/615744" pageId="0" pageNumber="55" start="Figure 40" startId="F17">
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">Figure 40.</emphasis>
<taxonomicName authorityName="Barr &amp; Shepard" authorityYear="2021" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus metatibialis" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="metatibialis">
<emphasis italics="true" pageId="0" pageNumber="55">Stetholus metatibialis</emphasis>
</taxonomicName>
sp. nov., holotype female, 3.9 mm long
<emphasis bold="true" pageId="0" pageNumber="55">A</emphasis>
dorsal habitus
<emphasis bold="true" pageId="0" pageNumber="55">B</emphasis>
ventral habitus
<emphasis bold="true" pageId="0" pageNumber="55">C</emphasis>
head, frontal view
<emphasis bold="true" pageId="0" pageNumber="55">D</emphasis>
head, ventral view.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1073.71843.figures41-42" httpUri="https://binary.pensoft.net/fig/615745" pageId="0" pageNumber="55" start="Figures 41, 42" startId="F18">
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">Figures 41, 42.</emphasis>
<taxonomicName authorityName="Barr &amp; Shepard" authorityYear="2021" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus woronora" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="woronora">
<emphasis italics="true" pageId="0" pageNumber="55">Stetholus woronora</emphasis>
</taxonomicName>
sp. nov., holotype male
<emphasis bold="true" pageId="0" pageNumber="55">41</emphasis>
habitus, 5.2 mm long
<emphasis bold="true" pageId="0" pageNumber="55">A</emphasis>
dorsal
<emphasis bold="true" pageId="0" pageNumber="55">B</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="55">42</emphasis>
male genitalia
<emphasis bold="true" pageId="0" pageNumber="55">A</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="55">B</emphasis>
lateral view
<emphasis bold="true" pageId="0" pageNumber="55">C</emphasis>
ventral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="description">
<paragraph pageId="0" pageNumber="55">Description</paragraph>
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">
(n = 127).
<emphasis italics="true" pageId="0" pageNumber="55">Body</emphasis>
</emphasis>
: Size 4.1-4.6 long, 1.6-1.8 wide (n = 21). Color dark brown to black dorsally and ventrally, but appearing lighter due to layer of dense, short, yellow setae; first two antennomeres, trochanters, basal 2/3-3/4 of femora yellow; apical antennomeres, coxae, tibiae, tarsi dark brown.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Head</emphasis>
: Punctures shallow, fine, evenly spaced; setae fine, yellow, recumbent to erect. Vertex with faint V- or U-shaped impression, open anteriorly, extending from near antennal bases to a distinct median impression; frontoclypeal suture arcuate. Antenna with eleven antennomeres; antennomere 1 longest, 3
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, curved; antennomere 2 spherical; both with long, curved setae; antennomere 3 ovoid; antennomeres 4-11 each subspherical, widening slightly towards antennal apex, forming a slender, elongate, almost moniliform, club. Eye finely-faceted, suboval at base, moderately protuberant; dorsal and posteroventral margin with fringe of long, curved, black setae. Clypeus convex, broadly rectangular, emarginate anteriorly, densely setose. Labrum rectangular, longer and slightly narrower than clypeus; setose; anterior margin weakly emarginate, with band of short, yellow setae; lateral margins each with a wide, dense band of long, curved setae. Mandible with two teeth, outermost acute apically, innermost truncate apically; prostheca shelf-like, very thin, apically with coarse, long setae; molar area large, moderately oval, surface striate; lateral edge basally with a partially free lobe with several thin setae. Maxillary palpus black, setose, with four palpomeres; palpomere 1 short, annular, with inner apical border spine-like; palpomere 2 twice as long as wide; palpomere 3 as long as 2, wider apically; palpomere 4 wide, ventral surface with a subcircular, concave, white sensory area angled obliquely from the apex to the base. Labial palpus black, glabrous, with three palpomeres; palpomeres 1 and 2 short, annular; palpomere 3 longer, wider, quadrate, apex truncate with an oval, concave, white sensory area.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Pronotum</emphasis>
: Shape generally trapezoidal, wider than long, widest at base; length 0.8-1.0 mm, width 1.2-1.3 mm; disc with distinct punctures evenly spaced ~ 1 diameter apart, deeper near lateral margins; densely setose, with short, fine, yellow setae and longer, coarser, dark setae. Anterior margin arcuate; anterior angles obsolete; lateral margins bisinuate; posterior angles 90°, blunt, depressed; posterior margin weakly trisinuate. Disc slightly convex; shallow, broadly V-shaped, transverse impression at apical 1/4-1/2; basal 1/2 with two faint to distinct, oblique, lateral depressions and two large, deep prescutellar foveae; pronotal margin inflated posterior to fovea.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Scutellar shield</emphasis>
: Cordate; posterior apex slightly raised; finely setose.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Elytron</emphasis>
: 3.3-3.6 mm long, 0.8-0.9 mm wide. Elytra conjointly ~ 2
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, widest near posterior 1/3; apices together forming a triangular notch between. Humerus prominently inflated, elytral base depressed medially; disc at 1/4-1/3 distance from base with a shallow depression from suture to stria 5. Disc with 10 punctate, weakly impressed striae, intervals flat; punctures very fine, spaced one diameter apart, obsolete apically; accessory basal stria between striae 1 and 2 long; stria 3 ending before apex; striae 4 and 5 joining before apex; striae 10 and 11 joining and ending before apex.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Metathoracic wings</emphasis>
: Macropterous.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Prosternum</emphasis>
: Very short anterior to procoxae; prosternal process moderately narrow, 3
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, margined, with a median longitudinal carina, apex narrowly rounded.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Mesoventrite</emphasis>
: Short; disc convex between mesocoxae with a deep mesoventral cavity to receive prosternal process.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Metaventrite</emphasis>
: Broadly rectangular; disc with a median, round concavity near anterior margin and a wide median depression covering posterior 1/2-2/3; discrimen extending from concavity to posterior margin; metakatepisternal suture distinct; disc with small, shallow, variably spaced punctures mostly obscured by dense, fine, recumbent, yellow setae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Legs</emphasis>
: Of similar lengths; each leg with femur and tibia subequal in length; tarsus slender, with tarsomere 5 slightly longer than tarsomeres 1-4 combined; claws simple, moderately large, sharply acute. Coxae dark brown, metacoxae deeply sulcate; femora yellow, dorsal surfaces of each with a narrow brown stripe, apical 1/4-1/3 dark brown; tibiae dark brown, mesotibiae with posterior surfaces nearly glabrous, shiny; metatibiae entirely setose, weakly arcuate.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Abdomen</emphasis>
: Five convex ventrites, each with a shallow depression near lateral margin; ventrite 1 with a margined, broadly triangular, intercoxal projection; ventrites 2-4 broadly rectangular, with lateral margins each produced to form a small lobe of varying size, largest on ventrites 3 and 4, which clasps the epipleuron; posterior border of ventrite 5 with a broadly rounded apex. Ventrites covered with shallow punctures spaced 1-2 diameters apart, mostly obscured by dense covering of yellow setae; setae longer at median 1/5 of ventrites 3-5.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Aedeagus</emphasis>
: Phallobase lightly sclerotized, open dorsally, fused to short parameres; penis very long, nearly as long as phallobase and parameres together (Fig.
<figureCitation captionStart="Figures 38, 39" captionStartId="F16" captionText="Figures 38, 39. Stetholus longipennis sp. nov., male 38 habitus, 4.2 mm long A dorsal B ventral 39 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures38-39" httpUri="https://binary.pensoft.net/fig/615743" pageId="0" pageNumber="55">39</figureCitation>
). In dorsal view (Fig.
<figureCitation captionStart="Figures 38, 39" captionStartId="F16" captionText="Figures 38, 39. Stetholus longipennis sp. nov., male 38 habitus, 4.2 mm long A dorsal B ventral 39 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures38-39" httpUri="https://binary.pensoft.net/fig/615743" pageId="0" pageNumber="55">39A</figureCitation>
), parameres with lateral margins sinuate, convergent from junction with phallobase to near apex then strongly divergent and curved laterally; median margins nearly parallel at basal 2/3 then strongly divergent; apices acute. Penis beyond paramere tips with lateral margins nearly parallel almost to acute apex; no visible corona; dorsal median longitudinal carina with darker sclerotization present; basal apophyses very long, nearly as long as phallobase, straight, very broad, blunt at tips. In lateral view (Fig.
<figureCitation captionStart="Figures 38, 39" captionStartId="F16" captionText="Figures 38, 39. Stetholus longipennis sp. nov., male 38 habitus, 4.2 mm long A dorsal B ventral 39 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures38-39" httpUri="https://binary.pensoft.net/fig/615743" pageId="0" pageNumber="55">39B</figureCitation>
), paramere triangular, moderately convex dorsally, nearly flat ventrally; apex broadly rounded to truncate with a few, small, irregular teeth; penis curved, tip ventrally directed. Fibula absent.
<emphasis bold="true" pageId="0" pageNumber="55">Ovipositor</emphasis>
: Moderately sclerotized; oval in outline, 2
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; baculum ~ 2
<normalizedToken originalValue="×">x</normalizedToken>
longer than gonocoxite; proximal gonocoxite subrectangular, wide and short; distal gonocoxite narrow and short, ~ 2
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, length equal to proximal gonocoxite length, median and lateral margins arcuate, together separate basally but contiguous medially to apices; stylus very narrow.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="variation">
<paragraph pageId="0" pageNumber="55">Variation.</paragraph>
<paragraph pageId="0" pageNumber="55">The females exhibit minor secondary sexual dimorphism with the lateral elytral margin slightly explanate at the posterior 1/4 laterad of stria 11; in males, stria 11 is just inside the lateral margin, which is not explanate. Measured specimens vary from 4.1-4.6 long and 1.6-1.8 wide (n = 21). The females are slightly larger than the males: females 4.3-4.6 mm long, 1.8 mm wide (n = 8); males 4.1-4.5 mm long, 1.6-1.8 mm wide (n = 13). The width of the prosternal process varies a bit between individuals. Otherwise, except for minor differences in the depth and extent of pronotal impressions, the specimens are quite uniform.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="etymology">
<paragraph pageId="0" pageNumber="55">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="55">
The specific epithet
<emphasis italics="true" pageId="0" pageNumber="55">longipennis</emphasis>
, an adjective in the nominative singular derived from the Latin
<emphasis italics="true" pageId="0" pageNumber="55">longi</emphasis>
(long) plus
<emphasis italics="true" pageId="0" pageNumber="55">pennis</emphasis>
(penis), refers to the male genitalia in which the length of the penis greatly exceeds the length of the parameres (Fig.
<figureCitation captionStart="Figures 38, 39" captionStartId="F16" captionText="Figures 38, 39. Stetholus longipennis sp. nov., male 38 habitus, 4.2 mm long A dorsal B ventral 39 male genitalia A dorsal view B lateral view C ventral view." figureDoi="10.3897/zookeys.1073.71843.figures38-39" httpUri="https://binary.pensoft.net/fig/615743" pageId="0" pageNumber="55">39</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="distribution">
<paragraph pageId="0" pageNumber="55">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="55">
<taxonomicName authorityName="Barr &amp; Shepard" authorityYear="2021" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus longipennis" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="longipennis">
<emphasis italics="true" pageId="0" pageNumber="55">Stetholus longipennis</emphasis>
</taxonomicName>
occurs in north and central Queensland, Australia (Fig.
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. Geographical distribution of species records 1 Australara glaisteri gen. nov., sp. nov. 2 Hydora laticeps 3 Ovolara australis 4 Ovolara lawrencei sp. nov. 5 Ovolara leai 6 Ovolara monteithi sp. nov. 7 Potamophilinus papuanus 8 Stetholus carinatus sp. nov. 9 Stetholus elongatus 10 Stetholus longipennis sp. nov. 11 Stetholus metatibialis sp. nov. 12 Stetholus woronora sp. nov." figureDoi="10.3897/zookeys.1073.71843.figures1-12" httpUri="https://binary.pensoft.net/fig/615728" pageId="0" pageNumber="55">10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="habitat">
<paragraph pageId="0" pageNumber="55">Habitat and behavior.</paragraph>
<paragraph pageId="0" pageNumber="55">
<taxonomicName authorityName="Barr &amp; Shepard" authorityYear="2021" class="Insecta" family="Elmidae" genus="Stetholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stetholus longipennis" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="longipennis">
<emphasis italics="true" pageId="0" pageNumber="55">Stetholus longipennis</emphasis>
</taxonomicName>
was collected by the authors from medium to large sand-bottomed streams with logs and debris, some with boulders, and a small river at elevations ranging from 5-417 m (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F10" captionText="Figure 27. Ovolara lawrencei sp. nov., type locality, and Stetholus longipennis sp. nov., collection locality: Emerald Creek, east of Mareeba, QLD, Australia (photograph courtesy of David Rentz, James Cook University, Smithfield, QLD)." figureDoi="10.3897/zookeys.1073.71843.figure27" httpUri="https://binary.pensoft.net/fig/615737" pageId="0" pageNumber="55">27</figureCitation>
). All had clear water which varied from warm to cool. The beetles were collected from logs and rocks in fast current or rapids, and from a spillway. The adults fly readily from the net, and also have been taken at lights. At the type locality, Hunters Creek, the stream was well-shaded, with many logs and much debris, and many
<taxonomicName lsidName="S. longipennis" pageId="0" pageNumber="55" rank="species" species="longipennis">
<emphasis italics="true" pageId="0" pageNumber="55">S. longipennis</emphasis>
</taxonomicName>
were concentrated on a concrete spillway below a bridge.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="associated byrrhoid taxa">
<paragraph pageId="0" pageNumber="55">Associated byrrhoid taxa.</paragraph>
<paragraph pageId="0" pageNumber="55">
<taxonomicName authorityName="Curtis" authorityYear="1830" class="Insecta" family="Elmidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="family">Elmidae</taxonomicName>
:
<taxonomicName authorityName="LeConte" authorityYear="1861" lsidName="" pageId="0" pageNumber="55" rank="subFamily" subFamily="Larainae">Larainae</taxonomicName>
:
<taxonomicName authorityName="Barr &amp; Shepard" authorityYear="2021" family="Elmidae" genus="Australara" kingdom="Animalia" lsidName="Australara glaisteri" order="Coleoptera" pageId="0" pageNumber="55" rank="species" species="glaisteri">
<emphasis italics="true" pageId="0" pageNumber="55">Australara glaisteri</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName authorityName="Barr &amp; Shepard" authorityYear="2021" class="Insecta" family="Elmidae" genus="Ovolara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ovolara lawrencei" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis italics="true" pageId="0" pageNumber="55">Ovolara lawrencei</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="O. leai" pageId="0" pageNumber="55" rank="species" species="leai">
<emphasis italics="true" pageId="0" pageNumber="55">O. leai</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. monteithi" pageId="0" pageNumber="55" rank="species" species="monteithi">
<emphasis italics="true" pageId="0" pageNumber="55">O. monteithi</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName authorityName="Sato" authorityYear="1973" class="Insecta" family="Elmidae" genus="Potamophilinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamophilinus papuanus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis italics="true" pageId="0" pageNumber="55">Potamophilinus papuanus</emphasis>
</taxonomicName>
;
<taxonomicName authorityName="Curtis" authorityYear="1830" lsidName="" pageId="0" pageNumber="55" rank="subFamily" subFamily="Elminae">Elminae</taxonomicName>
:
<taxonomicName authorityName="Carter &amp; Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Austrolimnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrolimnius" order="Diptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Austrolimnius</emphasis>
</taxonomicName>
spp.,
<taxonomicName baseAuthorityName="Carter" baseAuthorityYear="1926" class="Insecta" family="Elmidae" genus="Graphelmis" kingdom="Animalia" lsidName="Graphelmis pallidipes" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="pallidipes">
<emphasis italics="true" pageId="0" pageNumber="55">Graphelmis pallidipes</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Carter &amp; Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Kingolus" kingdom="Animalia" lsidName="Kingolus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Kingolus</emphasis>
</taxonomicName>
spp.,
<taxonomicName authorityName="Carter &amp; Zeck" authorityYear="1936" class="Insecta" family="Elmidae" genus="Notriolus" kingdom="Animalia" lsidName="Notriolus taylori" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="taylori">
<emphasis italics="true" pageId="0" pageNumber="55">Notriolus taylori</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Carter &amp; Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Notriolus" kingdom="Animalia" lsidName="Notriolus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Notriolus</emphasis>
</taxonomicName>
spp.,
<taxonomicName authorityName="Carter &amp; Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Simsonia" kingdom="Animalia" lsidName="Simsonia" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Simsonia</emphasis>
</taxonomicName>
sp.
<taxonomicName class="Insecta" family="Psephenidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="family">Psephenidae</taxonomicName>
:
<taxonomicName class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon basicollis" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="basicollis">
<emphasis italics="true" pageId="0" pageNumber="55">Sclerocyphon basicollis</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon minimus" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="minimus">
<emphasis italics="true" pageId="0" pageNumber="55">Sclerocyphon minimus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>