treatments-xml/data/03/BF/05/03BF0537640DFFF1C9BFFEA660D39186.xml

<document id="33FEE9AB5CE8DE3751568FB790885FD7" ID-DOI="10.5281/zenodo.276757" ID-GBIF-Dataset="f7c97ce5-a40f-4b2b-8a89-08a570cd64e5" ID-ISSN="1175-5326" ID-Zenodo-Dep="276757" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460235275132" checkinUser="plazi" docAuthor="Moraza, María L. &amp; Lindquist, Evert E." docDate="2011" docId="03BF0537640DFFF1C9BFFEA660D39186" docLanguage="en" docName="zt02758p025.pdf" docOrigin="Zootaxa 2758" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Fungiseius Moraza &amp; Lindquist, 2011, new genus" docType="treatment" docVersion="7" lastPageNumber="6" masterDocId="FF867D4F640EFFF4C928FFF363779526" masterDocTitle="A new genus of fungus-inhabiting blattisociid mites (Acari: Mesostigmata: Phytoseioidea) from Middle America, with a key to genera and subgenera of the subfamily Blattisociinae" masterLastPageNumber="25" masterPageNumber="1" pageNumber="4" updateTime="1698239449555" updateUser="plazi">
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<mods:title id="608F7AC8BAB5E53B1CB7A42C3C5E2500">A new genus of fungus-inhabiting blattisociid mites (Acari: Mesostigmata: Phytoseioidea) from Middle America, with a key to genera and subgenera of the subfamily Blattisociinae</mods:title>
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<mods:namePart id="E2C75B4D918E558B5FF5F956A15A5307">Moraza, María L.</mods:namePart>
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<mods:namePart id="BB2F3193F29EFE89BD4E43933E2BAEA8">Lindquist, Evert E.</mods:namePart>
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<mods:date id="A7740F3B6732C77A278E49ECF73EF04F">2011</mods:date>
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<treatment id="03BF0537640DFFF1C9BFFEA660D39186" ID-DOI="http://doi.org/10.5281/zenodo.6183323" ID-GBIF-Taxon="119388018" ID-Zenodo-Dep="6183323" LSID="urn:lsid:plazi:treatment:03BF0537640DFFF1C9BFFEA660D39186" httpUri="http://treatment.plazi.org/id/03BF0537640DFFF1C9BFFEA660D39186" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
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<taxonomicName id="4C16CFA2640DFFF7C9BFFEA662699449" ID-CoL="9BBXB" box="[151,286,341,367]" class="Arachnida" family="Blattisociidae" genus="Fungiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9626833640DFFF7C9BFFEA662699449" bold="true" box="[151,286,341,367]" italics="true" pageId="3" pageNumber="4">Fungiseius</emphasis>
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(
<figureCitation id="132DA8A4640DFFF7C9B7FE84626794A9" box="[159,272,375,399]" captionStart-0="FIGURES 1 – 8" captionStart-1="FIGURES 9 – 14" captionStart-10="FIGURES 59 – 60" captionStart-2="FIGURES 15 – 18" captionStart-3="FIGURES 19 – 24" captionStart-4="FIGURES 25 – 31" captionStart-5="FIGURES 32 – 37" captionStart-6="FIGURES 38 – 41" captionStart-7="FIGURES 42 – 47" captionStart-8="FIGURES 48 – 55" captionStart-9="FIGURES 56 – 58" captionStartId-0="6.[151,264,1633,1656]" captionStartId-1="8.[151,264,1897,1920]" captionStartId-10="17.[151,264,1978,2001]" captionStartId-2="9.[151,264,1939,1962]" captionStartId-3="10.[151,264,1498,1521]" captionStartId-4="11.[151,264,1201,1224]" captionStartId-5="12.[151,264,1939,1962]" captionStartId-6="14.[151,264,1922,1945]" captionStartId-7="15.[151,264,1747,1770]" captionStartId-8="16.[151,264,1591,1614]" captionStartId-9="17.[151,264,1189,1212]" captionTargetBox-0="[151,1436,217,1606]" captionTargetBox-1="[165,1426,401,1858]" captionTargetBox-10="[151,1436,1292,1954]" captionTargetBox-2="[151,1431,193,1890]" captionTargetBox-3="[162,1420,192,1472]" captionTargetBox-4="[166,1417,183,1174]" captionTargetBox-5="[151,1433,433,1911]" captionTargetBox-6="[151,1431,380,1900]" captionTargetBox-7="[151,1436,193,1723]" captionTargetBox-8="[170,1413,460,1558]" captionTargetBox-9="[172,1418,470,1171]" captionTargetId-0="figure@6.[151,1436,195,1616]" captionTargetId-1="figure@8.[165,1426,401,1880]" captionTargetId-10="figure@17.[151,1436,1292,1957]" captionTargetId-2="figure@9.[151,1436,193,1901]" captionTargetId-3="figure@10.[162,1422,173,1479]" captionTargetId-4="figure@11.[165,1425,183,1186]" captionTargetId-5="figure@12.[151,1436,433,1918]" captionTargetId-6="figure@14.[151,1436,367,1900]" captionTargetId-7="figure@15.[151,1436,193,1726]" captionTargetId-8="figure@16.[169,1413,445,1563]" captionTargetId-9="figure@17.[164,1424,470,1172]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionTargetPageId-10="17" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionTargetPageId-5="12" captionTargetPageId-6="14" captionTargetPageId-7="15" captionTargetPageId-8="16" captionTargetPageId-9="17" captionText-0="FIGURES 1 – 8. Fungiseius armatus n. sp., adult female: 1, gnathotectum; 2, subcapitulum; 3, corniculi, dorsal view; 4, labrum; 5, tritosternum, showing denticles at base of laciniae; 6, palpus, dorsolateral view; 7 a, detail of seta al of palpgenu; 7 b, detail of seta al of palpfemur; 8 a, chelicera, lateral view; 8 b, separate paraxial view of fixed digit." captionText-1="FIGURES 9 – 14. Fungiseius armatus n. sp., adult female: 9, idiosoma, dorsal aspect with detail of peritrematal shield (arrows indicate delineated strip of dorsal shield margins bearing some of r - marginal and all S - lateral setae); 10, detail of separate views of dorsal setae denoted and lyrifissure; 11 a – b, detail of ventral view of dorsal and peritrematal shields; 12, detail of exopodal IV with peritremal poroidotaxy at level of coxa IV; 13, detail of denoted dorsal setae; 14, detail of dorsal ornamentation." captionText-10="FIGURES 59 – 60. Photomicrographs of adult females in dorsoventral habitus view: 59, Fungiseius armatus n. sp.; 60, Fungiseius clavulisetis n. sp." captionText-2="FIGURES 15 – 18. Fungiseius armatus n. sp., adult female: 15, idiosoma, ventral aspect; 16, detail of peritrematal shield, fused to exopodal strips between legs II – III and III – IV; 17, detail of sternal shielding; 18 a, b, detail of spermatheca." captionText-3="FIGURES 19 – 24. Fungiseius armatus n. sp., adult female: 19 – 22, legs I – IV, excepting tarsi: 19, leg I excluding tarsus, laterodorsal view; 20, leg II, posterolateral view; 21, leg III, dorsal view; 22, leg IV, anterolateral view; 23, trochanter I, ventral view; 24, detail of denoted setae of tibia I." captionText-4="FIGURES 25 – 31. Fungiseius armatus n. sp., adult female: 25, tarsus I, anterolateral view; 26, detail of apical seta on tarsus I; 27, tarsus II, posterolateral view; 28, tarsus III, dorsal view; 29, tarsus IV, dorsal view; 30, detail of ventral aspect of tarsus IV with ventral process; 31, detail of lateral aspect of tarsus IV. Abbreviation: t p, tarsal process." captionText-5="FIGURES 32 – 37. Fungiseius armatus n. sp., adult male: 32, subcapitulum; 33, chelicerae, dorsolateral view; 34, chelicerae, detail of fixed digit with pilus dentilis; 35, chelicera, detail of movable digit and spermatodactyl; 36, detail of spermatodactyl; 37, idiosoma, ventral view." captionText-6="FIGURES 38 – 41. Fungiseius clavulisetis n. sp., adult female: 38, idiosoma, dorsal aspect (arrows indicate delineated strip of dorsal shield margins bearing r - marginal and S - lateral setae); 39, detail of denoted dorsal setae; 40, chelicera, detail of movable digit; 41, chelicera, dorsal view." captionText-7="FIGURES 42 – 47. Fungiseius clavulisetis n. sp., adult female: 42, tritosternum; 43, gnathotectum; 44, corniculi, dorsal view; 45, subcapitulum; 46, idiosoma, ventral aspect; 47, detail of spermatheca." captionText-8="FIGURES 48 – 55. Fungiseius clavulisetis n. sp., adult female, legs I – IV, excluding tarsus: 48, leg I, posterolateral view; 49, leg II, posterolateral view; 50, leg III, dorsal view; 51, leg IV, posterolateral view; 52, coxa I, ventral view; 53, coxa II, ventral view; 54, coxa III, ventral view; 55, coxa IV, ventral view." captionText-9="FIGURES 56 – 58. Fungiseius clavulisetis n. sp., adult female: 56, tarsus IV, dorsal view; 57, tarsus II, posterolateral view; 58, tarsus II, anterolateral view. Abbreviation: t p, tarsal process." httpUri-0="https://zenodo.org/record/276758/files/figure.png" httpUri-1="https://zenodo.org/record/276759/files/figure.png" httpUri-10="https://zenodo.org/record/276768/files/figure.png" httpUri-2="https://zenodo.org/record/276760/files/figure.png" httpUri-3="https://zenodo.org/record/276761/files/figure.png" httpUri-4="https://zenodo.org/record/276762/files/figure.png" httpUri-5="https://zenodo.org/record/276763/files/figure.png" httpUri-6="https://zenodo.org/record/276764/files/figure.png" httpUri-7="https://zenodo.org/record/276765/files/figure.png" httpUri-8="https://zenodo.org/record/276766/files/figure.png" httpUri-9="https://zenodo.org/record/276767/files/figure.png" pageId="3" pageNumber="4">Figs 1–60</figureCitation>
)
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<paragraph id="8BA9B421640DFFF7C9BFFE33610094F0" blockId="3.[151,631,447,470]" box="[151,631,447,470]" pageId="3" pageNumber="4">
<emphasis id="B9626833640DFFF7C9BFFE33625294F0" bold="true" box="[151,293,447,470]" pageId="3" pageNumber="4">
<typeStatus id="54AD0A83640DFFF7C9BFFE3363BB94F3" box="[151,204,448,469]" pageId="3" pageNumber="4">Type</typeStatus>
species:
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<taxonomicName id="4C16CFA2640DFFF7C803FE32628394F0" box="[299,500,449,470]" class="Arachnida" family="Blattisociidae" genus="Fungiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="armatus" status="sp. nov.">
<emphasis id="B9626833640DFFF7C803FE32628394F0" box="[299,500,449,470]" italics="true" pageId="3" pageNumber="4">Fungiseius armatus</emphasis>
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<emphasis id="B9626833640DFFF7C8D2FE33610094F0" bold="true" box="[506,631,447,470]" pageId="3" pageNumber="4">new species</emphasis>
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<subSubSection id="C30CE7AA640DFFF7C9BFFDF162FA9134" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BA9B421640DFFF7C9BFFDF162FA9134" blockId="3.[151,1436,514,2014]" pageId="3" pageNumber="4">
<emphasis id="B9626833640DFFF7C9BFFDF16266973D" bold="true" box="[151,273,514,539]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
Adults of 
<taxonomicName id="4C16CFA2640DFFF7C8A4FDF76170973D" box="[396,519,516,539]" class="Arachnida" family="Blattisociidae" genus="Fungiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9626833640DFFF7C8A4FDF76170973D" box="[396,519,516,539]" italics="true" pageId="3" pageNumber="4">Fungiseius</emphasis>
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are distinguished apomorphically from those of other blattisociine genera by the following attributes: dorsal shield with lateral margins bearing a delineated strip extending from setae 
<emphasis id="B9626833640DFFF7CC3BFDD5665B9718" box="[1299,1324,550,574]" italics="true" pageId="3" pageNumber="4">z2</emphasis>
to 
<emphasis id="B9626833640DFFF7CC78FDD4661A9718" box="[1360,1389,551,574]" italics="true" pageId="3" pageNumber="4">S5</emphasis>
and bearing some of 
<emphasis id="B9626833640DFFF7C870FDBF62159745" box="[344,354,588,611]" italics="true" pageId="3" pageNumber="4">r</emphasis>
-marginal setae anteriorly and the series of 
<emphasis id="B9626833640DFFF7CA7CFDB860159744" box="[852,866,587,610]" italics="true" pageId="3" pageNumber="4">S</emphasis>
- lateral setae posteriorly; sternal or sternitigenital shield with gland pore at apex of endopodal extensions between legs I and II; female with small ventrianal or anal shield; female with deep postgenital furrow; male and female with deep postanal furrow and lacking opisthogastric setae 
<emphasis id="B9626833640DFFF7C9FEFD45627697E8" box="[214,257,694,718]" italics="true" pageId="3" pageNumber="4">JV3</emphasis>
; hypostomatic setae 
<emphasis id="B9626833640DFFF7C8C6FD45616097E8" box="[494,535,694,718]" italics="true" pageId="3" pageNumber="4">hp2</emphasis>
minute; legs II greatly thickened, and with enlarged, curved, spine-like setae on dorsolateral surfaces of tarsus; genua III and IV with eight and nine setae respectively, lacking 
<emphasis id="B9626833640DFFF7CDE6FD2F667797D5" box="[1230,1280,731,755]" italics="true" pageId="3" pageNumber="4">pv -1</emphasis>
; tarsus I with one dorsobasal seta 
<emphasis id="B9626833640DFFF7C850FD0D62E39630" box="[376,404,766,790]" italics="true" pageId="3" pageNumber="4">pd</emphasis>
attenuate, erect; claws on legs I–IV with a basal swelling. Among the genera of Blattisociinae, adults of 
<taxonomicName id="4C16CFA2640DFFF7C81EFCD762C6961D" box="[310,433,804,827]" class="Arachnida" family="Blattisociidae" genus="Fungiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9626833640DFFF7C81EFCD762C6961D" box="[310,433,804,827]" italics="true" pageId="3" pageNumber="4">Fungiseius</emphasis>
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are strikingly similar to those of another fungus-inhabiting genus, 
<taxonomicName id="4C16CFA2640DFFF7CD8DFCD06654961D" box="[1189,1315,803,827]" class="Arachnida" family="Ascidae" genus="Hoploseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9626833640DFFF7CD8DFCD06654961D" box="[1189,1315,803,827]" italics="true" pageId="3" pageNumber="4">Hoploseius</emphasis>
</taxonomicName>
, in having legs II greatly thickened and provided with spine-like setae. In contrast to 
<taxonomicName id="4C16CFA2640DFFF7CAD1FCB467039678" box="[1017,1140,839,862]" class="Arachnida" family="Blattisociidae" genus="Fungiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9626833640DFFF7CAD1FCB467039678" box="[1017,1140,839,862]" italics="true" pageId="3" pageNumber="4">Fungiseius</emphasis>
</taxonomicName>
, leg II of 
<taxonomicName id="4C16CFA2640DFFF7CDC7FCB5661A9678" box="[1263,1389,838,862]" class="Arachnida" family="Ascidae" genus="Hoploseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9626833640DFFF7CDC7FCB5661A9678" box="[1263,1389,838,862]" italics="true" pageId="3" pageNumber="4">Hoploseius</emphasis>
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has enlarged spine-like setae on the ventral faces of the femur, genu, tibia and tarsus, and lacks enlarged curved setae on the dorsolateral face of the tarsus. Adults of 
<taxonomicName id="4C16CFA2640DFFF7CB86FC7D605B9680" box="[686,812,910,934]" class="Arachnida" family="Ascidae" genus="Hoploseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9626833640DFFF7CB86FC7D605B9680" box="[686,812,910,934]" italics="true" pageId="3" pageNumber="4">Hoploseius</emphasis>
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differ in other apomorphic ways from those of the new genus in having the apical margin of the fixed cheliceral digit with a curved row of small, rasping teeth; females with an expansive ventrianal shield; genu I lacking setae 
<emphasis id="B9626833640DFFF7CA12FC25601A96C8" box="[826,877,982,1006]" italics="true" pageId="3" pageNumber="4">pd -3</emphasis>
and 
<emphasis id="B9626833640DFFF7CA84FC2B60AA96C8" box="[940,989,982,1007]" italics="true" pageId="3" pageNumber="4">av -2</emphasis>
, and genua III-IV retaining seta 
<emphasis id="B9626833640DFFF7CC4BFC2B66E296C8" box="[1379,1429,982,1007]" italics="true" pageId="3" pageNumber="4">pv -1</emphasis>
, although lacking 
<emphasis id="B9626833640DFFF7C871FC0862F19135" box="[345,390,1019,1043]" italics="true" pageId="3" pageNumber="4">al -2</emphasis>
.
</paragraph>
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<paragraph id="8BA9B421640DFFF7C9EFFBED676790E4" blockId="3.[151,1436,514,2014]" pageId="3" pageNumber="4">
<emphasis id="B9626833640DFFF7C9EFFBED622E9111" bold="true" box="[199,345,1054,1079]" pageId="3" pageNumber="4">Description.</emphasis>
<emphasis id="B9626833640DFFF7C848FBED615A9110" box="[352,557,1054,1078]" italics="true" pageId="3" pageNumber="4">Idiosomal dorsum</emphasis>
. ADULT FEMALE AND MALE (
<figureCitation id="132DA8A4640DFFF7CA57FBED60BE9110" box="[895,969,1054,1078]" captionStart="FIGURES 9 – 14" captionStartId="8.[151,264,1897,1920]" captionTargetBox="[165,1426,401,1858]" captionTargetId="figure@8.[165,1426,401,1880]" captionTargetPageId="8" captionText="FIGURES 9 – 14. Fungiseius armatus n. sp., adult female: 9, idiosoma, dorsal aspect with detail of peritrematal shield (arrows indicate delineated strip of dorsal shield margins bearing some of r - marginal and all S - lateral setae); 10, detail of separate views of dorsal setae denoted and lyrifissure; 11 a – b, detail of ventral view of dorsal and peritrematal shields; 12, detail of exopodal IV with peritremal poroidotaxy at level of coxa IV; 13, detail of denoted dorsal setae; 14, detail of dorsal ornamentation." httpUri="https://zenodo.org/record/276759/files/figure.png" pageId="3" pageNumber="4">Figs 9</figureCitation>
, 
<figureCitation id="132DA8A4640DFFF7CAFEFBED60819110" box="[982,1014,1054,1078]" captionStart="FIGURES 38 – 41" captionStartId="14.[151,264,1922,1945]" captionTargetBox="[151,1431,380,1900]" captionTargetId="figure@14.[151,1436,367,1900]" captionTargetPageId="14" captionText="FIGURES 38 – 41. Fungiseius clavulisetis n. sp., adult female: 38, idiosoma, dorsal aspect (arrows indicate delineated strip of dorsal shield margins bearing r - marginal and S - lateral setae); 39, detail of denoted dorsal setae; 40, chelicera, detail of movable digit; 41, chelicera, dorsal view." httpUri="https://zenodo.org/record/276764/files/figure.png" pageId="3" pageNumber="4">38</figureCitation>
). Dorsal shield entire, without lateral incisions, well sclerotized and with delineated lateral rim extending from setae 
<emphasis id="B9626833640DFFF7CD19FBB0673D917D" box="[1073,1098,1091,1115]" italics="true" pageId="3" pageNumber="4">z2</emphasis>
to 
<emphasis id="B9626833640DFFF7CD5CFBB067E7917C" box="[1140,1168,1091,1114]" italics="true" pageId="3" pageNumber="4">S5</emphasis>
and bearing a variable number of 
<emphasis id="B9626833640DFFF7C83AFB9B626A9159" box="[274,285,1128,1151]" italics="true" pageId="3" pageNumber="4">s</emphasis>
or 
<emphasis id="B9626833640DFFF7C86AFB9B623A9159" box="[322,333,1128,1151]" italics="true" pageId="3" pageNumber="4">r</emphasis>
setae, and all 
<emphasis id="B9626833640DFFF7C8C4FB9561599158" box="[492,558,1126,1150]" italics="true" pageId="3" pageNumber="4">S1-S5</emphasis>
; surrounding soft integument smoothly striate. Dorsal shield with a maximum complement of 37 pairs of setae (
<figureCitation id="132DA8A4640DFFF7CB3AFB7961119184" box="[530,614,1162,1186]" captionStart="FIGURES 38 – 41" captionStartId="14.[151,264,1922,1945]" captionTargetBox="[151,1431,380,1900]" captionTargetId="figure@14.[151,1436,367,1900]" captionTargetPageId="14" captionText="FIGURES 38 – 41. Fungiseius clavulisetis n. sp., adult female: 38, idiosoma, dorsal aspect (arrows indicate delineated strip of dorsal shield margins bearing r - marginal and S - lateral setae); 39, detail of denoted dorsal setae; 40, chelicera, detail of movable digit; 41, chelicera, dorsal view." httpUri="https://zenodo.org/record/276764/files/figure.png" pageId="3" pageNumber="4">Fig. 38</figureCitation>
), including 22 podonotal (
<emphasis id="B9626833640DFFF7CABAFB7860B19185" box="[914,966,1163,1187]" italics="true" pageId="3" pageNumber="4">j1-j6</emphasis>
, 
<emphasis id="B9626833640DFFF7CAFDFB78676F9185" box="[981,1048,1163,1187]" italics="true" pageId="3" pageNumber="4">z1- z6</emphasis>
, 
<emphasis id="B9626833640DFFF7CD0FFB7867149185" box="[1063,1123,1163,1187]" italics="true" pageId="3" pageNumber="4">s1-s6</emphasis>
, 
<emphasis id="B9626833640DFFF7CD59FB7867DB9184" box="[1137,1196,1163,1187]" italics="true" pageId="3" pageNumber="4">r2-r5</emphasis>
) and 15 opisthonotal pairs (
<emphasis id="B9626833640DFFF7C9F5FB5D626B91E0" box="[221,284,1198,1222]" italics="true" pageId="3" pageNumber="4">J1-J5</emphasis>
, 
<emphasis id="B9626833640DFFF7C802FB5D621991E0" box="[298,366,1198,1222]" italics="true" pageId="3" pageNumber="4">Z1-Z5</emphasis>
, 
<emphasis id="B9626833640DFFF7C854FB5D62C991E0" box="[380,446,1198,1222]" italics="true" pageId="3" pageNumber="4">S1-S5</emphasis>
), or with complement reduced to 28 pairs (
<figureCitation id="132DA8A4640DFFF7CA8CFB5D609F91E0" box="[932,1000,1198,1222]" captionStart="FIGURES 9 – 14" captionStartId="8.[151,264,1897,1920]" captionTargetBox="[165,1426,401,1858]" captionTargetId="figure@8.[165,1426,401,1880]" captionTargetPageId="8" captionText="FIGURES 9 – 14. Fungiseius armatus n. sp., adult female: 9, idiosoma, dorsal aspect with detail of peritrematal shield (arrows indicate delineated strip of dorsal shield margins bearing some of r - marginal and all S - lateral setae); 10, detail of separate views of dorsal setae denoted and lyrifissure; 11 a – b, detail of ventral view of dorsal and peritrematal shields; 12, detail of exopodal IV with peritremal poroidotaxy at level of coxa IV; 13, detail of denoted dorsal setae; 14, detail of dorsal ornamentation." httpUri="https://zenodo.org/record/276759/files/figure.png" pageId="3" pageNumber="4">Fig. 9</figureCitation>
), including 17 podonotal (
<emphasis id="B9626833640DFFF7CC3AFB5D663091E0" box="[1298,1351,1198,1222]" italics="true" pageId="3" pageNumber="4">j1-j6</emphasis>
, 
<emphasis id="B9626833640DFFF7CC7DFB5D661991E0" box="[1365,1390,1198,1222]" italics="true" pageId="3" pageNumber="4">z1</emphasis>
, 
<emphasis id="B9626833640DFFF7CC54FB5D66E391E0" box="[1404,1428,1198,1222]" italics="true" pageId="3" pageNumber="4">z2</emphasis>
, 
<emphasis id="B9626833640DFFF7C9BFFB2063C791CD" box="[151,176,1235,1259]" italics="true" pageId="3" pageNumber="4">z4</emphasis>
, 
<emphasis id="B9626833640DFFF7C9E8FB2063AE91CC" box="[192,217,1235,1258]" italics="true" pageId="3" pageNumber="4">z5</emphasis>
, 
<emphasis id="B9626833640DFFF7C9C1FB20625491CD" box="[233,291,1235,1259]" italics="true" pageId="3" pageNumber="4">s1-s6</emphasis>
, 
<emphasis id="B9626833640DFFF7C81CFB20623A91CD" box="[308,333,1235,1259]" italics="true" pageId="3" pageNumber="4">r2</emphasis>
) and 11 opisthonotal pairs (
<emphasis id="B9626833640DFFF7CBBAFB2061DB91CC" box="[658,684,1235,1258]" italics="true" pageId="3" pageNumber="4">J5</emphasis>
, 
<emphasis id="B9626833640DFFF7CB94FB20607691CC" box="[700,769,1235,1259]" italics="true" pageId="3" pageNumber="4">Z1-Z5</emphasis>
, 
<emphasis id="B9626833640DFFF7CA39FB20602491CC" box="[785,851,1235,1259]" italics="true" pageId="3" pageNumber="4">S1-S5</emphasis>
); dorsal setae moderately short, smooth, of similar lengths, or with two pairs of posterior setae (
<emphasis id="B9626833640DFFF7CBBCFB0561AE9028" box="[660,729,1270,1294]" italics="true" pageId="3" pageNumber="4">Z4-Z5</emphasis>
) enlarged and modified. Dorsal shield with complement of 18 pairs of discernible pore-like structures (eight podonotal, ten opisthonotal), of which seven pairs (four podonotal, three opisthonotal) superficially appear secretory (gland pores) and 11 pairs (four podonotal, seven opisthonotal) non-secretory (poroids). Soft surrounding cuticle with five to eight pairs of 
<emphasis id="B9626833640DFFF7CACCFA97677E905D" box="[996,1033,1379,1403]" italics="true" pageId="3" pageNumber="4">r -R</emphasis>
marginal setae and pair of marginal poroids 
<emphasis id="B9626833640DFFF7C9DCFA75625E90B8" box="[244,297,1414,1438]" italics="true" pageId="3" pageNumber="4">idRp</emphasis>
. Peritrematal plates narrowly uniting with dorsal shield anteriorly at level of setae 
<emphasis id="B9626833640DFFF7CDC3FA75667390B8" box="[1259,1284,1414,1438]" italics="true" pageId="3" pageNumber="4">z1</emphasis>
or free there (
<figureCitation id="132DA8A4640DFFF7C9B7FA59638690E4" box="[159,241,1450,1474]" captionStart="FIGURES 9 – 14" captionStartId="8.[151,264,1897,1920]" captionTargetBox="[165,1426,401,1858]" captionTargetId="figure@8.[165,1426,401,1880]" captionTargetPageId="8" captionText="FIGURES 9 – 14. Fungiseius armatus n. sp., adult female: 9, idiosoma, dorsal aspect with detail of peritrematal shield (arrows indicate delineated strip of dorsal shield margins bearing some of r - marginal and all S - lateral setae); 10, detail of separate views of dorsal setae denoted and lyrifissure; 11 a – b, detail of ventral view of dorsal and peritrematal shields; 12, detail of exopodal IV with peritremal poroidotaxy at level of coxa IV; 13, detail of denoted dorsal setae; 14, detail of dorsal ornamentation." httpUri="https://zenodo.org/record/276759/files/figure.png" pageId="3" pageNumber="4">Fig. 11</figureCitation>
); peritremes well developed, reaching to level between setae 
<emphasis id="B9626833640DFFF7CA89FA5860CD90E5" box="[929,954,1451,1475]" italics="true" pageId="3" pageNumber="4">s1</emphasis>
and 
<emphasis id="B9626833640DFFF7CAD8FA58677E90E5" box="[1008,1033,1451,1475]" italics="true" pageId="3" pageNumber="4">z1</emphasis>
.
</paragraph>
<paragraph id="8BA9B421640DFFF0C9EFFA3D63B49419" blockId="3.[151,1436,514,2014]" lastBlockId="4.[151,1436,151,2013]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="B9626833640DFFF7C9EFFA3D62F590C1" box="[199,386,1486,1511]" italics="true" pageId="3" pageNumber="4">Idiosomal venter</emphasis>
. ADULT FEMALE (
<figureCitation id="132DA8A4640DFFF7CB79FA3D61DF90C0" box="[593,680,1486,1510]" captionStart="FIGURES 15 – 18" captionStartId="9.[151,264,1939,1962]" captionTargetBox="[151,1431,193,1890]" captionTargetId="figure@9.[151,1436,193,1901]" captionTargetPageId="9" captionText="FIGURES 15 – 18. Fungiseius armatus n. sp., adult female: 15, idiosoma, ventral aspect; 16, detail of peritrematal shield, fused to exopodal strips between legs II – III and III – IV; 17, detail of sternal shielding; 18 a, b, detail of spermatheca." httpUri="https://zenodo.org/record/276760/files/figure.png" pageId="3" pageNumber="4">Figs 15</figureCitation>
, 
<figureCitation id="132DA8A4640DFFF7CB9DFA3D61A290C0" box="[693,725,1486,1510]" captionStart="FIGURES 42 – 47" captionStartId="15.[151,264,1747,1770]" captionTargetBox="[151,1436,193,1723]" captionTargetId="figure@15.[151,1436,193,1726]" captionTargetPageId="15" captionText="FIGURES 42 – 47. Fungiseius clavulisetis n. sp., adult female: 42, tritosternum; 43, gnathotectum; 44, corniculi, dorsal view; 45, subcapitulum; 46, idiosoma, ventral aspect; 47, detail of spermatheca." httpUri="https://zenodo.org/record/276765/files/figure.png" pageId="3" pageNumber="4">46</figureCitation>
). Tritosternum with laciniae free for most of length, their fused bases with a series of denticles or smooth (
<figureCitation id="132DA8A4640DFFF7CB55FA0161B1932C" box="[637,710,1522,1546]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1633,1656]" captionTargetBox="[151,1436,217,1606]" captionTargetId="figure@6.[151,1436,195,1616]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Fungiseius armatus n. sp., adult female: 1, gnathotectum; 2, subcapitulum; 3, corniculi, dorsal view; 4, labrum; 5, tritosternum, showing denticles at base of laciniae; 6, palpus, dorsolateral view; 7 a, detail of seta al of palpgenu; 7 b, detail of seta al of palpfemur; 8 a, chelicera, lateral view; 8 b, separate paraxial view of fixed digit." httpUri="https://zenodo.org/record/276758/files/figure.png" pageId="3" pageNumber="4">Figs 5</figureCitation>
, 
<figureCitation id="132DA8A4640DFFF7CBFCFA016183932C" box="[724,756,1522,1546]" captionStart="FIGURES 42 – 47" captionStartId="15.[151,264,1747,1770]" captionTargetBox="[151,1436,193,1723]" captionTargetId="figure@15.[151,1436,193,1726]" captionTargetPageId="15" captionText="FIGURES 42 – 47. Fungiseius clavulisetis n. sp., adult female: 42, tritosternum; 43, gnathotectum; 44, corniculi, dorsal view; 45, subcapitulum; 46, idiosoma, ventral aspect; 47, detail of spermatheca." httpUri="https://zenodo.org/record/276765/files/figure.png" pageId="3" pageNumber="4">42</figureCitation>
). Ventral shields variably ornamented, well sclerotized. Presternal region ornamented with pair of weakly sclerotized plates consolidated with sternal shield. Sternal shield entire, with strongly developed endopodal extensions between coxae I–II and II–III, that between legs I–II with a gland pore apically where abutting peritrematal/exopodal shield; sternal shield with three pairs of setae and two pairs of lyrifissures; setae 
<emphasis id="B9626833640DFFF7C8EAF970629493BD" box="[450,483,1667,1691]" italics="true" pageId="3" pageNumber="4">st4</emphasis>
and poroids 
<date id="FFA892E1640DFFF7CB51F97061EC93BD" box="[633,667,1667,1691]" pageId="3" pageNumber="4">
<emphasis id="B9626833640DFFF7CB51F97061EC93BD" box="[633,667,1667,1691]" italics="true" pageId="3" pageNumber="4">iv3</emphasis>
</date>
on metasternal plates that are free or narrowly connected to sternal shield (
<figureCitation id="132DA8A4640DFFF7C9C6F955623D9398" box="[238,330,1702,1726]" captionStart="FIGURES 15 – 18" captionStartId="9.[151,264,1939,1962]" captionTargetBox="[151,1431,193,1890]" captionTargetId="figure@9.[151,1436,193,1901]" captionTargetPageId="9" captionText="FIGURES 15 – 18. Fungiseius armatus n. sp., adult female: 15, idiosoma, ventral aspect; 16, detail of peritrematal shield, fused to exopodal strips between legs II – III and III – IV; 17, detail of sternal shielding; 18 a, b, detail of spermatheca." httpUri="https://zenodo.org/record/276760/files/figure.png" pageId="3" pageNumber="4">Figs 15</figureCitation>
, 
<figureCitation id="132DA8A4640DFFF7C873F955620C9398" box="[347,379,1702,1726]" captionStart="FIGURES 42 – 47" captionStartId="15.[151,264,1747,1770]" captionTargetBox="[151,1436,193,1723]" captionTargetId="figure@15.[151,1436,193,1726]" captionTargetPageId="15" captionText="FIGURES 42 – 47. Fungiseius clavulisetis n. sp., adult female: 42, tritosternum; 43, gnathotectum; 44, corniculi, dorsal view; 45, subcapitulum; 46, idiosoma, ventral aspect; 47, detail of spermatheca." httpUri="https://zenodo.org/record/276765/files/figure.png" pageId="3" pageNumber="4">46</figureCitation>
). Endopodal strips between coxae III and IV free, well defined. Epigynal shield with its rounded, hyaline anterior margin between legs III abutting or overlapping posterior edge of sternal shield, its posterior margin truncate; setae 
<emphasis id="B9626833640DFFF7C8E0F91C629E9220" box="[456,489,1775,1798]" italics="true" pageId="3" pageNumber="4">st5</emphasis>
on shield’s lateral margins, paragenital poroids 
<date id="FFA892E1640DFFF7CD21F91C675C9220" box="[1033,1067,1775,1798]" pageId="3" pageNumber="4">
<emphasis id="B9626833640DFFF7CD21F91C675C9220" box="[1033,1067,1775,1798]" italics="true" pageId="3" pageNumber="4">iv5</emphasis>
</date>
on soft cuticle; deep postgenital furrow present with small postgenital platelets usually discernible. Opisthosomatic venter with well-developed metapodal platelets, nine pairs of setae (
<emphasis id="B9626833640DFFF7CB7EF8C561F69268" box="[598,641,1846,1870]" italics="true" pageId="3" pageNumber="4">JV1</emphasis>
, 
<emphasis id="B9626833640DFFF7CBA7F8C561CD9268" box="[655,698,1846,1870]" italics="true" pageId="3" pageNumber="4">JV2</emphasis>
, 
<emphasis id="B9626833640DFFF7CBE0F8C561839268" box="[712,756,1846,1870]" italics="true" pageId="3" pageNumber="4">JV4</emphasis>
, 
<emphasis id="B9626833640DFFF7CA29F8C4605A9268" box="[769,813,1847,1870]" italics="true" pageId="3" pageNumber="4">JV5</emphasis>
, 
<emphasis id="B9626833640DFFF7CA12F8C560D69268" box="[826,929,1846,1870]" italics="true" pageId="3" pageNumber="4">ZV1-ZV5</emphasis>
), and three pairs of poroids, 
<emphasis id="B9626833640DFFF7CDC9F8C466749268" box="[1249,1283,1847,1870]" italics="true" pageId="3" pageNumber="4">ivp</emphasis>
conspicuous; setae 
<emphasis id="B9626833640DFFF7C9FFF8A862749255" box="[215,259,1883,1907]" italics="true" pageId="3" pageNumber="4">JV3</emphasis>
absent. Opisthogaster with either small ventrianal shield bearing three pairs of ventral setae (
<emphasis id="B9626833640DFFF7CC06F8A8662E9255" box="[1326,1369,1883,1907]" italics="true" pageId="3" pageNumber="4">JV1</emphasis>
, 
<emphasis id="B9626833640DFFF7CC41F8A866E29255" box="[1385,1429,1883,1907]" italics="true" pageId="3" pageNumber="4">JV2</emphasis>
, 
<emphasis id="B9626833640DFFF7C9BFF88D63B192B0" box="[151,198,1918,1942]" italics="true" pageId="3" pageNumber="4">ZV2</emphasis>
) or oval anal shield, with paranal setae inserted at mid level of anus, and more elongate postanal seta; shield with a pair of gland pores 
<emphasis id="B9626833640DFFF7C8E1F8506286929D" box="[457,497,1955,1979]" italics="true" pageId="3" pageNumber="4">gv3</emphasis>
on posterolateral margins, and a well-developed cribrum behind level of postanal seta; deep postanal furrow present. Peritrematal shield connecting with exopodal strips behind coxae IV, with two poroids and one gland pore in area behind stigma, and with poroid (
<emphasis id="B9626833640AFFF0CA8FFF6B60BD9596" box="[935,970,152,176]" italics="true" pageId="4" pageNumber="5">ip1</emphasis>
) and two weakly developed gland pores (
<emphasis id="B9626833640AFFF0C988FF4F63BD95F2" box="[160,202,188,212]" italics="true" pageId="4" pageNumber="5">gp1</emphasis>
, 
<emphasis id="B9626833640AFFF0C9F1FF4F627595F2" box="[217,258,188,212]" italics="true" pageId="4" pageNumber="5">gp2</emphasis>
) along length of peritreme (
<figureCitation id="132DA8A4640AFFF0CB6BFF4F61FF95F2" box="[579,648,188,212]" captionStart="FIGURES 9 – 14" captionStartId="8.[151,264,1897,1920]" captionTargetBox="[165,1426,401,1858]" captionTargetId="figure@8.[165,1426,401,1880]" captionTargetPageId="8" captionText="FIGURES 9 – 14. Fungiseius armatus n. sp., adult female: 9, idiosoma, dorsal aspect with detail of peritrematal shield (arrows indicate delineated strip of dorsal shield margins bearing some of r - marginal and all S - lateral setae); 10, detail of separate views of dorsal setae denoted and lyrifissure; 11 a – b, detail of ventral view of dorsal and peritrematal shields; 12, detail of exopodal IV with peritremal poroidotaxy at level of coxa IV; 13, detail of denoted dorsal setae; 14, detail of dorsal ornamentation." httpUri="https://zenodo.org/record/276759/files/figure.png" pageId="4" pageNumber="5">Fig. 9</figureCitation>
); exopodal strip continuous alongside peritrematal shield by coxae I– IV, with extensions between bases of coxae I–II, II–III and III–IV. Spermathecal apparatus of phytoseioid-type, but sometimes with minor duct difficult to discern emanating from embolus near base of sclerotized calyx (
<figureCitation id="132DA8A4640AFFF0CC16FEF766E0943A" box="[1342,1431,260,284]" captionStart="FIGURES 15 – 18" captionStartId="9.[151,264,1939,1962]" captionTargetBox="[151,1431,193,1890]" captionTargetId="figure@9.[151,1436,193,1901]" captionTargetPageId="9" captionText="FIGURES 15 – 18. Fungiseius armatus n. sp., adult female: 15, idiosoma, ventral aspect; 16, detail of peritrematal shield, fused to exopodal strips between legs II – III and III – IV; 17, detail of sternal shielding; 18 a, b, detail of spermatheca." httpUri="https://zenodo.org/record/276760/files/figure.png" pageId="4" pageNumber="5">Figs 18</figureCitation>
, 
<figureCitation id="132DA8A4640AFFF0C9BFFED463C09419" box="[151,183,295,319]" captionStart="FIGURES 42 – 47" captionStartId="15.[151,264,1747,1770]" captionTargetBox="[151,1436,193,1723]" captionTargetId="figure@15.[151,1436,193,1726]" captionTargetPageId="15" captionText="FIGURES 42 – 47. Fungiseius clavulisetis n. sp., adult female: 42, tritosternum; 43, gnathotectum; 44, corniculi, dorsal view; 45, subcapitulum; 46, idiosoma, ventral aspect; 47, detail of spermatheca." httpUri="https://zenodo.org/record/276765/files/figure.png" pageId="4" pageNumber="5">47</figureCitation>
).
</paragraph>
<paragraph id="8BA9B421640AFFF0C9EFFEBF665C971A" blockId="4.[151,1436,151,2013]" pageId="4" pageNumber="5">
ADULT MALE (
<figureCitation id="132DA8A4640AFFF0C844FEBF62C89442" box="[364,447,332,356]" captionStart="FIGURES 32 – 37" captionStartId="12.[151,264,1939,1962]" captionTargetBox="[151,1433,433,1911]" captionTargetId="figure@12.[151,1436,433,1918]" captionTargetPageId="12" captionText="FIGURES 32 – 37. Fungiseius armatus n. sp., adult male: 32, subcapitulum; 33, chelicerae, dorsolateral view; 34, chelicerae, detail of fixed digit with pilus dentilis; 35, chelicera, detail of movable digit and spermatodactyl; 36, detail of spermatodactyl; 37, idiosoma, ventral view." httpUri="https://zenodo.org/record/276763/files/figure.png" pageId="4" pageNumber="5">Fig. 37</figureCitation>
). Form of tritosternum and presternal area as in female. Ventral shields ornamented with reticula, well sclerotized. Sternitigenital shield with five pairs of setae and three pairs of poroids, and with endopodal extensions between coxae I–II, II–III, and partially fused with those between coxae III–IV (
<figureCitation id="132DA8A4640AFFF0CD93FE676679948A" box="[1211,1294,404,428]" captionStart="FIGURES 32 – 37" captionStartId="12.[151,264,1939,1962]" captionTargetBox="[151,1433,433,1911]" captionTargetId="figure@12.[151,1436,433,1918]" captionTargetPageId="12" captionText="FIGURES 32 – 37. Fungiseius armatus n. sp., adult male: 32, subcapitulum; 33, chelicerae, dorsolateral view; 34, chelicerae, detail of fixed digit with pilus dentilis; 35, chelicera, detail of movable digit and spermatodactyl; 36, detail of spermatodactyl; 37, idiosoma, ventral view." httpUri="https://zenodo.org/record/276763/files/figure.png" pageId="4" pageNumber="5">Fig. 37</figureCitation>
). Opisthosomatic venter with one or two pairs of metapodal platelets; soft integument with two or three pairs of setae and three pairs of poroids. Ventrianal shield large, narrowed at level of anal region, with four pairs of opisthogastric setae (
<emphasis id="B9626833640AFFF0C988FDF363BC973E" box="[160,203,512,536]" italics="true" pageId="4" pageNumber="5">JV1</emphasis>
, 
<emphasis id="B9626833640AFFF0C9F1FDF36272973E" box="[217,261,512,536]" italics="true" pageId="4" pageNumber="5">JV2</emphasis>
, 
<emphasis id="B9626833640AFFF0C83AFDF36237973E" box="[274,320,512,536]" italics="true" pageId="4" pageNumber="5">ZV1</emphasis>
, 
<emphasis id="B9626833640AFFF0C866FDF3620A973E" box="[334,381,512,536]" italics="true" pageId="4" pageNumber="5">ZV2</emphasis>
) and the three circumanal setae; attributes of circumanal setae, cribrum, and deep postanal furrow as in female; setae 
<emphasis id="B9626833640AFFF0C8B6FDD762BD971A" box="[414,458,548,572]" italics="true" pageId="4" pageNumber="5">JV3</emphasis>
, 
<emphasis id="B9626833640AFFF0C8F0FDD76149971A" box="[472,574,548,572]" italics="true" pageId="4" pageNumber="5">ZV3-ZV5</emphasis>
absent. Posteriorly, peritrematal shield and peritreme as in female.
</paragraph>
<paragraph id="8BA9B421640AFFF0C9EFFDBB60BB96E1" blockId="4.[151,1436,151,2013]" pageId="4" pageNumber="5">
<taxonomicName id="4C16CFA2640AFFF0C9EFFDBB62219746" box="[199,342,584,608]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="5" phylum="Nematoda" rank="genus">
<emphasis id="B9626833640AFFF0C9EFFDBB62219746" box="[199,342,584,608]" italics="true" pageId="4" pageNumber="5">Gnathosoma</emphasis>
</taxonomicName>
. FEMALE AND MALE. Gnathotectum with convex, denticulate anterior margin (
<figureCitation id="132DA8A4640AFFF0CDEBFDB4667E9779" box="[1219,1289,583,607]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1633,1656]" captionTargetBox="[151,1436,217,1606]" captionTargetId="figure@6.[151,1436,195,1616]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Fungiseius armatus n. sp., adult female: 1, gnathotectum; 2, subcapitulum; 3, corniculi, dorsal view; 4, labrum; 5, tritosternum, showing denticles at base of laciniae; 6, palpus, dorsolateral view; 7 a, detail of seta al of palpgenu; 7 b, detail of seta al of palpfemur; 8 a, chelicera, lateral view; 8 b, separate paraxial view of fixed digit." httpUri="https://zenodo.org/record/276758/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
, 
<figureCitation id="132DA8A4640AFFF0CC3FFDB466409779" box="[1303,1335,583,607]" captionStart="FIGURES 42 – 47" captionStartId="15.[151,264,1747,1770]" captionTargetBox="[151,1436,193,1723]" captionTargetId="figure@15.[151,1436,193,1726]" captionTargetPageId="15" captionText="FIGURES 42 – 47. Fungiseius clavulisetis n. sp., adult female: 42, tritosternum; 43, gnathotectum; 44, corniculi, dorsal view; 45, subcapitulum; 46, idiosoma, ventral aspect; 47, detail of spermatheca." httpUri="https://zenodo.org/record/276765/files/figure.png" pageId="4" pageNumber="5">43</figureCitation>
). Chelicerae large (
<figureCitation id="132DA8A4640AFFF0C83DFD9F622E97A2" box="[277,345,620,644]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1633,1656]" captionTargetBox="[151,1436,217,1606]" captionTargetId="figure@6.[151,1436,195,1616]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Fungiseius armatus n. sp., adult female: 1, gnathotectum; 2, subcapitulum; 3, corniculi, dorsal view; 4, labrum; 5, tritosternum, showing denticles at base of laciniae; 6, palpus, dorsolateral view; 7 a, detail of seta al of palpgenu; 7 b, detail of seta al of palpfemur; 8 a, chelicera, lateral view; 8 b, separate paraxial view of fixed digit." httpUri="https://zenodo.org/record/276758/files/figure.png" pageId="4" pageNumber="5">Fig. 8</figureCitation>
), or small (
<figureCitation id="132DA8A4640AFFF0C8F5FD9F614697A2" box="[477,561,620,644]" captionStart="FIGURES 38 – 41" captionStartId="14.[151,264,1922,1945]" captionTargetBox="[151,1431,380,1900]" captionTargetId="figure@14.[151,1436,367,1900]" captionTargetPageId="14" captionText="FIGURES 38 – 41. Fungiseius clavulisetis n. sp., adult female: 38, idiosoma, dorsal aspect (arrows indicate delineated strip of dorsal shield margins bearing r - marginal and S - lateral setae); 39, detail of denoted dorsal setae; 40, chelicera, detail of movable digit; 41, chelicera, dorsal view." httpUri="https://zenodo.org/record/276764/files/figure.png" pageId="4" pageNumber="5">Fig. 41</figureCitation>
), without any conspicuous process along antiaxial or paraxial lateral surfaces basal to the digits; fixed digit with tiny pilus dentilis and row of small teeth along distal half of masticatory surface and one or two small rasping teeth (gabelzähne) subapically; movable cheliceral digit with three coarse teeth; arthrodial envelope margin smooth; movable digit of male unidentate; spermatodactyl digitiform (
<figureCitation id="132DA8A4640AFFF0CC2EFD2466E797C9" box="[1286,1424,727,751]" captionStart="FIGURES 32 – 37" captionStartId="12.[151,264,1939,1962]" captionTargetBox="[151,1433,433,1911]" captionTargetId="figure@12.[151,1436,433,1918]" captionTargetPageId="12" captionText="FIGURES 32 – 37. Fungiseius armatus n. sp., adult male: 32, subcapitulum; 33, chelicerae, dorsolateral view; 34, chelicerae, detail of fixed digit with pilus dentilis; 35, chelicera, detail of movable digit and spermatodactyl; 36, detail of spermatodactyl; 37, idiosoma, ventral view." httpUri="https://zenodo.org/record/276763/files/figure.png" pageId="4" pageNumber="5">Figs 35, 36</figureCitation>
). Deutosternum with seven transverse rows of denticles of similar width (
<figureCitation id="132DA8A4640AFFF0CAF2FD0F67599632" box="[986,1070,764,788]" captionStart="FIGURES 42 – 47" captionStartId="15.[151,264,1747,1770]" captionTargetBox="[151,1436,193,1723]" captionTargetId="figure@15.[151,1436,193,1726]" captionTargetPageId="15" captionText="FIGURES 42 – 47. Fungiseius clavulisetis n. sp., adult female: 42, tritosternum; 43, gnathotectum; 44, corniculi, dorsal view; 45, subcapitulum; 46, idiosoma, ventral aspect; 47, detail of spermatheca." httpUri="https://zenodo.org/record/276765/files/figure.png" pageId="4" pageNumber="5">Fig. 45</figureCitation>
) or progressively wider basally (
<figureCitation id="132DA8A4640AFFF0C9B7FCEC63919611" box="[159,230,799,823]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1633,1656]" captionTargetBox="[151,1436,217,1606]" captionTargetId="figure@6.[151,1436,195,1616]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Fungiseius armatus n. sp., adult female: 1, gnathotectum; 2, subcapitulum; 3, corniculi, dorsal view; 4, labrum; 5, tritosternum, showing denticles at base of laciniae; 6, palpus, dorsolateral view; 7 a, detail of seta al of palpgenu; 7 b, detail of seta al of palpfemur; 8 a, chelicera, lateral view; 8 b, separate paraxial view of fixed digit." httpUri="https://zenodo.org/record/276758/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
), rows multidenticulate, all rows connected, or basal row free. Corniculi normal in form, well-separated from base to apex, shorter and stouter in male (
<figureCitation id="132DA8A4640AFFF0CB8EFCB7618D967A" box="[678,762,836,860]" captionStart="FIGURES 32 – 37" captionStartId="12.[151,264,1939,1962]" captionTargetBox="[151,1433,433,1911]" captionTargetId="figure@12.[151,1436,433,1918]" captionTargetPageId="12" captionText="FIGURES 32 – 37. Fungiseius armatus n. sp., adult male: 32, subcapitulum; 33, chelicerae, dorsolateral view; 34, chelicerae, detail of fixed digit with pilus dentilis; 35, chelicera, detail of movable digit and spermatodactyl; 36, detail of spermatodactyl; 37, idiosoma, ventral view." httpUri="https://zenodo.org/record/276763/files/figure.png" pageId="4" pageNumber="5">Fig. 32</figureCitation>
); internal malae highly variable in form (
<figureCitation id="132DA8A4640AFFF0CDEFFCB76678967A" box="[1223,1295,836,860]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1633,1656]" captionTargetBox="[151,1436,217,1606]" captionTargetId="figure@6.[151,1436,195,1616]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Fungiseius armatus n. sp., adult female: 1, gnathotectum; 2, subcapitulum; 3, corniculi, dorsal view; 4, labrum; 5, tritosternum, showing denticles at base of laciniae; 6, palpus, dorsolateral view; 7 a, detail of seta al of palpgenu; 7 b, detail of seta al of palpfemur; 8 a, chelicera, lateral view; 8 b, separate paraxial view of fixed digit." httpUri="https://zenodo.org/record/276758/files/figure.png" pageId="4" pageNumber="5">Figs 2</figureCitation>
, 
<figureCitation id="132DA8A4640AFFF0CC33FCB7664C967A" box="[1307,1339,836,860]" captionStart="FIGURES 42 – 47" captionStartId="15.[151,264,1747,1770]" captionTargetBox="[151,1436,193,1723]" captionTargetId="figure@15.[151,1436,193,1726]" captionTargetPageId="15" captionText="FIGURES 42 – 47. Fungiseius clavulisetis n. sp., adult female: 42, tritosternum; 43, gnathotectum; 44, corniculi, dorsal view; 45, subcapitulum; 46, idiosoma, ventral aspect; 47, detail of spermatheca." httpUri="https://zenodo.org/record/276765/files/figure.png" pageId="4" pageNumber="5">45</figureCitation>
). Subcapitular setae smooth, 
<emphasis id="B9626833640AFFF0C8A0FC9B62C696A6" box="[392,433,872,896]" italics="true" pageId="4" pageNumber="5">hp1</emphasis>
and 
<emphasis id="B9626833640AFFF0C8C1FC9B616396A6" box="[489,532,872,896]" italics="true" pageId="4" pageNumber="5">hp3</emphasis>
similar in length and longer than 
<emphasis id="B9626833640AFFF0CABAFC9A60DB96A6" box="[914,940,873,896]" italics="true" pageId="4" pageNumber="5">cp</emphasis>
, lateral 
<emphasis id="B9626833640AFFF0CD20FC9B674596A6" box="[1032,1074,872,896]" italics="true" pageId="4" pageNumber="5">hp2</emphasis>
minute. Palpi with normal setation as described for 
<taxonomicName id="4C16CFA2640AFFF0C8A0FC7F628F9682" box="[392,504,908,932]" class="Arachnida" infraOrder="Gamasina" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="5" phylum="Arthropoda" rank="infraOrder">Gamasina</taxonomicName>
by 
<bibRefCitation id="EF87C9D0640AFFF0CB0EFC7E61B79682" author="Evans" box="[550,704,908,932]" pageId="4" pageNumber="23" refString="Evans, G. O. (1964) Some observations on the chaetotaxy of the pedipalps in the Mesostigmata (Acari). Annals and Magazine of Natural History, Series 13, 6, 513 - 527." type="journal article" year="1964">Evans (1964)</bibRefCitation>
; palpfemoral seta 
<emphasis id="B9626833640AFFF0CABAFC7F60DF9682" box="[914,936,908,932]" italics="true" pageId="4" pageNumber="5">al</emphasis>
and palpgenual setae 
<emphasis id="B9626833640AFFF0CD8EFC7F67A49682" box="[1190,1235,908,932]" italics="true" pageId="4" pageNumber="5">al -1</emphasis>
and 
<emphasis id="B9626833640AFFF0CC24FC7F664D9682" box="[1292,1338,908,932]" italics="true" pageId="4" pageNumber="5">al -2</emphasis>
more or less spatulate; palptarsal apotele two-tined, with spatulate tips (
<figureCitation id="132DA8A4640AFFF0CA72FC5C60C996E1" box="[858,958,943,967]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1633,1656]" captionTargetBox="[151,1436,217,1606]" captionTargetId="figure@6.[151,1436,195,1616]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Fungiseius armatus n. sp., adult female: 1, gnathotectum; 2, subcapitulum; 3, corniculi, dorsal view; 4, labrum; 5, tritosternum, showing denticles at base of laciniae; 6, palpus, dorsolateral view; 7 a, detail of seta al of palpgenu; 7 b, detail of seta al of palpfemur; 8 a, chelicera, lateral view; 8 b, separate paraxial view of fixed digit." httpUri="https://zenodo.org/record/276758/files/figure.png" pageId="4" pageNumber="5">Figs 6, 7</figureCitation>
).
</paragraph>
<paragraph id="8BA9B421640AFFF0C9EFFC26674F905E" blockId="4.[151,1436,151,2013]" pageId="4" pageNumber="5">
<emphasis id="B9626833640AFFF0C9EFFC26638B96CA" box="[199,252,981,1004]" italics="true" pageId="4" pageNumber="5">Legs</emphasis>
. FEMALE AND MALE. Legs I to IV with pretarsi bearing paired small claws with basal swelling, paradactyli and rounded pulvillae (
<figureCitation id="132DA8A4640AFFF0C8E3FC0461539129" box="[459,548,1015,1039]" captionStart="FIGURES 25 – 31" captionStartId="11.[151,264,1201,1224]" captionTargetBox="[166,1417,183,1174]" captionTargetId="figure@11.[165,1425,183,1186]" captionTargetPageId="11" captionText="FIGURES 25 – 31. Fungiseius armatus n. sp., adult female: 25, tarsus I, anterolateral view; 26, detail of apical seta on tarsus I; 27, tarsus II, posterolateral view; 28, tarsus III, dorsal view; 29, tarsus IV, dorsal view; 30, detail of ventral aspect of tarsus IV with ventral process; 31, detail of lateral aspect of tarsus IV. Abbreviation: t p, tarsal process." httpUri="https://zenodo.org/record/276762/files/figure.png" pageId="4" pageNumber="5">Figs 30</figureCitation>
, 
<figureCitation id="132DA8A4640AFFF0CB19FC0461269129" box="[561,593,1015,1039]" captionStart="FIGURES 56 – 58" captionStartId="17.[151,264,1189,1212]" captionTargetBox="[172,1418,470,1171]" captionTargetId="figure@17.[164,1424,470,1172]" captionTargetPageId="17" captionText="FIGURES 56 – 58. Fungiseius clavulisetis n. sp., adult female: 56, tarsus IV, dorsal view; 57, tarsus II, posterolateral view; 58, tarsus II, anterolateral view. Abbreviation: t p, tarsal process." httpUri="https://zenodo.org/record/276767/files/figure.png" pageId="4" pageNumber="5">57</figureCitation>
). Legs II enlarged, much stouter than other legs (
<figureCitation id="132DA8A4640AFFF0CDAAFC0467AD9129" box="[1154,1242,1015,1039]" captionStart="FIGURES 19 – 24" captionStartId="10.[151,264,1498,1521]" captionTargetBox="[162,1420,192,1472]" captionTargetId="figure@10.[162,1422,173,1479]" captionTargetPageId="10" captionText="FIGURES 19 – 24. Fungiseius armatus n. sp., adult female: 19 – 22, legs I – IV, excepting tarsi: 19, leg I excluding tarsus, laterodorsal view; 20, leg II, posterolateral view; 21, leg III, dorsal view; 22, leg IV, anterolateral view; 23, trochanter I, ventral view; 24, detail of denoted setae of tibia I." httpUri="https://zenodo.org/record/276761/files/figure.png" pageId="4" pageNumber="5">Figs 20</figureCitation>
, 
<figureCitation id="132DA8A4640AFFF0CDC0FC0466719129" box="[1256,1286,1015,1039]" captionStart="FIGURES 48 – 55" captionStartId="16.[151,264,1591,1614]" captionTargetBox="[170,1413,460,1558]" captionTargetId="figure@16.[169,1413,445,1563]" captionTargetPageId="16" captionText="FIGURES 48 – 55. Fungiseius clavulisetis n. sp., adult female, legs I – IV, excluding tarsus: 48, leg I, posterolateral view; 49, leg II, posterolateral view; 50, leg III, dorsal view; 51, leg IV, posterolateral view; 52, coxa I, ventral view; 53, coxa II, ventral view; 54, coxa III, ventral view; 55, coxa IV, ventral view." httpUri="https://zenodo.org/record/276766/files/figure.png" pageId="4" pageNumber="5">49</figureCitation>
, 
<figureCitation id="132DA8A4640AFFF0CC3BFC0466299129" box="[1299,1374,1015,1039]" captionStart="FIGURES 59 – 60" captionStartId="17.[151,264,1978,2001]" captionTargetBox="[151,1436,1292,1954]" captionTargetId="figure@17.[151,1436,1292,1957]" captionTargetPageId="17" captionText="FIGURES 59 – 60. Photomicrographs of adult females in dorsoventral habitus view: 59, Fungiseius armatus n. sp.; 60, Fungiseius clavulisetis n. sp." httpUri="https://zenodo.org/record/276768/files/figure.png" pageId="4" pageNumber="5">59, 60</figureCitation>
), and with strongly thickened, claw-like setae on dorsal and lateral surfaces of tarsus (
<figureCitation id="132DA8A4640AFFF0CD3AFBEF671F9112" box="[1042,1128,1052,1076]" captionStart="FIGURES 25 – 31" captionStartId="11.[151,264,1201,1224]" captionTargetBox="[166,1417,183,1174]" captionTargetId="figure@11.[165,1425,183,1186]" captionTargetPageId="11" captionText="FIGURES 25 – 31. Fungiseius armatus n. sp., adult female: 25, tarsus I, anterolateral view; 26, detail of apical seta on tarsus I; 27, tarsus II, posterolateral view; 28, tarsus III, dorsal view; 29, tarsus IV, dorsal view; 30, detail of ventral aspect of tarsus IV with ventral process; 31, detail of lateral aspect of tarsus IV. Abbreviation: t p, tarsal process." httpUri="https://zenodo.org/record/276762/files/figure.png" pageId="4" pageNumber="5">Figs 27</figureCitation>
, 
<figureCitation id="132DA8A4640AFFF0CD5CFBEF67E39112" box="[1140,1172,1052,1076]" captionStart="FIGURES 56 – 58" captionStartId="17.[151,264,1189,1212]" captionTargetBox="[172,1418,470,1171]" captionTargetId="figure@17.[164,1424,470,1172]" captionTargetPageId="17" captionText="FIGURES 56 – 58. Fungiseius clavulisetis n. sp., adult female: 56, tarsus IV, dorsal view; 57, tarsus II, posterolateral view; 58, tarsus II, anterolateral view. Abbreviation: t p, tarsal process." httpUri="https://zenodo.org/record/276767/files/figure.png" pageId="4" pageNumber="5">57</figureCitation>
). Legs II to IV with tarsus (excluding pretarsus) at most two times (1.3–1.5 for II, 1.7– 2.1 for III–IV) as long as tibia. Distal margins of coxae I–IV without prominent serrations or spur-like processes. Tarsus I with sensilla 
<emphasis id="B9626833640AFFF0CD73FB966711915A" box="[1115,1126,1125,1148]" italics="true" pageId="4" pageNumber="5">s</emphasis>
inconspicuously lanceolate (
<figureCitation id="132DA8A4640AFFF0C9B7FB74638491B9" box="[159,243,1159,1183]" captionStart="FIGURES 25 – 31" captionStartId="11.[151,264,1201,1224]" captionTargetBox="[166,1417,183,1174]" captionTargetId="figure@11.[165,1425,183,1186]" captionTargetPageId="11" captionText="FIGURES 25 – 31. Fungiseius armatus n. sp., adult female: 25, tarsus I, anterolateral view; 26, detail of apical seta on tarsus I; 27, tarsus II, posterolateral view; 28, tarsus III, dorsal view; 29, tarsus IV, dorsal view; 30, detail of ventral aspect of tarsus IV with ventral process; 31, detail of lateral aspect of tarsus IV. Abbreviation: t p, tarsal process." httpUri="https://zenodo.org/record/276762/files/figure.png" pageId="4" pageNumber="5">Fig. 26</figureCitation>
), and without markedly elongated setae apically; one solid, dorsobasal seta long, erect (
<figureCitation id="132DA8A4640AFFF0CDFBFB74665091B9" box="[1235,1319,1159,1183]" captionStart="FIGURES 25 – 31" captionStartId="11.[151,264,1201,1224]" captionTargetBox="[166,1417,183,1174]" captionTargetId="figure@11.[165,1425,183,1186]" captionTargetPageId="11" captionText="FIGURES 25 – 31. Fungiseius armatus n. sp., adult female: 25, tarsus I, anterolateral view; 26, detail of apical seta on tarsus I; 27, tarsus II, posterolateral view; 28, tarsus III, dorsal view; 29, tarsus IV, dorsal view; 30, detail of ventral aspect of tarsus IV with ventral process; 31, detail of lateral aspect of tarsus IV. Abbreviation: t p, tarsal process." httpUri="https://zenodo.org/record/276762/files/figure.png" pageId="4" pageNumber="5">Fig. 25</figureCitation>
). Tarsi II– IV with apical setal processes 
<emphasis id="B9626833640AFFF0C8C1FB5F616A91E2" box="[489,541,1196,1220]" italics="true" pageId="4" pageNumber="5">ad -1</emphasis>
, 
<emphasis id="B9626833640AFFF0CB02FB5F612991E2" box="[554,606,1196,1220]" italics="true" pageId="4" pageNumber="5">pd -1</emphasis>
long, as long as pretarsi (to the base of claws), and with acutely triangular apical process ventrally. Complement of setae on segments of legs I-II-III-IV normal for Blattisociinae as presented by 
<bibRefCitation id="EF87C9D0640AFFF0C992FB076293902A" author="Lindquist" box="[186,484,1268,1292]" pageId="4" pageNumber="24" refString="Lindquist, E. E. &amp; Evans, G. O. (1965) Taxonomic concepts in the Ascidae, with a modified setal nomenclature for the idiosoma of the Gamasina (Acarina: Mesostigmata). Memoirs of the Entomological Society of Canada, 47, 1 - 64." type="journal article" year="1965">Lindquist &amp; Evans (1965)</bibRefCitation>
: coxae, 2-2-2-1; trochanters, (4-6)-5-5-5; femora, 12 (2 2/1 3/2 2) - 11 (1 2/1 3/2 2) - 6 (1 2/1 1/0 1) - 6 (1 2/1 1/0 1); genua, (2 3/1 or 2 3/1 2) - (2 3/1 2/0 2) - (2 2/1 2/0 1) - (2 2/1 2 or 3/0 1); tibiae, (2 3/1 3/1 or 2 2) - (2 2/1 2/1 2) - (2 1/1 2/1 1) - (2 1/1 3/1 2); genua II and III each lacking 
<emphasis id="B9626833640AFFF0CDA8FACE67C59072" box="[1152,1202,1340,1364]" italics="true" pageId="4" pageNumber="5">pv -1</emphasis>
(
<figureCitation id="132DA8A4640AFFF0CDE9FACF66499072" box="[1217,1342,1340,1364]" captionStart="FIGURES 19 – 24" captionStartId="10.[151,264,1498,1521]" captionTargetBox="[162,1420,192,1472]" captionTargetId="figure@10.[162,1422,173,1479]" captionTargetPageId="10" captionText="FIGURES 19 – 24. Fungiseius armatus n. sp., adult female: 19 – 22, legs I – IV, excepting tarsi: 19, leg I excluding tarsus, laterodorsal view; 20, leg II, posterolateral view; 21, leg III, dorsal view; 22, leg IV, anterolateral view; 23, trochanter I, ventral view; 24, detail of denoted setae of tibia I." httpUri="https://zenodo.org/record/276761/files/figure.png" pageId="4" pageNumber="5">Figs 19-22</figureCitation>
, 
<figureCitation id="132DA8A4640AFFF0CC62FACF66F89072" box="[1354,1423,1340,1364]" captionStart="FIGURES 48 – 55" captionStartId="16.[151,264,1591,1614]" captionTargetBox="[170,1413,460,1558]" captionTargetId="figure@16.[169,1413,445,1563]" captionTargetPageId="16" captionText="FIGURES 48 – 55. Fungiseius clavulisetis n. sp., adult female, legs I – IV, excluding tarsus: 48, leg I, posterolateral view; 49, leg II, posterolateral view; 50, leg III, dorsal view; 51, leg IV, posterolateral view; 52, coxa I, ventral view; 53, coxa II, ventral view; 54, coxa III, ventral view; 55, coxa IV, ventral view." httpUri="https://zenodo.org/record/276766/files/figure.png" pageId="4" pageNumber="5">48-51</figureCitation>
). Leg setae collectively smooth. Legs of male without dimorphically modified setae.
</paragraph>
</subSubSection>
<subSubSection id="C30CE7AA640AFFF0C9EFFA7767D090C2" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BA9B421640AFFF0C9EFFA7767D090C2" blockId="4.[151,1436,151,2013]" pageId="4" pageNumber="5">
<emphasis id="B9626833640AFFF0C9EFFA77623990BA" bold="true" box="[199,334,1412,1436]" pageId="4" pageNumber="5">Etymology.</emphasis>
The name of the genus is a Latinized combination of the term 
<emphasis id="B9626833640AFFF0CD3FFA77671590BA" box="[1047,1122,1412,1436]" italics="true" pageId="4" pageNumber="5">fungus</emphasis>
, meaning fungus, and 
<emphasis id="B9626833640AFFF0CC4CFA7666EB90BA" box="[1380,1436,1413,1436]" italics="true" pageId="4" pageNumber="5">seius</emphasis>
or 
<emphasis id="B9626833640AFFF0C99CFA5A639B90E6" box="[180,236,1449,1472]" italics="true" pageId="4" pageNumber="5">sejus</emphasis>
, a Roman surname commonly used by authors to form names for genera of mesostigmatic mites. The name is masculine in gender, and indicates the habitat from which these mites have been collected.
</paragraph>
</subSubSection>
<subSubSection id="C30CE7AA640AFFF0C9EFFA1C66179376" pageId="4" pageNumber="5" type="distribution">
<paragraph id="8BA9B421640AFFF0C9EFFA1C66179376" blockId="4.[151,1436,151,2013]" pageId="4" pageNumber="5">
<emphasis id="B9626833640AFFF0C9EFFA1C6172932E" bold="true" box="[199,517,1519,1544]" pageId="4" pageNumber="5">Distribution and habitats.</emphasis>
This genus is based on material associated with perennial fungal growths on decaying logs in two widely separated, subtropical to tropical New World localities, one from pine-oak forest south of the Isthmus of Tehuantepec in 
<collectingCountry id="F301F4B1640AFFF0CB25F9CB61129376" box="[525,613,1592,1616]" name="Mexico" pageId="4" pageNumber="5">Mexico</collectingCountry>
, and one from a locality in coastal lowland rainforest of 
<collectingCountry id="F301F4B1640AFFF0CDF7F9CB662B9376" box="[1247,1372,1592,1616]" name="Costa Rica" pageId="4" pageNumber="5">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30CE7AA640AFFF1C9EFF9AF60D39186" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="discussion">
<paragraph id="8BA9B421640AFFF1C9EFF9AF60AA943A" blockId="4.[151,1436,151,2013]" lastBlockId="5.[151,1436,151,1184]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="B9626833640AFFF0C9EFF9AF624C9352" bold="true" box="[199,315,1628,1652]" pageId="4" pageNumber="5">Remarks.</emphasis>
The genus 
<taxonomicName id="4C16CFA2640AFFF0C894F9AE61409352" box="[444,567,1629,1652]" class="Arachnida" family="Blattisociidae" genus="Fungiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9626833640AFFF0C894F9AE61409352" box="[444,567,1629,1652]" italics="true" pageId="4" pageNumber="5">Fungiseius</emphasis>
</taxonomicName>
is based on their adults having several apomorphies, some of which appear to be unique among the known taxa of 
<taxonomicName id="4C16CFA2640AFFF0CB6BF973619593BE" box="[579,738,1664,1688]" class="Arachnida" family="Blattisociidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Blattisociidae</taxonomicName>
: (1) dorsal shield with delineated lateral rim extending from setae 
<emphasis id="B9626833640AFFF0C9FFF9576398939A" box="[215,239,1700,1724]" italics="true" pageId="4" pageNumber="5">z2</emphasis>
to 
<emphasis id="B9626833640AFFF0C83EF9566245939A" box="[278,306,1701,1724]" italics="true" pageId="4" pageNumber="5">S5</emphasis>
and bearing 
<emphasis id="B9626833640AFFF0C8EEF95762A8939A" box="[454,479,1700,1724]" italics="true" pageId="4" pageNumber="5">z2</emphasis>
or 
<emphasis id="B9626833640AFFF0CB2FF9576157939A" box="[519,544,1700,1724]" italics="true" pageId="4" pageNumber="5">s2</emphasis>
, 
<emphasis id="B9626833640AFFF0CB07F957611D939A" box="[559,618,1700,1724]" italics="true" pageId="4" pageNumber="5">r2-r5</emphasis>
, and 
<emphasis id="B9626833640AFFF0CB82F957619C939A" box="[682,747,1700,1724]" italics="true" pageId="4" pageNumber="5">S1-S5</emphasis>
; (2) apex of endopodal extension of sternal or sternitigenital shield between legs I and II with a gland pore; (3) female with deep postgenital furrow; (4) male and female with deep postanal furrow; (5) opisthogastric setae 
<emphasis id="B9626833640AFFF0CB9CF91F61979222" box="[692,736,1772,1796]" italics="true" pageId="4" pageNumber="5">JV3</emphasis>
absent; (6) hypostomatic setae 
<emphasis id="B9626833640AFFF0CD7DF91F67059222" box="[1109,1138,1772,1796]" italics="true" pageId="4" pageNumber="5">h2</emphasis>
minute; (7) legs II much stouter than other legs; (8) strongly thickened claw-like setae on tarsi of legs II to IV, especially leg II; (9) genua of legs III–IV lacking seta 
<emphasis id="B9626833640AFFF0C882F8C662AB926A" box="[426,476,1844,1868]" italics="true" pageId="4" pageNumber="5">pv -1</emphasis>
. Attributes (1), (3), (4) and (8) appear to be unique to this genus. Although attribute (1) is superficially similar to a condition found in 
<taxonomicName id="4C16CFA2640AFFF0CBE1F8AB60299256" box="[713,862,1880,1904]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9626833640AFFF0CBE1F8AB60299256" box="[713,862,1880,1904]" italics="true" pageId="4" pageNumber="5">Orthadenella</emphasis>
</taxonomicName>
, the latter genus does not belong to this subfamily, in view of its females not having an apparently phytoseioid-type spermathecal system (
<bibRefCitation id="EF87C9D0640AFFF0CD54F88F622B929E" author="Athias-Henriot" pageId="4" pageNumber="23" refString="Athias-Henriot, C. (1973) Observations sur les genres Neojordensia et Orthadenella n. g. en Europe occidentale (Gamasides, Dermanyssina, Ascidae). Acarologia, 15, 18 - 32." type="journal article" year="1973">Athias-Henriot 1973; see discussion below</bibRefCitation>
). Also, the delineated rim of attribute (1) in 
<taxonomicName id="4C16CFA2640AFFF0CA7AF85260BA929E" box="[850,973,1953,1976]" class="Arachnida" family="Blattisociidae" genus="Fungiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9626833640AFFF0CA7AF85260BA929E" box="[850,973,1953,1976]" italics="true" pageId="4" pageNumber="5">Fungiseius</emphasis>
</taxonomicName>
is unique in carrying lateral setae 
<emphasis id="B9626833640AFFF0CC7CF85366E2929E" box="[1364,1429,1952,1976]" italics="true" pageId="4" pageNumber="5">S1-S5</emphasis>
, leaving the 
<emphasis id="B9626833640AFFF0C832F836625C92FA" box="[282,299,1989,2012]" italics="true" pageId="4" pageNumber="5">R</emphasis>
- marginals on soft cuticle, while the rim found in 
<taxonomicName id="4C16CFA2640AFFF0CA77F837608392FA" box="[863,1012,1988,2012]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9626833640AFFF0CA77F837608392FA" box="[863,1012,1988,2012]" italics="true" pageId="4" pageNumber="5">Orthadenella</emphasis>
</taxonomicName>
and among genera of the ascoid family 
<taxonomicName id="4C16CFA2640BFFF1C995FF6462269589" box="[189,337,151,175]" class="Arachnida" family="Ascidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Melicharidae</taxonomicName>
differs in bearing most or all of the 
<emphasis id="B9626833640BFFF1CBDEFF6A60709596" box="[758,775,153,176]" italics="true" pageId="5" pageNumber="6">R</emphasis>
- marginal setae, with the laterals inserted mediad the rim (personal observations). Attribute (7) is shared only with 
<taxonomicName id="4C16CFA2640BFFF1CA31FF4F60E095F2" box="[793,919,188,212]" class="Arachnida" family="Ascidae" genus="Hoploseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9626833640BFFF1CA31FF4F60E095F2" box="[793,919,188,212]" italics="true" pageId="5" pageNumber="6">Hoploseius</emphasis>
</taxonomicName>
, whose members are also restricted to bracket fungi. Attributes (2) and (6) are uncertainly unique in that their presence or absence among other taxa of this and related families has not been confirmed, while (5) and (9) are homoplastic.
</paragraph>
<paragraph id="8BA9B421640BFFF1C9EDFED461F694D2" blockId="5.[151,1436,151,1184]" pageId="5" pageNumber="6">
The apomorphic attribute of a “phytoseioid-type” sperm access system in adult females, on which the superfamily Phytoseioidea is based (
<bibRefCitation id="EF87C9D0640BFFF1C8DBFEBF61A99442" author="Athias-Henriot" box="[499,734,332,356]" pageId="5" pageNumber="23" refString="Athias-Henriot, C. (1968) L'appareil d'insemination laelapoide (Acariens anactinotriches: Laelapoidea). Premieres observations. Possibilite d'emploi a des fins taxonomiques. Bulletin Scientifique de Bourgogne, 25, 229 - 274." type="journal article" year="1968">Athias-Henriot 1968</bibRefCitation>
, 
<bibRefCitation id="EF87C9D0640BFFF1CBC1FEBF60539442" author="Athias-Henriot" box="[745,804,332,356]" pageId="5" pageNumber="23" refString="Athias-Henriot, C. (1971) Nouvelles notes sur les Amblyseiini (Gamasides Podospermiques, Phytoseiidae). 1. La depilation des genuaux et tibias des pattes. Acarologia, 13, 4 - 15." type="journal article" year="1971">1971</bibRefCitation>
; 
<bibRefCitation id="EF87C9D0640BFFF1CA18FEBF67399442" author="Alberti" box="[816,1102,332,356]" pageId="5" pageNumber="23" refString="Alberti, G. &amp; Di Palma, A. (2002) Fine structure of the phytoseiid-type sperm access system (Acari, Gamasida, Phytoseiidae). In: Bernini, F., Nannelli, R. &amp; de Lillo, E. (Eds.), Acarid phylogeny and evolution. Adaptations in mites and ticks. Proceedings IV EURAAC Symposium, Siena. Kluwer Academic Publishers, Dordrecht, pp. 241 - 252." type="book chapter" year="2002">
Alberti &amp; 
<collectingCountry id="F301F4B1640BFFF1CA8BFEBF60C89442" box="[931,959,332,356]" name="Iceland" pageId="5" pageNumber="6">Di</collectingCountry>
Palma 2002
</bibRefCitation>
; 
<bibRefCitation id="EF87C9D0640BFFF1CD72FEBF66319442" author="Lindquist" box="[1114,1350,332,356]" pageId="5" pageNumber="24" refString="Lindquist, E. E., Krantz, G. W. &amp; Walter, D. E. (2009) Order Mesostigmata. In: Krantz, G. W. &amp; Walter, D. E. (Eds.), A Manual of Acarology, Third Edition. Texas Tech University Press, Lubbock, pp. 124 - 232." type="book chapter" year="2009">
Lindquist 
<emphasis id="B9626833640BFFF1CDE4FEBE678B9442" box="[1228,1276,332,356]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
. 2009
</bibRefCitation>
), is difficult to observe in both species of 
<taxonomicName id="4C16CFA2640BFFF1CB18FE8261DC94AE" box="[560,683,369,392]" class="Arachnida" family="Blattisociidae" genus="Fungiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9626833640BFFF1CB18FE8261DC94AE" box="[560,683,369,392]" italics="true" pageId="5" pageNumber="6">Fungiseius</emphasis>
</taxonomicName>
, due to the reduced size and relatively weak sclerotization of the calyx, such that discernment of a minor duct emanating from the embolus is problematical. However, other, more readily observable attributes detailed by 
<bibRefCitation id="EF87C9D0640BFFF1CB73FE4460F894E9" author="Lindquist" box="[603,911,439,463]" pageId="5" pageNumber="24" refString="Lindquist E. E. &amp; Moraza, M. L. (2010) Revised diagnosis of the family Blattisociidae (Acari: Mesostigmata: Phytoseioidea), with a key to its genera and description of a new fungus-inhabiting genus from Costa Rica. Zootaxa, 2479, 1 - 21." type="journal article" year="2010">Lindquist &amp; Moraza (2010)</bibRefCitation>
as diagnostic for the family 
<taxonomicName id="4C16CFA2640BFFF1CDE3FE44661294E9" box="[1227,1381,439,463]" class="Arachnida" family="Blattisociidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Blattisociidae</taxonomicName>
support placement of 
<taxonomicName id="4C16CFA2640BFFF1C84BFE2E62A994D2" box="[355,478,477,500]" class="Arachnida" family="Blattisociidae" genus="Fungiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9626833640BFFF1C84BFE2E62A994D2" box="[355,478,477,500]" italics="true" pageId="5" pageNumber="6">Fungiseius</emphasis>
</taxonomicName>
in this family.
</paragraph>
<paragraph id="8BA9B421640BFFF1C9EDFDF3616D9136" blockId="5.[151,1436,151,1184]" pageId="5" pageNumber="6">
Among the genera of Blattisociinae, adults of 
<taxonomicName id="4C16CFA2640BFFF1CBF8FDF2603C973E" box="[720,843,513,536]" class="Arachnida" family="Blattisociidae" genus="Fungiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9626833640BFFF1CBF8FDF2603C973E" box="[720,843,513,536]" italics="true" pageId="5" pageNumber="6">Fungiseius</emphasis>
</taxonomicName>
are strikingly similar to those of 
<taxonomicName id="4C16CFA2640BFFF1CDEFFDF36632973E" box="[1223,1349,512,536]" class="Arachnida" family="Ascidae" genus="Hoploseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9626833640BFFF1CDEFFDF36632973E" box="[1223,1349,512,536]" italics="true" pageId="5" pageNumber="6">Hoploseius</emphasis>
</taxonomicName>
, whose members also dwell in bracket fungi, in having greatly thickened legs II. However, this appears to be an interesting case of convergence. Legs II of 
<taxonomicName id="4C16CFA2640BFFF1CB25FDBB61FC9746" box="[525,651,584,608]" class="Arachnida" family="Ascidae" genus="Hoploseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9626833640BFFF1CB25FDBB61FC9746" box="[525,651,584,608]" italics="true" pageId="5" pageNumber="6">Hoploseius</emphasis>
</taxonomicName>
bear strongly thickened, spine-like setae on the ventral faces of the femur, genu, tibia and tarsus; these setae are apposed to one another, apparently for grasping structures, most probably those of the drosophilid flies used for phoresy (
<bibRefCitation id="EF87C9D0640BFFF1CBC9FD7C60E69781" author="Lindquist" box="[737,913,655,679]" pageId="5" pageNumber="24" refString="Lindquist, E. E. (1963) A taxonomic review of the genus Hoploseius Berlese (Acarina: Blattisociidae). The Canadian Entomologist, 95 (11), 1175 - 1185." type="journal article" year="1963">Lindquist 1963</bibRefCitation>
). 
<collectingCountry id="F301F4B1640BFFF1CA8DFD6360B3978E" box="[933,964,656,680]" name="American Samoa" pageId="5" pageNumber="6">As</collectingCountry>
adult males are similarly armed, but have not been found to be phoretic, the grasping function may also be used by both sexes in copulation. In contrast, legs II of both sexes in 
<taxonomicName id="4C16CFA2640BFFF1C85AFD2A629A97D6" box="[370,493,729,752]" class="Arachnida" family="Blattisociidae" genus="Fungiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9626833640BFFF1C85AFD2A629A97D6" box="[370,493,729,752]" italics="true" pageId="5" pageNumber="6">Fungiseius</emphasis>
</taxonomicName>
have slender setae on the ventral faces of the femur and genu; strongly spine-like setae are restricted to the tarsus, primarily on the dorsal and lateral faces. The latter setae are claw-like, not apposable, and probably are used for scraping or burrowing. Other than a within-genus tendency towards hypotrichy among some of the leg segments, which is subject to considerable homoplasy, members of these two genera do not share other apomorphic attributes that would indicate a sister-group relationship. Even the loss of seta 
<emphasis id="B9626833640BFFF1CC6CFC9A660296A6" box="[1348,1397,872,896]" italics="true" pageId="5" pageNumber="6">pv -1</emphasis>
on genua III and IV in 
<taxonomicName id="4C16CFA2640BFFF1C85DFC7E62879682" box="[373,496,909,932]" class="Arachnida" family="Blattisociidae" genus="Fungiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9626833640BFFF1C85DFC7E62879682" box="[373,496,909,932]" italics="true" pageId="5" pageNumber="6">Fungiseius</emphasis>
</taxonomicName>
differs from the losses of setae 
<emphasis id="B9626833640BFFF1CA7CFC7F60F59682" box="[852,898,908,932]" italics="true" pageId="5" pageNumber="6">al -2</emphasis>
and 
<emphasis id="B9626833640BFFF1CA90FC7E609D9682" box="[952,1002,908,932]" italics="true" pageId="5" pageNumber="6">av -1</emphasis>
on genu III and of 
<emphasis id="B9626833640BFFF1CDEBFC7F67819682" box="[1219,1270,908,932]" italics="true" pageId="5" pageNumber="6">pd -3</emphasis>
, 
<emphasis id="B9626833640BFFF1CC2CFC7F66469682" box="[1284,1329,908,932]" italics="true" pageId="5" pageNumber="6">al -2</emphasis>
, and 
<emphasis id="B9626833640BFFF1CC47FC7F66EB9682" box="[1391,1436,908,932]" italics="true" pageId="5" pageNumber="6">pl -1</emphasis>
on genu IV among some species in 
<taxonomicName id="4C16CFA2640BFFF1CB04FC4360AA96E1" authority="Lindquist &amp; Evans 1965" authorityName="Lindquist &amp; Evans" authorityYear="1965" box="[556,989,943,968]" class="Arachnida" family="Ascidae" genus="Hoploseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9626833640BFFF1CB04FC4361DD96EE" box="[556,682,944,968]" italics="true" pageId="5" pageNumber="6">Hoploseius</emphasis>
(
<bibRefCitation id="EF87C9D0640BFFF1CB91FC5C60A396E1" author="Lindquist" box="[697,980,943,968]" pageId="5" pageNumber="24" refString="Lindquist, E. E. &amp; Evans, G. O. (1965) Taxonomic concepts in the Ascidae, with a modified setal nomenclature for the idiosoma of the Gamasina (Acarina: Mesostigmata). Memoirs of the Entomological Society of Canada, 47, 1 - 64." type="journal article" year="1965">Lindquist &amp; Evans 1965</bibRefCitation>
)
</taxonomicName>
. The affinity of 
<taxonomicName id="4C16CFA2640BFFF1CDBDFC43666496EE" box="[1173,1299,944,968]" class="Arachnida" family="Ascidae" genus="Hoploseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9626833640BFFF1CDBDFC43666496EE" box="[1173,1299,944,968]" italics="true" pageId="5" pageNumber="6">Hoploseius</emphasis>
</taxonomicName>
with 
<taxonomicName id="4C16CFA2640BFFF1CC7CFC4263B896CA" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9626833640BFFF1CC7CFC4263B896CA" italics="true" pageId="5" pageNumber="6">Lasioseius</emphasis>
</taxonomicName>
, based on the tricarinate form of at least some of the dorsal shield setae and the strongly formed ventrianal shield, is distinct from 
<taxonomicName id="4C16CFA2640BFFF1C8B0FC0A61649136" box="[408,531,1017,1040]" class="Arachnida" family="Blattisociidae" genus="Fungiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9626833640BFFF1C8B0FC0A61649136" box="[408,531,1017,1040]" italics="true" pageId="5" pageNumber="6">Fungiseius</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA9B421640BFFF1C9EDFBEF60D39186" blockId="5.[151,1436,151,1184]" pageId="5" pageNumber="6">
Among other genera of Blattisociinae, adult females of 
<taxonomicName id="4C16CFA2640BFFF1CA66FBEE60BE9112" box="[846,969,1053,1076]" class="Arachnida" family="Blattisociidae" genus="Fungiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9626833640BFFF1CA66FBEE60BE9112" box="[846,969,1053,1076]" italics="true" pageId="5" pageNumber="6">Fungiseius</emphasis>
</taxonomicName>
are similar to those of 
<taxonomicName id="4C16CFA2640BFFF1CDF6FBEF661E9112" box="[1246,1385,1052,1076]" class="Arachnida" family="Blattisociidae" genus="Aceodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9626833640BFFF1CDF6FBEF661E9112" box="[1246,1385,1052,1076]" italics="true" pageId="5" pageNumber="6">Aceodromus</emphasis>
</taxonomicName>
and 
<emphasis id="B9626833640BFFF1C9BFFBB36260917E" box="[151,279,1088,1112]" italics="true" pageId="5" pageNumber="6">Opilioseius</emphasis>
in having a reduced ventrianal shield or an anal shield, and in the spermathecal apparatus having a weakly sclerotized calyx. These attributes are subject to considerable homoplasy among other genera of this subfamily, especially the species-rich 
<taxonomicName id="4C16CFA2640BFFF1CB31FB7A61F89186" box="[537,655,1161,1184]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9626833640BFFF1CB31FB7A61F89186" box="[537,655,1161,1184]" italics="true" pageId="5" pageNumber="6">Lasioseius</emphasis>
</taxonomicName>
(personal observations).
</paragraph>
</subSubSection>
</treatment>
</document>