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<document id="F0AEF280CEE3920870B93CE2616B3C44" ID-DOI="10.1111/j.1096-3642.2008.00466.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5445358" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630458879514" checkinUser="felipe" docAuthor="Biju, S. D. &amp; Bossuyt, Franky" docDate="2009" docId="03F5945FFFEA1E03FF31FE34FEFDC327" docLanguage="en" docName="j.1096-3642.2008.00466.x.pdf" docOrigin="Zoological Journal of the Linnean Society 155 (2)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00466.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Philautus wynaadensis Jerdon 1853" docType="treatment" docVersion="3" lastPageNumber="440" masterDocId="FFCCEC27FFAB1E41FFE9FFFDFF8CC65A" masterDocTitle="Systematics and phylogeny of Philautus Gistel, 1848 (Anura, Rhacophoridae) in the Western Ghats of India, with descriptions of 12 new species" masterLastPageNumber="444" masterPageNumber="374" pageNumber="439" updateTime="1699278541911" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="1723C8315D4EFE2107B1E54A4021CC81">Systematics and phylogeny of Philautus Gistel, 1848 (Anura, Rhacophoridae) in the Western Ghats of India, with descriptions of 12 new species</mods:title>
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<mods:namePart id="33F7702C57805969D7578A14428E145A">Biju, S. D.</mods:namePart>
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<mods:namePart id="E2D43118BE2D352677E57487268F05AD">Bossuyt, Franky</mods:namePart>
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<mods:title id="722822B4AF6447582E30437951C54748">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="6BCF9431D0251636578D3CDE12A124E1">2009</mods:date>
<mods:detail id="44658D54FCFFE5F56A477D621DB6098B" type="pubDate">
<mods:number id="969515F891C3186DFDDB9E2FD0239072">2009-02-28</mods:number>
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<mods:detail id="E021D912DA8D213C5FA6F70C81CDC206" type="volume">
<mods:number id="32FE92F39E2843DC06A4692E77126135">155</mods:number>
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<mods:number id="54AB68B48906D1682D33678E3F9EEFBA">2</mods:number>
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<mods:start id="101DFDC2912B7A7D871BB096E74A2295">374</mods:start>
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<mods:url id="17F4FC690C7F77DD483BD252031EE199">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00466.x</mods:url>
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<mods:classification id="DCA44B606159D0FD9C3A876F4760D0A1">journal article</mods:classification>
<mods:identifier id="7E4A0EB5254FFF301DE3F981CE4232FF" type="DOI">10.1111/j.1096-3642.2008.00466.x</mods:identifier>
<mods:identifier id="795FAC990927A209D7C6B26790FA42F1" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="A1EDBC382A9930ECA882E2EBE576D87A" type="Zenodo-Dep">5445358</mods:identifier>
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<treatment id="03F5945FFFEA1E03FF31FE34FEFDC327" LSID="urn:lsid:plazi:treatment:03F5945FFFEA1E03FF31FE34FEFDC327" httpUri="http://treatment.plazi.org/id/03F5945FFFEA1E03FF31FE34FEFDC327" lastPageId="66" lastPageNumber="440" pageId="65" pageNumber="439">
<subSubSection id="C34676C2FFEA1E00FF31FE34FD22C784" box="[216,686,456,478]" pageId="65" pageNumber="439" type="nomenclature">
<paragraph id="8BE32549FFEA1E00FF31FE34FD22C784" blockId="65.[216,686,456,478]" box="[216,686,456,478]" pageId="65" pageNumber="439">
<heading id="D0AB9225FFEA1E00FF31FE34FD22C784" box="[216,686,456,478]" centered="true" fontSize="9" level="2" pageId="65" pageNumber="439" reason="2">
<taxonomicName id="4C5C5ECAFFEA1E00FF31FE34FD22C784" ID-CoL="4G5MB" authority="JERDON, 1853" authorityName="Jerdon" authorityYear="1853" box="[216,686,456,478]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="65" pageNumber="393" phylum="Chordata" rank="species" species="wynaadensis">
<emphasis id="B928F95BFFEA1E00FF31FE34FD8CC786" box="[216,512,457,478]" italics="true" pageId="65" pageNumber="439">PHILAUTUS WYNAADENSIS</emphasis>
<bibRefCitation id="EFCD58B8FFEA1E00FDE3FE34FD22C784" author="Jerdon TC" box="[522,686,456,478]" pageId="65" pageNumber="439" pagination="1486 - 1488" refId="ref43806" refString="Jerdon TC. 1853. Catalogue of reptiles inhabiting the Peninsula of India (continued from p. 479). Journal of Asiatic Society of Bengal 1870: 1486 - 1488." type="journal article" year="1853">JERDON, 1853</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34676C2FFEA1E03FF25FE12FEFDC327" lastPageId="66" lastPageNumber="440" pageId="65" pageNumber="439" type="description">
<paragraph id="8BE32549FFEA1E00FF25FE12FD36C45F" blockId="65.[204,698,495,517]" box="[204,698,495,517]" pageId="65" pageNumber="439">
(
<figureCitation id="136739CCFFEA1E00FF3CFE12FEADC45E" box="[213,289,495,516]" captionStart="Figure 2" captionStartId="6.[164,243,1531,1550]" captionTargetBox="[167,1442,195,1498]" captionTargetId="figure-103@6.[164,1444,193,1501]" captionTargetPageId="6" captionText="Figure 2. Maximum-likelihood phylogram (GTR + G + I; –Ln L = 14 763.97) for the mitochondrial DNA data set of 1488 bp of 34 taxa of Western Ghats Philautus and five outgroup species. Numbers above branches indicate nonparametric bootstrap values under maximum parsimony (MP). Numbers below branches indicate Bayesian posterior probabilities. *Values of less than 50." figureDoi="http://doi.org/10.5281/zenodo.5445362" httpUri="https://zenodo.org/record/5445362/files/figure.png" pageId="65" pageNumber="439">FIGS 2</figureCitation>
,
<figureCitation id="136739CCFFEA1E00FED9FE12FECCC45E" box="[304,320,495,516]" captionStart="Figure 9" captionStartId="19.[144,223,1824,1843]" captionTargetBox="[402,1165,1216,1791]" captionTargetId="figure-382@19.[399,1167,1214,1794]" captionTargetPageId="19" captionText="Figure 9. Morphometric distinction between males of three Philautus species: -, Philautus kani sp. nov.; O, Philautus wynaadensis; Z, Philautus amboli sp. nov. The ratio of head width (HW)/head length (HL) is plotted against snout–vent length SVL (see Table 2)." figureDoi="http://doi.org/10.5281/zenodo.5445376" httpUri="https://zenodo.org/record/5445376/files/figure.png" pageId="65" pageNumber="439">9</figureCitation>
,
<figureCitation id="136739CCFFEA1E00FEA6FE12FE0EC45F" box="[335,386,495,517]" captionStart="Figure 59" captionStartId="61.[144,223,1842,1861]" captionTargetBox="[164,1401,196,1811]" captionTargetId="figure-0@61.[162,1404,194,1813]" captionTargetPageId="61" captionText="Figure 59. Distribution map. A, Philautus tinniens; B, Philautus travancoricus; C, Philautus tuberohumerus; D, Philautus wynaadensis." figureDoi="http://doi.org/10.5281/zenodo.5445480" httpUri="https://zenodo.org/record/5445480/files/figure.png" pageId="65" pageNumber="439">59D</figureCitation>
,
<figureCitation id="136739CCFFEA1E00FE79FE12FE6EC45F" box="[400,482,495,517]" captionStart="Figure 64" captionStartId="65.[806,885,1486,1505]" captionTargetBox="[810,1418,196,1457]" captionTargetId="figure-452@65.[810,1418,196,1457]" captionTargetPageId="65" captionText="Figure 64. Philautus wynaadensis: A, specimen (BNHS 4556) from Kalpetta; B, specimen (BNHS 4554) from Sulthanbathery; C, specimen (BNHS 4552) from Kalpetta." figureDoi="http://doi.org/10.5281/zenodo.5445492" httpUri="https://zenodo.org/record/5445492/files/figure.png" pageId="65" pageNumber="439">64A–C</figureCitation>
,
<figureCitation id="136739CCFFEA1E00FE19FE12FD11C45E" box="[496,669,495,517]" captionStart="Figure 65" captionStartId="66.[164,243,1026,1045]" captionTargetBox="[168,777,195,996]" captionTargetId="figure-580@66.[168,777,195,996]" captionTargetPageId="66" captionText="Figure 65. A, holotype (ZMB 3057) of Ixalus leucorhinus; B, holotype (BMNH 1947.2.6.21) of Ixalus nasutus." figureDoi="http://doi.org/10.5281/zenodo.5445495" httpUri="https://zenodo.org/record/5445495/files/figure.png" pageId="65" pageNumber="439">65A, B; TABLE</figureCitation>
2)
</paragraph>
<paragraph id="8BE32549FFEA1E00FF79FDEBFDF5C433" blockId="65.[144,758,533,617]" pageId="65" pageNumber="439">
<emphasis id="B928F95BFFEA1E00FF79FDEBFEBAC470" box="[144,310,533,555]" italics="true" pageId="65" pageNumber="439">Type material:</emphasis>
<materialsCitation id="3B342F14FFEA1E00FEA9FDEBFDF5C433" collectionCode="SVL" country="India" location="Sultan's Battery'" municipality="Wayanad" pageId="65" pageNumber="439" specimenCode="MNHN 1999.5596" specimenCount="1" stateProvince="Kerala" typeStatus="neotype">
<typeStatus id="54E79BEBFFEA1E00FEA9FDEBFE2DC471" box="[320,417,534,555]" pageId="65" pageNumber="439" type="neotype">Neotype</typeStatus>
,
<specimenCode id="DBFA8D32FFEA1E00FE46FDEBFD09C471" box="[431,645,533,555]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="65" pageNumber="439">MNHN 1999.5596</specimenCode>
, an adult male,
<collectionCode id="ED4DBD8CFFEA1E00FF08FDC9FE9FC413" box="[225,275,564,585]" pageId="65" pageNumber="439">SVL</collectionCode>
<quantity id="4CA488ACFFEA1E00FECCFDC9FE06C413" box="[293,394,564,585]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="65" pageNumber="439" unit="mm" value="28.3">28.3 mm</quantity>
, from ‘
<location id="8E837392FFEA1E00FE07FDC9FD37C413" LSID="urn:lsid:plazi:treatment:03F5945FFFEA1E03FF31FE34FEFDC327:8E837392FFEA1E00FE07FDC9FD37C413" box="[494,699,564,586]" country="India" municipality="Wayanad" name="Sultan's Battery'" pageId="65" pageNumber="439" stateProvince="Kerala">Sultan’s Battery’</location>
(i.e. Sulthanbathery),
<collectingMunicipality id="6B87BF33FFEA1E00FEB4FDAEFE46C433" box="[349,458,595,617]" pageId="65" pageNumber="439">Wayanad</collectingMunicipality>
,
<collectingRegion id="4998EBABFFEA1E00FE3EFDAEFDABC433" box="[471,551,595,617]" country="India" name="Kerala" pageId="65" pageNumber="439">Kerala</collectingRegion>
,
<collectingCountry id="F34B65D9FFEA1E00FDDCFDAEFDF9C433" box="[565,629,595,617]" name="India" pageId="65" pageNumber="439">India</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE32549FFEA1E00FF79FD6DFDF5C565" blockId="65.[144,759,656,831]" pageId="65" pageNumber="439">
<emphasis id="B928F95BFFEA1E00FF79FD6DFE10C4FF" box="[144,412,656,677]" italics="true" pageId="65" pageNumber="439">Other material studied:</emphasis>
<materialsCitation id="3B342F14FFEA1E00FE4DFD6DFDD3C4FC" box="[420,607,656,678]" collectionCode="BNHS" pageId="65" pageNumber="439" specimenCode="BNHS 4551" specimenCount="1">
<specimenCode id="DBFA8D32FFEA1E00FE4DFD6DFDA1C4FC" box="[420,557,656,678]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="65" pageNumber="439">BNHS 4551</specimenCode>
and
</materialsCitation>
<materialsCitation id="3B342F14FFEA1E00FD8EFD6DFE8CC49F" collectionCode="BNHS" pageId="65" pageNumber="439" specimenCode="BNHS 4552" specimenCount="1" specimenCount-adult="1">
<specimenCode id="DBFA8D32FFEA1E00FD8EFD6DFD7FC4FC" box="[615,755,656,678]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="65" pageNumber="439">BNHS 4552</specimenCode>
,
<specimenCount id="9D5AEEC0FFEA1E00FF79FD52FE8CC49F" box="[144,256,687,709]" pageId="65" pageNumber="439" type="adult">two adult</specimenCount>
</materialsCitation>
males, and
<materialsCitation id="3B342F14FFEA1E00FE7FFD52FDA8C49E" box="[406,548,686,708]" collectionCode="BNHS" pageId="65" pageNumber="439" specimenCode="BNHS 4556" specimenCount="1">
<specimenCode id="DBFA8D32FFEA1E00FE7FFD52FDA8C49E" box="[406,548,686,708]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="65" pageNumber="439">BNHS 4556</specimenCode>
</materialsCitation>
, an adult female, from Kalpetta;
<materialsCitation id="3B342F14FFEA1E00FEA7FD30FD94C4B9" box="[334,536,717,739]" collectionCode="BNHS" pageId="65" pageNumber="439" specimenCode="BNHS 4553" specimenCount="1">
<specimenCode id="DBFA8D32FFEA1E00FEA7FD30FE52C4B9" box="[334,478,717,739]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="65" pageNumber="439">BNHS 4553</specimenCode>
and
</materialsCitation>
<materialsCitation id="3B342F14FFEA1E00FDCEFD30FF40C558" collectionCode="BNHS" pageId="65" pageNumber="439" specimenCode="BNHS 4554" specimenCount="1" specimenCount-adult="1">
<specimenCode id="DBFA8D32FFEA1E00FDCEFD30FD37C4B9" box="[551,699,717,739]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="65" pageNumber="439">BNHS 4554</specimenCode>
,
<specimenCount id="9D5AEEC0FFEA1E00FD24FD33FF40C558" pageId="65" pageNumber="439" type="adult">two adult</specimenCount>
</materialsCitation>
males, from Sulthanbathery;
<materialsCitation id="3B342F14FFEA1E00FDB3FD11FD7EC558" box="[602,754,748,770]" collectionCode="BNHS" pageId="65" pageNumber="439" specimenCode="BNHS 4555" specimenCount="1">
<specimenCode id="DBFA8D32FFEA1E00FDB3FD11FD7EC558" box="[602,754,748,770]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="65" pageNumber="439">BNHS 4555</specimenCode>
</materialsCitation>
, an adult male, from Mananthavady;
<materialsCitation id="3B342F14FFEA1E00FDD6FCF6FD41C57A" box="[575,717,778,800]" collectionCode="BNHS" pageId="65" pageNumber="439" specimenCode="BNHS 4550" specimenCount="1">
<specimenCode id="DBFA8D32FFEA1E00FDD6FCF6FD41C57A" box="[575,717,778,800]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="65" pageNumber="439">BNHS 4550</specimenCode>
</materialsCitation>
, an adult male, from Mettupalayam (
<tableCitation id="C6DE10F2FFEA1E00FDFFFCD4FDE0C565" box="[534,620,809,831]" captionStart="Table 2" captionStartId="10.[113,177,400,421]" captionText="Table 2. Morphometric measurements (in mm) of the specimens used in this study" pageId="65" pageNumber="439">Table 2</tableCitation>
).
</paragraph>
<paragraph id="8BE32549FFEA1E00FF79FC9AFD42C26E" blockId="65.[144,759,870,1168]" pageId="65" pageNumber="439">
<emphasis id="B928F95BFFEA1E00FF79FC9AFDBCC521" box="[144,560,870,892]" italics="true" pageId="65" pageNumber="439">
Diagnosis:
<taxonomicName id="4C5C5ECAFFEA1E00FEF3FC9BFDBCC521" authorityName="Jerdon" authorityYear="1853" box="[282,560,870,891]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="65" pageNumber="439" phylum="Chordata" rank="species" species="wynaadensis">Philautus wynaadensis</taxonomicName>
</emphasis>
can be distinguished from known congeners by the following combination of characters: (1) medium adult size (SVL 25.7 ±
<quantity id="4CA488ACFFEA1E00FF35FC3FFEBFC582" box="[220,307,962,984]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="65" pageNumber="439" unit="mm" value="1.5">1.5 mm</quantity>
,
<emphasis id="B928F95BFFEA1E00FEA2FC3EFED3C582" box="[331,351,963,984]" italics="true" pageId="65" pageNumber="439">N</emphasis>
= 7, male; SVL
<quantity id="4CA488ACFFEA1E00FDDFFC3FFD17C582" box="[566,667,962,984]" metricMagnitude="-2" metricUnit="m" metricValue="2.7199999999999998" pageId="65" pageNumber="439" unit="mm" value="27.2">27.2 mm</quantity>
,
<emphasis id="B928F95BFFEA1E00FD5AFC3EFD4BC582" box="[691,711,963,984]" italics="true" pageId="65" pageNumber="439">N</emphasis>
= 1, female); (2) body rather slender; (3) snout subelliptical in ventral view; (4) upper two-thirds of tympanum dark black; (5) dorsum with spinular projections.
</paragraph>
<paragraph id="8BE32549FFEA1E00FF41FBC0FE7BC2CA" blockId="65.[144,759,870,1168]" pageId="65" pageNumber="439">
<taxonomicName id="4C5C5ECAFFEA1E00FF41FBC0FE4DC208" authorityName="Jerdon" authorityYear="1853" box="[168,449,1085,1106]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="65" pageNumber="439" phylum="Chordata" rank="species" species="wynaadensis">
<emphasis id="B928F95BFFEA1E00FF41FBC0FE4DC208" box="[168,449,1085,1106]" italics="true" pageId="65" pageNumber="439">Philautus wynaadensis</emphasis>
</taxonomicName>
can be confused with
<taxonomicName id="4C5C5ECAFFEA1E00FF79FBA1FE8DC22A" authority="Biju &amp; Bossuyt, 2009" authorityName="Biju &amp; Bossuyt" authorityYear="2009" box="[144,257,1115,1137]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="65" pageNumber="439" phylum="Chordata" rank="species" species="amboli" status="sp. nov.">
<emphasis id="B928F95BFFEA1E00FF79FBA1FE8DC22A" box="[144,257,1115,1137]" italics="true" pageId="65" pageNumber="439">P. amboli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B4420FFEA1E00FEF8FBA1FEE2C22B" box="[273,366,1116,1137]" pageId="65" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C5C5ECAFFEA1E00FE5EFBA1FD81C22A" authority="Biju &amp; Bossuyt, 2009" authorityName="Biju &amp; Bossuyt" authorityYear="2009" box="[439,525,1115,1137]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="65" pageNumber="439" phylum="Chordata" rank="species" species="kani" status="sp. nov.">
<emphasis id="B928F95BFFEA1E00FE5EFBA1FD81C22A" box="[439,525,1115,1137]" italics="true" pageId="65" pageNumber="439">P. kani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B4420FFEA1E00FDF5FBA1FDF6C22B" box="[540,634,1116,1137]" pageId="65" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
(see both those species for comparisons).
</paragraph>
<paragraph id="8BE32549FFEA1E00FF79FB45FF52C351" blockId="65.[144,759,1207,1291]" pageId="65" pageNumber="439">
<emphasis id="B928F95BFFEA1E00FF79FB45FE4BC297" box="[144,455,1207,1229]" italics="true" pageId="65" pageNumber="439">
Description of the
<typeStatus id="54E79BEBFFEA1E00FE85FB45FE4DC297" box="[364,449,1208,1229]" pageId="65" pageNumber="439" type="neotype">neotype</typeStatus>
:
</emphasis>
A detailed description of the
<typeStatus id="54E79BEBFFEA1E00FF2FFB2BFE93C2B1" box="[198,287,1238,1259]" pageId="65" pageNumber="439" type="neotype">neotype</typeStatus>
was published in
<bibRefCitation id="EFCD58B8FFEA1E00FDFFFB2BFF56C351" author="Bossuyt F &amp; Dubois A" pageId="65" pageNumber="439" pagination="1 - 112" refId="ref42998" refString="Bossuyt F, Dubois A. 2001. A review of the frog genus Philautus Gistel, 1848 (Amphibia, Anura, Ranidae, Rhacophorinae). Zeylanica 6: 1 - 112." type="journal article" year="2001">Bossuyt &amp; Dubois (2001)</bibRefCitation>
.
</paragraph>
<paragraph id="8BE32549FFEA1E00FF79FACEFD28C135" blockId="65.[144,759,1330,1903]" pageId="65" pageNumber="439">
<emphasis id="B928F95BFFEA1E00FF79FACEFE8CC312" box="[144,256,1331,1352]" italics="true" pageId="65" pageNumber="439">Variation:</emphasis>
<materialsCitation id="3B342F14FFEA1E00FEE5FACFFD2CC135" collectionCode="BNHS" pageId="65" pageNumber="439" specimenCode="BNHS 4556, BNHS 4554, BNHS 4552" specimenCount="1" typeStatus="neotype">
Measurements of
<specimenCount id="9D5AEEC0FFEA1E00FE08FACFFD10C312" box="[481,668,1330,1352]" pageId="65" pageNumber="439" type="generic">eight specimens</specimenCount>
including the
<typeStatus id="54E79BEBFFEA1E00FF1DFAACFEC1C33C" box="[244,333,1361,1382]" pageId="65" pageNumber="439" type="neotype">neotype</typeStatus>
are given in
<tableCitation id="C6DE10F2FFEA1E00FE17FAACFDDFC33C" box="[510,595,1361,1383]" captionStart="Table 2" captionStartId="10.[113,177,400,421]" captionText="Table 2. Morphometric measurements (in mm) of the specimens used in this study" pageId="65" pageNumber="439">Table 2</tableCitation>
. This species shows some colour variation, even within the same population, from uniform grey to brownish or reddish grey.
<specimenCode id="DBFA8D32FFEA1E00FF38FA50FED2C398" box="[209,350,1452,1474]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="65" pageNumber="439">BNHS 4556</specimenCode>
(
<figureCitation id="136739CCFFEA1E00FE9BFA51FE5BC398" box="[370,471,1452,1474]" captionStart="Figure 64" captionStartId="65.[806,885,1486,1505]" captionTargetBox="[810,1418,196,1457]" captionTargetId="figure-452@65.[810,1418,196,1457]" captionTargetPageId="65" captionText="Figure 64. Philautus wynaadensis: A, specimen (BNHS 4556) from Kalpetta; B, specimen (BNHS 4554) from Sulthanbathery; C, specimen (BNHS 4552) from Kalpetta." figureDoi="http://doi.org/10.5281/zenodo.5445492" httpUri="https://zenodo.org/record/5445492/files/figure.png" pageId="65" pageNumber="439">Fig. 64A</figureCitation>
): dorsum grey–brown, a dark-brown band from the half portion of upper eyelid to vent, snout grey–brown, upper two-thirds of tympanum dark black, upper half of iris light brownish, lower half dark brownish; lateral side light brownish, either side of snout including loreal and tympanic regions dark-brownish black, limbs light brown with dark cross bands; ventral side light creamy white with dark spots;
<specimenCode id="DBFA8D32FFEA1E00FE98F95CFD89C0ED" box="[369,517,1697,1719]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="65" pageNumber="439">BNHS 4554</specimenCode>
(
<figureCitation id="136739CCFFEA1E00FDF6F95CFD0EC0ED" box="[543,642,1697,1719]" captionStart="Figure 64" captionStartId="65.[806,885,1486,1505]" captionTargetBox="[810,1418,196,1457]" captionTargetId="figure-452@65.[810,1418,196,1457]" captionTargetPageId="65" captionText="Figure 64. Philautus wynaadensis: A, specimen (BNHS 4556) from Kalpetta; B, specimen (BNHS 4554) from Sulthanbathery; C, specimen (BNHS 4552) from Kalpetta." figureDoi="http://doi.org/10.5281/zenodo.5445492" httpUri="https://zenodo.org/record/5445492/files/figure.png" pageId="65" pageNumber="439">Fig. 64B</figureCitation>
): dorsum reddish brown with a faint ‘X’ marking, loreal and tympanic regions light grey, upper two-thirds of tympanum dark black, limbs light-reddish brown with faint cross bands; ventral side uniform white with faint minute spots;
<specimenCode id="DBFA8D32FFEA1E00FE97F8C7FD87C10A" box="[382,523,1850,1872]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="65" pageNumber="439">BNHS 4552</specimenCode>
(
<figureCitation id="136739CCFFEA1E00FDC9F8C7FD0FC10A" box="[544,643,1850,1872]" captionStart="Figure 64" captionStartId="65.[806,885,1486,1505]" captionTargetBox="[810,1418,196,1457]" captionTargetId="figure-452@65.[810,1418,196,1457]" captionTargetPageId="65" captionText="Figure 64. Philautus wynaadensis: A, specimen (BNHS 4556) from Kalpetta; B, specimen (BNHS 4554) from Sulthanbathery; C, specimen (BNHS 4552) from Kalpetta." figureDoi="http://doi.org/10.5281/zenodo.5445492" httpUri="https://zenodo.org/record/5445492/files/figure.png" pageId="65" pageNumber="439">Fig. 64C</figureCitation>
): uniform grey dorsum with faint cross bands on limbs
</materialsCitation>
.
</paragraph>
<caption id="DF2375C1FFEA1E00FCCFFA33FA02C046" ID-DOI="http://doi.org/10.5281/zenodo.5445492" ID-Zenodo-Dep="5445492" httpUri="https://zenodo.org/record/5445492/files/figure.png" pageId="65" pageNumber="439" startId="65.[806,885,1486,1505]" targetBox="[810,1418,196,1457]" targetPageId="65">
<paragraph id="8BE32549FFEA1E00FCCFFA33FA02C046" blockId="65.[806,1422,1486,1564]" pageId="65" pageNumber="439">
<emphasis id="B928F95BFFEA1E00FCCFFA33FC10C3BB" bold="true" box="[806,924,1486,1505]" pageId="65" pageNumber="439">Figure 64.</emphasis>
<taxonomicName id="4C5C5ECAFFEA1E00FC41FA33FB2DC3BB" authorityName="Jerdon" authorityYear="1853" box="[936,1185,1486,1505]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="65" pageNumber="439" phylum="Chordata" rank="species" species="wynaadensis">
<emphasis id="B928F95BFFEA1E00FC41FA33FB2DC3BB" box="[936,1185,1486,1505]" italics="true" pageId="65" pageNumber="439">Philautus wynaadensis</emphasis>
</taxonomicName>
: A, specimen (BNHS 4556) from Kalpetta; B, specimen (BNHS 4554) from Sulthanbathery; C, specimen (BNHS 4552) from Kalpetta.
</paragraph>
</caption>
<paragraph id="8BE32549FFEA1E03FCCFF999FE2EC2C5" blockId="65.[806,1421,1636,1903]" lastBlockId="66.[164,779,1131,1183]" lastPageId="66" lastPageNumber="440" pageId="65" pageNumber="439">
<emphasis id="B928F95BFFEA1E00FCCFF999FB44C023" box="[806,1224,1636,1657]" italics="true" pageId="65" pageNumber="439">
Distribution:
<taxonomicName id="4C5C5ECAFFEA1E00FC56F999FB44C023" authorityName="Jerdon" authorityYear="1853" box="[959,1224,1636,1657]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="65" pageNumber="439" phylum="Chordata" rank="species" species="wynaadensis">Philautus wynaadensis</taxonomicName>
</emphasis>
is widely distributed in the Western Ghats, especially north of the Palghat Gap, with its southern limit reaching up to Periyar (South of the Palghat Gap). This species was collected from nine localities: Mananthavady, Sulthanbathery, Kalpetta, Palakkad, Parambikulam- Puliyarapadam, and Mettupalayam (NPG); and Thrissur, Meladoor, and Periyar (SPG) (
<figureCitation id="136739CCFFEA1E00FACDF8C7FA05C10A" box="[1316,1417,1850,1872]" captionStart="Figure 58" captionStartId="60.[826,905,1853,1872]" captionTargetBox="[830,1439,1440,1824]" captionTargetId="figure-683@60.[830,1439,1440,1824]" captionTargetPageId="60" captionText="Figure 58. Philautus tinniens: specimen (BNHS 4495) from Udhagamandalam." figureDoi="http://doi.org/10.5281/zenodo.5445478" httpUri="https://zenodo.org/record/5445478/files/figure.png" pageId="65" pageNumber="439">Fig. 58D</figureCitation>
,
<tableCitation id="C6DE10F2FFEA1E00FCCFF8A4FCF4C135" box="[806,888,1881,1903]" captionStart="Table 1" captionStartId="2.[164,228,200,219]" captionText="Table 1. Collection localities that are discussed in the text. Localities are arranged by state" pageId="65" pageNumber="439">Table 1</tableCitation>
). It is one of the most common
<taxonomicName id="4C5C5ECAFFEA1E03FB15F8A4FF45C2DB" authority="in the" authorityName="IN THE" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" lastPageId="66" lastPageNumber="440" order="Anura" pageId="65" pageNumber="427" phylum="Chordata" rank="genus">
<emphasis id="B928F95BFFEA1E00FB15F8A4FAE0C134" box="[1276,1388,1881,1902]" italics="true" pageId="65" pageNumber="439">Philautus</emphasis>
in the
</taxonomicName>
study region, and is dominant in wayside vegetation and urban areas.
</paragraph>
<caption id="DF2375C1FFE91E03FF4DFBFFFD44C268" ID-DOI="http://doi.org/10.5281/zenodo.5445495" ID-Zenodo-Dep="5445495" httpUri="https://zenodo.org/record/5445495/files/figure.png" pageId="66" pageNumber="440" startId="66.[164,243,1026,1045]" targetBox="[168,777,195,996]" targetPageId="66">
<paragraph id="8BE32549FFE91E03FF4DFBFFFD44C268" blockId="66.[164,778,1026,1075]" pageId="66" pageNumber="440">
<emphasis id="B928F95BFFE91E03FF4DFBFFFE95C24F" bold="true" box="[164,281,1026,1045]" pageId="66" pageNumber="440">Figure 65.</emphasis>
A, holotype (ZMB 3057) of
<taxonomicName id="4C5C5ECAFFE91E03FDABFBFFFC88C24F" authorityName="Lichtenstein &amp; Martens" authorityYear="1856" box="[578,772,1026,1045]" class="Amphibia" family="Rhacophoridae" genus="Ixalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="440" phylum="Chordata" rank="species" species="leucorhinus">
<emphasis id="B928F95BFFE91E03FDABFBFFFC88C24F" box="[578,772,1026,1045]" italics="true" pageId="66" pageNumber="440">Ixalus leucorhinus</emphasis>
</taxonomicName>
; B, holotype (BMNH 1947.2.6.21) of
<taxonomicName id="4C5C5ECAFFE91E03FDCCFBE2FD48C268" authorityName="Gunther" authorityYear="1869" box="[549,708,1055,1074]" class="Amphibia" family="Rhacophoridae" genus="Ixalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="440" phylum="Chordata" rank="species" species="nasutus">Ixalus nasutus</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BE32549FFE91E03FF4DFB33FEFDC327" blockId="66.[164,779,1230,1405]" pageId="66" pageNumber="440">
<emphasis id="B928F95BFFE91E03FF4DFB33FDA2C2B9" box="[164,558,1230,1251]" italics="true" pageId="66" pageNumber="440">
Remarks:
<taxonomicName id="4C5C5ECAFFE91E03FEF7FB33FDA2C2B9" authorityName="Jerdon" authorityYear="1853" box="[286,558,1230,1251]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="66" pageNumber="440" phylum="Chordata" rank="species" species="wynaadensis">Philautus wynaadensis</taxonomicName>
</emphasis>
was discussed as I
<emphasis id="B928F95BFFE91E03FF47FB10FF67C358" box="[174,235,1261,1282]" italics="true" pageId="66" pageNumber="440">xalus</emphasis>
<taxonomicName id="4C5C5ECAFFE91E03FF14FB10FC87C358" authority="Lichtenstein &amp; Martens, 1856" authorityName="Lichtenstein &amp; Martens" authorityYear="1856" box="[253,779,1260,1283]" class="Amphibia" family="Rhacophoridae" genus="Ixalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="440" phylum="Chordata" rank="species" species="leucorhinus">
<emphasis id="B928F95BFFE91E03FF14FB10FE0EC358" box="[253,386,1261,1282]" italics="true" pageId="66" pageNumber="440">leucorhinus</emphasis>
Lichtenstein &amp; Martens, 1856
</taxonomicName>
(BMNH 72.4.17257a–d,
<specimenCount id="9D5AEEC0FFE91E03FE20FAF6FDCFC37B" box="[457,579,1291,1313]" pageId="66" pageNumber="440" type="adult">four adult</specimenCount>
males from ‘N. Canara’; BMNH 74.4.29.587 adult male, BMNH 74.4.29.495 adult female from ‘Malabar’) by
<bibRefCitation id="EFCD58B8FFE91E03FF4DFA9AFEE1C327" author="Boulenger GA" box="[164,365,1383,1405]" pageId="66" pageNumber="440" refId="ref43127" refString="Boulenger GA. 1882. Catalogue of the Batrachia Salientia s. Ecaudata in the collection of the British Museum. London: Taylor &amp; Francis." type="book" year="1882">Boulenger (1882)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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