treatments-xml/data/4A/A9/D9/4AA9D981254655039E4C77AD3F1D809F.xml

<document ID-DOI="http://dx.doi.org/10.3897/alpento.6.79598" ID-Pensoft-Pub="2535-0889-6-7" ID-Pensoft-UUID="A48E84FA8FF753CF979C8BAEE7950B6D" ID-ZooBank="75DB5DB6C3E54B3C9B5C15722723ED3B" LSID="urn:lsid:zoobank.org:act:CB562F9F-95AD-4537-ACA5-305AA84CB53C" ModsDocID="2535-0889-6-7" checkinTime="1652223128647" checkinUser="pensoft" docAuthor="Balke, Michael, Bilton, David T., Garcia, Mauricio, Viloria, Angel L., Villastrigo, Adrian &amp; Hendrich, Lars" docDate="2022" docId="4AA9D981254655039E4C77AD3F1D809F" docLanguage="en" docName="AlpEntomol 6: 7-12" docOrigin="Alpine Entomology 6" docPubDate="2022-05-10" docSource="http://dx.doi.org/10.3897/alpento.6.79598" docTitle="Liodessus meridensis Balke &amp; Bilton &amp; García &amp; Viloria &amp; Villastrigo &amp; Hendrich 2022, sp. nov." docType="treatment" docUuid="CB562F9F-95AD-4537-ACA5-305AA84CB53C" docUuidSource="ZooBank" docVersion="1" id="A48E84FA8FF753CF979C8BAEE7950B6D" lastPageNumber="7" masterDocId="A48E84FA8FF753CF979C8BAEE7950B6D" masterDocTitle="Two new high Andean species of Liodessus diving beetles from Venezuela (Coleoptera, Dytiscidae, Bidessini)" masterLastPageNumber="12" masterPageNumber="7" pageId="0" pageNumber="7" updateTime="1652223128647" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Two new high Andean species of Liodessus diving beetles from Venezuela (Coleoptera, Dytiscidae, Bidessini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Balke, Michael</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
<mods:affiliation>SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany and GeoBioCenter, Ludwig Maximilians University, Munich, Germany</mods:affiliation>
<mods:nameIdentifier type="email">balke.m@snsb.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bilton, David T.</mods:namePart>
<mods:affiliation>School of Biological and Marine Sciences, University of Plymouth, Drake Circus, Plymouth, UK &amp; Department of Zoology, University of Johannesburg, PO Box 524, Auckland Park, 2006 Johannesburg, South Africa</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Garcia, Mauricio</mods:namePart>
<mods:affiliation>Centro de Investigaciones Biologicas, Facultad de Humanidades y Educacion, Universidad del Zulia, Apartado 526, Maracaibo A- 4001, Estado Zulia, Venezuela</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Viloria, Angel L.</mods:namePart>
<mods:affiliation>Centro de Ecologia, Instituto Venezolano de Investigaciones Cientificas (IVIC), km 11 carretera Panamericana, Altos de Pipe, edo. Miranda 1204, Venezuela</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Villastrigo, Adrian</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3531-0821</mods:nameIdentifier>
<mods:affiliation>SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hendrich, Lars</mods:namePart>
<mods:affiliation>SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany</mods:affiliation>
</mods:name>
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<mods:titleInfo>
<mods:title>Alpine Entomology</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-05-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>6</mods:number>
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<mods:extent unit="page">
<mods:start>7</mods:start>
<mods:end>12</mods:end>
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<mods:url>http://dx.doi.org/10.3897/alpento.6.79598</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/alpento.6.79598</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:CB562F9F-95AD-4537-ACA5-305AA84CB53C" httpUri="http://treatment.plazi.org/id/4AA9D981254655039E4C77AD3F1D809F" lastPageNumber="7" pageId="0" pageNumber="7">
<subSubSection pageId="0" pageNumber="7" type="nomenclature">
<paragraph pageId="0" pageNumber="7">
<taxonomicName LSID="http://zoobank.org/CB562F9F-95AD-4537-ACA5-305AA84CB53C" authority="Balke &amp; Bilton &amp; García &amp; Viloria &amp; Villastrigo &amp; Hendrich, 2022" authorityName="Balke &amp; Bilton &amp; García &amp; Viloria &amp; Villastrigo &amp; Hendrich" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus meridensis" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="species" species="meridensis" status="sp. nov.">Liodessus meridensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="7">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="7" type="description">
<paragraph pageId="0" pageNumber="7">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Liodessus spp. nov. L. meridensis (A); L. venezuelanus (B). Scale bar: 2 mm." figureDoi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7">Figs 1A</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Liodessus spp. nov. Detail of head in dorsal view. L. meridensis (A, B); L. venezuelanus (C, D)." figureDoi="10.3897/alpento.6.79598.figure2" httpUri="https://binary.pensoft.net/fig/682025" pageId="0" pageNumber="7">, 2A, B</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Liodessus spp. nov. Median lobe ventral and lateral views; parameres outer and inner surfaces, L. meridensis (A); L. venezuelanus (B). Scale bar: 0.5 mm." figureDoi="10.3897/alpento.6.79598.figure3" httpUri="https://binary.pensoft.net/fig/682026" pageId="0" pageNumber="7">, 3A</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitat of Liodessus spp. nov. near Gavidia, Venenzuela, where both new species were collected (collecting event VZ 10 - 0123 - 03 A). Overview (A); detail of shallow water at the edge (B)." figureDoi="10.3897/alpento.6.79598.figure4" httpUri="https://binary.pensoft.net/fig/682027" pageId="0" pageNumber="7">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="7" type="type locality">
<paragraph pageId="0" pageNumber="7">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="7">
Laguna de 
<normalizedToken originalValue="Mucabají">Mucabaji</normalizedToken>
, 
<normalizedToken originalValue="Mérida">Merida</normalizedToken>
, Venezuela.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="7" type="type material">
<paragraph pageId="0" pageNumber="7">Type material.</paragraph>
<paragraph pageId="0" pageNumber="7">
<emphasis bold="true" italics="true" pageId="0" pageNumber="7">Holotype</emphasis>
: Male: Venezuela; 
<normalizedToken originalValue="Mérida">Merida</normalizedToken>
, Laguna de 
<normalizedToken originalValue="Mucabají">Mucabaji</normalizedToken>
, 3,500 m, i.2004, 8.7964, -70.8255, 
<normalizedToken originalValue="García">Garcia</normalizedToken>
&amp; Balke (MIZA). 
<emphasis bold="true" italics="true" pageId="0" pageNumber="7">Paratypes</emphasis>
: 13 exs same label data as holotype (MIZA, ZSM); 47 exs Venezuela: 
<normalizedToken originalValue="Mérida">Merida</normalizedToken>
, ca. 5 km E Gavidia, 3,600 m, 23.I.2012, 
<geoCoordinate degrees="8.6895" direction="north" orientation="latitude" precision="5" value="8.6895">8.6895</geoCoordinate>
, 
<geoCoordinate degrees="70.8817" direction="west" orientation="longitude" precision="5" value="-70.8817">-70.8817</geoCoordinate>
, Short, Arias &amp; Gustafson, lagoon margin, VZ12-0123-03A (MIZA, SEMC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="7" type="description">
<paragraph pageId="0" pageNumber="7">Description.</paragraph>
<paragraph pageId="0" pageNumber="7">
Habitus with only slight discontinuity between pronotum and elytron (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Liodessus spp. nov. L. meridensis (A); L. venezuelanus (B). Scale bar: 2 mm." figureDoi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7">1A</figureCitation>
), pronotum widest before base and more parallel sided laterally (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Liodessus spp. nov. L. meridensis (A); L. venezuelanus (B). Scale bar: 2 mm." figureDoi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7">1A</figureCitation>
). Total length (TL) 2.1 mm; length without head (TL-h) 1.9 mm; maximum width (MW) 0.9 mm.
</paragraph>
<caption doi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="7">
<emphasis bold="true" pageId="0" pageNumber="7">Figure 1.</emphasis>
Dorsal habitus of 
<taxonomicName authorityName="Guignot" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="7">Liodessus</emphasis>
</taxonomicName>
spp. nov. 
<taxonomicName genus="L." lsidName="L. meridensis" pageId="0" pageNumber="7" rank="species" species="meridensis">
<emphasis italics="true" pageId="0" pageNumber="7">L. meridensis</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="7">A</emphasis>
); 
<taxonomicName genus="L." lsidName="L. venezuelanus" pageId="0" pageNumber="7" rank="species" species="venezuelanus">
<emphasis italics="true" pageId="0" pageNumber="7">L. venezuelanus</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="7">B</emphasis>
). Scale bar: 2 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="7">
<emphasis bold="true" pageId="0" pageNumber="7">
<emphasis italics="true" pageId="0" pageNumber="7">Color</emphasis>
.
</emphasis>
Very dark brown to blackish dorsally and ventrally, basal antennomeres, base of meso- and metatibia contrastingly of lighter color (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Liodessus spp. nov. L. meridensis (A); L. venezuelanus (B). Scale bar: 2 mm." figureDoi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7">1A</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Liodessus spp. nov. Detail of head in dorsal view. L. meridensis (A, B); L. venezuelanus (C, D)." figureDoi="10.3897/alpento.6.79598.figure2" httpUri="https://binary.pensoft.net/fig/682025" pageId="0" pageNumber="7">2A, B</figureCitation>
).
</paragraph>
<caption doi="10.3897/alpento.6.79598.figure2" httpUri="https://binary.pensoft.net/fig/682025" pageId="0" pageNumber="7" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="7">
<emphasis bold="true" pageId="0" pageNumber="7">Figure 2.</emphasis>
<taxonomicName authorityName="Guignot" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="7">Liodessus</emphasis>
</taxonomicName>
spp. nov. Detail of head in dorsal view. 
<taxonomicName genus="L." lsidName="L. meridensis" pageId="0" pageNumber="7" rank="species" species="meridensis">
<emphasis italics="true" pageId="0" pageNumber="7">L. meridensis</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="7">A, B</emphasis>
); 
<taxonomicName genus="L." lsidName="L. venezuelanus" pageId="0" pageNumber="7" rank="species" species="venezuelanus">
<emphasis italics="true" pageId="0" pageNumber="7">L. venezuelanus</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="7">C, D</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="7">
<emphasis bold="true" pageId="0" pageNumber="7">
<emphasis italics="true" pageId="0" pageNumber="7">Surface sculpture</emphasis>
.
</emphasis>
Head with few larger and smaller setiferous punctures; with distinct microreticulation except on middle of head between the eyes (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Liodessus spp. nov. L. meridensis (A); L. venezuelanus (B). Scale bar: 2 mm." figureDoi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7">1A</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Liodessus spp. nov. Detail of head in dorsal view. L. meridensis (A, B); L. venezuelanus (C, D)." figureDoi="10.3897/alpento.6.79598.figure2" httpUri="https://binary.pensoft.net/fig/682025" pageId="0" pageNumber="7">2A, B</figureCitation>
). Pronotum and elytron shiny; with moderately dense and coarse setiferous punctation (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Liodessus spp. nov. L. meridensis (A); L. venezuelanus (B). Scale bar: 2 mm." figureDoi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7">1A</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="7">
<emphasis bold="true" pageId="0" pageNumber="7">
<emphasis italics="true" pageId="0" pageNumber="7">Structures</emphasis>
.
</emphasis>
Antenna stout. Head without occipital line; with rounded clypeus. Pronotum with distinct lateral beads; with distinct long and deep basal striae (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Liodessus spp. nov. L. meridensis (A); L. venezuelanus (B). Scale bar: 2 mm." figureDoi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7">1A</figureCitation>
). Elytron with short basal stria; without sutural line; without basal epipleural transverse carina. Metathoracic wings not examined in holotype (in paratype: short, about half the length of elytron).
</paragraph>
<paragraph pageId="0" pageNumber="7">
<emphasis bold="true" pageId="0" pageNumber="7">
<emphasis italics="true" pageId="0" pageNumber="7">Genitalia</emphasis>
.
</emphasis>
Median lobe of aedeagus of long, simple curvature in lateral view, more or less gradually narrowing towards tip; also gradually narrowing towards a narrow tip in ventral view; parameres of simple &quot; 
<taxonomicName authorityName="Sharp" authorityYear="1880" lsidName="" pageId="0" pageNumber="7" rank="tribe" tribe="Bidessini">Bidessini</taxonomicName>
&quot; type, two-segmented (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Liodessus spp. nov. Median lobe ventral and lateral views; parameres outer and inner surfaces, L. meridensis (A); L. venezuelanus (B). Scale bar: 0.5 mm." figureDoi="10.3897/alpento.6.79598.figure3" httpUri="https://binary.pensoft.net/fig/682026" pageId="0" pageNumber="7">3A</figureCitation>
).
</paragraph>
<caption doi="10.3897/alpento.6.79598.figure3" httpUri="https://binary.pensoft.net/fig/682026" pageId="0" pageNumber="7" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="7">
<emphasis bold="true" pageId="0" pageNumber="7">Figure 3.</emphasis>
<taxonomicName authorityName="Guignot" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="7">Liodessus</emphasis>
</taxonomicName>
spp. nov. Median lobe ventral and lateral views; parameres outer and inner surfaces, 
<taxonomicName genus="L." lsidName="L. meridensis" pageId="0" pageNumber="7" rank="species" species="meridensis">
<emphasis italics="true" pageId="0" pageNumber="7">L. meridensis</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="7">A</emphasis>
); 
<taxonomicName genus="L." lsidName="L. venezuelanus" pageId="0" pageNumber="7" rank="species" species="venezuelanus">
<emphasis italics="true" pageId="0" pageNumber="7">L. venezuelanus</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="7">B</emphasis>
). Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="7">
<emphasis bold="true" pageId="0" pageNumber="7">Female.</emphasis>
Pronotum and elytra microreticulate, dull (specimens from Gavidia); shiny in the ones from Laguna de 
<normalizedToken originalValue="Mucabají">Mucabaji</normalizedToken>
.
</paragraph>
<paragraph pageId="0" pageNumber="7">
<emphasis bold="true" pageId="0" pageNumber="7">
<emphasis italics="true" pageId="0" pageNumber="7">Variation</emphasis>
.
</emphasis>
Size variation of the paratypes is TL 2.1-2.3 mm; TL-h 1.8-2.0 mm; MW 0.9-1.1 mm. The elytral striae are hardly noticeable in a few paratypes.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="7" type="etymology">
<paragraph pageId="0" pageNumber="7">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="7">
Named after the state of 
<normalizedToken originalValue="Mérida">Merida</normalizedToken>
, in which the collecting localities are situated. The name is an adjective in the nominative singular.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="7" type="comparisons">
<paragraph pageId="0" pageNumber="7">Comparative notes.</paragraph>
<paragraph pageId="0" pageNumber="7">
Distinguished from the other species of 
<taxonomicName authorityName="Guignot" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="7">Liodessus</emphasis>
</taxonomicName>
by the diagnostic combination of the following features: Species from Venezuela; beetle smaller (TL 2.1-2.3 mm); very dark brown to blackish; head without occipital line; pronotal and elytral striae present; wings reduced considerably; shape of median lobe (narrow and 
<normalizedToken originalValue="“pointed”">&quot;pointed&quot;</normalizedToken>
in lateral and ventral view, with its particular shape).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="7" type="bold platform">
<paragraph pageId="0" pageNumber="7">BOLD platform.</paragraph>
<paragraph pageId="0" pageNumber="7">
We provided 1 entry to our BOLD 
<taxonomicName authorityName="Guignot" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="7">Liodessus</emphasis>
</taxonomicName>
project. The species is well delineated from all other 
<taxonomicName authorityName="Guignot" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="7">Liodessus</emphasis>
</taxonomicName>
, with an uncorrected p-distance of around 9% or more.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="7" type="distribution">
<paragraph pageId="0" pageNumber="7">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="7">
Known from around 
<normalizedToken originalValue="Mérida">Merida</normalizedToken>
, Venezuela.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="7" type="habitat">
<paragraph pageId="0" pageNumber="7">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="7">
Collected from shallow water at the edge of high-altitude ponds (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitat of Liodessus spp. nov. near Gavidia, Venenzuela, where both new species were collected (collecting event VZ 10 - 0123 - 03 A). Overview (A); detail of shallow water at the edge (B)." figureDoi="10.3897/alpento.6.79598.figure4" httpUri="https://binary.pensoft.net/fig/682027" pageId="0" pageNumber="7">4</figureCitation>
).
</paragraph>
<caption doi="10.3897/alpento.6.79598.figure4" httpUri="https://binary.pensoft.net/fig/682027" pageId="0" pageNumber="7" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="7">
<emphasis bold="true" pageId="0" pageNumber="7">Figure 4.</emphasis>
Habitat of 
<taxonomicName authorityName="Guignot" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="7">Liodessus</emphasis>
</taxonomicName>
spp. nov. near Gavidia, Venenzuela, where both new species were collected (collecting event VZ10-0123-03A). Overview (
<emphasis bold="true" pageId="0" pageNumber="7">A</emphasis>
); detail of shallow water at the edge (
<emphasis bold="true" pageId="0" pageNumber="7">B</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>