456 lines
54 KiB
XML
456 lines
54 KiB
XML
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<mods:title id="8959EC97FE716981D76A1A97EBA43AE7">Revision of the enigmatic crocodyliform Elosuchus felixi de Lapparent de Broin, 2002 from the Lower-Upper Cretaceous boundary of Niger: potential evidence for an early origin of the clade Dyrosauridae</mods:title>
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<mods:namePart id="4FC7051C5FEA3868E6EBD2F8BFBA2C72">Young, Mark T.</mods:namePart>
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<mods:namePart id="877192A702718412B2F38BE72F41885B">Hastings, Alexander K.</mods:namePart>
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<mods:namePart id="205DC1315D8FF0E10280293984142636">Allain, Ronan</mods:namePart>
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<mods:namePart id="99E547249E30F8DB8569219794C8FA0D">Smith, Thomas J.</mods:namePart>
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<treatment id="03AEC940FF8AF506FBC6D2BDFA65FD2F" ID-DOI="http://doi.org/10.5281/zenodo.5710905" ID-GBIF-Taxon="190222482" ID-Zenodo-Dep="5710905" LSID="urn:lsid:plazi:treatment:03AEC940FF8AF506FBC6D2BDFA65FD2F" httpUri="http://treatment.plazi.org/id/03AEC940FF8AF506FBC6D2BDFA65FD2F" lastPageId="6" lastPageNumber="383" pageId="4" pageNumber="381">
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<subSubSection id="C31D2BDDFF8AF504FBC6D2BDFB53FB1E" box="[1028,1207,1185,1211]" pageId="4" pageNumber="381" type="nomenclature">
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<paragraph id="8BB87856FF8AF504FBC6D2BDFB53FB1E" blockId="4.[1028,1207,1185,1211]" box="[1028,1207,1185,1211]" pageId="4" pageNumber="381">
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<heading id="D0F0CF3AFF8AF504FBC6D2BDFB53FB1E" box="[1028,1207,1185,1211]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="381" reason="2">
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<taxonomicName id="4C0703D5FF8AF504FBC6D2BDFB53FB1E" authority="SP." authorityName="SP." box="[1028,1207,1185,1211]" class="Reptilia" family="Pholidosauridae" genus="Elosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" pageNumber="381" phylum="Chordata" rank="genus">
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<emphasis id="B973A444FF8AF504FBC6D2BDFB68FB1D" box="[1028,1164,1189,1210]" italics="true" pageId="4" pageNumber="381">ELOSUCHUS</emphasis>
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SP.
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D2BDDFF8AF504FCE8D2CAFC72FAE6" pageId="4" pageNumber="381" type="reference_group">
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<paragraph id="8BB87856FF8AF504FCE8D2CAFC74FAA3" blockId="4.[810,1425,1234,1286]" pageId="4" pageNumber="381">
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<treatmentCitation id="0AA65E47FF8AF504FCE8D2CAFA75FB42" author="de Lapparent de Broin F" box="[810,1425,1234,1256]" page="283" pageId="4" pageNumber="381" year="2002">
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<taxonomicName id="4C0703D5FF8AF504FCE8D2CAFA75FB42" authority="de Lapparent de Broin, 2002: 283" authorityName="de Lapparent de Broin" authorityPageNumber="283" authorityYear="2002" box="[810,1425,1234,1256]" class="Reptilia" family="Pholidosauridae" genus="Elosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" pageNumber="381" phylum="Chordata" rank="species" species="felixi">
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<emphasis id="B973A444FF8AF504FCE8D2CAFC00FB42" box="[810,996,1234,1255]" italics="true" pageId="4" pageNumber="381">Elosuchus felixi</emphasis>
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<bibRefCitation id="EF9605A7FF8AF504FC30D2CAFA75FB42" author="de Lapparent de Broin F" box="[1010,1425,1234,1256]" pageId="4" pageNumber="381" pagination="275 - 285" refId="ref17195" refString="de Lapparent de Broin F. 2002. Elosuchus, a new genus of crocodile from the Lower Cretaceous of the North of Africa. Comptes Rendus Palevol 1: 275 - 285." type="journal article" year="2002">de Lapparent de Broin, 2002: 283</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(
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<emphasis id="B973A444FF8AF504FCF0D2E9FC64FAA3" box="[818,896,1265,1286]" italics="true" pageId="4" pageNumber="381">partim</emphasis>
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).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BB87856FF8AF504FCE8D336FC72FAE6" blockId="4.[810,1425,1326,1409]" box="[810,918,1326,1347]" pageId="4" pageNumber="381">
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<emphasis id="B973A444FF8AF504FCE8D336FC72FAE6" box="[810,918,1326,1347]" italics="true" pageId="4" pageNumber="381">Specimen</emphasis>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C31D2BDDFF8AF506FCE8D355FEFBF895" lastPageId="6" lastPageNumber="383" pageId="4" pageNumber="381" type="description">
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<paragraph id="8BB87856FF8AF504FCE8D355FCB2FA24" blockId="4.[810,1425,1326,1409]" pageId="4" pageNumber="381">MNHN.F INA 30, an incomplete and broken premaxilla.</paragraph>
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<paragraph id="8BB87856FF8AF504FCE8D3B0FC61FA18" blockId="4.[810,1348,1448,1501]" box="[810,901,1448,1469]" pageId="4" pageNumber="381">
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<emphasis id="B973A444FF8AF504FCE8D3B0FC61FA18" box="[810,901,1448,1469]" italics="true" pageId="4" pageNumber="381">Locality</emphasis>
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</paragraph>
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<paragraph id="8BB87856FF8AF504FCE8D3DFFAA0FA78" blockId="4.[810,1348,1448,1501]" box="[810,1348,1479,1501]" pageId="4" pageNumber="381">
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West of In Abangharit,
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<collectingRegion id="49C3B6B4FF8AF504FBF9D3DFFB0BFA78" box="[1083,1263,1479,1501]" country="Niger" name="Agadez" pageId="4" pageNumber="381">Agadez District</collectingRegion>
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,
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<collectingCountry id="F31038C6FF8AF504FB3ED3DFFAA4FA78" box="[1276,1344,1479,1501]" name="Niger" pageId="4" pageNumber="381">Niger</collectingCountry>
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.
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</paragraph>
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<paragraph id="8BB87856FF8AF504FCE8D01CFC03F9BC" blockId="4.[810,1424,1539,1653]" box="[810,999,1539,1561]" pageId="4" pageNumber="381">
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<emphasis id="B973A444FF8AF504FCE8D01CFC03F9BC" box="[810,999,1539,1561]" italics="true" pageId="4" pageNumber="381">Horizon and age</emphasis>
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</paragraph>
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<paragraph id="8BB87856FF8AF504FCE8D03AFC5CF9D0" blockId="4.[810,1424,1539,1653]" pageId="4" pageNumber="381">Echkar Formation, Tegama Series. Late Albian (Lower Cretaceous) to early Cenomanian (Upper Cretaceous).</paragraph>
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<paragraph id="8BB87856FF8AF504FCE8D085FC48F917" blockId="4.[810,1425,1693,1897]" box="[810,940,1693,1714]" pageId="4" pageNumber="381">
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<emphasis id="B973A444FF8AF504FCE8D085FC48F917" box="[810,940,1693,1714]" italics="true" pageId="4" pageNumber="381">Description</emphasis>
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</paragraph>
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<paragraph id="8BB87856FF8AF505FCE8D0A3FE51FDEE" blockId="4.[810,1425,1693,1897]" lastBlockId="5.[166,781,199,1260]" lastPageId="5" lastPageNumber="382" pageId="4" pageNumber="381">
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The lateral surfaces of MNHN.F INA 30 are convex, whereas the anterior margin is more vertically oriented. There is a raised rim around the external nares, which is most prominent at the anterior margin of the narial opening (
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<figureCitation id="133C64D3FF8AF504FB98D12DFB7BF8EE" box="[1114,1183,1845,1867]" captionStart="Figure 3" captionStartId="5.[166,245,1850,1869]" captionTargetBox="[174,1436,1307,1816]" captionTargetId="figure-648@5.[174,1438,1307,1816]" captionTargetPageId="5" captionText="Figure 3. Premaxilla MNHN.F INA 30, referred to Elosuchus sp.: A, photograph in dorsal view; B, photograph in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5710913" httpUri="https://zenodo.org/record/5710913/files/figure.png" pageId="4" pageNumber="381">Fig. 3</figureCitation>
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). Along this anterior margin there is no dorsal projection, although this could be because of the presence of the raised narial rim. The loss of this projection is considered to be a defining characteristic of
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<taxonomicName id="4C0703D5FF8BF505FE28D71CFECBFE9D" authority="(Fortier et al., 2011)" baseAuthorityName="Fortier" baseAuthorityYear="2011" class="Reptilia" family="Pholidosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="382" phylum="Chordata" rank="family">
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Pholidosauridae (
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<bibRefCitation id="EF9605A7FF8BF505FD7ED71CFEC3FE9D" author="Fortier D & Perea D & Schultz C" pageId="5" pageNumber="382" pagination="272" refId="ref16232" refString="Fortier D, Perea D, Schultz C. 2011. Redescription and phylogenetic relationships of Meridiosaurus vallisparadisi, a pholidosaurid from the Late Jurassic of Uruguay. Zoological Journal of the Linnean Society 163: S 257 - S 272." type="journal article" year="2011">
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Fortier
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<emphasis id="B973A444FF8BF505FF64D73BFF04FE92" box="[166,224,290,312]" italics="true" pageId="5" pageNumber="382">et al.</emphasis>
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, 2011
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</bibRefCitation>
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)
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</taxonomicName>
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, with the premaxilla anterior to the external nares being convex in most pholidosaurids (e.g.
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<bibRefCitation id="EF9605A7FF8BF505FF64D747FEC8FED0" author="Mook CC" box="[166,300,351,373]" pageId="5" pageNumber="382" pagination="1 - 11" refId="ref17689" refString="Mook CC. 1934. A new species of Teleorhinus from the Benton Shales. American Museum Novitates 702: 1 - 11." type="journal article" year="1934">Mook, 1934</bibRefCitation>
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;
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<bibRefCitation id="EF9605A7FF8BF505FEFAD778FE0DFED0" author="Wu X-C & Russell AP & Cumbaa SL" box="[312,489,351,373]" pageId="5" pageNumber="382" pagination="492 - 514" refId="ref18460" refString="Wu X-C, Russell AP, Cumbaa SL. 2001. Terminonaris (Archosauria: Crocodyliformes): new material from Saskatchewan, Canada, and comments on its phylogenetic relationships. Journal of Vertebrate Paleontology 21: 492 - 514." type="journal article" year="2001">
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Wu
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<emphasis id="B973A444FF8BF505FEA5D778FE45FED1" box="[359,417,351,373]" italics="true" pageId="5" pageNumber="382">et al.</emphasis>
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||
, 2001
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF8BF505FE37D778FD4BFED0" author="Hua S & Buffetaut E & Legall C & Rogron P" box="[501,687,351,373]" pageId="5" pageNumber="382" pagination="503 - 513" refId="ref16531" refString="Hua S, Buffetaut E, Legall C, Rogron P. 2007. Oceanosuchus boecensis n. gen, n. sp., a marine pholidosaurid (Crocodylia, Mesosuchia) from the Lower Cenomanian of Normandy (western France). Bulletin de la Societe Geologique de France 178: 503 - 513." type="journal article" year="2007">
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Hua
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<emphasis id="B973A444FF8BF505FDECD778FD8CFED1" box="[558,616,351,373]" italics="true" pageId="5" pageNumber="382">et al.</emphasis>
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||
, 2007
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||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF8BF505FD79D778FEC9FE36" author="Lepage Y & Buffetaut E & Hua S & Martin JE & Tabouelle J" pageId="5" pageNumber="382" pagination="5 - 152" refId="ref17292" refString="Lepage Y, Buffetaut E, Hua S, Martin JE, Tabouelle J. 2008. Catalogue descriptif, anatomique, geologique et historique des fossiles presentes<65>a l'exposition " Les Crocodiliens fossiles de Normandie " (6 novembre - 14 decembre 2008). Bulletin de la Societe Geologique de Normandie et des Amis du Museum du Havre 95: 5 - 152." type="journal article" year="2008">
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Lepage
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<emphasis id="B973A444FF8BF505FF64D767FF04FE36" box="[166,224,382,404]" italics="true" pageId="5" pageNumber="382">et al.</emphasis>
|
||
, 2008
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF8BF505FEFCD766FDC5FE31" author="Fortier D & Perea D & Schultz C" box="[318,545,382,404]" pageId="5" pageNumber="382" pagination="272" refId="ref16232" refString="Fortier D, Perea D, Schultz C. 2011. Redescription and phylogenetic relationships of Meridiosaurus vallisparadisi, a pholidosaurid from the Late Jurassic of Uruguay. Zoological Journal of the Linnean Society 163: S 257 - S 272." type="journal article" year="2011">
|
||
Fortier
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||
<emphasis id="B973A444FF8BF505FE58D767FE30FE36" box="[410,468,382,404]" italics="true" pageId="5" pageNumber="382">et al.</emphasis>
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||
, 2011
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||
</bibRefCitation>
|
||
); however, a dorsal projection is present on the most well-preserved specimen of
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<taxonomicName id="4C0703D5FF8BF505FECBD7A4FE4FFE75" baseAuthorityName="Lavocat" baseAuthorityYear="1955" box="[265,427,443,465]" class="Reptilia" family="Pholidosauridae" genus="Elosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="382" phylum="Chordata" rank="species" species="cherifiensis">
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<emphasis id="B973A444FF8BF505FECBD7A4FE4FFE75" box="[265,427,443,465]" italics="true" pageId="5" pageNumber="382">E. cherifiensis</emphasis>
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</taxonomicName>
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(MNHN.F SAM 129) and in well-preserved isolated premaxillae (MNHN.F MRS 334). Therefore, this characteristic may define a subset of pholidosaurids, rather than an unambiguous apomorphy of the clade.
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</paragraph>
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<paragraph id="8BB87856FF8BF505FF7DD44CFE17FBB3" blockId="5.[166,781,199,1260]" pageId="5" pageNumber="382">
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Ornamentation on the dorsal surface of MNHN.F INA 30 is composed of numerous irregularly distributed pits that vary in size and shape. On the lateral and anterior surfaces of the premaxilla, the ornamentation becomes less pitted with a more irregular array of grooves and raised ridges. The posterior margins of the premaxillae are not preserved, and both posterior processes are missing. Thus we cannot determine the nature of the premaxilla
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<emphasis id="B973A444FF8BF505FE04D55FFE36FCFB" box="[454,466,839,862]" italics="true" pageId="5" pageNumber="382">–</emphasis>
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maxilla contact or the premaxillary constriction (i.e. the narrowing of the premaxilla immediately posterior to the point of maximum width). Moreover, because of the fragmentary nature of the premaxillae, and the post-mortem surface damage it experienced, the medial (i.e. non-alveolar) palatal surfaces are poorly preserved.
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</paragraph>
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<paragraph id="8BB87856FF8BF505FF7DD206FB5AFD02" blockId="5.[166,781,199,1260]" lastBlockId="5.[829,1444,199,1260]" pageId="5" pageNumber="382">
|
||
Fortunately, the dorsal and lateral surfaces are better preserved. There is an undivided, dorsallyfacing external naris (
|
||
<figureCitation id="133C64D3FF8BF505FE67D243FE0DFBD7" box="[421,489,1115,1138]" captionStart="Figure 3" captionStartId="5.[166,245,1850,1869]" captionTargetBox="[174,1436,1307,1816]" captionTargetId="figure-648@5.[174,1438,1307,1816]" captionTargetPageId="5" captionText="Figure 3. Premaxilla MNHN.F INA 30, referred to Elosuchus sp.: A, photograph in dorsal view; B, photograph in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5710913" httpUri="https://zenodo.org/record/5710913/files/figure.png" pageId="5" pageNumber="382">Fig. 3</figureCitation>
|
||
). As a result of imperfect preservation, it is hard to discern how extensive the fossa within the narial opening would have been. Moreover, because of the fragmentary nature of the premaxilla we cannot ascertain the former
|
||
<emphasis id="B973A444FF8BF505FD7CD2CEFCE9FB4E" box="[702,781,1238,1259]" italics="true" pageId="5" pageNumber="382">in vivo</emphasis>
|
||
shape, although it is likely to be very similar to that of
|
||
<taxonomicName id="4C0703D5FF8BF505FC99D6FEFA8FFF59" authority="(de Lapparent de Broin, 2002)" baseAuthorityName="de Lapparent de Broin" baseAuthorityYear="2002" box="[859,1387,229,252]" class="Reptilia" family="Pholidosauridae" genus="Elosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="382" phylum="Chordata" rank="species" species="cherifiensis">
|
||
<emphasis id="B973A444FF8BF505FC99D6FEFC19FF5F" box="[859,1021,229,251]" italics="true" pageId="5" pageNumber="382">E. cherifiensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9605A7FF8BF505FBCCD6FDFA87FF59" author="de Lapparent de Broin F" box="[1038,1379,229,252]" pageId="5" pageNumber="382" pagination="275 - 285" refId="ref17195" refString="de Lapparent de Broin F. 2002. Elosuchus, a new genus of crocodile from the Lower Cretaceous of the North of Africa. Comptes Rendus Palevol 1: 275 - 285." type="journal article" year="2002">de Lapparent de Broin, 2002</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The posteromedial section of the narial margin is not preserved, and the preserved premaxillary fragments do not fully contact one another along the midline posterior to the external nares. Thus, it is likely that MNHN.F INA 30 shared the same condition as
|
||
<taxonomicName id="4C0703D5FF8BF505FCFFD786FC3BFE17" baseAuthorityName="de Lapparent de Broin" baseAuthorityYear="2002" box="[829,991,413,435]" class="Reptilia" family="Pholidosauridae" genus="Elosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="382" phylum="Chordata" rank="species" species="cherifiensis">
|
||
<emphasis id="B973A444FF8BF505FCFFD786FC3BFE17" box="[829,991,413,435]" italics="true" pageId="5" pageNumber="382">E. cherifiensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9605A7FF8BF505FBC0D785FA44FE17" author="de Lapparent de Broin F" box="[1026,1440,413,435]" pageId="5" pageNumber="382" pagination="275 - 285" refId="ref17195" refString="de Lapparent de Broin F. 2002. Elosuchus, a new genus of crocodile from the Lower Cretaceous of the North of Africa. Comptes Rendus Palevol 1: 275 - 285." type="journal article" year="2002">de Lapparent de Broin, 2002</bibRefCitation>
|
||
; MNHN.F SAM 129), that the anterior processes of the nasals prevented the premaxillae from suturing along the rostral midline, and that the nasals contributed to the posterior margin of the external nares. The maximal width of the premaxilla of MNHN.F INA 30 was mostly level with the P3 and P4 alveoli, as in
|
||
<taxonomicName id="4C0703D5FF8BF505FBDFD46BFB5BFD2D" baseAuthorityName="de Lapparent de Broin" baseAuthorityYear="2002" box="[1053,1215,627,648]" class="Reptilia" family="Pholidosauridae" genus="Elosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="382" phylum="Chordata" rank="species" species="cherifiensis">
|
||
<emphasis id="B973A444FF8BF505FBDFD46BFB5BFD2D" box="[1053,1215,627,648]" italics="true" pageId="5" pageNumber="382">E. cherifiensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9605A7FF8BF505FB14D46BFC21FD03" author="de Lapparent de Broin F" pageId="5" pageNumber="382" pagination="275 - 285" refId="ref17195" refString="de Lapparent de Broin F. 2002. Elosuchus, a new genus of crocodile from the Lower Cretaceous of the North of Africa. Comptes Rendus Palevol 1: 275 - 285." type="journal article" year="2002">de Lapparent de Broin, 2002</bibRefCitation>
|
||
; MNHN.F SAM 129).
|
||
</paragraph>
|
||
<paragraph id="8BB87856FF8BF505FC94D4A8FAD2FB0A" blockId="5.[829,1444,199,1260]" pageId="5" pageNumber="382">
|
||
The right premaxilla bears five alveoli, whereas the fifth alveolus is not preserved in the left premaxilla (
|
||
<figureCitation id="133C64D3FF8BF505FC87D4F5FC6EFCA6" box="[837,906,749,771]" captionStart="Figure 3" captionStartId="5.[166,245,1850,1869]" captionTargetBox="[174,1436,1307,1816]" captionTargetId="figure-648@5.[174,1438,1307,1816]" captionTargetPageId="5" captionText="Figure 3. Premaxilla MNHN.F INA 30, referred to Elosuchus sp.: A, photograph in dorsal view; B, photograph in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5710913" httpUri="https://zenodo.org/record/5710913/files/figure.png" pageId="5" pageNumber="382">Fig. 3</figureCitation>
|
||
). This matches the premaxillary tooth count of five seen in specimens referred to
|
||
<taxonomicName id="4C0703D5FF8BF505FB0ED514FA8AFC84" baseAuthorityName="de Lapparent de Broin" baseAuthorityYear="2002" box="[1228,1390,780,801]" class="Reptilia" family="Pholidosauridae" genus="Elosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="382" phylum="Chordata" rank="species" species="cherifiensis">
|
||
<emphasis id="B973A444FF8BF505FB0ED514FA8AFC84" box="[1228,1390,780,801]" italics="true" pageId="5" pageNumber="382">E. cherifiensis</emphasis>
|
||
</taxonomicName>
|
||
(not four as stated by
|
||
<bibRefCitation id="EF9605A7FF8BF505FBCCD532FA87FCE5" author="de Lapparent de Broin F" box="[1038,1379,810,832]" pageId="5" pageNumber="382" pagination="275 - 285" refId="ref17195" refString="de Lapparent de Broin F. 2002. Elosuchus, a new genus of crocodile from the Lower Cretaceous of the North of Africa. Comptes Rendus Palevol 1: 275 - 285." type="journal article" year="2002">de Lapparent de Broin, 2002</bibRefCitation>
|
||
). The first two alveoli (P1 and P2) of both premaxillae are aligned in the same coronal plane, and are oriented vertically and slightly posteriorly. The first two alveoli of both premaxillae form a row of four small and tightly spaced alveoli. These would have created the premaxillary overhang of the dentary seen in other pholidosaurids (e.g.
|
||
<bibRefCitation id="EF9605A7FF8BF505FBE8D218FB57FBB0" author="Mook CC" box="[1066,1203,1024,1046]" pageId="5" pageNumber="382" pagination="1 - 6" refId="ref17668" refString="Mook CC. 1933. Skull characters of Teleorhinus browni Osborn. American Museum Novitates 603: 1 - 6." type="journal article" year="1933">Mook, 1933</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9605A7FF8BF505FB00D218FB1FFBB0" author="Mook CC" box="[1218,1275,1024,1045]" pageId="5" pageNumber="382" pagination="1 - 11" refId="ref17689" refString="Mook CC. 1934. A new species of Teleorhinus from the Benton Shales. American Museum Novitates 702: 1 - 11." type="journal article" year="1934">1934</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF8BF505FAC8D218FC92FB91" author="Sereno PC & Larsson HCE & Sidor CA & Gado B" pageId="5" pageNumber="382" pagination="1516 - 1519" refId="ref18058" refString="Sereno PC, Larsson HCE, Sidor CA, Gado B. 2001. The giant crocodyliform Sarcosuchus from the Cretaceous of Africa. Science 294: 1516 - 1519." type="journal article" year="2001">
|
||
Sereno
|
||
<emphasis id="B973A444FF8BF505FAA1D219FA79FBB0" box="[1379,1437,1024,1046]" italics="true" pageId="5" pageNumber="382">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF8BF505FC41D207FBD0FB91" author="Wu X-C & Russell AP & Cumbaa SL" box="[899,1076,1055,1076]" pageId="5" pageNumber="382" pagination="492 - 514" refId="ref18460" refString="Wu X-C, Russell AP, Cumbaa SL. 2001. Terminonaris (Archosauria: Crocodyliformes): new material from Saskatchewan, Canada, and comments on its phylogenetic relationships. Journal of Vertebrate Paleontology 21: 492 - 514." type="journal article" year="2001">
|
||
Wu
|
||
<emphasis id="B973A444FF8BF505FC70D207FC08FB91" box="[946,1004,1055,1076]" italics="true" pageId="5" pageNumber="382">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF8BF505FB83D207FB18FB91" author="Hua S & Buffetaut E & Legall C & Rogron P" box="[1089,1276,1055,1076]" pageId="5" pageNumber="382" pagination="503 - 513" refId="ref16531" refString="Hua S, Buffetaut E, Legall C, Rogron P. 2007. Oceanosuchus boecensis n. gen, n. sp., a marine pholidosaurid (Crocodylia, Mesosuchia) from the Lower Cenomanian of Normandy (western France). Bulletin de la Societe Geologique de France 178: 503 - 513." type="journal article" year="2007">
|
||
Hua
|
||
<emphasis id="B973A444FF8BF505FBB8D207FB50FB91" box="[1146,1204,1055,1076]" italics="true" pageId="5" pageNumber="382">et al.</emphasis>
|
||
, 2007
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF8BF505FACBD207FC92FBF7" author="Lepage Y & Buffetaut E & Hua S & Martin JE & Tabouelle J" pageId="5" pageNumber="382" pagination="5 - 152" refId="ref17292" refString="Lepage Y, Buffetaut E, Hua S, Martin JE, Tabouelle J. 2008. Catalogue descriptif, anatomique, geologique et historique des fossiles presentes<65>a l'exposition " Les Crocodiliens fossiles de Normandie " (6 novembre - 14 decembre 2008). Bulletin de la Societe Geologique de Normandie et des Amis du Museum du Havre 95: 5 - 152." type="journal article" year="2008">
|
||
Lepage
|
||
<emphasis id="B973A444FF8BF505FAA1D207FA79FB91" box="[1379,1437,1055,1076]" italics="true" pageId="5" pageNumber="382">et al.</emphasis>
|
||
, 2008
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF8BF505FC45D225FB8EFBF7" author="Fortier D & Perea D & Schultz C" box="[903,1130,1085,1107]" pageId="5" pageNumber="382" pagination="272" refId="ref16232" refString="Fortier D, Perea D, Schultz C. 2011. Redescription and phylogenetic relationships of Meridiosaurus vallisparadisi, a pholidosaurid from the Late Jurassic of Uruguay. Zoological Journal of the Linnean Society 163: S 257 - S 272." type="journal article" year="2011">
|
||
Fortier
|
||
<emphasis id="B973A444FF8BF505FC26D226FBFAFBF7" box="[996,1054,1085,1107]" italics="true" pageId="5" pageNumber="382">et al.</emphasis>
|
||
, 2011
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF8BF505FBB9D225FA84FBF6" author="Martin JE & Lauprasert K & Buffetaut E & Liard R & Suteethorn V" box="[1147,1376,1085,1107]" pageId="5" pageNumber="382" pagination="757 - 769" refId="ref17495" refString="Martin JE, Lauprasert K, Buffetaut E, Liard R, Suteethorn V. 2014. A large pholidosaurid in the Phu Kradung Formation of north-eastern Thailand. Palaeontology 57: 757 - 769." type="journal article" year="2014">
|
||
Martin
|
||
<emphasis id="B973A444FF8BF505FB1AD226FAF6FBF7" box="[1240,1298,1085,1107]" italics="true" pageId="5" pageNumber="382">et al.</emphasis>
|
||
, 2014
|
||
</bibRefCitation>
|
||
). The shape, and how many premaxillary alveoli contribute to this overhang, vary amongst pholidosaurids and are potentially phylogenetically informative.
|
||
</paragraph>
|
||
<paragraph id="8BB87856FF8BF506FC94D2AFFD0CFD02" blockId="5.[829,1444,199,1260]" lastBlockId="6.[147,762,199,1535]" lastPageId="6" lastPageNumber="383" pageId="5" pageNumber="382">
|
||
The P3 and P4 alveoli are separated from P1 and P2 alveoli by large foramina, just as in the specimens referred to
|
||
<taxonomicName id="4C0703D5FF88F506FEE1D6D0FE21FF79" baseAuthorityName="de Lapparent de Broin" baseAuthorityYear="2002" box="[291,453,199,221]" class="Reptilia" family="Pholidosauridae" genus="Elosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="383" phylum="Chordata" rank="species" species="cherifiensis">
|
||
<emphasis id="B973A444FF88F506FEE1D6D0FE21FF79" box="[291,453,199,221]" italics="true" pageId="6" pageNumber="383">E. cherifiensis</emphasis>
|
||
</taxonomicName>
|
||
(MNHN.F SAM 129 and MRS 334; NHMUK PV R 36828;
|
||
<bibRefCitation id="EF9605A7FF88F506FDEFD6FDFEF9FEBF" author="de Lapparent de Broin F" pageId="6" pageNumber="383" pagination="275 - 285" refId="ref17195" refString="de Lapparent de Broin F. 2002. Elosuchus, a new genus of crocodile from the Lower Cretaceous of the North of Africa. Comptes Rendus Palevol 1: 275 - 285." type="journal article" year="2002">de Lapparent de Broin, 2002</bibRefCitation>
|
||
). These large, paired foramina appear to be for the reception of the enlarged first dentary tooth crowns. Similar notches (or in some cases just depressions) are also present on the palatal surface of the premaxillae of other pholidosaurids (e.g.
|
||
<bibRefCitation id="EF9605A7FF88F506FDACD766FD12FE31" author="Mook CC" box="[622,758,382,404]" pageId="6" pageNumber="383" pagination="1 - 6" refId="ref17668" refString="Mook CC. 1933. Skull characters of Teleorhinus browni Osborn. American Museum Novitates 603: 1 - 6." type="journal article" year="1933">Mook, 1933</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF88F506FF51D785FE88FE17" author="Sereno PC & Larsson HCE & Sidor CA & Gado B" box="[147,364,413,435]" pageId="6" pageNumber="383" pagination="1516 - 1519" refId="ref18058" refString="Sereno PC, Larsson HCE, Sidor CA, Gado B. 2001. The giant crocodyliform Sarcosuchus from the Cretaceous of Africa. Science 294: 1516 - 1519." type="journal article" year="2001">
|
||
Sereno
|
||
<emphasis id="B973A444FF88F506FF28D786FEC0FE17" box="[234,292,413,435]" italics="true" pageId="6" pageNumber="383">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF88F506FEBAD785FDB0FE17" author="Lepage Y & Buffetaut E & Hua S & Martin JE & Tabouelle J" box="[376,596,413,435]" pageId="6" pageNumber="383" pagination="5 - 152" refId="ref17292" refString="Lepage Y, Buffetaut E, Hua S, Martin JE, Tabouelle J. 2008. Catalogue descriptif, anatomique, geologique et historique des fossiles presentes<65>a l'exposition " Les Crocodiliens fossiles de Normandie " (6 novembre - 14 decembre 2008). Bulletin de la Societe Geologique de Normandie et des Amis du Museum du Havre 95: 5 - 152." type="journal article" year="2008">
|
||
Lepage
|
||
<emphasis id="B973A444FF88F506FE10D786FDE8FE17" box="[466,524,413,435]" italics="true" pageId="6" pageNumber="383">et al.</emphasis>
|
||
, 2008
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF88F506FDA3D785FF28FE77" author="Fortier D & Perea D & Schultz C" pageId="6" pageNumber="383" pagination="272" refId="ref16232" refString="Fortier D, Perea D, Schultz C. 2011. Redescription and phylogenetic relationships of Meridiosaurus vallisparadisi, a pholidosaurid from the Late Jurassic of Uruguay. Zoological Journal of the Linnean Society 163: S 257 - S 272." type="journal article" year="2011">
|
||
Fortier
|
||
<emphasis id="B973A444FF88F506FD7BD786FD17FE17" box="[697,755,413,435]" italics="true" pageId="6" pageNumber="383">et al.</emphasis>
|
||
, 2011
|
||
</bibRefCitation>
|
||
); however, in
|
||
<taxonomicName id="4C0703D5FF88F506FE53D7A3FDD3FE75" authorityName="Mones" authorityYear="1980" box="[401,567,443,464]" class="Reptilia" family="Pholidosauridae" genus="Meridiosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="383" phylum="Chordata" rank="genus">
|
||
<emphasis id="B973A444FF88F506FE53D7A3FDD3FE75" box="[401,567,443,464]" italics="true" pageId="6" pageNumber="383">Meridiosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0703D5FF88F506FD90D7A3FD17FE75" box="[594,755,443,464]" class="Reptilia" family="Pholidosauridae" genus="Oceanosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="383" phylum="Chordata" rank="genus">
|
||
<emphasis id="B973A444FF88F506FD90D7A3FD17FE75" box="[594,755,443,464]" italics="true" pageId="6" pageNumber="383">Oceanosuchus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0703D5FF88F506FF51D7C2FEC6FE4A" authorityName="de Broin & Taquet" authorityYear="1966" box="[147,290,474,495]" class="Reptilia" family="Crocodylidae" genus="Sarcosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="383" phylum="Chordata" rank="genus">
|
||
<emphasis id="B973A444FF88F506FF51D7C2FEC6FE4A" box="[147,290,474,495]" italics="true" pageId="6" pageNumber="383">Sarcosuchus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0703D5FF88F506FEA2D7C3FE1FFE55" authorityName="Osborn" authorityYear="1904" box="[352,507,475,496]" class="Reptilia" family="Teleosauridae" genus="Terminonaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="383" phylum="Chordata" rank="genus">
|
||
<emphasis id="B973A444FF88F506FEA2D7C3FE1FFE55" box="[352,507,475,496]" italics="true" pageId="6" pageNumber="383">Terminonaris</emphasis>
|
||
</taxonomicName>
|
||
these notches are posterior to the P1 and P2 alveoli (or just the P2 alveoli), and are ventral to the external nares (
|
||
<bibRefCitation id="EF9605A7FF88F506FDA8D40FFD12FD89" author="Mook CC" box="[618,758,535,557]" pageId="6" pageNumber="383" pagination="1 - 6" refId="ref17668" refString="Mook CC. 1933. Skull characters of Teleorhinus browni Osborn. American Museum Novitates 603: 1 - 6." type="journal article" year="1933">Mook, 1933</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF88F506FF51D42EFE88FDEE" author="Sereno PC & Larsson HCE & Sidor CA & Gado B" box="[147,364,566,587]" pageId="6" pageNumber="383" pagination="1516 - 1519" refId="ref18058" refString="Sereno PC, Larsson HCE, Sidor CA, Gado B. 2001. The giant crocodyliform Sarcosuchus from the Cretaceous of Africa. Science 294: 1516 - 1519." type="journal article" year="2001">
|
||
Sereno
|
||
<emphasis id="B973A444FF88F506FF28D42EFEC0FDEE" box="[234,292,566,587]" italics="true" pageId="6" pageNumber="383">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF88F506FEBAD42EFDB0FDEE" author="Lepage Y & Buffetaut E & Hua S & Martin JE & Tabouelle J" box="[376,596,566,587]" pageId="6" pageNumber="383" pagination="5 - 152" refId="ref17292" refString="Lepage Y, Buffetaut E, Hua S, Martin JE, Tabouelle J. 2008. Catalogue descriptif, anatomique, geologique et historique des fossiles presentes<65>a l'exposition " Les Crocodiliens fossiles de Normandie " (6 novembre - 14 decembre 2008). Bulletin de la Societe Geologique de Normandie et des Amis du Museum du Havre 95: 5 - 152." type="journal article" year="2008">
|
||
Lepage
|
||
<emphasis id="B973A444FF88F506FE10D42EFDE8FDEE" box="[466,524,566,587]" italics="true" pageId="6" pageNumber="383">et al.</emphasis>
|
||
, 2008
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF88F506FDA3D42EFF29FDCF" author="Fortier D & Perea D & Schultz C" pageId="6" pageNumber="383" pagination="272" refId="ref16232" refString="Fortier D, Perea D, Schultz C. 2011. Redescription and phylogenetic relationships of Meridiosaurus vallisparadisi, a pholidosaurid from the Late Jurassic of Uruguay. Zoological Journal of the Linnean Society 163: S 257 - S 272." type="journal article" year="2011">
|
||
Fortier
|
||
<emphasis id="B973A444FF88F506FD7BD42EFD17FDEE" box="[697,755,566,587]" italics="true" pageId="6" pageNumber="383">et al.</emphasis>
|
||
, 2011
|
||
</bibRefCitation>
|
||
). The position and very large size of these foraminae, and the separation of the premaxillary alveoli into distinct units, are autapomorphies of
|
||
<taxonomicName id="4C0703D5FF88F506FDAFD489FD05FD03" box="[621,737,657,678]" class="Reptilia" family="Pholidosauridae" genus="Elosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="383" phylum="Chordata" rank="genus">
|
||
<emphasis id="B973A444FF88F506FDAFD489FD05FD03" box="[621,737,657,678]" italics="true" pageId="6" pageNumber="383">Elosuchus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF7828DEFF8BF505FF64D122FF3AF8CF" ID-DOI="http://doi.org/10.5281/zenodo.5710913" ID-Zenodo-Dep="5710913" httpUri="https://zenodo.org/record/5710913/files/figure.png" pageId="5" pageNumber="382" startId="5.[166,245,1850,1869]" targetBox="[174,1436,1307,1816]" targetPageId="5">
|
||
<paragraph id="8BB87856FF8BF505FF64D122FF3AF8CF" blockId="5.[166,1444,1850,1898]" pageId="5" pageNumber="382">
|
||
<emphasis id="B973A444FF8BF505FF64D122FEF4F8E8" bold="true" box="[166,272,1850,1869]" pageId="5" pageNumber="382">Figure 3.</emphasis>
|
||
Premaxilla MNHN.F INA 30, referred to
|
||
<taxonomicName id="4C0703D5FF8BF505FD0ED122FC84F8E8" authority="sp." authorityName="SP." box="[716,864,1850,1869]" class="Reptilia" family="Pholidosauridae" genus="Elosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="381" phylum="Chordata" rank="genus">
|
||
<emphasis id="B973A444FF8BF505FD0ED122FCD3F8E8" box="[716,823,1850,1869]" italics="true" pageId="5" pageNumber="382">Elosuchus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
: A, photograph in dorsal view; B, photograph in ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB87856FF88F506FF6ED4A8FEE3FA5A" blockId="6.[147,762,199,1535]" pageId="6" pageNumber="383">
|
||
As with
|
||
<taxonomicName id="4C0703D5FF88F506FEDAD4A9FE52FD60" baseAuthorityName="de Lapparent de Broin" baseAuthorityYear="2002" box="[280,438,688,710]" class="Reptilia" family="Pholidosauridae" genus="Elosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="383" phylum="Chordata" rank="species" species="cherifiensis">
|
||
<emphasis id="B973A444FF88F506FEDAD4A9FE52FD60" box="[280,438,688,710]" italics="true" pageId="6" pageNumber="383">E. cherifiensis</emphasis>
|
||
</taxonomicName>
|
||
, the P3 and P4 alveoli of MNHN.F INA 30 are considerably larger than the P1 and P2 alveoli, and are in approximately the same sagittal plane. The P3 and P4 alveoli are widely separated from one another, with a deep interalveolar space created by their high alveolar rims. The left premaxilla preserves the P5 alveolus posteromedial to the P4 alveolus. There is no deep interalveolar space between these alveoli, and the gap between the P4 and P5 alveoli is proportionally smaller than the P3
|
||
<emphasis id="B973A444FF88F506FD93D5DAFDB9FC7C" box="[593,605,962,985]" italics="true" pageId="6" pageNumber="383">–</emphasis>
|
||
P4 interalveolar space. The P5 alveolus is smaller than the P1
|
||
<emphasis id="B973A444FF88F506FD12D5F8FD38FC52" box="[720,732,992,1015]" italics="true" pageId="6" pageNumber="383">–</emphasis>
|
||
P4 alveoli. Also, it is oriented posteriorly. The posteriorly oriented P5 and its posteromedial position relative to the P4 alveolus (rather than posterolateral) are characteristics that MNHN.F INA 30 shares with
|
||
<taxonomicName id="4C0703D5FF88F506FD5BD244FF3DFB2A" baseAuthorityName="de Lapparent de Broin" baseAuthorityYear="2002" class="Reptilia" family="Pholidosauridae" genus="Elosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="383" phylum="Chordata" rank="species" species="cherifiensis">
|
||
<emphasis id="B973A444FF88F506FD5BD244FF3DFB2A" italics="true" pageId="6" pageNumber="383">E. cherifiensis</emphasis>
|
||
</taxonomicName>
|
||
(MNHN.F SAM 129;
|
||
<bibRefCitation id="EF9605A7FF88F506FE20D262FF29FB0A" author="de Lapparent de Broin F" pageId="6" pageNumber="383" pagination="275 - 285" refId="ref17195" refString="de Lapparent de Broin F. 2002. Elosuchus, a new genus of crocodile from the Lower Cretaceous of the North of Africa. Comptes Rendus Palevol 1: 275 - 285." type="journal article" year="2002">de Lapparent de Broin, 2002</bibRefCitation>
|
||
). The P5 alveoli of most other pholidosaurids are posterolateral to the P4 alveoli (e.g.
|
||
<bibRefCitation id="EF9605A7FF88F506FDE2D2AFFD47FB68" author="Mook CC" box="[544,675,1207,1229]" pageId="6" pageNumber="383" pagination="1 - 11" refId="ref17689" refString="Mook CC. 1934. A new species of Teleorhinus from the Benton Shales. American Museum Novitates 702: 1 - 11." type="journal article" year="1934">Mook, 1934</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF88F506FD6ED2A0FEF0FB4E" author="Sereno PC & Larsson HCE & Sidor CA & Gado B" pageId="6" pageNumber="383" pagination="1516 - 1519" refId="ref18058" refString="Sereno PC, Larsson HCE, Sidor CA, Gado B. 2001. The giant crocodyliform Sarcosuchus from the Cretaceous of Africa. Science 294: 1516 - 1519." type="journal article" year="2001">
|
||
Sereno
|
||
<emphasis id="B973A444FF88F506FF51D2CFFF28FB4E" box="[147,204,1238,1260]" italics="true" pageId="6" pageNumber="383">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF88F506FEE0D2CEFE37FB4E" author="Wu X-C & Russell AP & Cumbaa SL" box="[290,467,1238,1260]" pageId="6" pageNumber="383" pagination="492 - 514" refId="ref18460" refString="Wu X-C, Russell AP, Cumbaa SL. 2001. Terminonaris (Archosauria: Crocodyliformes): new material from Saskatchewan, Canada, and comments on its phylogenetic relationships. Journal of Vertebrate Paleontology 21: 492 - 514." type="journal article" year="2001">
|
||
Wu
|
||
<emphasis id="B973A444FF88F506FE90D2CFFE6FFB4E" box="[338,395,1238,1260]" italics="true" pageId="6" pageNumber="383">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF88F506FE22D2CEFD7FFB4E" author="Hua S & Buffetaut E & Legall C & Rogron P" box="[480,667,1238,1260]" pageId="6" pageNumber="383" pagination="503 - 513" refId="ref16531" refString="Hua S, Buffetaut E, Legall C, Rogron P. 2007. Oceanosuchus boecensis n. gen, n. sp., a marine pholidosaurid (Crocodylia, Mesosuchia) from the Lower Cenomanian of Normandy (western France). Bulletin de la Societe Geologique de France 178: 503 - 513." type="journal article" year="2007">
|
||
Hua
|
||
<emphasis id="B973A444FF88F506FDD8D2CFFDB7FB4E" box="[538,595,1238,1260]" italics="true" pageId="6" pageNumber="383">et al.</emphasis>
|
||
, 2007
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF88F506FD6BD2CEFEF6FAAF" author="Lepage Y & Buffetaut E & Hua S & Martin JE & Tabouelle J" pageId="6" pageNumber="383" pagination="5 - 152" refId="ref17292" refString="Lepage Y, Buffetaut E, Hua S, Martin JE, Tabouelle J. 2008. Catalogue descriptif, anatomique, geologique et historique des fossiles presentes<65>a l'exposition " Les Crocodiliens fossiles de Normandie " (6 novembre - 14 decembre 2008). Bulletin de la Societe Geologique de Normandie et des Amis du Museum du Havre 95: 5 - 152." type="journal article" year="2008">
|
||
Lepage
|
||
<emphasis id="B973A444FF88F506FF51D2EDFF28FAAC" box="[147,204,1268,1290]" italics="true" pageId="6" pageNumber="383">et al.</emphasis>
|
||
, 2008
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9605A7FF88F506FEDCD2ECFE17FAAE" author="Fortier D & Perea D & Schultz C" box="[286,499,1268,1291]" pageId="6" pageNumber="383" pagination="272" refId="ref16232" refString="Fortier D, Perea D, Schultz C. 2011. Redescription and phylogenetic relationships of Meridiosaurus vallisparadisi, a pholidosaurid from the Late Jurassic of Uruguay. Zoological Journal of the Linnean Society 163: S 257 - S 272." type="journal article" year="2011">
|
||
Fortier
|
||
<emphasis id="B973A444FF88F506FEB6D2EDFE49FAAC" box="[372,429,1268,1290]" italics="true" pageId="6" pageNumber="383">et al.</emphasis>
|
||
, 2011
|
||
</bibRefCitation>
|
||
); however, the P5 alveoli of
|
||
<taxonomicName id="4C0703D5FF88F506FF16D30BFD28FA8C" authority="(Martin et al., 2014)" baseAuthorityName="Martin" baseAuthorityYear="2014" box="[212,716,1299,1321]" class="Reptilia" family="Pholidosauridae" genus="Chalawan" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="383" phylum="Chordata" rank="species" species="thailandicus">
|
||
<emphasis id="B973A444FF88F506FF16D30BFE3BFA8D" box="[212,479,1299,1320]" italics="true" pageId="6" pageNumber="383">Chalawan thailandicus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9605A7FF88F506FE2CD30BFD21FA8C" author="Martin JE & Lauprasert K & Buffetaut E & Liard R & Suteethorn V" box="[494,709,1299,1321]" pageId="6" pageNumber="383" pagination="757 - 769" refId="ref17495" refString="Martin JE, Lauprasert K, Buffetaut E, Liard R, Suteethorn V. 2014. A large pholidosaurid in the Phu Kradung Formation of north-eastern Thailand. Palaeontology 57: 757 - 769." type="journal article" year="2014">
|
||
Martin
|
||
<emphasis id="B973A444FF88F506FD85D30CFD64FA8D" box="[583,640,1299,1321]" italics="true" pageId="6" pageNumber="383">et al.</emphasis>
|
||
, 2014
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
are also in a posteromedial position. Thus,
|
||
<emphasis id="B973A444FF88F506FD98D32AFD45FAE2" box="[602,673,1330,1351]" italics="true" pageId="6" pageNumber="383">contra</emphasis>
|
||
<bibRefCitation id="EF9605A7FF88F506FD69D32AFEFFFAC3" author="Fortier D & Perea D & Schultz C" pageId="6" pageNumber="383" pagination="272" refId="ref16232" refString="Fortier D, Perea D, Schultz C. 2011. Redescription and phylogenetic relationships of Meridiosaurus vallisparadisi, a pholidosaurid from the Late Jurassic of Uruguay. Zoological Journal of the Linnean Society 163: S 257 - S 272." type="journal article" year="2011">
|
||
Fortier
|
||
<emphasis id="B973A444FF88F506FF51D349FF28FAC0" box="[147,204,1360,1382]" italics="true" pageId="6" pageNumber="383">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
, the posterolateral position of the fifth premaxillary tooth is not an unambiguous synapomorphy of a monophyletic
|
||
<taxonomicName id="4C0703D5FF88F506FE49D395FDA7FA06" box="[395,579,1421,1443]" class="Reptilia" family="Pholidosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="383" phylum="Chordata" rank="family">Pholidosauridae</taxonomicName>
|
||
, although the medial ‘migration’ of these alveoli could be a shared characteristic of a more inclusive clade within pholidosaurids.
|
||
</paragraph>
|
||
<paragraph id="8BB87856FF88F506FEC8D02EFDC2F941" blockId="6.[195,714,1585,1764]" pageId="6" pageNumber="383">
|
||
TETHYSUCHIA
|
||
<bibRefCitation id="EF9605A7FF88F506FE72D02BFD67F9EE" author="Buffetaut E" box="[432,643,1585,1612]" pageId="6" pageNumber="383" pagination="1 - 88" refId="ref15691" refString="Buffetaut E. 1982. Radiation evolutive, paleoecologie et biogeographie des crocodiliens mesosuchiens. Memoires de la Societe Geologique de France 142: 1 - 88." type="journal article" year="1982">BUFFETAUT, 1982</bibRefCitation>
|
||
(
|
||
<emphasis id="B973A444FF88F506FED4D04FFEBBF9C2" box="[278,351,1623,1639]" italics="true" pageId="6" pageNumber="383">SENSU</emphasis>
|
||
ANDRADE
|
||
<emphasis id="B973A444FF88F506FE1CD040FDC3F9CF" box="[478,551,1617,1642]" italics="true" pageId="6" pageNumber="383">ET AL.</emphasis>
|
||
, 2011)
|
||
<taxonomicName id="4C0703D5FF88F506FEEBD068FE04F922" authority="Young & Hastings & Allain & Smith, 2017" authorityName="Young & Hastings & Allain & Smith" authorityYear="2017" box="[297,480,1648,1672]" class="Reptilia" family="Peirosauridae" genus="Fortignathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neosuchia" pageId="6" pageNumber="383" phylum="Chordata" rank="genus" status="gen. nov.">FORTIGNATHUS</taxonomicName>
|
||
<taxonomicNameLabel id="A240193FFF88F506FE28D06EFD80F92C" box="[490,612,1646,1673]" pageId="6" pageNumber="383" rank="genus">
|
||
<emphasis id="B973A444FF88F506FE28D06EFDB8F922" bold="true" box="[490,604,1646,1673]" pageId="6" pageNumber="383">GEN. NOV</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
http://ZOOBANK.ORG/URN:LSID:ZOOBANK.ORG: ACT:
|
||
<uuid id="FFA14283FF88F506FEE5D0B3FDC2F941" pageId="6" pageNumber="383">0F0CCD3A-2831-40DA- A05C- E29E30524889</uuid>
|
||
</paragraph>
|
||
<paragraph id="8BB87856FF88F506FF51D103FEFBF895" blockId="6.[147,762,1819,1901]" box="[147,287,1819,1840]" pageId="6" pageNumber="383">
|
||
<emphasis id="B973A444FF88F506FF51D103FEFBF895" box="[147,287,1819,1840]" italics="true" pageId="6" pageNumber="383">
|
||
<typeStatus id="54BCC6F4FF88F506FF51D103FF2DF895" box="[147,201,1819,1840]" pageId="6" pageNumber="383">Type</typeStatus>
|
||
species
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D2BDDFF88F506FF51D121FBDFFF5E" pageId="6" pageNumber="383" type="reference_group">
|
||
<paragraph id="8BB87856FF88F506FF51D121FD1DF8C8" blockId="6.[147,762,1819,1901]" pageId="6" pageNumber="383">
|
||
<taxonomicName id="4C0703D5FF88F506FF51D121FD79F8EB" authority="de Lapparent de Broin, 2002" authorityName="de Lapparent de Broin" authorityYear="2002" box="[147,669,1849,1871]" class="Reptilia" family="Pholidosauridae" genus="Elosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="383" phylum="Chordata" rank="species" species="felixi">
|
||
<emphasis id="B973A444FF88F506FF51D121FEACF8EB" box="[147,328,1849,1870]" italics="true" pageId="6" pageNumber="383">Elosuchus felixi</emphasis>
|
||
<bibRefCitation id="EF9605A7FF88F506FE93D121FD79F8EB" author="de Lapparent de Broin F" box="[337,669,1849,1871]" pageId="6" pageNumber="383" pagination="275 - 285" refId="ref17195" refString="de Lapparent de Broin F. 2002. Elosuchus, a new genus of crocodile from the Lower Cretaceous of the North of Africa. Comptes Rendus Palevol 1: 275 - 285." type="journal article" year="2002">de Lapparent de Broin, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
(following recommendation 67B of the ICZN code). Now
|
||
</paragraph>
|
||
<paragraph id="8BB87856FF88F506FCE8D6DFFBDFFF5E" blockId="6.[810,1424,199,252]" pageId="6" pageNumber="383">
|
||
referred to as
|
||
<taxonomicName id="4C0703D5FF88F506FC1ED6DFFC53FF59" authority="(de Lapparent de Broin, 2002)" authorityName="Young & Hastings & Allain & Smith" authorityYear="2017" baseAuthorityName="de Lapparent de Broin" baseAuthorityYear="2002" class="Reptilia" family="Peirosauridae" genus="Fortignathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neosuchia" pageId="6" pageNumber="383" phylum="Chordata" rank="species" species="felixi" status="comb. nov.">
|
||
<emphasis id="B973A444FF88F506FC1ED6DFFB51FF79" box="[988,1205,199,220]" italics="true" pageId="6" pageNumber="383">Fortignathus felixi</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9605A7FF88F506FB08D6DFFC54FF59" author="de Lapparent de Broin F" pageId="6" pageNumber="383" pagination="275 - 285" refId="ref17195" refString="de Lapparent de Broin F. 2002. Elosuchus, a new genus of crocodile from the Lower Cretaceous of the North of Africa. Comptes Rendus Palevol 1: 275 - 285." type="journal article" year="2002">de Lapparent de Broin, 2002</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A240193FFF88F506FC02D6FDFBDFFF5E" box="[960,1083,229,251]" pageId="6" pageNumber="383" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D2BDDFF88F506FCE8D73BFA65FD2F" pageId="6" pageNumber="383" type="etymology">
|
||
<paragraph id="8BB87856FF88F506FCE8D73BFC45FE9D" blockId="6.[810,1425,291,465]" box="[810,929,291,312]" pageId="6" pageNumber="383">
|
||
<emphasis id="B973A444FF88F506FCE8D73BFC45FE9D" box="[810,929,291,312]" italics="true" pageId="6" pageNumber="383">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB87856FF88F506FCE8D759FC14FE74" blockId="6.[810,1425,291,465]" pageId="6" pageNumber="383">
|
||
‘Strong jaws’. Derived from the Latin for strong or powerful (
|
||
<emphasis id="B973A444FF88F506FC5DD778FC3EFED0" box="[927,986,352,373]" italics="true" pageId="6" pageNumber="383">fortis</emphasis>
|
||
) and the Latinized form of the Greek for jaws (
|
||
<emphasis id="B973A444FF88F506FC6AD766FBE0FE36" box="[936,1028,382,403]" italics="true" pageId="6" pageNumber="383">gnathus</emphasis>
|
||
). Named for the robustness of MNHN.F INA 21 (a referred specimen from a very large individual).
|
||
</paragraph>
|
||
<paragraph id="8BB87856FF88F506FCE8D7E1FC7EFDAB" blockId="6.[810,1404,505,557]" box="[810,922,505,526]" pageId="6" pageNumber="383">
|
||
<emphasis id="B973A444FF88F506FCE8D7E1FC7EFDAB" box="[810,922,505,526]" italics="true" pageId="6" pageNumber="383">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB87856FF88F506FCE8D40FFA98FD89" blockId="6.[810,1404,505,557]" box="[810,1404,535,557]" pageId="6" pageNumber="383">Same as the only known species (monotypic genus).</paragraph>
|
||
<paragraph id="8BB87856FF88F506FCF8D46BFA65FD2F" blockId="6.[826,1409,623,681]" box="[826,1409,623,650]" pageId="6" pageNumber="383">
|
||
<taxonomicName id="4C0703D5FF88F506FCF8D46BFA65FD2F" box="[826,1409,623,650]" pageId="6" pageNumber="383">
|
||
<emphasis id="B973A444FF88F506FCF8D46BFBA1FD22" bold="true" box="[826,1093,627,648]" italics="true" pageId="6" pageNumber="383">FORTIGNATHUS FELIXI</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9605A7FF88F506FB9AD46FFA65FD2F" author="de Lapparent de Broin F" box="[1112,1409,624,650]" pageId="6" pageNumber="383" pagination="275 - 285" refId="ref17195" refString="de Lapparent de Broin F. 2002. Elosuchus, a new genus of crocodile from the Lower Cretaceous of the North of Africa. Comptes Rendus Palevol 1: 275 - 285." type="journal article" year="2002">DE LAPPARENT DE BROIN,</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |