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<document id="855FE5F720B5264ECC4AAEE764DE23A9" ID-DOI="10.11646/zootaxa.3709.4.1" ID-GBIF-Dataset="b3a7f752-0509-4e01-868d-97e4ab16245e" ID-ISSN="1175-5326" ID-PMC="PMC3914756" ID-PubMed="24511221" ID-Zenodo-Dep="249070" ID-ZooBank="6F6CC130-6A51-4FC0-8E3C-A82B598FFB49" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460673920785" checkinUser="plazi" docAuthor="Herler, Juergen, Bogorodsky, Sergey V. &amp; Suzuki, Toshiyuki" docDate="2013" docId="03E187891928AD543AEBFA21EF06FE34" docLanguage="en" docName="zt03709p329.pdf" docOrigin="Zootaxa 3709 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gobiodon irregularis Herler, Bogorodsky &amp; Suzuki, 2013, sp. nov." docType="treatment" docUuid="94A22A23-C6E4-46BB-A8E8-195B83D61431" docUuidSource="ZooBank" docVersion="10" lastPageNumber="313" masterDocId="FFD8FFF11920AD593A7CFF86EB3DFFE4" masterDocTitle="Four new species of coral gobies (Teleostei: Gobiidae: Gobiodon), with comments on their relationships within the genus" masterLastPageNumber="329" masterPageNumber="301" pageNumber="309" updateTime="1698349206470" updateUser="plazi">
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<mods:title id="8860AFC98BCF9364A782A68B6FDFB396">Four new species of coral gobies (Teleostei: Gobiidae: Gobiodon), with comments on their relationships within the genus</mods:title>
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<taxonomicName id="4C484D1C1928AD513AEBFA21EAABFA25" ID-CoL="3GPCY" box="[151,406,1447,1473]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="irregularis" status="sp. nov.">
<emphasis id="B93CEA8D1928AD513AEBFA21EAABFA25" bold="true" box="[151,406,1447,1473]" italics="true" pageId="8" pageNumber="309">Gobiodon irregularis</emphasis>
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, 
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<paragraph id="8BF7369F1928AD513AEBFA4CEA59FA07" blockId="8.[151,588,1447,1543]" box="[151,356,1482,1507]" pageId="8" pageNumber="309">Rufous Coralgoby</paragraph>
<paragraph id="8BF7369F1928AD513AEBFA68E971F9E3" blockId="8.[151,588,1447,1543]" box="[151,588,1518,1543]" pageId="8" pageNumber="309">
<figureCitation id="13732A1A1928AD513AEBFA68EBD9F9E3" box="[151,228,1518,1543]" captionStart="FIGURE 3" captionStartId="9.[151,250,1796,1818]" captionTargetBox="[151,1436,766,1774]" captionTargetId="figure@9.[151,1436,766,1774]" captionTargetPageId="9" captionText="FIGURE 3. Gobiodon irregularis sp. nov. A. freshly collected adult, holotype, NMW 95078, 30.1 mm SL, Dahab, Egypt, Red Sea. B. freshly collected subadult, CH 232 - 41 b- 057, 22.3 mm SL, Dahab, Egypt, Red Sea. Photos by J. Herler." httpUri="https://zenodo.org/record/249073/files/figure.png" pageId="8" pageNumber="309">Figs. 3</figureCitation>
, 
<figureCitation id="13732A1A1928AD513A8DFA69EA3CF9E3" box="[241,257,1519,1543]" captionStart="FIGURE 4" captionStartId="10.[151,250,1841,1863]" captionTargetBox="[151,1436,918,1819]" captionTargetId="figure@10.[151,1436,918,1819]" captionTargetPageId="10" captionText="FIGURE 4. Gobiodon irregularis sp. nov. A. freshly collected adult, paratype MNHN 2006 - 1699, 32.3 mm SL, Dahab, Egypt, Red Sea; B. freshly collected adult, CH 232 - 41 - 020, 31.2 mm SL, Dahab, Egypt, Red Sea; C. freshly collected adult, SMF uncatalogued, 27.6 mm SL, Farasan Archipelago, Red Sea; D. freshly collected subadult, paratype NMW 95565, 23.4 mm SL, Dahab, Egypt, Red Sea; E. alive subadult, PMR VP 3201, 23.5 mm SL, Sharm el Moya, Egypt, Red Sea; F. freshly collected juvenile, SAIAB 70430, 15 mm SL, Rodrigues, western Indian Ocean. Photos by J. Herler (A, B, D), S. V. Bogorodsky (C, E), P. Heemstra, © SAIAB (F)." httpUri="https://zenodo.org/record/249074/files/figure.png" pageId="8" pageNumber="309">4</figureCitation>
, 
<figureCitation id="13732A1A1928AD513B71FA69EA14F9E3" box="[269,297,1519,1543]" captionStart="FIGURE 10" captionStartId="23.[151,250,1629,1651]" captionTargetBox="[224,1369,736,1597]" captionTargetId="figure@23.[209,1384,724,1615]" captionTargetPageId="23" captionText="FIGURE 10. First two principal components of a PCA of Procrustes coordinates of 15 landmarks and 6 semi-landmarks put on 111 specimens of Gobiodon. The holotype (MNHN A- 4015) of G. unicolor (Castelnau, 1873) is marked by a red circle. RS = Red Sea. For species abbreviations see Table 1." httpUri="https://zenodo.org/record/249080/files/figure.png" pageId="8" pageNumber="309">10</figureCitation>
&amp; 
<figureCitation id="13732A1A1928AD513B31FA69EA56F9E3" box="[333,363,1519,1543]" captionStart="FIGURE 11" captionStartId="25.[151,250,1851,1873]" captionTargetBox="[358,1210,187,1822]" captionTargetId="figure@25.[348,1247,178,1829]" captionTargetPageId="25" captionText="FIGURE 11. Neighbor-joining tree of 23 taxa of Gobiodon and 2 Paragobiodon species (as nearest neighbour at genus level) based on combined sequences of partial 12 S and 16 S rRNA mitochondrial genes. According to a model test in MEGA 5.05, a K 2 P-model with a gamma-parameter of 0.23 and pairwise deletions of gaps / missing data were selected. Numbers at nodes are bootstrap values (1000 replicates); values &lt;50 are not shown. Bar below tree shows substitutions per site. The 4 new species are marked with a box. For abbreviations of species and localities see Table 1. Undescribed species are given with references in parentheses: JH = according to Herler et al. (2009), PM = according to Munday et al. (1999)." httpUri="https://zenodo.org/record/249081/files/figure.png" pageId="8" pageNumber="309">11</figureCitation>
; 
<tableCitation id="C6CA03241928AD513B0AFA68EAE9F9E3" box="[374,468,1518,1543]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionTargetBox="[159,1431,362,1898]" captionTargetPageId="4" captionText="TABLE 1. GenBank accession numbers (GB No.) and references for published sequences (12 S and 16 S rRNA) of 25 taxa (4 new species are underlined) of Gobiodon (G.) and Paragobiodon (P.). Localities: GBR = Great Barrier Reef, Mal = Maldives, PNG = Papua New Guinea, RS = Red Sea, Tai = Taiwan. Abbreviations (PM) and (JH) refer to the first demonstration of sequenced but undescribed species, Munday et al. (1999) and Herler and Hilgers (2009), respectively. Superscript numbers in locality code indicate type status: a: Paratype MNHN 2012 - 0111, b: Paratype BMNH 2006.10. 6.1, c: Paratype NMW 95565." httpUri="http://table.plazi.org/id/DF3766171924AD5D3AEBFF1EE90BFEAB" pageId="8" pageNumber="309" tableUuid="DF3766171924AD5D3AEBFF1EE90BFEAB">Tables 1</tableCitation>
, 
<tableCitation id="C6CA03241928AD513B9EFA69EACFF9E3" box="[482,498,1519,1543]" captionStart="TABLE 4" captionStartId="11.[151,239,584,607]" captionTargetBox="[159,1412,701,867]" captionTargetPageId="11" captionText="TABLE 4. Body proportions of holotype and three adult paratypes of Gobiodon irregularis sp. nov. from the northern Red Sea. The juvenile paratype NMW 95566 (SL = 16.1 mm) is excluded here. Values are proportions of standard length (SL) and head length (last five measurements), respectively, means and the first standard deviation (SD)." httpUri="http://table.plazi.org/id/DF376617192BAD523AEBFDCEEFC9FD45" pageId="8" pageNumber="309" tableUuid="DF376617192BAD523AEBFDCEEFC9FD45">4</tableCitation>
, 
<tableCitation id="C6CA03241928AD513B82FA68E931F9E3" box="[510,524,1518,1543]" captionStart="TABLE 5" captionStartId="12.[151,239,152,175]" captionTargetBox="[159,1330,237,610]" captionTargetPageId="12" captionText="TABLE 5. Fin counts of holotype and four paratypes of Gobiodon irregularis sp. nov. from the northern Red Sea. d = damaged." httpUri="http://table.plazi.org/id/DF376617192CAD553AEBFF1EEBC2FF35" pageId="8" pageNumber="309" tableUuid="DF376617192CAD553AEBFF1EEBC2FF35">5</tableCitation>
&amp; 
<tableCitation id="C6CA03241928AD51384CFA69E971F9E3" box="[560,588,1519,1543]" captionStart="TABLE 10" captionStartId="21.[151,239,670,693]" captionTargetBox="[159,1393,234,607]" captionTargetPageId="21" captionText="TABLE 10. Selection of genetic (p -) distances from pairwise comparisons of specimens of G. bilineatus sp. nov., G. irregularis sp. nov., G. ater sp. nov. and G. fuscoruber sp. nov. from different regions (GBR: Great Barrier Reef, Jap: Japan, Mal: Maldives, RS: Red Sea, Tai: Taiwan). For comparison of intra- and interspecific genetic distances across the genus, other species from different regions as well as “ G. unicolor ” (sensu Munday et al. (1999) and Harold et al. (2008 )) from the GBR are included. Italic numbers indicate intraspecific distances. Bold values indicate intraspecific distances of more than 2 %. For species abbreviations and Genbank accession numbers see Table 1." httpUri="http://table.plazi.org/id/DF3766171935AD4C3AEBFD18EF0CFCB1" pageId="8" pageNumber="309" tableUuid="DF3766171935AD4C3AEBFD18EF0CFCB1">10</tableCitation>
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<taxonomicName id="4C484D1C1928AD513AEBF9B0EBC1F9A8" ID-CoL="4PQ8" box="[151,252,1590,1612]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="309" phylum="Chordata" rank="genus">
<emphasis id="B93CEA8D1928AD513AEBF9B0EBC1F9A8" box="[151,252,1590,1612]" italics="true" pageId="8" pageNumber="309">Gobiodon</emphasis>
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sp. 1 
<bibRefCitation id="EFD94B6E1928AD513B44F9B0E913F9A8" author="Herler" box="[312,558,1590,1612]" pageId="8" pageNumber="309" refString="Herler, J. &amp; Hilgers, H. (2005) A synopsis of coral and coral rock associated gobiid fishes from the Gulf of Aqaba, northern Red Sea. aqua, Journal of Ichthyology and Aquatic Biology, 10, 103 - 132." type="journal article" year="2005">Herler and Hilgers 2005</bibRefCitation>
(part): 120, Figs. 13a, b; 
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Herler 
<emphasis id="B93CEA8D1928AD51390BF9BEE89EF9A8" box="[887,931,1590,1613]" italics="true" pageId="8" pageNumber="309">et al</emphasis>
. (2009)
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: 733
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.
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<typeStatus id="54F3883D1928AD513AEBF9FFEA3BF976" box="[151,262,1657,1682]" pageId="8" pageNumber="309" type="holotype">Holotype</typeStatus>
.
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95078, 
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SL, Gulf of Aqaba, 
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, Dahab, “Islands” (
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, 
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), 
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, coll. J. Herler, 
<date id="FFF6105F1928AD513BFEF918E963F952" box="[386,606,1693,1718]" pageId="8" pageNumber="309" value="2005-11-13">13 November 2005</date>
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<typeStatus id="54F3883D1928AD513ABBF947EA7CF93E" box="[199,321,1729,1754]" pageId="8" pageNumber="309" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Four specimens: 
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95565, 
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SL, Gulf of Aqaba, 
<collectingCountry id="F35F760F1928AD513E54F944EF53F93E" box="[1064,1134,1730,1754]" name="Egypt" pageId="8" pageNumber="309">Egypt</collectingCountry>
, Dahab, “Napoleon Reef” (
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, 
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), 
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, coll. J. Herler, 
<date id="FFF6105F1928AD51388CF960E8F3F91A" box="[752,974,1765,1790]" pageId="8" pageNumber="309" value="2005-11-18">18 November 2005</date>
. 
<collectionCode id="ED59AE5A1928AD5139A4F960EF22F919" box="[984,1055,1766,1789]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="309">NMW</collectionCode>
95566 (
<collectingCountry id="F35F760F1928AD513E01F960EF9BF91A" box="[1149,1190,1766,1790]" name="Switzerland" pageId="8" pageNumber="309">CH</collectingCountry>
232-41-019 in 
<bibRefCitation id="EFD94B6E1928AD513F28F963EA6DF8C6" author="Herler" pageId="8" pageNumber="309" refString="Herler, J. &amp; Hilgers, H. (2005) A synopsis of coral and coral rock associated gobiid fishes from the Gulf of Aqaba, northern Red Sea. aqua, Journal of Ichthyology and Aquatic Biology, 10, 103 - 132." type="journal article" year="2005">Herler &amp; Hilgers, 2005</bibRefCitation>
), 
<quantity id="4CB09B7A1928AD513B1FF88FEAFAF8C5" box="[355,455,1801,1826]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="8" pageNumber="309" unit="mm" value="16.1">16.1 mm</quantity>
SL, 
<quantity id="4CB09B7A1928AD513B80F88CE915F8C5" box="[508,552,1802,1826]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="309" unit="m" value="3.0">3 m</quantity>
, 
<date id="FFF6105F1928AD513848F88CE983F8C6" box="[564,702,1802,1826]" pageId="8" pageNumber="309" value="2004-05-04">4 May 2004</date>
, other data same as 
<typeStatus id="54F3883D1928AD5139E1F88FEF3EF8C6" box="[925,1027,1801,1826]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
. 
<collectionCode id="ED59AE5A1928AD513E71F88CEF5FF8C5" LSID="urn:lsid:biocol.org:col:34988" box="[1037,1122,1802,1825]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="309">MNHN</collectionCode>
2006–1699, 
<quantity id="4CB09B7A1928AD513E89F88CEE64F8C5" box="[1269,1369,1802,1826]" metricMagnitude="-2" metricUnit="m" metricValue="3.2299999999999995" pageId="8" pageNumber="309" unit="mm" value="32.3">32.3 mm</quantity>
SL, 
<quantity id="4CB09B7A1928AD513FF2F88FEB92F8A1" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="309" unit="m" value="5.0">5 m</quantity>
, 
<date id="FFF6105F1928AD513AC7F8A8EAA8F8A2" box="[187,405,1837,1862]" pageId="8" pageNumber="309" value="2005-11-19">19 November 2005</date>
, other data same as 
<typeStatus id="54F3883D1928AD513808F8ABE9E7F8A2" box="[628,730,1837,1862]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
. 
<collectionCode id="ED59AE5A1928AD513898F8A8E805F8A1" box="[740,824,1838,1861]" pageId="8" pageNumber="309">BMNH</collectionCode>
2006.10.6.1, 
<quantity id="4CB09B7A1928AD5139AEF8ABEF05F8A1" box="[978,1080,1837,1862]" metricMagnitude="-2" metricUnit="m" metricValue="2.96" pageId="8" pageNumber="309" unit="mm" value="29.6">29.6 mm</quantity>
, 
<quantity id="4CB09B7A1928AD513E38F8A8EF4CF8A1" box="[1092,1137,1838,1862]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="309" unit="m" value="3.0">3 m</quantity>
, 
<date id="FFF6105F1928AD513E00F8A8EE6AF8A2" box="[1148,1367,1837,1862]" pageId="8" pageNumber="309" value="2005-11-17">17 November 2005</date>
, other data same as 
<typeStatus id="54F3883D1928AD513B57F8D7EAACF88E" box="[299,401,1873,1898]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BF7369F1928AD503ABBF8F3EE4AFF30" blockId="8.[151,1436,1657,2006]" lastBlockId="9.[151,1436,151,717]" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309">
<emphasis id="B93CEA8D1928AD513ABBF8F3EA86F86A" bold="true" box="[199,443,1909,1934]" pageId="8" pageNumber="309">Additional material.</emphasis>
<taxonomicName id="4C484D1C1928AD513BB8F8F3E989F869" box="[452,692,1909,1933]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B93CEA8D1928AD513BB8F8F3E989F869" box="[452,692,1909,1933]" italics="true" pageId="8" pageNumber="309">Gobiodon irregularis</emphasis>
</taxonomicName>
: 
<collectingCountry id="F35F760F1928AD5138B8F8F0E9D6F86A" box="[708,747,1910,1934]" name="Switzerland" pageId="8" pageNumber="309">CH</collectingCountry>
232-41b-057, 
<quantity id="4CB09B7A1928AD5139E8F8F0E8C7F869" box="[916,1018,1910,1934]" metricMagnitude="-2" metricUnit="m" metricValue="2.23" pageId="8" pageNumber="309" unit="mm" value="22.3">22.3 mm</quantity>
SL, 
<collectingCountry id="F35F760F1928AD513E4EF8F0EF45F86A" box="[1074,1144,1910,1934]" name="Egypt" pageId="8" pageNumber="309">Egypt</collectingCountry>
, Dahab, coll J. Herler, 
<date id="FFF6105F1928AD513FFCF8F0EA36F856" pageId="8" pageNumber="309" value="2012-05-13">13 May 2012</date>
; 
<collectingCountry id="F35F760F1928AD513B6BF81CEA03F856" box="[279,318,1946,1970]" name="Switzerland" pageId="8" pageNumber="309">CH</collectingCountry>
232-41-020, 
<quantity id="4CB09B7A1928AD513BA9F81CE905F855" box="[469,568,1946,1970]" metricMagnitude="-2" metricUnit="m" metricValue="3.12" pageId="8" pageNumber="309" unit="mm" value="31.2">31.2 mm</quantity>
SL, 
<collectingCountry id="F35F760F1928AD513810F81CE98FF856" box="[620,690,1946,1970]" name="Egypt" pageId="8" pageNumber="309">Egypt</collectingCountry>
, Dahab, coll J. Herler, 0 
<date id="FFF6105F1928AD5139C2F81CEF75F856" box="[958,1096,1946,1970]" pageId="8" pageNumber="309" value="2004-05-04">4 May 2004</date>
; 
<collectionCode id="ED59AE5A1928AD513E2FF81CEFB6F856" LSID="urn:lsid:biocol.org:col:34838" box="[1107,1163,1946,1970]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="309">SMF</collectionCode>
, uncatalogued, 
<quantity id="4CB09B7A1928AD513F45F81FEEA6F855" box="[1337,1435,1945,1970]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" pageId="8" pageNumber="309" unit="mm" value="27.6">27.6 mm</quantity>
SL, 
<collectingCountry id="F35F760F1928AD513AB4F83BEA5CF832" box="[200,353,1981,2006]" name="Saudi Arabia" pageId="8" pageNumber="309">Saudi Arabia</collectingCountry>
, Farasan Archipelago, coll. S.V. Bogorodsky, 
<date id="FFF6105F1928AD513909F838EF78F832" box="[885,1093,1981,2006]" pageId="8" pageNumber="309" value="2012-02-27">27 February 2012</date>
; 
<collectionCode id="ED59AE5A1928AD513E2FF838EFB3F831" box="[1107,1166,1982,2005]" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="8" pageNumber="309">PMR</collectionCode>
<collectionCode id="ED59AE5A1928AD513EE4F838EF83F832" box="[1176,1214,1982,2006]" pageId="8" pageNumber="309">VP</collectionCode>
3201, 
<quantity id="4CB09B7A1928AD513F78F83BEE56F831" box="[1284,1387,1981,2006]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="8" pageNumber="309" unit="mm" value="23.5">23.5 mm</quantity>
SL, 
<collectingCountry id="F35F760F1929AD503AEBFF1EEBE0FF54" box="[151,221,152,176]" name="Egypt" pageId="9" pageNumber="310">Egypt</collectingCountry>
, Sharm el Sheikh, Sharm el Moya, coll. S.V. Bogorodsky, 
<date id="FFF6105F1929AD50391BFF1EE8C3FF54" box="[871,1022,152,176]" pageId="9" pageNumber="310" value="2011-06-30">30 June 2011</date>
. 
<taxonomicName id="4C484D1C1929AD503E74FF1EEF45FF54" box="[1032,1144,152,176]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="310" phylum="Chordata" rank="genus">
<emphasis id="B93CEA8D1929AD503E74FF1EEF45FF54" box="[1032,1144,152,176]" italics="true" pageId="9" pageNumber="310">Gobiodon</emphasis>
</taxonomicName>
RW sp. 1 (identified as 
<taxonomicName id="4C484D1C1929AD503FF8FF1EEA32FF30" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="310" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B93CEA8D1929AD503FF8FF1EEA32FF30" italics="true" pageId="9" pageNumber="310">G. irregularis</emphasis>
</taxonomicName>
): 
<collectionCode id="ED59AE5A1929AD503B5BFF3AEA46FF30" LSID="urn:lsid:biocol.org:col:34984" box="[295,379,188,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="9" pageNumber="310">SAIAB</collectionCode>
70430, juvenile, 
<quantity id="4CB09B7A1929AD503842FF3AE9B0FF30" box="[574,653,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="310" unit="mm" value="15.0">15 mm</quantity>
SL, Indian Ocean, Rodrigues, coll. P. Heemstra, 
<date id="FFF6105F1929AD503EC8FF3AEE4EFF30" box="[1204,1395,188,213]" pageId="9" pageNumber="310" value="2001-10-22">22 October 2001</date>
.
</paragraph>
<paragraph id="8BF7369F1929AD503ABBFF59EAB2FE49" blockId="9.[151,1436,151,717]" pageId="9" pageNumber="310">
<emphasis id="B93CEA8D1929AD503ABBFF59EAE3FF1C" bold="true" box="[199,478,223,248]" pageId="9" pageNumber="310">Comparative material.</emphasis>
<taxonomicName id="4C484D1C1929AD503B97FF66EF9FFF1C" authority="Winterbottom &amp; Emery 1986" authorityName="Winterbottom &amp; Emery" authorityYear="1986" box="[491,1186,223,248]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="310" phylum="Chordata" rank="species" species="fulvus">
<emphasis id="B93CEA8D1929AD503B97FF66E966FF1C" box="[491,603,224,248]" italics="true" pageId="9" pageNumber="310">Gobiodon</emphasis>
cf. 
<emphasis id="B93CEA8D1929AD5038EFFF66E9EBFF1C" box="[659,726,224,248]" italics="true" pageId="9" pageNumber="310">fulvus</emphasis>
(sensu 
<bibRefCitation id="EFD94B6E1929AD503944FF59EFA4FF1C" author="Winterbottom" box="[824,1177,223,248]" pageId="9" pageNumber="310" refString="Winterbottom, R. &amp; Emery, A. R. (1986) Review of the gobioid fishes of the Chagos Archipelago, central Indian Ocean. Life Sciences Contributions, Royal Ontario Museum, Canada, 142, 1 - 82." type="journal article" year="1986">Winterbottom &amp; Emery 1986</bibRefCitation>
)
</taxonomicName>
: uncatalogued (
<collectionCode id="ED59AE5A1929AD503F21FF66EEA1FF1C" LSID="urn:lsid:biocol.org:col:15536" box="[1373,1436,224,248]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="9" pageNumber="310">DNA</collectionCode>
sample), 
<quantity id="4CB09B7A1929AD503A81FE82EA5DFEF8" box="[253,352,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="2.3899999999999997" pageId="9" pageNumber="310" unit="mm" value="23.9">23.9 mm</quantity>
SL, 
<collectingCountry id="F35F760F1929AD503BE8FE82EAC2FEF9" box="[404,511,260,285]" name="Maldives" pageId="9" pageNumber="310">Maldives</collectingCountry>
, Makunudu, coll. J. Herler, 
<date id="FFF6105F1929AD50394FFE82E8DDFEF8" box="[819,992,260,285]" pageId="9" pageNumber="310" value="2007-03-17">17 March 2007</date>
; uncatalogued (
<collectionCode id="ED59AE5A1929AD503EF2FE82EFF0FEF8" LSID="urn:lsid:biocol.org:col:15536" box="[1166,1229,260,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="9" pageNumber="310">DNA</collectionCode>
sample), 
<quantity id="4CB09B7A1929AD503F45FE82EEA1FEF8" box="[1337,1436,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="2.37" pageId="9" pageNumber="310" unit="mm" value="23.7">23.7 mm</quantity>
SL, southern 
<collectingCountry id="F35F760F1929AD503B53FEA1EAB9FEA4" box="[303,388,295,320]" name="Taiwan" pageId="9" pageNumber="310">Taiwan</collectingCountry>
, Kenting, coll. J. Herler, 0 
<date id="FFF6105F1929AD5038CCFEA1E840FEA4" box="[688,893,295,320]" pageId="9" pageNumber="310" value="2008-12-06">6 December 2008</date>
; 
<collectionCode id="ED59AE5A1929AD5039F5FEAEE8E2FEA4" box="[905,991,296,320]" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Sam Noble Oklahoma Museum of Natural History" pageId="9" pageNumber="310">OMNH</collectionCode>
P40259 (
<figureCitation id="13732A1A1929AD503E31FEA1EFADFEA4" box="[1101,1168,295,320]" captionStart="FIGURE 9" captionStartId="22.[151,250,1660,1682]" captionTargetBox="[151,1436,181,1640]" captionTargetId="figure@22.[151,1436,181,1641]" captionTargetPageId="22" captionText="FIGURE 9. Comparative material. Freshly collected adults: Gobiodon cf. bilineatus sp. nov. A. uncatalogued, 28.9 mm SL, Kagi Island, Maldives. B. uncatalogued, 22.9 mm SL, Kenting, southern Taiwan. C. Gobiodon prolixus, ROM 73338, holotype, male 26.2 mm SL, Nha Trang, Vietnam. D. Gobiodon sp. D sensu Munday et al. (1999), uncatalogued, 29.9 mm SL, Hembadhu Island, Maldives E. Gobiodon quinquestrigatus, uncatalogued, 28.7 mm SL, Kenting, southern Taiwan; F. Gobiodon cf. fulvus (sensu Winterbottom &amp; Emery 1986), OMNH P 40259, 20.8 mm SL, Iriomote Island, Japan. G. Gobiodon oculolineatus, OMNH P 40260, 22.6 mm SL, Okinawa Island, Japan. H. Gobiodon ceramensis, OMNH P 34042, 35 mm SL, Ryukyu Island, Japan. Gobiodon unicolor, preserved holotype, MNHN A- 4015, 31 mm SL, I. entire specimen. J. close-up of black opercular spot (arrow) on the right side. Photos by J. Herler (A, B, D, E, I, J), R. Winterbottom (C), T. Suzuki (F-H)." httpUri="https://zenodo.org/record/249079/files/figure.png" pageId="9" pageNumber="310">Fig. 9</figureCitation>
F), 
<quantity id="4CB09B7A1929AD503EC5FEAEEE23FEDB" box="[1209,1310,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="2.08" pageId="9" pageNumber="310" unit="mm" value="20.8">20.8 mm</quantity>
SL, 
<collectingCountry id="F35F760F1929AD503F29FEAEEEA5FEDB" box="[1365,1432,296,319]" name="Japan" pageId="9" pageNumber="310">Japan</collectingCountry>
, Ryukyu Islands, Iriomote Island, coll. T. Suzuki and M. Suzuki, 
<date id="FFF6105F1929AD5039F1FECAEF76FE80" box="[909,1099,332,356]" pageId="9" pageNumber="310" value="2004-08-22">22 August 2004</date>
. 
<taxonomicName id="4C484D1C1929AD503E24FECAEBEEFE6C" authority="Wu 1972" authorityName="Wu" authorityYear="1972" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="310" phylum="Chordata" rank="species" species="oculolineatus">
<emphasis id="B93CEA8D1929AD503E24FECAEE57FE80" box="[1112,1386,332,356]" italics="true" pageId="9" pageNumber="310">Gobiodon oculolineatus</emphasis>
Wu 1972
</taxonomicName>
: 
<collectionCode id="ED59AE5A1929AD503A9DFEF6EA0BFE6C" box="[225,310,368,392]" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Sam Noble Oklahoma Museum of Natural History" pageId="9" pageNumber="310">OMNH</collectionCode>
P40260 (
<figureCitation id="13732A1A1929AD503BD4FEE9EAD3FE6C" box="[424,494,367,392]" captionStart="FIGURE 9" captionStartId="22.[151,250,1660,1682]" captionTargetBox="[151,1436,181,1640]" captionTargetId="figure@22.[151,1436,181,1641]" captionTargetPageId="22" captionText="FIGURE 9. Comparative material. Freshly collected adults: Gobiodon cf. bilineatus sp. nov. A. uncatalogued, 28.9 mm SL, Kagi Island, Maldives. B. uncatalogued, 22.9 mm SL, Kenting, southern Taiwan. C. Gobiodon prolixus, ROM 73338, holotype, male 26.2 mm SL, Nha Trang, Vietnam. D. Gobiodon sp. D sensu Munday et al. (1999), uncatalogued, 29.9 mm SL, Hembadhu Island, Maldives E. Gobiodon quinquestrigatus, uncatalogued, 28.7 mm SL, Kenting, southern Taiwan; F. Gobiodon cf. fulvus (sensu Winterbottom &amp; Emery 1986), OMNH P 40259, 20.8 mm SL, Iriomote Island, Japan. G. Gobiodon oculolineatus, OMNH P 40260, 22.6 mm SL, Okinawa Island, Japan. H. Gobiodon ceramensis, OMNH P 34042, 35 mm SL, Ryukyu Island, Japan. Gobiodon unicolor, preserved holotype, MNHN A- 4015, 31 mm SL, I. entire specimen. J. close-up of black opercular spot (arrow) on the right side. Photos by J. Herler (A, B, D, E, I, J), R. Winterbottom (C), T. Suzuki (F-H)." httpUri="https://zenodo.org/record/249079/files/figure.png" pageId="9" pageNumber="310">Fig. 9</figureCitation>
G), 
<quantity id="4CB09B7A1929AD503860FEE9E9BEFE63" box="[540,643,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="2.26" pageId="9" pageNumber="310" unit="mm" value="22.6">22.6 mm</quantity>
SL, 
<collectingCountry id="F35F760F1929AD5038C3FEF6E83FFE63" box="[703,770,368,391]" name="Japan" pageId="9" pageNumber="310">Japan</collectingCountry>
, Ryukyu Islands, Okinawa Island, coll. T. Suzuki and M. Suzuki, 0 
<date id="FFF6105F1929AD503B7CFE12EAB6FE49" box="[256,395,404,429]" pageId="9" pageNumber="310" value="1996-05-04">4 May 1996</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C35265141929AD503ABBFE31E885FD29" pageId="9" pageNumber="310" type="diagnosis">
<paragraph id="8BF7369F1929AD503ABBFE31E885FD29" blockId="9.[151,1436,151,717]" pageId="9" pageNumber="310">
<emphasis id="B93CEA8D1929AD503ABBFE31EA7CFE34" bold="true" box="[199,321,439,464]" pageId="9" pageNumber="310">Diagnosis.</emphasis>
Dorsal-fin rays VI + I,10-11 (usually 11); anal-fin rays I,9-10 (usually 9); head and body naked; body relatively elongate (depth 35.1–39% SL) and compressed; head slightly pointed in juveniles, becoming more rounded in adults; snout sometimes with a slight hump above upper lip; no groove between isthmus and interopercle; caudal peduncle slender (depth 14.5–15.8% SL); caudal fin relatively long (22.6–24.6% SL). Juveniles and subadults greenish or brownish green with up to seven red bars with bluish interspaces on head and pectoral-fin base, three anteriormost bars also run across eye; nape and dorsal part of body with irregular red lines and small spots; lines and dots usually vanish in adults of more than 
<quantity id="4CB09B7A1929AD5039DEFD16E8E7FD43" box="[930,986,656,680]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="310" unit="cm" value="3.0">3 cm</quantity>
TL; body becomes uniformly brown or reddish brown, usually with remnants of the orbital and suborbital bars.
</paragraph>
</subSubSection>
<caption id="DF3766171929AD503AEBF882EFC4F8DC" httpUri="https://zenodo.org/record/249073/files/figure.png" pageId="9" pageNumber="310" targetBox="[151,1436,766,1774]" targetPageId="9">
<paragraph id="8BF7369F1929AD503AEBF882EFC4F8DC" blockId="9.[151,1436,1796,1848]" pageId="9" pageNumber="310">
<emphasis id="B93CEA8D1929AD503AEBF882EA2EF8FD" bold="true" box="[151,275,1796,1818]" pageId="9" pageNumber="310">FIGURE 3.</emphasis>
<taxonomicName id="4C484D1C1929AD503B65F882EACCF8FE" box="[281,497,1796,1818]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="310" phylum="Chordata" rank="species" species="irregularis" status="sp. nov.">
<emphasis id="B93CEA8D1929AD503B65F882EACCF8FE" box="[281,497,1796,1818]" italics="true" pageId="9" pageNumber="310">Gobiodon irregularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57F61929AD503B84F883E97BF8FE" box="[504,582,1797,1818]" pageId="9" pageNumber="310" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B93CEA8D1929AD503830F882E958F8FD" bold="true" box="[588,613,1796,1817]" pageId="9" pageNumber="310">A.</emphasis>
freshly collected adult, holotype, NMW 95078, 30.1 mm SL, Dahab, Egypt, Red Sea. 
<emphasis id="B93CEA8D1929AD503AB4F8A5EBE2F8DC" bold="true" box="[200,223,1827,1848]" pageId="9" pageNumber="310">B.</emphasis>
freshly collected subadult, CH 232-41b-057, 22.3 mm SL, Dahab, Egypt, Red Sea. Photos by J. Herler.
</paragraph>
</caption>
<subSubSection id="C35265141929AD523ABBF8E3EA5CFDFC" lastPageId="11" lastPageNumber="312" pageId="9" pageNumber="310" type="description">
<paragraph id="8BF7369F1929AD533ABBF8E3E926FDFE" blockId="9.[151,1437,1893,2026]" lastBlockId="10.[151,1436,151,864]" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310">
<emphasis id="B93CEA8D1929AD503ABBF8E3EA6CF89A" bold="true" box="[199,337,1893,1918]" pageId="9" pageNumber="310">Description</emphasis>
(based on 5 
<typeStatus id="54F3883D1929AD503B9AF8E0E91CF89A" box="[486,545,1894,1918]" pageId="9" pageNumber="310">types</typeStatus>
and several non-type specimens (for osteology)). Head and body compressed. Head relatively deep, posterior part of body slender, caudal peduncle low. Body proportions and meristics for 
<typeStatus id="54F3883D1929AD503F1DF80CEEA1F846" box="[1377,1436,1930,1954]" pageId="9" pageNumber="310">types</typeStatus>
are provided in 
<tableCitation id="C6CA03241929AD503B31F82BEA91F822" box="[333,428,1965,1990]" captionStart="TABLE 4" captionStartId="11.[151,239,584,607]" captionTargetBox="[159,1412,701,867]" captionTargetPageId="11" captionText="TABLE 4. Body proportions of holotype and three adult paratypes of Gobiodon irregularis sp. nov. from the northern Red Sea. The juvenile paratype NMW 95566 (SL = 16.1 mm) is excluded here. Values are proportions of standard length (SL) and head length (last five measurements), respectively, means and the first standard deviation (SD)." httpUri="http://table.plazi.org/id/DF376617192BAD523AEBFDCEEFC9FD45" pageId="9" pageNumber="310" tableUuid="DF376617192BAD523AEBFDCEEFC9FD45">Tables 4</tableCitation>
and 
<tableCitation id="C6CA03241929AD503B9AF82BEACBF822" box="[486,502,1965,1990]" captionStart="TABLE 5" captionStartId="12.[151,239,152,175]" captionTargetBox="[159,1330,237,610]" captionTargetPageId="12" captionText="TABLE 5. Fin counts of holotype and four paratypes of Gobiodon irregularis sp. nov. from the northern Red Sea. d = damaged." httpUri="http://table.plazi.org/id/DF376617192CAD553AEBFF1EEBC2FF35" pageId="9" pageNumber="310" tableUuid="DF376617192CAD553AEBFF1EEBC2FF35">5</tableCitation>
, respectively. Dorsal-fin rays VI + I,10– 
<emphasis id="B93CEA8D1929AD5039BAF82BE8E2F822" bold="true" box="[966,991,1965,1990]" pageId="9" pageNumber="310">11</emphasis>
(10:1, 11:4); anal-fin rays I, 
<emphasis id="B93CEA8D1929AD503F5BF82BEE08F822" bold="true" box="[1319,1333,1965,1990]" pageId="9" pageNumber="310">9</emphasis>
–10 (9:4, 10:1); pectoral-fin rays 20 (n = 5); pelvic-fin rays I,5 (all specimens); caudal fin with 15–17 segmented and branched rays; disc short (not reaching anus) and cup-shaped with significant frenum between spines. First dorsal fin as high as D 
<quantity id="4CB09B7A192AAD533B31FF3AEA44FF31" box="[333,377,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="311" unit="in" value="2.0">2 in</quantity>
juveniles and somewhat triangular, but lower than D 
<quantity id="4CB09B7A192AAD5339ABFF3AEF3EFF31" box="[983,1027,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="311" unit="in" value="2.0">2 in</quantity>
adults. No scales. Gill opening less wide than pectoral-fin base, ending ventrally in opposite of 3rd or 4th lower pectoral-fin ray. Gill rakers 0–3 + 7–8. No groove between interopercle and isthmus. Mouth terminal, relatively straight. Upper jaw reaching to about below anterior margin of orbit or to mid-orbit. Upper lip slightly curved. One outer row of up to 10 larger, slightly recurved teeth in upper and lower jaw, and increasing in size towards symphysis. Several rows of small, slender and recurved teeth in both jaws behind the outer row. Lower jaw with a pair of large, postsymphysial canines on each side, one often much smaller or absent, probably due to tooth loss and replacement. Anterior and posterior nasal openings at the end of short tubes. Head sensory canals typical for 
<taxonomicName id="4C484D1C192AAD5339BCFE3DEEAAFE37" authority="Winterbottom &amp; Harold 2005" authorityName="Winterbottom &amp; Harold" authorityYear="2005" box="[960,1431,442,467]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="311" phylum="Chordata" rank="genus">
<emphasis id="B93CEA8D192AAD5339BCFE3DEF0DFE37" box="[960,1072,443,467]" italics="true" pageId="10" pageNumber="311">Gobiodon</emphasis>
(
<bibRefCitation id="EFD94B6E192AAD533E42FE3CEEADFE37" author="Winterbottom" box="[1086,1424,442,467]" pageId="10" pageNumber="311" refString="Winterbottom, R. &amp; Harold, A. S. (2005) Gobiodon prolixus, a new species of gobiid fish (Teleostei: Perciformes: Gobiidae) from the Indo-west Pacific. Proceedings of the Biological Society of Washington, 118 (3), 582 - 589." type="journal article" year="2005">Winterbottom &amp; Harold 2005</bibRefCitation>
)
</taxonomicName>
, with anterior oculoscapular (pores NA (paired), AI, PI (unpaired), SO, 
<collectingCountry id="F35F760F192AAD5339CCFE59E8E5FE13" box="[944,984,479,503]" name="Austria" pageId="10" pageNumber="311">AO</collectingCountry>
and IT (paired)) and preopercular (three pores on each side) canals present.
</paragraph>
<paragraph id="8BF7369F192AAD533ABBFDA0E881FCBB" blockId="10.[151,1436,151,864]" pageId="10" pageNumber="311">
<emphasis id="B93CEA8D192AAD533ABBFDA0EAADFDDB" bold="true" box="[199,400,550,575]" pageId="10" pageNumber="311">Life colouration.</emphasis>
Juveniles greenish with seven broad red bars on head and pectoral-fin base (
<figureCitation id="13732A1A192AAD533E82FDA0EE7EFDDB" box="[1278,1347,550,575]" captionStart="FIGURE 3" captionStartId="9.[151,250,1796,1818]" captionTargetBox="[151,1436,766,1774]" captionTargetId="figure@9.[151,1436,766,1774]" captionTargetPageId="9" captionText="FIGURE 3. Gobiodon irregularis sp. nov. A. freshly collected adult, holotype, NMW 95078, 30.1 mm SL, Dahab, Egypt, Red Sea. B. freshly collected subadult, CH 232 - 41 b- 057, 22.3 mm SL, Dahab, Egypt, Red Sea. Photos by J. Herler." httpUri="https://zenodo.org/record/249073/files/figure.png" pageId="10" pageNumber="311">Fig. 3</figureCitation>
B): first from orbit to upper lip, second and third through orbit and across cheek, fourth to sixth, wider and somewhat wavy, across opercle, seventh wide across pectoral-fin base; interspaces bluish or greyish; red bars become irregular and wavy on postorbital area; nape and dorsal half of body with red vermiculations; both dorsal fins sometimes with narrow bright bluish band along the base; banded pattern of internal pigment along vertebral column; median fins greenish. Subadults brownish green or brown with small red spots and irregular short lines on nape and dorsal part of body; only the three more distinct red suborbital head bars remain visible (
<figureCitation id="13732A1A192AAD53398AFD78EF04FCF3" box="[1014,1081,766,791]" captionStart="FIGURE 4" captionStartId="10.[151,250,1841,1863]" captionTargetBox="[151,1436,918,1819]" captionTargetId="figure@10.[151,1436,918,1819]" captionTargetPageId="10" captionText="FIGURE 4. Gobiodon irregularis sp. nov. A. freshly collected adult, paratype MNHN 2006 - 1699, 32.3 mm SL, Dahab, Egypt, Red Sea; B. freshly collected adult, CH 232 - 41 - 020, 31.2 mm SL, Dahab, Egypt, Red Sea; C. freshly collected adult, SMF uncatalogued, 27.6 mm SL, Farasan Archipelago, Red Sea; D. freshly collected subadult, paratype NMW 95565, 23.4 mm SL, Dahab, Egypt, Red Sea; E. alive subadult, PMR VP 3201, 23.5 mm SL, Sharm el Moya, Egypt, Red Sea; F. freshly collected juvenile, SAIAB 70430, 15 mm SL, Rodrigues, western Indian Ocean. Photos by J. Herler (A, B, D), S. V. Bogorodsky (C, E), P. Heemstra, © SAIAB (F)." httpUri="https://zenodo.org/record/249074/files/figure.png" pageId="10" pageNumber="311">Fig. 4</figureCitation>
B–E). Adults uniformly reddish brown or brown, including fins (
<figureCitation id="13732A1A192AAD533860FCA4E953FCDF" box="[540,622,802,827]" captionStart="FIGURE 3" captionStartId="9.[151,250,1796,1818]" captionTargetBox="[151,1436,766,1774]" captionTargetId="figure@9.[151,1436,766,1774]" captionTargetPageId="9" captionText="FIGURE 3. Gobiodon irregularis sp. nov. A. freshly collected adult, holotype, NMW 95078, 30.1 mm SL, Dahab, Egypt, Red Sea. B. freshly collected subadult, CH 232 - 41 b- 057, 22.3 mm SL, Dahab, Egypt, Red Sea. Photos by J. Herler." httpUri="https://zenodo.org/record/249073/files/figure.png" pageId="10" pageNumber="311">Figs. 3</figureCitation>
A, 4A), nape and upper half of body covered with dark-brown dots; remnants of orbital bars, extending onto suborbital area, may be visible.
</paragraph>
<caption id="DF376617192AAD533AEBF8B7EA93F804" httpUri="https://zenodo.org/record/249074/files/figure.png" pageId="10" pageNumber="311" targetBox="[151,1436,918,1819]" targetPageId="10">
<paragraph id="8BF7369F192AAD533AEBF8B7EA93F804" blockId="10.[151,1436,1841,2016]" pageId="10" pageNumber="311">
<emphasis id="B93CEA8D192AAD533AEBF8B7EA2FF8A2" bold="true" box="[151,274,1841,1863]" pageId="10" pageNumber="311">FIGURE 4.</emphasis>
<taxonomicName id="4C484D1C192AAD533B64F8B7EACCF8A3" box="[280,497,1841,1863]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="311" phylum="Chordata" rank="species" species="irregularis" status="sp. nov.">
<emphasis id="B93CEA8D192AAD533B64F8B7EACCF8A3" box="[280,497,1841,1863]" italics="true" pageId="10" pageNumber="311">Gobiodon irregularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57F6192AAD533B8AF8B4E978F8A3" box="[502,581,1842,1863]" pageId="10" pageNumber="311" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B93CEA8D192AAD533836F8B7E95EF8A2" bold="true" box="[586,611,1841,1862]" pageId="10" pageNumber="311">A.</emphasis>
freshly collected adult, paratype MNHN 2006-1699, 32.3 mm SL, Dahab, Egypt, Red Sea; 
<emphasis id="B93CEA8D192AAD533A80F8D6EA2EF881" bold="true" box="[252,275,1872,1893]" pageId="10" pageNumber="311">B.</emphasis>
freshly collected adult, CH 232-41-020, 31.2 mm SL, Dahab, Egypt, Red Sea; 
<emphasis id="B93CEA8D192AAD533E28F8D6EF51F882" bold="true" box="[1108,1132,1872,1894]" pageId="10" pageNumber="311">C.</emphasis>
freshly collected adult, SMF uncatalogued, 27.6 mm SL, Farasan Archipelago, Red Sea; 
<emphasis id="B93CEA8D192AAD53388AF8E9E832F860" bold="true" box="[758,783,1903,1924]" pageId="10" pageNumber="311">D.</emphasis>
freshly collected subadult, paratype NMW 95565, 23.4 mm SL, Dahab, Egypt, Red Sea; 
<emphasis id="B93CEA8D192AAD533BE9F80BEA91F846" bold="true" box="[405,428,1933,1954]" pageId="10" pageNumber="311">E.</emphasis>
alive subadult, PMR VP3201, 23.5 mm SL, Sharm el Moya, Egypt, Red Sea; 
<emphasis id="B93CEA8D192AAD533EABF80BEFD4F846" bold="true" box="[1239,1257,1933,1954]" pageId="10" pageNumber="311">F.</emphasis>
freshly collected juvenile, SAIAB 70430, 15 mm SL, Rodrigues, western Indian Ocean. Photos by J. Herler (A, B, D), S.V. Bogorodsky (C, E), P. Heemstra, © SAIAB (F).
</paragraph>
</caption>
<paragraph id="8BF7369F192BAD523ABBFF11E99BFF31" blockId="11.[151,1436,151,536]" pageId="11" pageNumber="312">
<emphasis id="B93CEA8D192BAD523ABBFF11EAEAFF54" bold="true" box="[199,471,151,176]" pageId="11" pageNumber="312">Preserved colouration.</emphasis>
Specimens uniformly light or dark brown. Bars and dots in juveniles and subadults may be retained. Bars through eyes not visible.
</paragraph>
<paragraph id="8BF7369F192BAD523ABBFF59EF3BFE6C" blockId="11.[151,1436,151,536]" pageId="11" pageNumber="312">
<emphasis id="B93CEA8D192BAD523ABBFF59EA92FF1C" bold="true" box="[199,431,223,248]" pageId="11" pageNumber="312">Molecular genetics.</emphasis>
The present analysis includes the sequences of the specimen with the original number 
<emphasis id="B93CEA8D192BAD523FF8FF66EEA8FF13" box="[1412,1429,224,247]" italics="true" pageId="11" pageNumber="312">G</emphasis>
. sp. 1_GA1 (
<bibRefCitation id="EFD94B6E192BAD523B63FE82EAD5FEF8" author="Herler" box="[287,488,260,285]" pageId="11" pageNumber="312" refString="Herler, J., Koblmueller, S. &amp; Sturmbauer, C. (2009) Phylogenetic relationships of coral-associated gobies (Teleostei, Gobiidae) from the Red Sea based on mitochondrial DNA data. Marine Biology, 156, 725 - 739. http: // dx. doi. org / 10.1007 / s 00227 - 008 - 1124 - 7" type="journal article" year="2009">
Herler 
<emphasis id="B93CEA8D192BAD523B13FE83EA9DFEF8" box="[367,416,260,284]" italics="true" pageId="11" pageNumber="312">et al</emphasis>
. 2009
</bibRefCitation>
) from the Gulf of Aqaba, Red Sea as well as of one of the 
<typeStatus id="54F3883D192BAD523E07FE83EFDBFEF9" box="[1147,1254,261,285]" pageId="11" pageNumber="312" type="paratype">paratypes</typeStatus>
(NMW 95565). See 
<tableCitation id="C6CA0324192BAD523AB0FEA1EA19FEA4" box="[204,292,295,320]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionTargetBox="[159,1431,362,1898]" captionTargetPageId="4" captionText="TABLE 1. GenBank accession numbers (GB No.) and references for published sequences (12 S and 16 S rRNA) of 25 taxa (4 new species are underlined) of Gobiodon (G.) and Paragobiodon (P.). Localities: GBR = Great Barrier Reef, Mal = Maldives, PNG = Papua New Guinea, RS = Red Sea, Tai = Taiwan. Abbreviations (PM) and (JH) refer to the first demonstration of sequenced but undescribed species, Munday et al. (1999) and Herler and Hilgers (2009), respectively. Superscript numbers in locality code indicate type status: a: Paratype MNHN 2012 - 0111, b: Paratype BMNH 2006.10. 6.1, c: Paratype NMW 95565." httpUri="http://table.plazi.org/id/DF3766171924AD5D3AEBFF1EE90BFEAB" pageId="11" pageNumber="312" tableUuid="DF3766171924AD5D3AEBFF1EE90BFEAB">Table 1</tableCitation>
for Genbank accession numbers. The genetically closest species to 
<taxonomicName id="4C484D1C192BAD523E28FEAEEFCDFEA4" box="[1108,1264,296,320]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="312" phylum="Chordata" rank="species" species="irregularis" status="sp. nov.">
<emphasis id="B93CEA8D192BAD523E28FEAEEFCDFEA4" box="[1108,1264,296,320]" italics="true" pageId="11" pageNumber="312">G. irregularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57F6192BAD523E80FEAEEE64FEDB" box="[1276,1369,296,319]" pageId="11" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
is 
<taxonomicName id="4C484D1C192BAD523FF8FEAEE99EFE80" authority="Wu, 1972 (Duchene et al. 2013)" authorityName="Wu, 1972 (Duchene et al." authorityYear="2013" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="312" phylum="Chordata" rank="species" species="oculolineatus">
<emphasis id="B93CEA8D192BAD523FF8FEAEEA13FE80" italics="true" pageId="11" pageNumber="312">G. oculolineatus</emphasis>
Wu, 1972 (
<bibRefCitation id="EFD94B6E192BAD523BCEFECAE9A1FE80" author="Duchene" box="[434,668,332,357]" pageId="11" pageNumber="312" refString="Duchene, D., Klanten, S. O., Munday, P. L., Herler, J. &amp; van Herwerden, L. (2013) Phylogenetic evidence for recent diversification of obligate coral-dwelling gobies compared with their host corals. Molecular Phylogenetics and Evolution, 69: 123 - 132. http: // dx. doi. org / 10.1016 / j. ympev. 2013.04.033" type="journal article" year="2013">
Duchene 
<emphasis id="B93CEA8D192BAD523863FECBE96CFE80" box="[543,593,332,356]" italics="true" pageId="11" pageNumber="312">et al</emphasis>
. 2013
</bibRefCitation>
)
</taxonomicName>
. The 
<emphasis id="B93CEA8D192BAD52389EFECBE9CDFE80" box="[738,752,333,356]" italics="true" pageId="11" pageNumber="312">p</emphasis>
-distance based on 12S and 16S rRNA comparisons between these two species is 0.013 (1.3% genetic difference; Herler, unpublished data).
</paragraph>
<paragraph id="8BF7369F192BAD523ABBFE12EA5CFDFC" blockId="11.[151,1436,151,536]" pageId="11" pageNumber="312">
<emphasis id="B93CEA8D192BAD523ABBFE12EA17FE49" bold="true" box="[199,298,404,429]" pageId="11" pageNumber="312">Habitat.</emphasis>
<taxonomicName id="4C484D1C192BAD523B4DFE12E91DFE48" box="[305,544,404,428]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="312" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B93CEA8D192BAD523B4DFE12E91DFE48" box="[305,544,404,428]" italics="true" pageId="11" pageNumber="312">Gobiodon irregularis</emphasis>
</taxonomicName>
displays a generalised habitat selection and occupies a great range of 
<taxonomicName id="4C484D1C192BAD523F4EFE12EEA1FE4F" box="[1330,1436,404,427]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="11" pageNumber="312" phylum="Cnidaria" rank="genus">
<emphasis id="B93CEA8D192BAD523F4EFE12EEA1FE4F" box="[1330,1436,404,427]" italics="true" pageId="11" pageNumber="312">Acropora</emphasis>
</taxonomicName>
corals. It occurs in deeper water regions (lower reef slope and fore reef areas) and is most common in corals such as 
<taxonomicName id="4C484D1C192BAD523AEBFE5AEABDFE10" box="[151,384,476,500]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="11" pageNumber="312" phylum="Cnidaria" rank="species" species="samoensis">
<emphasis id="B93CEA8D192BAD523AEBFE5AEABDFE10" box="[151,384,476,500]" italics="true" pageId="11" pageNumber="312">Acropora samoensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C484D1C192BAD523BEEFE5AEAC6FE10" box="[402,507,476,500]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="11" pageNumber="312" phylum="Cnidaria" rank="species" species="valida">
<emphasis id="B93CEA8D192BAD523BEEFE5AEAC6FE10" box="[402,507,476,500]" italics="true" pageId="11" pageNumber="312">A. valida</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C484D1C192BAD523847FE5AE999FE10" box="[571,676,476,500]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="11" pageNumber="312" phylum="Cnidaria" rank="species" species="secale">
<emphasis id="B93CEA8D192BAD523847FE5AE999FE10" box="[571,676,476,500]" italics="true" pageId="11" pageNumber="312">A. secale</emphasis>
</taxonomicName>
but was also observed in 
<taxonomicName id="4C484D1C192BAD52399CFE5AEF44FE10" box="[992,1145,476,500]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="11" pageNumber="312" phylum="Cnidaria" rank="species" species="eurystoma">
<emphasis id="B93CEA8D192BAD52399CFE5AEF44FE10" box="[992,1145,476,500]" italics="true" pageId="11" pageNumber="312">A. eurystoma</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C484D1C192BAD523EC4FE5AEE70FE10" box="[1208,1357,476,500]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="11" pageNumber="312" phylum="Cnidaria" rank="species" species="pharaonis">
<emphasis id="B93CEA8D192BAD523EC4FE5AEE70FE10" box="[1208,1357,476,500]" italics="true" pageId="11" pageNumber="312">A. pharaonis</emphasis>
</taxonomicName>
in the northern Red Sea.
</paragraph>
</subSubSection>
<caption id="DF376617192BAD523AEBFDCEEFC9FD45" ID-Table-UUID="DF376617192BAD523AEBFDCEEFC9FD45" httpUri="http://table.plazi.org/id/DF376617192BAD523AEBFDCEEFC9FD45" pageId="11" pageNumber="312" targetBox="[159,1412,701,867]" targetIsTable="true" targetPageId="11">
<paragraph id="8BF7369F192BAD523AEBFDCEEFC9FD45" blockId="11.[151,1436,583,673]" pageId="11" pageNumber="312">
<emphasis id="B93CEA8D192BAD523AEBFDCEEA37FDBB" bold="true" box="[151,266,584,607]" pageId="11" pageNumber="312">TABLE 4.</emphasis>
Body proportions of holotype and three adult paratypes of 
<taxonomicName id="4C484D1C192BAD5239EFFDC1EF44FDBB" box="[915,1145,583,607]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="312" phylum="Chordata" rank="species" species="irregularis" status="sp. nov.">
<emphasis id="B93CEA8D192BAD5239EFFDC1EF44FDBB" box="[915,1145,583,607]" italics="true" pageId="11" pageNumber="312">Gobiodon irregularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57F6192BAD523EFDFDCEEFE8FDBB" box="[1153,1237,584,607]" pageId="11" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
from the northern Red Sea. The juvenile paratype NMW 95566 (SL = 16.1 mm) is excluded here. Values are proportions of standard length (SL) and head length (last five measurements), respectively, means and the first standard deviation (SD).
</paragraph>
</caption>
<paragraph id="8BF7369F192BAD523AE3FD5AEE6AFC87" blockId="11.[159,1412,701,1960]" pageId="11" pageNumber="312">
<table id="F948C43F192B52A63AE3FD3BEEB9FC87" box="[159,1412,701,867]" gridcols="5" gridrows="3" pageId="11" pageNumber="312">
<tr id="357834DD192B52A63AE3FD3BEEB9FCF4" box="[159,1412,701,784]" gridrow="0" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3FD3BEAF2FCF4" box="[159,463,701,784]" gridcol="0" gridrow="0" pageId="11" pageNumber="312">Status Coll.No.</th>
<th id="76A95DA1192B52A63863FD3BE814FCF4" box="[543,809,701,784]" gridcol="1" gridrow="0" pageId="11" pageNumber="312">Holotype Paratype NMW 95078 NMW 95565</th>
<th id="76A95DA1192B52A639FEFD3BE8CDFCF4" box="[898,1008,701,784]" gridcol="2" gridrow="0" pageId="11" pageNumber="312">Paratype MNHN 2006-1699</th>
<th id="76A95DA1192B52A63E49FD3BEF90FCF4" box="[1077,1197,701,784]" gridcol="3" gridrow="0" pageId="11" pageNumber="312">Paratype BMNH 2006.10.6.2</th>
<th id="76A95DA1192B52A63E88FD3BEEB9FCF4" box="[1268,1412,701,784]" gridcol="4" gridrow="0" pageId="11" pageNumber="312">MEAN (±SD)</th>
</tr>
<tr id="357834DD192B52A63AE3FCA2EEB9FCDE" box="[159,1412,804,826]" gridrow="1" pageId="11" pageNumber="312" rowspan-4="1">
<th id="76A95DA1192B52A63AE3FCA2EAF2FCDE" box="[159,463,804,826]" gridcol="0" gridrow="1" pageId="11" pageNumber="312">SL (mm)</th>
<td id="76A95DA1192B52A63863FCA2E814FCDE" box="[543,809,804,826]" gridcol="1" gridrow="1" pageId="11" pageNumber="312">30.1 23.4</td>
<td id="76A95DA1192B52A639FEFCA2E8CDFCDE" box="[898,1008,804,826]" gridcol="2" gridrow="1" pageId="11" pageNumber="312">32.3</td>
<td id="76A95DA1192B52A63E49FCA2EF90FCDE" box="[1077,1197,804,826]" gridcol="3" gridrow="1" pageId="11" pageNumber="312">29.6</td>
</tr>
<tr id="357834DD192B52A63AE3FCCBEEB9FC87" box="[159,1412,845,867]" gridrow="2" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3FCCBEAF2FC87" box="[159,463,845,867]" gridcol="0" gridrow="2" pageId="11" pageNumber="312">Snout to first dorsal-fin origin</th>
<td id="76A95DA1192B52A63863FCCBE814FC87" box="[543,809,845,867]" gridcol="1" gridrow="2" pageId="11" pageNumber="312">37.9 35.4</td>
<td id="76A95DA1192B52A639FEFCCBE8CDFC87" box="[898,1008,845,867]" gridcol="2" gridrow="2" pageId="11" pageNumber="312">36.9</td>
<td id="76A95DA1192B52A63E49FCCBEF90FC87" box="[1077,1197,845,867]" gridcol="3" gridrow="2" pageId="11" pageNumber="312">38.1</td>
<td id="76A95DA1192B52A63E88FCCBEEB9FC87" box="[1268,1412,845,867]" gridcol="4" gridrow="2" pageId="11" pageNumber="312">37.0 (1.3)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF7369F192BAD523AE3FCF0EE6AF86E" blockId="11.[159,1412,701,1960]" pageId="11" pageNumber="312">
<table id="F948C43F192B52A63AE3FCF0EEB9F84C" box="[159,1412,886,1960]" gridcols="5" gridrows="22" pageId="11" pageNumber="312">
<tr id="357834DD192B52A63AE3FCF0EEB9FC4F" box="[159,1412,886,939]" gridrow="0" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3FCF0E976FC4F" box="[159,587,886,939]" gridcol="0" gridrow="0" pageId="11" pageNumber="312">First dorsal-fin origin to second 24.7 dorsal-fin origin</th>
<th id="76A95DA1192B52A638ACFCF0E9C0FC4F" box="[720,765,886,939]" gridcol="1" gridrow="0" pageId="11" pageNumber="312">23.4</th>
<th id="76A95DA1192B52A639FEFCF0E893FC4F" box="[898,942,886,939]" gridcol="2" gridrow="0" pageId="11" pageNumber="312">24.3</th>
<th id="76A95DA1192B52A63E48FCF0EF5CFC4F" box="[1076,1121,886,939]" gridcol="3" gridrow="0" pageId="11" pageNumber="312">24.3</th>
<th id="76A95DA1192B52A63E88FCF0EEB9FC4F" box="[1268,1412,886,939]" gridcol="4" gridrow="0" pageId="11" pageNumber="312">24.1 (0.6)</th>
</tr>
<tr id="357834DD192B52A63AE3FC38EEB9FC17" box="[159,1412,958,1011]" gridrow="1" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3FC38E976FC17" box="[159,587,958,1011]" gridcol="0" gridrow="1" pageId="11" pageNumber="312">Second dorsal-fin origin to anal-fin 33.8 origin</th>
<td id="76A95DA1192B52A638ACFC38E9C0FC17" box="[720,765,958,1011]" gridcol="1" gridrow="1" pageId="11" pageNumber="312">31.7</td>
<td id="76A95DA1192B52A639FEFC38E893FC17" box="[898,942,958,1011]" gridcol="2" gridrow="1" pageId="11" pageNumber="312">31.5</td>
<td id="76A95DA1192B52A63E48FC38EF5CFC17" box="[1076,1121,958,1011]" gridcol="3" gridrow="1" pageId="11" pageNumber="312">35.1</td>
<td id="76A95DA1192B52A63E88FC38EEB9FC17" box="[1268,1412,958,1011]" gridcol="4" gridrow="1" pageId="11" pageNumber="312">33.0 (1.7)</td>
</tr>
<tr id="357834DD192B52A63AE3FB80EEB9FBF8" box="[159,1412,1030,1052]" gridrow="2" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3FB80E976FBF8" box="[159,587,1030,1052]" gridcol="0" gridrow="2" pageId="11" pageNumber="312">Pelvic-fin origin to anal-fin origin 25.3</th>
<td id="76A95DA1192B52A638ACFB80E9C0FBF8" box="[720,765,1030,1052]" gridcol="1" gridrow="2" pageId="11" pageNumber="312">26.4</td>
<td id="76A95DA1192B52A639FEFB80E893FBF8" box="[898,942,1030,1052]" gridcol="2" gridrow="2" pageId="11" pageNumber="312">26.7</td>
<td id="76A95DA1192B52A63E48FB80EF5CFBF8" box="[1076,1121,1030,1052]" gridcol="3" gridrow="2" pageId="11" pageNumber="312">25.8</td>
<td id="76A95DA1192B52A63E88FB80EEB9FBF8" box="[1268,1412,1030,1052]" gridcol="4" gridrow="2" pageId="11" pageNumber="312">26.1 (0.6)</td>
</tr>
<tr id="357834DD192B52A63AE3FBA9EEB9FBA1" box="[159,1412,1071,1093]" gridrow="3" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3FBA9E976FBA1" box="[159,587,1071,1093]" gridcol="0" gridrow="3" pageId="11" pageNumber="312">Snout to pelvic-fin origin 37.6</th>
<td id="76A95DA1192B52A638ACFBA9E9C0FBA1" box="[720,765,1071,1093]" gridcol="1" gridrow="3" pageId="11" pageNumber="312">38.0</td>
<td id="76A95DA1192B52A639FEFBA9E893FBA1" box="[898,942,1071,1093]" gridcol="2" gridrow="3" pageId="11" pageNumber="312">36.5</td>
<td id="76A95DA1192B52A63E48FBA9EF5CFBA1" box="[1076,1121,1071,1093]" gridcol="3" gridrow="3" pageId="11" pageNumber="312">37.6</td>
<td id="76A95DA1192B52A63E88FBA9EEB9FBA1" box="[1268,1412,1071,1093]" gridcol="4" gridrow="3" pageId="11" pageNumber="312">37.4 (0.6)</td>
</tr>
<tr id="357834DD192B52A63AE3FBDFEEB9FB69" box="[159,1412,1113,1165]" gridrow="4" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3FBDFE976FB69" box="[159,587,1113,1165]" gridcol="0" gridrow="4" pageId="11" pageNumber="312">First dorsal-fin origin to pelvic-fin 40.0 origin</th>
<td id="76A95DA1192B52A638ACFBDFE9C0FB69" box="[720,765,1113,1165]" gridcol="1" gridrow="4" pageId="11" pageNumber="312">35.1</td>
<td id="76A95DA1192B52A639FEFBDFE893FB69" box="[898,942,1113,1165]" gridcol="2" gridrow="4" pageId="11" pageNumber="312">37.5</td>
<td id="76A95DA1192B52A63E48FBDFEF5CFB69" box="[1076,1121,1113,1165]" gridcol="3" gridrow="4" pageId="11" pageNumber="312">39.2</td>
<td id="76A95DA1192B52A63E88FBDFEEB9FB69" box="[1268,1412,1113,1165]" gridcol="4" gridrow="4" pageId="11" pageNumber="312">37.9 (2.2)</td>
</tr>
<tr id="357834DD192B52A63AE3FB27EEB9FB31" box="[159,1412,1185,1237]" gridrow="5" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3FB27E976FB31" box="[159,587,1185,1237]" gridcol="0" gridrow="5" pageId="11" pageNumber="312">First dorsal-fin origin to anal-fin 44.3 origin</th>
<td id="76A95DA1192B52A638ACFB27E9C0FB31" box="[720,765,1185,1237]" gridcol="1" gridrow="5" pageId="11" pageNumber="312">43.0</td>
<td id="76A95DA1192B52A639FEFB27E893FB31" box="[898,942,1185,1237]" gridcol="2" gridrow="5" pageId="11" pageNumber="312">43.6</td>
<td id="76A95DA1192B52A63E48FB27EF5CFB31" box="[1076,1121,1185,1237]" gridcol="3" gridrow="5" pageId="11" pageNumber="312">45.8</td>
<td id="76A95DA1192B52A63E88FB27EEB9FB31" box="[1268,1412,1185,1237]" gridcol="4" gridrow="5" pageId="11" pageNumber="312">44.2 (1.2)</td>
</tr>
<tr id="357834DD192B52A63AE3FB6FEEB9FAF9" box="[159,1412,1257,1309]" gridrow="6" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3FB6FE976FAF9" box="[159,587,1257,1309]" gridcol="0" gridrow="6" pageId="11" pageNumber="312">Pelvic-fin origin to second dorsal- 43.0 fin origin</th>
<td id="76A95DA1192B52A638ACFB6FE9C0FAF9" box="[720,765,1257,1309]" gridcol="1" gridrow="6" pageId="11" pageNumber="312">39.2</td>
<td id="76A95DA1192B52A639FEFB6FE893FAF9" box="[898,942,1257,1309]" gridcol="2" gridrow="6" pageId="11" pageNumber="312">40.5</td>
<td id="76A95DA1192B52A63E48FB6FEF5CFAF9" box="[1076,1121,1257,1309]" gridcol="3" gridrow="6" pageId="11" pageNumber="312">42.9</td>
<td id="76A95DA1192B52A63E88FB6FEEB9FAF9" box="[1268,1412,1257,1309]" gridcol="4" gridrow="6" pageId="11" pageNumber="312">41.4 (1.9)</td>
</tr>
<tr id="357834DD192B52A63AE3FAB7EEB9FAA3" box="[159,1412,1329,1351]" gridrow="7" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3FAB7E976FAA3" box="[159,587,1329,1351]" gridcol="0" gridrow="7" pageId="11" pageNumber="312">Head length 31.1</th>
<td id="76A95DA1192B52A638ACFAB7E9C0FAA3" box="[720,765,1329,1351]" gridcol="1" gridrow="7" pageId="11" pageNumber="312">28.7</td>
<td id="76A95DA1192B52A639FEFAB7E893FAA3" box="[898,942,1329,1351]" gridcol="2" gridrow="7" pageId="11" pageNumber="312">30.2</td>
<td id="76A95DA1192B52A63E48FAB7EF5CFAA3" box="[1076,1121,1329,1351]" gridcol="3" gridrow="7" pageId="11" pageNumber="312">30.9</td>
<td id="76A95DA1192B52A63E88FAB7EEB9FAA3" box="[1268,1412,1329,1351]" gridcol="4" gridrow="7" pageId="11" pageNumber="312">30.2 (1.1)</td>
</tr>
<tr id="357834DD192B52A63AE3FADCEEB9FA94" box="[159,1412,1370,1392]" gridrow="8" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3FADCE976FA94" box="[159,587,1370,1392]" gridcol="0" gridrow="8" pageId="11" pageNumber="312">Head depth 34.2</th>
<td id="76A95DA1192B52A638ACFADCE9C0FA94" box="[720,765,1370,1392]" gridcol="1" gridrow="8" pageId="11" pageNumber="312">30.9</td>
<td id="76A95DA1192B52A639FEFADCE893FA94" box="[898,942,1370,1392]" gridcol="2" gridrow="8" pageId="11" pageNumber="312">31.2</td>
<td id="76A95DA1192B52A63E48FADCEF5CFA94" box="[1076,1121,1370,1392]" gridcol="3" gridrow="8" pageId="11" pageNumber="312">34.5</td>
<td id="76A95DA1192B52A63E88FADCEEB9FA94" box="[1268,1412,1370,1392]" gridcol="4" gridrow="8" pageId="11" pageNumber="312">32.7 (1.9)</td>
</tr>
<tr id="357834DD192B52A63AE3FA02EEB9FA7E" box="[159,1412,1412,1434]" gridrow="9" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3FA02E976FA7E" box="[159,587,1412,1434]" gridcol="0" gridrow="9" pageId="11" pageNumber="312">Body depth 39.0</th>
<td id="76A95DA1192B52A638ACFA02E9C0FA7E" box="[720,765,1412,1434]" gridcol="1" gridrow="9" pageId="11" pageNumber="312">35.1</td>
<td id="76A95DA1192B52A639FEFA02E893FA7E" box="[898,942,1412,1434]" gridcol="2" gridrow="9" pageId="11" pageNumber="312">35.7</td>
<td id="76A95DA1192B52A63E48FA02EF5CFA7E" box="[1076,1121,1412,1434]" gridcol="3" gridrow="9" pageId="11" pageNumber="312">38.4</td>
<td id="76A95DA1192B52A63E88FA02EEB9FA7E" box="[1268,1412,1412,1434]" gridcol="4" gridrow="9" pageId="11" pageNumber="312">37.0 (1.9)</td>
</tr>
<tr id="357834DD192B52A63AE3FA2BEEB9FA27" box="[159,1412,1453,1475]" gridrow="10" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3FA2BE976FA27" box="[159,587,1453,1475]" gridcol="0" gridrow="10" pageId="11" pageNumber="312">Pelvic-fin length 18.1</th>
<td id="76A95DA1192B52A638ACFA2BE9C0FA27" box="[720,765,1453,1475]" gridcol="1" gridrow="10" pageId="11" pageNumber="312">17.3</td>
<td id="76A95DA1192B52A639FEFA2BE893FA27" box="[898,942,1453,1475]" gridcol="2" gridrow="10" pageId="11" pageNumber="312">16.5</td>
<td id="76A95DA1192B52A63E48FA2BEF5CFA27" box="[1076,1121,1453,1475]" gridcol="3" gridrow="10" pageId="11" pageNumber="312">18.3</td>
<td id="76A95DA1192B52A63E88FA2BEEB9FA27" box="[1268,1412,1453,1475]" gridcol="4" gridrow="10" pageId="11" pageNumber="312">17.6 (0.8)</td>
</tr>
<tr id="357834DD192B52A63AE3FA50EEB9FA08" box="[159,1412,1494,1516]" gridrow="11" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3FA50E976FA08" box="[159,587,1494,1516]" gridcol="0" gridrow="11" pageId="11" pageNumber="312">Anal-fin length 22.8</th>
<td id="76A95DA1192B52A638ACFA50E9C0FA08" box="[720,765,1494,1516]" gridcol="1" gridrow="11" pageId="11" pageNumber="312">22.5</td>
<td id="76A95DA1192B52A639FEFA50E893FA08" box="[898,942,1494,1516]" gridcol="2" gridrow="11" pageId="11" pageNumber="312">21.9</td>
<td id="76A95DA1192B52A63E48FA50EF5CFA08" box="[1076,1121,1494,1516]" gridcol="3" gridrow="11" pageId="11" pageNumber="312">24.1</td>
<td id="76A95DA1192B52A63E88FA50EEB9FA08" box="[1268,1412,1494,1516]" gridcol="4" gridrow="11" pageId="11" pageNumber="312">22.8 (0.9)</td>
</tr>
<tr id="357834DD192B52A63AE3FA79EEB9F9F1" box="[159,1412,1535,1557]" gridrow="12" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3FA79E976F9F1" box="[159,587,1535,1557]" gridcol="0" gridrow="12" pageId="11" pageNumber="312">Second dorsal-fin length 32.3</th>
<td id="76A95DA1192B52A638ACFA79E9C0F9F1" box="[720,765,1535,1557]" gridcol="1" gridrow="12" pageId="11" pageNumber="312">31.5</td>
<td id="76A95DA1192B52A639FEFA79E893F9F1" box="[898,942,1535,1557]" gridcol="2" gridrow="12" pageId="11" pageNumber="312">29.4</td>
<td id="76A95DA1192B52A63E48FA79EF5CF9F1" box="[1076,1121,1535,1557]" gridcol="3" gridrow="12" pageId="11" pageNumber="312">32.7</td>
<td id="76A95DA1192B52A63E88FA79EEB9F9F1" box="[1268,1412,1535,1557]" gridcol="4" gridrow="12" pageId="11" pageNumber="312">31.5 (1.5)</td>
</tr>
<tr id="357834DD192B52A63AE3F9AFEEB9F9DB" box="[159,1412,1577,1599]" gridrow="13" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3F9AFE976F9DB" box="[159,587,1577,1599]" gridcol="0" gridrow="13" pageId="11" pageNumber="312">Caudal-fin length 24.6</th>
<td id="76A95DA1192B52A638ACF9AFE9C0F9DB" box="[720,765,1577,1599]" gridcol="1" gridrow="13" pageId="11" pageNumber="312">24.2</td>
<td id="76A95DA1192B52A639FEF9AFE893F9DB" box="[898,942,1577,1599]" gridcol="2" gridrow="13" pageId="11" pageNumber="312">22.6</td>
<td id="76A95DA1192B52A63E48F9AFEF5CF9DB" box="[1076,1121,1577,1599]" gridcol="3" gridrow="13" pageId="11" pageNumber="312">24.3</td>
<td id="76A95DA1192B52A63E88F9AFEEB9F9DB" box="[1268,1412,1577,1599]" gridcol="4" gridrow="13" pageId="11" pageNumber="312">23.9 (0.9)</td>
</tr>
<tr id="357834DD192B52A63AE3F9D4EEB9F98C" box="[159,1412,1618,1640]" gridrow="14" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3F9D4E976F98C" box="[159,587,1618,1640]" gridcol="0" gridrow="14" pageId="11" pageNumber="312">Pectoral-fin length 26.5</th>
<td id="76A95DA1192B52A638ACF9D4E9C0F98C" box="[720,765,1618,1640]" gridcol="1" gridrow="14" pageId="11" pageNumber="312">24.7</td>
<td id="76A95DA1192B52A639FEF9D4E893F98C" box="[898,942,1618,1640]" gridcol="2" gridrow="14" pageId="11" pageNumber="312">24.9</td>
<td id="76A95DA1192B52A63E48F9D4EF5CF98C" box="[1076,1121,1618,1640]" gridcol="3" gridrow="14" pageId="11" pageNumber="312">25.8</td>
<td id="76A95DA1192B52A63E88F9D4EEB9F98C" box="[1268,1412,1618,1640]" gridcol="4" gridrow="14" pageId="11" pageNumber="312">25.5 (0.9)</td>
</tr>
<tr id="357834DD192B52A63AE3F9FAEEB9F976" box="[159,1412,1660,1682]" gridrow="15" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3F9FAE976F976" box="[159,587,1660,1682]" gridcol="0" gridrow="15" pageId="11" pageNumber="312">Caudal-peduncle length 21.8</th>
<td id="76A95DA1192B52A638ACF9FAE9C0F976" box="[720,765,1660,1682]" gridcol="1" gridrow="15" pageId="11" pageNumber="312">21.5</td>
<td id="76A95DA1192B52A639FEF9FAE893F976" box="[898,942,1660,1682]" gridcol="2" gridrow="15" pageId="11" pageNumber="312">22.1</td>
<td id="76A95DA1192B52A63E48F9FAEF5CF976" box="[1076,1121,1660,1682]" gridcol="3" gridrow="15" pageId="11" pageNumber="312">22.5</td>
<td id="76A95DA1192B52A63E88F9FAEEB9F976" box="[1268,1412,1660,1682]" gridcol="4" gridrow="15" pageId="11" pageNumber="312">22.0 (0.4)</td>
</tr>
<tr id="357834DD192B52A63AE3F923EEB9F95F" box="[159,1412,1701,1723]" gridrow="16" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3F923E976F95F" box="[159,587,1701,1723]" gridcol="0" gridrow="16" pageId="11" pageNumber="312">Caudal-peduncle depth 15.4</th>
<td id="76A95DA1192B52A638ACF923E9C0F95F" box="[720,765,1701,1723]" gridcol="1" gridrow="16" pageId="11" pageNumber="312">14.5</td>
<td id="76A95DA1192B52A639FEF923E893F95F" box="[898,942,1701,1723]" gridcol="2" gridrow="16" pageId="11" pageNumber="312">14.6</td>
<td id="76A95DA1192B52A63E48F923EF5CF95F" box="[1076,1121,1701,1723]" gridcol="3" gridrow="16" pageId="11" pageNumber="312">15.8</td>
<td id="76A95DA1192B52A63E88F923EEB9F95F" box="[1268,1412,1701,1723]" gridcol="4" gridrow="16" pageId="11" pageNumber="312">15.1 (0.6)</td>
</tr>
<tr id="357834DD192B52A63AE3F948EEB9F900" box="[159,1412,1742,1764]" gridrow="17" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3F948E976F900" box="[159,587,1742,1764]" gridcol="0" gridrow="17" pageId="11" pageNumber="312">Interorbital width 21.3</th>
<td id="76A95DA1192B52A638ACF948E9C0F900" box="[720,765,1742,1764]" gridcol="1" gridrow="17" pageId="11" pageNumber="312">15.6</td>
<td id="76A95DA1192B52A639FEF948E893F900" box="[898,942,1742,1764]" gridcol="2" gridrow="17" pageId="11" pageNumber="312">18.1</td>
<td id="76A95DA1192B52A63E48F948EF5CF900" box="[1076,1121,1742,1764]" gridcol="3" gridrow="17" pageId="11" pageNumber="312">17.5</td>
<td id="76A95DA1192B52A63E88F948EEB9F900" box="[1268,1412,1742,1764]" gridcol="4" gridrow="17" pageId="11" pageNumber="312">18.1 (2.4)</td>
</tr>
<tr id="357834DD192B52A63AE3F971EEB9F8E9" box="[159,1412,1783,1805]" gridrow="18" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3F971E976F8E9" box="[159,587,1783,1805]" gridcol="0" gridrow="18" pageId="11" pageNumber="312">Horizontal eye diameter 23.3</th>
<td id="76A95DA1192B52A638ACF971E9C0F8E9" box="[720,765,1783,1805]" gridcol="1" gridrow="18" pageId="11" pageNumber="312">25.6</td>
<td id="76A95DA1192B52A639FEF971E893F8E9" box="[898,942,1783,1805]" gridcol="2" gridrow="18" pageId="11" pageNumber="312">24.3</td>
<td id="76A95DA1192B52A63E48F971EF5CF8E9" box="[1076,1121,1783,1805]" gridcol="3" gridrow="18" pageId="11" pageNumber="312">23.5</td>
<td id="76A95DA1192B52A63E88F971EEB9F8E9" box="[1268,1412,1783,1805]" gridcol="4" gridrow="18" pageId="11" pageNumber="312">24.2 (1.0)</td>
</tr>
<tr id="357834DD192B52A63AE3F8A7EEB9F8D3" box="[159,1412,1825,1847]" gridrow="19" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3F8A7E976F8D3" box="[159,587,1825,1847]" gridcol="0" gridrow="19" pageId="11" pageNumber="312">Snout length 31.0</th>
<td id="76A95DA1192B52A638ACF8A7E9C0F8D3" box="[720,765,1825,1847]" gridcol="1" gridrow="19" pageId="11" pageNumber="312">29.6</td>
<td id="76A95DA1192B52A639FEF8A7E893F8D3" box="[898,942,1825,1847]" gridcol="2" gridrow="19" pageId="11" pageNumber="312">32.9</td>
<td id="76A95DA1192B52A63E48F8A7EF5CF8D3" box="[1076,1121,1825,1847]" gridcol="3" gridrow="19" pageId="11" pageNumber="312">32.4</td>
<td id="76A95DA1192B52A63E88F8A7EEB9F8D3" box="[1268,1412,1825,1847]" gridcol="4" gridrow="19" pageId="11" pageNumber="312">31.5 (1.5)</td>
</tr>
<tr id="357834DD192B52A63AE3F8CCEEB9F884" box="[159,1412,1866,1888]" gridrow="20" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3F8CCE976F884" box="[159,587,1866,1888]" gridcol="0" gridrow="20" pageId="11" pageNumber="312">Upper-jaw length 36.6</th>
<td id="76A95DA1192B52A638ACF8CCE9C0F884" box="[720,765,1866,1888]" gridcol="1" gridrow="20" pageId="11" pageNumber="312">38.2</td>
<td id="76A95DA1192B52A639FEF8CCE893F884" box="[898,942,1866,1888]" gridcol="2" gridrow="20" pageId="11" pageNumber="312">34.4</td>
<td id="76A95DA1192B52A63E48F8CCEF5CF884" box="[1076,1121,1866,1888]" gridcol="3" gridrow="20" pageId="11" pageNumber="312">34.7</td>
<td id="76A95DA1192B52A63E88F8CCEEB9F884" box="[1268,1412,1866,1888]" gridcol="4" gridrow="20" pageId="11" pageNumber="312">36.0 (1.8)</td>
</tr>
<tr id="357834DD192B52A63AE3F8F2EEB9F84C" box="[159,1412,1908,1960]" gridrow="21" pageId="11" pageNumber="312">
<th id="76A95DA1192B52A63AE3F8F2E976F84C" box="[159,587,1908,1960]" gridcol="0" gridrow="21" pageId="11" pageNumber="312">First dorsal spine to first pectoral 58.6 ray</th>
<td id="76A95DA1192B52A638ACF8F2E9C0F84C" box="[720,765,1908,1960]" gridcol="1" gridrow="21" pageId="11" pageNumber="312">54.0</td>
<td id="76A95DA1192B52A639FEF8F2E893F84C" box="[898,942,1908,1960]" gridcol="2" gridrow="21" pageId="11" pageNumber="312">53.2</td>
<td id="76A95DA1192B52A63E48F8F2EF5CF84C" box="[1076,1121,1908,1960]" gridcol="3" gridrow="21" pageId="11" pageNumber="312">56.1</td>
<td id="76A95DA1192B52A63E88F8F2EEB9F84C" box="[1268,1412,1908,1960]" gridcol="4" gridrow="21" pageId="11" pageNumber="312">55.5 (2.4)</td>
</tr>
</table>
</paragraph>
<caption id="DF376617192CAD553AEBFF1EEBC2FF35" ID-Table-UUID="DF376617192CAD553AEBFF1EEBC2FF35" httpUri="http://table.plazi.org/id/DF376617192CAD553AEBFF1EEBC2FF35" pageId="12" pageNumber="313" targetBox="[159,1330,237,610]" targetIsTable="true" targetPageId="12">
<paragraph id="8BF7369F192CAD553AEBFF1EEBC2FF35" blockId="12.[151,1436,151,209]" pageId="12" pageNumber="313">
<emphasis id="B93CEA8D192CAD553AEBFF1EEA37FF4B" bold="true" box="[151,266,152,175]" pageId="12" pageNumber="313">TABLE 5.</emphasis>
Fin counts of holotype and four paratypes of 
<taxonomicName id="4C484D1C192CAD55397CFF11E8D8FF4B" box="[768,997,151,175]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="irregularis" status="sp. nov.">
<emphasis id="B93CEA8D192CAD55397CFF11E8D8FF4B" box="[768,997,151,175]" italics="true" pageId="12" pageNumber="313">Gobiodon irregularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F57F6192CAD553991FF1EEF7CFF4B" box="[1005,1089,152,175]" pageId="12" pageNumber="313" rank="species">sp. nov.</taxonomicNameLabel>
from the northern Red Sea. d = damaged.
</paragraph>
</caption>
<paragraph id="8BF7369F192CAD553AE3FE8AEE36FD86" pageId="12" pageNumber="313">
<table id="F948C43F192C52A63AE3FF6BEE0FFD86" box="[159,1330,237,610]" gridcols="5" gridrows="8" pageId="12" pageNumber="313">
<tr id="357834DD192C52A63AE3FF6BEE0FFEA4" box="[159,1330,237,320]" gridrow="0" pageId="12" pageNumber="313">
<th id="76A95DA1192C52A63AE3FF6BEA0EFEA4" box="[159,307,237,320]" gridcol="0" gridrow="0" pageId="12" pageNumber="313">Status Coll. No.</th>
<th id="76A95DA1192C52A63BEFFF6BE927FEA4" box="[403,538,237,320]" gridcol="1" gridrow="0" pageId="12" pageNumber="313">Holotype NMW 95078</th>
<th id="76A95DA1192C52A63829FF6BE9E6FEA4" box="[597,731,237,320]" gridcol="2" gridrow="0" pageId="12" pageNumber="313">Paratype NMW 95565</th>
<th id="76A95DA1192C52A6396BFF6BE8A3FEA4" box="[791,926,237,320]" gridcol="3" gridrow="0" pageId="12" pageNumber="313">Paratype NMW 95566</th>
<th id="76A95DA1192C52A639A5FF6BEE0FFEA4" box="[985,1330,237,320]" gridcol="4" gridrow="0" pageId="12" pageNumber="313">Paratype Paratype MNHN 2006-1699 BMNH 2006-10.6.2</th>
</tr>
<tr id="357834DD192C52A63AE3FED2EE0FFE8E" box="[159,1330,340,362]" gridrow="1" pageId="12" pageNumber="313">
<th id="76A95DA1192C52A63AE3FED2EA0EFE8E" box="[159,307,340,362]" gridcol="0" gridrow="1" pageId="12" pageNumber="313">D1</th>
<td id="76A95DA1192C52A63BEFFED2E927FE8E" box="[403,538,340,362]" gridcol="1" gridrow="1" pageId="12" pageNumber="313">VI</td>
<td id="76A95DA1192C52A63829FED2E9E6FE8E" box="[597,731,340,362]" gridcol="2" gridrow="1" pageId="12" pageNumber="313">VI</td>
<td id="76A95DA1192C52A6396BFED2E8A3FE8E" box="[791,926,340,362]" gridcol="3" gridrow="1" pageId="12" pageNumber="313">VI</td>
<td id="76A95DA1192C52A639A5FED2EE0FFE8E" box="[985,1330,340,362]" gridcol="4" gridrow="1" pageId="12" pageNumber="313">VI VI</td>
</tr>
<tr id="357834DD192C52A63AE3FEFBEE0FFE77" box="[159,1330,381,403]" gridrow="2" pageId="12" pageNumber="313">
<th id="76A95DA1192C52A63AE3FEFBEA0EFE77" box="[159,307,381,403]" gridcol="0" gridrow="2" pageId="12" pageNumber="313">D2</th>
<td id="76A95DA1192C52A63BEFFEFBE927FE77" box="[403,538,381,403]" gridcol="1" gridrow="2" pageId="12" pageNumber="313">11</td>
<td id="76A95DA1192C52A63829FEFBE9E6FE77" box="[597,731,381,403]" gridcol="2" gridrow="2" pageId="12" pageNumber="313">11</td>
<td id="76A95DA1192C52A6396BFEFBE8A3FE77" box="[791,926,381,403]" gridcol="3" gridrow="2" pageId="12" pageNumber="313">11</td>
<td id="76A95DA1192C52A639A5FEFBEE0FFE77" box="[985,1330,381,403]" gridcol="4" gridrow="2" pageId="12" pageNumber="313">10 11</td>
</tr>
<tr id="357834DD192C52A63AE3FE20EE0FFE58" box="[159,1330,422,444]" gridrow="3" pageId="12" pageNumber="313">
<th id="76A95DA1192C52A63AE3FE20EA0EFE58" box="[159,307,422,444]" gridcol="0" gridrow="3" pageId="12" pageNumber="313">A</th>
<td id="76A95DA1192C52A63BEFFE20E927FE58" box="[403,538,422,444]" gridcol="1" gridrow="3" pageId="12" pageNumber="313">9</td>
<td id="76A95DA1192C52A63829FE20E9E6FE58" box="[597,731,422,444]" gridcol="2" gridrow="3" pageId="12" pageNumber="313">10</td>
<td id="76A95DA1192C52A6396BFE20E8A3FE58" box="[791,926,422,444]" gridcol="3" gridrow="3" pageId="12" pageNumber="313">9</td>
<td id="76A95DA1192C52A639A5FE20EE0FFE58" box="[985,1330,422,444]" gridcol="4" gridrow="3" pageId="12" pageNumber="313">9 9</td>
</tr>
<tr id="357834DD192C52A63AE3FE49EE0FFE01" box="[159,1330,463,485]" gridrow="4" pageId="12" pageNumber="313">
<th id="76A95DA1192C52A63AE3FE49EA0EFE01" box="[159,307,463,485]" gridcol="0" gridrow="4" pageId="12" pageNumber="313">C (segmented)</th>
<td id="76A95DA1192C52A63BEFFE49E927FE01" box="[403,538,463,485]" gridcol="1" gridrow="4" pageId="12" pageNumber="313">16</td>
<td id="76A95DA1192C52A63829FE49E9E6FE01" box="[597,731,463,485]" gridcol="2" gridrow="4" pageId="12" pageNumber="313">17</td>
<td id="76A95DA1192C52A6396BFE49E8A3FE01" box="[791,926,463,485]" gridcol="3" gridrow="4" pageId="12" pageNumber="313">d</td>
<td id="76A95DA1192C52A639A5FE49EE0FFE01" box="[985,1330,463,485]" gridcol="4" gridrow="4" pageId="12" pageNumber="313">15 16</td>
</tr>
<tr id="357834DD192C52A63AE3FE7FEE0FFDEB" box="[159,1330,505,527]" gridrow="5" pageId="12" pageNumber="313">
<th id="76A95DA1192C52A63AE3FE7FEA0EFDEB" box="[159,307,505,527]" gridcol="0" gridrow="5" pageId="12" pageNumber="313">C (branched)</th>
<td id="76A95DA1192C52A63BEFFE7FE927FDEB" box="[403,538,505,527]" gridcol="1" gridrow="5" pageId="12" pageNumber="313">17</td>
<td id="76A95DA1192C52A63829FE7FE9E6FDEB" box="[597,731,505,527]" gridcol="2" gridrow="5" pageId="12" pageNumber="313">17</td>
<td id="76A95DA1192C52A6396BFE7FE8A3FDEB" box="[791,926,505,527]" gridcol="3" gridrow="5" pageId="12" pageNumber="313">d</td>
<td id="76A95DA1192C52A639A5FE7FEE0FFDEB" box="[985,1330,505,527]" gridcol="4" gridrow="5" pageId="12" pageNumber="313">17 17</td>
</tr>
<tr id="357834DD192C52A63AE3FDA4EE0FFDDC" box="[159,1330,546,568]" gridrow="6" pageId="12" pageNumber="313">
<th id="76A95DA1192C52A63AE3FDA4EA0EFDDC" box="[159,307,546,568]" gridcol="0" gridrow="6" pageId="12" pageNumber="313">P</th>
<td id="76A95DA1192C52A63BEFFDA4E927FDDC" box="[403,538,546,568]" gridcol="1" gridrow="6" pageId="12" pageNumber="313">20</td>
<td id="76A95DA1192C52A63829FDA4E9E6FDDC" box="[597,731,546,568]" gridcol="2" gridrow="6" pageId="12" pageNumber="313">20</td>
<td id="76A95DA1192C52A6396BFDA4E8A3FDDC" box="[791,926,546,568]" gridcol="3" gridrow="6" pageId="12" pageNumber="313">20</td>
<td id="76A95DA1192C52A639A5FDA4EE0FFDDC" box="[985,1330,546,568]" gridcol="4" gridrow="6" pageId="12" pageNumber="313">20 20</td>
</tr>
<tr id="357834DD192C52A63AE3FDCAEE0FFD86" box="[159,1330,588,610]" gridrow="7" pageId="12" pageNumber="313">
<th id="76A95DA1192C52A63AE3FDCAEA0EFD86" box="[159,307,588,610]" gridcol="0" gridrow="7" pageId="12" pageNumber="313">V</th>
<td id="76A95DA1192C52A63BEFFDCAE927FD86" box="[403,538,588,610]" gridcol="1" gridrow="7" pageId="12" pageNumber="313">I/5 + I/5</td>
<td id="76A95DA1192C52A63829FDCAE9E6FD86" box="[597,731,588,610]" gridcol="2" gridrow="7" pageId="12" pageNumber="313">I/5 + I/5</td>
<td id="76A95DA1192C52A6396BFDCAE8A3FD86" box="[791,926,588,610]" gridcol="3" gridrow="7" pageId="12" pageNumber="313">I/5 + I/5</td>
<td id="76A95DA1192C52A639A5FDCAEE0FFD86" box="[985,1330,588,610]" gridcol="4" gridrow="7" pageId="12" pageNumber="313">I/5 + I/5 I/5 + I/5</td>
</tr>
</table>
</paragraph>
<subSubSection id="C3526514192CAD553ABBFD27E859FCC1" pageId="12" pageNumber="313" type="distribution">
<paragraph id="8BF7369F192CAD553ABBFD27E859FCC1" blockId="12.[151,1436,673,2022]" pageId="12" pageNumber="313">
<emphasis id="B93CEA8D192CAD553ABBFD27EA5DFD5E" bold="true" box="[199,352,673,698]" pageId="12" pageNumber="313">Distribution.</emphasis>
This species was found in the Gulf of Aqaba, northern Red Sea main basin (near Marsa Alam, 
<collectingCountry id="F35F760F192CAD553AEBFD40EBE7FD3A" box="[151,218,710,734]" name="Egypt" pageId="12" pageNumber="313">Egypt</collectingCountry>
and Al Wajh, 
<collectingCountry id="F35F760F192CAD553B06FD42E933FD39" box="[378,526,708,733]" name="Saudi Arabia" pageId="12" pageNumber="313">Saudi Arabia</collectingCountry>
) and in the southern Red Sea (Dahlak Archipelago, 
<collectingCountry id="F35F760F192CAD553E2EFD42EF9FFD39" box="[1106,1186,708,733]" name="Eritrea" pageId="12" pageNumber="313">Eritrea</collectingCountry>
; Farasan Archipelago, 
<collectingCountry id="F35F760F192CAD553AEBFD6FEA12FCE6" box="[151,303,745,770]" name="Saudi Arabia" pageId="12" pageNumber="313">Saudi Arabia</collectingCountry>
). It is also known from Rodrigues in the western Indian Ocean (identified as 
<taxonomicName id="4C484D1C192CAD553EEBFD6FEE3AFCE5" box="[1175,1287,745,769]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="genus">
<emphasis id="B93CEA8D192CAD553EEBFD6FEE3AFCE5" box="[1175,1287,745,769]" italics="true" pageId="12" pageNumber="313">Gobiodon</emphasis>
</taxonomicName>
RW sp. 1 by Richard Winterbottom from a photo by Phil Heemstra; 
<figureCitation id="13732A1A192CAD55397EFC8AE878FCC1" box="[770,837,780,805]" captionStart="FIGURE 4" captionStartId="10.[151,250,1841,1863]" captionTargetBox="[151,1436,918,1819]" captionTargetId="figure@10.[151,1436,918,1819]" captionTargetPageId="10" captionText="FIGURE 4. Gobiodon irregularis sp. nov. A. freshly collected adult, paratype MNHN 2006 - 1699, 32.3 mm SL, Dahab, Egypt, Red Sea; B. freshly collected adult, CH 232 - 41 - 020, 31.2 mm SL, Dahab, Egypt, Red Sea; C. freshly collected adult, SMF uncatalogued, 27.6 mm SL, Farasan Archipelago, Red Sea; D. freshly collected subadult, paratype NMW 95565, 23.4 mm SL, Dahab, Egypt, Red Sea; E. alive subadult, PMR VP 3201, 23.5 mm SL, Sharm el Moya, Egypt, Red Sea; F. freshly collected juvenile, SAIAB 70430, 15 mm SL, Rodrigues, western Indian Ocean. Photos by J. Herler (A, B, D), S. V. Bogorodsky (C, E), P. Heemstra, © SAIAB (F)." httpUri="https://zenodo.org/record/249074/files/figure.png" pageId="12" pageNumber="313">Fig. 4</figureCitation>
F).
</paragraph>
</subSubSection>
<subSubSection id="C3526514192CAD553ABBFCB7EAA2FC68" pageId="12" pageNumber="313" type="etymology">
<paragraph id="8BF7369F192CAD553ABBFCB7EAA2FC68" blockId="12.[151,1436,673,2022]" pageId="12" pageNumber="313">
<emphasis id="B93CEA8D192CAD553ABBFCB7EA73FCAE" bold="true" box="[199,334,817,842]" pageId="12" pageNumber="313">Etymology.</emphasis>
This species is named “ 
<taxonomicName id="4C484D1C192CAD553827FCB7E9EEFCAE" box="[603,723,817,842]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="irregularis">irregularis</taxonomicName>
” for its variable colouration, in particular its irregular red wavy lines on the head and upper body in juveniles and subadults. Suggested common name: Rufous Coralgoby (colored red-brown when adult).
</paragraph>
</subSubSection>
<subSubSection id="C3526514192CAD543ABBFC12EF06FE34" lastPageId="13" lastPageNumber="314" pageId="12" pageNumber="313" type="discussion">
<paragraph id="8BF7369F192CAD543ABBFC12EF06FE34" blockId="12.[151,1436,673,2022]" lastBlockId="13.[151,1436,151,464]" lastPageId="13" lastPageNumber="314" pageId="12" pageNumber="313">
<emphasis id="B93CEA8D192CAD553ABBFC12EA06FC49" bold="true" box="[199,315,916,941]" pageId="12" pageNumber="313">Remarks.</emphasis>
The confusion of this species with 
<taxonomicName id="4C484D1C192CAD553895FC13E847FC49" box="[745,890,917,941]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="bilineatus">
<emphasis id="B93CEA8D192CAD553895FC13E847FC49" box="[745,890,917,941]" italics="true" pageId="12" pageNumber="313">G. bilineatus</emphasis>
</taxonomicName>
and grouping with 
<taxonomicName id="4C484D1C192CAD553E15FC13EFE4FC49" box="[1129,1241,917,941]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="genus">
<emphasis id="B93CEA8D192CAD553E15FC13EFE4FC49" box="[1129,1241,917,941]" italics="true" pageId="12" pageNumber="313">Gobiodon</emphasis>
</taxonomicName>
sp. 1 (
<bibRefCitation id="EFD94B6E192CAD553F4DFC12EA16FC36" author="Herler" pageId="12" pageNumber="313" refString="Herler, J. &amp; Hilgers, H. (2005) A synopsis of coral and coral rock associated gobiid fishes from the Gulf of Aqaba, northern Red Sea. aqua, Journal of Ichthyology and Aquatic Biology, 10, 103 - 132." type="journal article" year="2005">Herler &amp; Hilgers 2005</bibRefCitation>
) was mainly caused by the very similar adult colouration, although the body shape (in particular the rounder head and deeper caudal peduncle in 
<taxonomicName id="4C484D1C192CAD5538F2FC5BE826FC11" box="[654,795,989,1013]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="bilineatus">
<emphasis id="B93CEA8D192CAD5538F2FC5BE826FC11" box="[654,795,989,1013]" italics="true" pageId="12" pageNumber="313">G. bilineatus</emphasis>
</taxonomicName>
) and lines on the head are different in adult fishes. Also, D2, A and P fin-ray counts are on average higher in 
<taxonomicName id="4C484D1C192CAD553889FB87E8B3FBFD" box="[757,910,1025,1049]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B93CEA8D192CAD553889FB87E8B3FBFD" box="[757,910,1025,1049]" italics="true" pageId="12" pageNumber="313">G. irregularis</emphasis>
</taxonomicName>
, but the overlapping range of values does not permit definite identification. As confirmed by mitochondrial DNA investigations, the two species are genetically very distinct; the same holds true for the juvenile and subadult colouration of both. Although body shape differences are recognizable, geometric morphometric analysis failed to reveal statistically significant shape differences between the two species (MANOVA on the first 5 principal components: p = 0.08), but this may merely reflect the very low sample number, especially of 
<taxonomicName id="4C484D1C192CAD553895FB33E847FB29" box="[745,890,1205,1229]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="bilineatus">
<emphasis id="B93CEA8D192CAD553895FB33E847FB29" box="[745,890,1205,1229]" italics="true" pageId="12" pageNumber="313">G. bilineatus</emphasis>
</taxonomicName>
. Discrimination between 
<taxonomicName id="4C484D1C192CAD553ED5FB33EE78FB29" box="[1193,1349,1205,1229]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B93CEA8D192CAD553ED5FB33EF87FB28" box="[1193,1210,1205,1228]" italics="true" pageId="12" pageNumber="313">G</emphasis>
. 
<emphasis id="B93CEA8D192CAD553EB1FB33EE78FB29" box="[1229,1349,1205,1229]" italics="true" pageId="12" pageNumber="313">irregularis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C484D1C192CAD553FF8FB33EA38FB15" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="bilineatus">
<emphasis id="B93CEA8D192CAD553FF8FB33EA38FB15" italics="true" pageId="12" pageNumber="313">G. bilineatus</emphasis>
</taxonomicName>
can possibly be based on life colouration, at least among Red Sea specimens. Juveniles and subadults may be distinguished by the five thin blue lines on the head and the uniformly coloured body in 
<taxonomicName id="4C484D1C192CAD553F78FB7BEEA9FAF1" box="[1284,1428,1277,1301]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="bilineatus">
<emphasis id="B93CEA8D192CAD553F78FB7BEEA9FAF1" box="[1284,1428,1277,1301]" italics="true" pageId="12" pageNumber="313">G. bilineatus</emphasis>
</taxonomicName>
, versus the broader irregular red bars with bluish/greyish interspaces below the eye and the irregular lines and dots on the body of 
<taxonomicName id="4C484D1C192CAD553B38FAC0EAE1FAB9" box="[324,476,1349,1373]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B93CEA8D192CAD553B38FAC0EA68FAB9" box="[324,341,1350,1373]" italics="true" pageId="12" pageNumber="313">G</emphasis>
. 
<emphasis id="B93CEA8D192CAD553B18FAC3EAE1FAB9" box="[356,476,1349,1373]" italics="true" pageId="12" pageNumber="313">irregularis</emphasis>
</taxonomicName>
. In adults, lines on the head or bars may vanish in both species, but there are always two bright bluish lines running across the eye in 
<taxonomicName id="4C484D1C192CAD553890FAEFE842FA65" box="[748,895,1385,1409]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="bilineatus">
<emphasis id="B93CEA8D192CAD553890FAEFE842FA65" box="[748,895,1385,1409]" italics="true" pageId="12" pageNumber="313">G. bilineatus</emphasis>
</taxonomicName>
, which are far less distinct in 
<taxonomicName id="4C484D1C192CAD553E84FAEFEEA9FA65" box="[1272,1428,1385,1409]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B93CEA8D192CAD553E84FAEFEE34FA64" box="[1272,1289,1385,1408]" italics="true" pageId="12" pageNumber="313">G</emphasis>
. 
<emphasis id="B93CEA8D192CAD553F60FAEFEEA9FA65" box="[1308,1428,1385,1409]" italics="true" pageId="12" pageNumber="313">irregularis</emphasis>
</taxonomicName>
. Moreover, body colouration is bright orange to orange-red in 
<taxonomicName id="4C484D1C192CAD553932FA08E8E1FA41" box="[846,988,1421,1445]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="bilineatus">
<emphasis id="B93CEA8D192CAD553932FA08E8E1FA41" box="[846,988,1421,1445]" italics="true" pageId="12" pageNumber="313">G. bilineatus</emphasis>
</taxonomicName>
and rather brownish to red-brown in 
<taxonomicName id="4C484D1C192CAD553FF8FA08EA32FA2D" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B93CEA8D192CAD553FF8FA08EEA8FA41" box="[1412,1429,1422,1445]" italics="true" pageId="12" pageNumber="313">G</emphasis>
. 
<emphasis id="B93CEA8D192CAD553AEBFA37EA32FA2D" box="[151,271,1457,1481]" italics="true" pageId="12" pageNumber="313">irregularis</emphasis>
</taxonomicName>
, but this varies: the Indian Ocean/western Pacific populations, which are genetically very close to Red Sea 
<taxonomicName id="4C484D1C192CAD553ABBFA50EA69FA09" box="[199,340,1493,1517]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="bilineatus">
<emphasis id="B93CEA8D192CAD553ABBFA50EA69FA09" box="[199,340,1493,1517]" italics="true" pageId="12" pageNumber="313">G. bilineatus</emphasis>
</taxonomicName>
(
<figureCitation id="13732A1A192CAD553B19FA53EA8AFA0A" box="[357,439,1493,1518]" captionStart="FIGURE 11" captionStartId="25.[151,250,1851,1873]" captionTargetBox="[358,1210,187,1822]" captionTargetId="figure@25.[348,1247,178,1829]" captionTargetPageId="25" captionText="FIGURE 11. Neighbor-joining tree of 23 taxa of Gobiodon and 2 Paragobiodon species (as nearest neighbour at genus level) based on combined sequences of partial 12 S and 16 S rRNA mitochondrial genes. According to a model test in MEGA 5.05, a K 2 P-model with a gamma-parameter of 0.23 and pairwise deletions of gaps / missing data were selected. Numbers at nodes are bootstrap values (1000 replicates); values &lt;50 are not shown. Bar below tree shows substitutions per site. The 4 new species are marked with a box. For abbreviations of species and localities see Table 1. Undescribed species are given with references in parentheses: JH = according to Herler et al. (2009), PM = according to Munday et al. (1999)." httpUri="https://zenodo.org/record/249081/files/figure.png" pageId="12" pageNumber="313">Fig. 11</figureCitation>
), have a darker body (
<figureCitation id="13732A1A192CAD5538C8FA53E9CAFA0A" box="[692,759,1493,1518]" captionStart="FIGURE 9" captionStartId="22.[151,250,1660,1682]" captionTargetBox="[151,1436,181,1640]" captionTargetId="figure@22.[151,1436,181,1641]" captionTargetPageId="22" captionText="FIGURE 9. Comparative material. Freshly collected adults: Gobiodon cf. bilineatus sp. nov. A. uncatalogued, 28.9 mm SL, Kagi Island, Maldives. B. uncatalogued, 22.9 mm SL, Kenting, southern Taiwan. C. Gobiodon prolixus, ROM 73338, holotype, male 26.2 mm SL, Nha Trang, Vietnam. D. Gobiodon sp. D sensu Munday et al. (1999), uncatalogued, 29.9 mm SL, Hembadhu Island, Maldives E. Gobiodon quinquestrigatus, uncatalogued, 28.7 mm SL, Kenting, southern Taiwan; F. Gobiodon cf. fulvus (sensu Winterbottom &amp; Emery 1986), OMNH P 40259, 20.8 mm SL, Iriomote Island, Japan. G. Gobiodon oculolineatus, OMNH P 40260, 22.6 mm SL, Okinawa Island, Japan. H. Gobiodon ceramensis, OMNH P 34042, 35 mm SL, Ryukyu Island, Japan. Gobiodon unicolor, preserved holotype, MNHN A- 4015, 31 mm SL, I. entire specimen. J. close-up of black opercular spot (arrow) on the right side. Photos by J. Herler (A, B, D, E, I, J), R. Winterbottom (C), T. Suzuki (F-H)." httpUri="https://zenodo.org/record/249079/files/figure.png" pageId="12" pageNumber="313">Fig. 9</figureCitation>
A, B). Genetically, the closest relative of 
<taxonomicName id="4C484D1C192CAD553EB7FA50EE5FFA09" box="[1227,1378,1493,1517]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B93CEA8D192CAD553EB7FA50EFE1FA09" box="[1227,1244,1494,1517]" italics="true" pageId="12" pageNumber="313">G</emphasis>
. 
<emphasis id="B93CEA8D192CAD553E96FA53EE5FFA09" box="[1258,1378,1493,1517]" italics="true" pageId="12" pageNumber="313">irregularis</emphasis>
</taxonomicName>
is 
<taxonomicName id="4C484D1C192CAD553FF8FA50E916F9F5" authority="Duchene et al. 2013" authorityName="Duchene et al." authorityYear="2013" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="oculolineatus">
<emphasis id="B93CEA8D192CAD553FF8FA50EA13F9F5" italics="true" pageId="12" pageNumber="313">G. oculolineatus</emphasis>
(
<bibRefCitation id="EFD94B6E192CAD553B40FA7FE919F9F5" author="Duchene" box="[316,548,1529,1554]" pageId="12" pageNumber="313" refString="Duchene, D., Klanten, S. O., Munday, P. L., Herler, J. &amp; van Herwerden, L. (2013) Phylogenetic evidence for recent diversification of obligate coral-dwelling gobies compared with their host corals. Molecular Phylogenetics and Evolution, 69: 123 - 132. http: // dx. doi. org / 10.1016 / j. ympev. 2013.04.033" type="journal article" year="2013">
Duchene 
<emphasis id="B93CEA8D192CAD553BD4FA7CEAE4F9F5" box="[424,473,1529,1553]" italics="true" pageId="12" pageNumber="313">et al</emphasis>
. 2013
</bibRefCitation>
)
</taxonomicName>
. In fact, the two species have a genetic distance of &lt;2%, which suggests intrarather than interspecific variation. However, their colour pattern is distinct, in that 
<taxonomicName id="4C484D1C192CAD553E35F998EE3DF9D1" box="[1097,1280,1565,1589]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="oculolineatus">
<emphasis id="B93CEA8D192CAD553E35F998EE3DF9D1" box="[1097,1280,1565,1589]" italics="true" pageId="12" pageNumber="313">G. oculolineatus</emphasis>
</taxonomicName>
has only two bars across the eye and suborbital area, similar to the first two in 
<taxonomicName id="4C484D1C192CAD5539F2F9C7EF1AF9BD" box="[910,1063,1601,1625]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B93CEA8D192CAD5539F2F9C7EF1AF9BD" box="[910,1063,1601,1625]" italics="true" pageId="12" pageNumber="313">G. irregularis</emphasis>
</taxonomicName>
, but with a wide, conspicuously coloured (dark red to brown) interspace. In addition, 
<taxonomicName id="4C484D1C192CAD55396EF9E0E8F0F999" box="[786,973,1637,1661]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="oculolineatus">
<emphasis id="B93CEA8D192CAD55396EF9E0E8F0F999" box="[786,973,1637,1661]" italics="true" pageId="12" pageNumber="313">G. oculolineatus</emphasis>
</taxonomicName>
has no lines on the body, not even as juveniles. Therefore, we consider the two species as distinct, similar to other closely related species pairs, such as 
<emphasis id="B93CEA8D192CAD553AEBF928EB95F921" box="[151,168,1710,1733]" italics="true" pageId="12" pageNumber="313">G</emphasis>
. sp. D sensu 
<bibRefCitation id="EFD94B6E192CAD553B36F92BE97CF922" box="[330,577,1709,1734]" pageId="12" pageNumber="313" refString="Munday, P. L., Harold, A. S. &amp; Winterbottom, R. (1999) Guide to coral-dwelling gobies, genus Gobiodon (Gobiidae), from Papua New Guinea and the Great Barrier Reef. Revue Francaise d'Aquariologie Herpetologie, 26, 53 - 58." type="journal article">
Munday 
<emphasis id="B93CEA8D192CAD553BCDF928EADBF921" box="[433,486,1709,1733]" italics="true" pageId="12" pageNumber="313">et al</emphasis>
. (1999)
</bibRefCitation>
and 
<taxonomicName id="4C484D1C192CAD553802F928E861F921" box="[638,860,1710,1733]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="quinquestrigatus">
<emphasis id="B93CEA8D192CAD553802F928E861F921" box="[638,860,1710,1733]" italics="true" pageId="12" pageNumber="313">G. quinquestrigatus</emphasis>
</taxonomicName>
(P. Munday, personal communication). 
<taxonomicName id="4C484D1C192CAD553F50F92BEA32F90D" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B93CEA8D192CAD553F50F92BEA32F90D" italics="true" pageId="12" pageNumber="313">Gobiodon irregularis</emphasis>
</taxonomicName>
is also a close relative of 
<taxonomicName id="4C484D1C192CAD553845F957E84DF90E" authority="Playfair 1867" authorityName="Playfair" authorityYear="1867" box="[569,880,1745,1770]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="reticulatus">
<emphasis id="B93CEA8D192CAD553845F957E9EDF90D" box="[569,720,1745,1769]" italics="true" pageId="12" pageNumber="313">G. reticulatus</emphasis>
Playfair 1867
</taxonomicName>
(
<figureCitation id="13732A1A192CAD5539FDF957E8EEF90D" box="[897,979,1745,1770]" captionStart="FIGURE 11" captionStartId="25.[151,250,1851,1873]" captionTargetBox="[358,1210,187,1822]" captionTargetId="figure@25.[348,1247,178,1829]" captionTargetPageId="25" captionText="FIGURE 11. Neighbor-joining tree of 23 taxa of Gobiodon and 2 Paragobiodon species (as nearest neighbour at genus level) based on combined sequences of partial 12 S and 16 S rRNA mitochondrial genes. According to a model test in MEGA 5.05, a K 2 P-model with a gamma-parameter of 0.23 and pairwise deletions of gaps / missing data were selected. Numbers at nodes are bootstrap values (1000 replicates); values &lt;50 are not shown. Bar below tree shows substitutions per site. The 4 new species are marked with a box. For abbreviations of species and localities see Table 1. Undescribed species are given with references in parentheses: JH = according to Herler et al. (2009), PM = according to Munday et al. (1999)." httpUri="https://zenodo.org/record/249081/files/figure.png" pageId="12" pageNumber="313">Fig. 11</figureCitation>
). Similarities with the latter exist in finray counts but also in the head colour pattern of subadult 
<taxonomicName id="4C484D1C192CAD55395FF970E887F8E9" box="[803,954,1781,1805]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B93CEA8D192CAD55395FF970E809F8E9" box="[803,820,1782,1805]" italics="true" pageId="12" pageNumber="313">G</emphasis>
. 
<emphasis id="B93CEA8D192CAD55393EF973E887F8E9" box="[834,954,1781,1805]" italics="true" pageId="12" pageNumber="313">irregularis</emphasis>
</taxonomicName>
. The two species can be discriminated not only genetically but also based on life colouration in the field: 
<taxonomicName id="4C484D1C192CAD553926F89FE8CDF8D5" box="[858,1008,1817,1841]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="reticulatus">
<emphasis id="B93CEA8D192CAD553926F89FE8CDF8D5" box="[858,1008,1817,1841]" italics="true" pageId="12" pageNumber="313">G. reticulatus</emphasis>
</taxonomicName>
has large spots on the body and 5 or 6 rather regular and broad bluish head bars, whereas the small spots in juvenile and subadult 
<taxonomicName id="4C484D1C192CAD553EAFF8B8EE53F8B1" box="[1235,1390,1853,1877]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B93CEA8D192CAD553EAFF8B8EFD9F8B1" box="[1235,1252,1854,1877]" italics="true" pageId="12" pageNumber="313">G</emphasis>
. 
<emphasis id="B93CEA8D192CAD553E8AF8BBEE53F8B1" box="[1270,1390,1853,1877]" italics="true" pageId="12" pageNumber="313">irregularis</emphasis>
</taxonomicName>
are restricted to the upper half of the body and adults have only two less distinct orbital bars. They also differ in their habitat requirements in that 
<taxonomicName id="4C484D1C192CAD553B9FF800E940F879" box="[483,637,1925,1949]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="reticulatus">
<emphasis id="B93CEA8D192CAD553B9FF800E940F879" box="[483,637,1925,1949]" italics="true" pageId="12" pageNumber="313">G. reticulatus</emphasis>
</taxonomicName>
usually lives in deep water (up to 
<quantity id="4CB09B7A192CAD553E58F800EF5FF87A" box="[1060,1122,1926,1950]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="313" unit="m" value="30.0">30 m</quantity>
) and only occupies a few, particular 
<taxonomicName id="4C484D1C192CAD553B75F82FEA49F824" box="[265,372,1961,1984]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="12" pageNumber="313" phylum="Cnidaria" rank="genus">
<emphasis id="B93CEA8D192CAD553B75F82FEA49F824" box="[265,372,1961,1984]" italics="true" pageId="12" pageNumber="313">Acropora</emphasis>
</taxonomicName>
species which are rarely used by other species (see 
<bibRefCitation id="EFD94B6E192CAD5539C9F82FEFF9F826" author="Herler" box="[949,1220,1961,1986]" pageId="12" pageNumber="313" refString="Herler, J. &amp; Hilgers, H. (2005) A synopsis of coral and coral rock associated gobiid fishes from the Gulf of Aqaba, northern Red Sea. aqua, Journal of Ichthyology and Aquatic Biology, 10, 103 - 132." type="journal article" year="2005">Herler &amp; Hilgers (2005)</bibRefCitation>
and 
<bibRefCitation id="EFD94B6E192CAD553E86F82FEA00F802" author="Dirnwoeber" pageId="12" pageNumber="313" refString="Dirnwoeber, M. &amp; Herler, J. (2007) Microhabitat specialisation and ecological consequences for coral gobies of the genus Gobiodon in the Gulf of Aqaba, northern Red Sea. Marine Ecology Progress Series, 342, 265 - 275." type="journal article" year="2007">Dirnwoeber &amp; Herler (2007))</bibRefCitation>
. Another species with bright lines on the head as in 
<taxonomicName id="4C484D1C192CAD5539FCF848EF31F801" box="[896,1036,1997,2021]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="bilineatus">
<emphasis id="B93CEA8D192CAD5539FCF848EF31F801" box="[896,1036,1997,2021]" italics="true" pageId="12" pageNumber="313">G. bilineatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C484D1C192CAD553E66F848EF8CF801" box="[1050,1201,1997,2021]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B93CEA8D192CAD553E66F848EF8CF801" box="[1050,1201,1997,2021]" italics="true" pageId="12" pageNumber="313">G. irregularis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C484D1C192CAD553E9BF848EEA1F801" box="[1255,1436,1997,2021]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="313" phylum="Chordata" rank="species" species="oculolineatus">
<emphasis id="B93CEA8D192CAD553E9BF848EEA1F801" box="[1255,1436,1997,2021]" italics="true" pageId="12" pageNumber="313">G. oculolineatus</emphasis>
</taxonomicName>
is 
<taxonomicName id="4C484D1C192DAD543ACEFF1EE9FBFF54" authority="Winterbottom &amp; Emery (1986)" authorityName="Winterbottom &amp; Emery" authorityYear="1986" box="[178,710,151,176]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="314" phylum="Chordata" rank="species" species="fulvus">
<emphasis id="B93CEA8D192DAD543ACEFF1EEA28FF54" box="[178,277,152,176]" italics="true" pageId="13" pageNumber="314">G. fulvus</emphasis>
sensu 
<bibRefCitation id="EFD94B6E192DAD543B1FFF11E9FBFF54" author="Winterbottom" box="[355,710,151,176]" pageId="13" pageNumber="314" refString="Winterbottom, R. &amp; Emery, A. R. (1986) Review of the gobioid fishes of the Chagos Archipelago, central Indian Ocean. Life Sciences Contributions, Royal Ontario Museum, Canada, 142, 1 - 82." type="journal article" year="1986">Winterbottom &amp; Emery (1986)</bibRefCitation>
</taxonomicName>
. This species, however, is rarely referred to in the literature and the name 
<taxonomicName id="4C484D1C192DAD543B71FF3AEA49FF30" box="[269,372,188,212]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="314" phylum="Chordata" rank="species" species="fulvus">
<emphasis id="B93CEA8D192DAD543B71FF3AEA49FF30" box="[269,372,188,212]" italics="true" pageId="13" pageNumber="314">G. fulvus</emphasis>
</taxonomicName>
seems commonly applied to a species which does not agree with Herre’s (1927) original description (he mentioned a black opercular spot, no headlines and a pale orange body, all of which contradict the species recorded by 
<bibRefCitation id="EFD94B6E192DAD543BF8FE82E9C0FEF9" author="Winterbottom" box="[388,765,260,285]" pageId="13" pageNumber="314" refString="Winterbottom, R. &amp; Emery, A. R. (1986) Review of the gobioid fishes of the Chagos Archipelago, central Indian Ocean. Life Sciences Contributions, Royal Ontario Museum, Canada, 142, 1 - 82." type="journal article" year="1986">Winterbottom and Emery (1986)</bibRefCitation>
and the specimens from the 
<collectingCountry id="F35F760F192DAD543E28FE82EF82FEF9" box="[1108,1215,260,285]" name="Maldives" pageId="13" pageNumber="314">Maldives</collectingCountry>
, 
<collectingCountry id="F35F760F192DAD543EB1FE83EE30FEF8" box="[1229,1293,261,284]" name="Japan" pageId="13" pageNumber="314">Japan</collectingCountry>
and 
<collectingCountry id="F35F760F192DAD543F36FE82EEA1FEF9" box="[1354,1436,260,285]" name="Taiwan" pageId="13" pageNumber="314">Taiwan</collectingCountry>
examined herein). However, 
<taxonomicName id="4C484D1C192DAD543B9FFEAEE8C6FEA4" authority="Winterbottom &amp; Emery (1986)" authorityName="Winterbottom &amp; Emery" authorityYear="1986" box="[483,1019,295,320]" class="Actinopterygii" family="Gobiidae" genus="Gobiodon" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="314" phylum="Chordata" rank="species" species="fulvus">
<emphasis id="B93CEA8D192DAD543B9FFEAEE97AFEA4" box="[483,583,296,320]" italics="true" pageId="13" pageNumber="314">G. fulvus</emphasis>
sensu 
<bibRefCitation id="EFD94B6E192DAD5438E4FEA1E8C6FEA4" author="Winterbottom" box="[664,1019,295,320]" pageId="13" pageNumber="314" refString="Winterbottom, R. &amp; Emery, A. R. (1986) Review of the gobioid fishes of the Chagos Archipelago, central Indian Ocean. Life Sciences Contributions, Royal Ontario Museum, Canada, 142, 1 - 82." type="journal article" year="1986">Winterbottom &amp; Emery (1986)</bibRefCitation>
</taxonomicName>
can be distinguished from the three species mentioned above by a very dark body, two distinct bright blue lines with black borders across the eye that extend obliquely to the posterior edge of the preopercle, and distinct white bands along the D2 and A base (also with black borders). The fin-base bands may be hardly visible in life (JH, personal observation) and therefore this species may be most easily mistaken in the field with the species mentioned above.
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</subSubSection>
</treatment>
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