136 lines
16 KiB
XML
136 lines
16 KiB
XML
<document id="5972A254AF24AC308CE6C3D379B124F7" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFDCFFDF6194FF670571F252" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus quinarius Perkins 2021, new species" docType="treatment" docVersion="4" lastPageNumber="33" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="33" updateTime="1698978858223" updateUser="plazi">
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<mods:titleInfo id="A83619D84643C0B177E2F92517C2E96D">
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<mods:title id="BECD0A2ACA259275B8F9761DA819D3E3">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
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<mods:name id="01940573ACA0D4C7947824C596F6AAA9" type="personal">
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<mods:roleTerm id="AB275B5B6256CA6877CA90F72671EEB0">Author</mods:roleTerm>
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<mods:namePart id="F218A314449B4893AC204E5BB0240168">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="10E55204E43ECFBF3D25EC43F01EDCD2">text</mods:typeOfResource>
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<mods:title id="F506AE926938C20297B8B7BDB51CE741">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B2461B8F2B049CD63A0642D77E5AE27C">
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<mods:date id="601FA59238BB7B847691AAF6DE4EE252">2021</mods:date>
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<mods:detail id="DF7368DB2B54E37D739AE52DA039C838" type="pubDate">
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<mods:number id="D33D591F7DE330EEC72EED92F06F4FA1">2021-06-28</mods:number>
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<mods:detail id="F36A00AA1C06A5C286C6E84A0B311707" type="volume">
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<mods:number id="AF6F6264B19862C67B1D2CA0B6158375">4994</mods:number>
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<mods:detail id="745827B3FD8E30ACFEA1B160C974AF70" type="issue">
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<mods:number id="D960A1770B08270FB02453C0849D0286">1</mods:number>
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<mods:start id="1E95E1988A241CFE0735B9767BE7EEB1">1</mods:start>
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<mods:end id="9D933DE99FCC894415A9FA877221EF14">93</mods:end>
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<mods:classification id="AB63521CE3F9FFFCE35B4662712B4329">journal article</mods:classification>
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<mods:identifier id="BA20909D77EEFE2D714DE19C77001542" type="DOI">10.11646/zootaxa.4994.1.1</mods:identifier>
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<mods:identifier id="269644F46DD532E07592185A71DCC935" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A187DEFFDCFFDF6194FF670571F252" ID-DOI="http://doi.org/10.5281/zenodo.5043976" ID-Zenodo-Dep="5043976" LSID="urn:lsid:plazi:treatment:03A187DEFFDCFFDF6194FF670571F252" httpUri="http://treatment.plazi.org/id/03A187DEFFDCFFDF6194FF670571F252" lastPageNumber="33" pageId="32" pageNumber="33">
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<subSubSection id="C3126543FFDCFFDF6194FF670667F7B6" box="[151,568,151,178]" pageId="32" pageNumber="33" type="nomenclature">
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<paragraph id="8BB736C8FFDCFFDF6194FF670667F7B6" blockId="32.[151,568,151,214]" box="[151,568,151,178]" pageId="32" pageNumber="33">
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<heading id="D0FF81A4FFDCFFDF6194FF670667F7B6" bold="true" box="[151,568,151,178]" fontSize="11" level="2" pageId="32" pageNumber="33" reason="4">
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<emphasis id="B97CEADAFFDCFFDF6194FF670667F7B6" bold="true" box="[151,568,151,178]" pageId="32" pageNumber="33">
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<taxonomicName id="4C084D4BFFDCFFDF6194FF6705C6F7B6" ID-CoL="8Y4R7" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,409,151,178]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="quinarius" status="sp. nov.">
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<emphasis id="B97CEADAFFDCFFDF6194FF6705C6F7B6" bold="true" box="[151,409,151,178]" italics="true" pageId="32" pageNumber="33">Hydrochus quinarius</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24F57A1FFDCFFDF60ABFF670667F7B6" box="[424,568,151,177]" pageId="32" pageNumber="33" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFDCFFDF6194FF4B0568F7D1" box="[151,311,187,214]" pageId="32" pageNumber="33" type="description">
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<paragraph id="8BB736C8FFDCFFDF6194FF4B0568F7D1" blockId="32.[151,568,151,214]" box="[151,311,187,214]" pageId="32" pageNumber="33">
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<figureCitation id="13332A4DFFDCFFDF6194FF4B04BBF7D2" box="[151,228,187,213]" captionStart="FIGURE 3" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[169,1397,189,1912]" captionTargetId="figure-28@8.[154,1433,181,1919]" captionTargetPageId="8" captionText="FIGURE 3. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043269" httpUri="https://zenodo.org/record/5043269/files/figure.png" pageId="32" pageNumber="33">Figs. 3</figureCitation>
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,
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<figureCitation id="13332A4DFFDCFFDF61F2FF4B0550F7D2" box="[241,271,187,213]" captionStart="FIGURES 32–33" captionStartId="68.[151,264,1944,1968]" captionTargetBox="[163,1427,196,1890]" captionTargetId="figure-28@68.[151,1435,181,1920]" captionTargetPageId="68" captionText="FIGURES 32–33. Habitus and male genitalia of holotypes. 32. Hydrochus quinarius n. sp. 33. Hydrochus baloghi Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043316" httpUri="https://zenodo.org/record/5043316/files/figure.png" pageId="32" pageNumber="33">32</figureCitation>
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,
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<figureCitation id="13332A4DFFDCFFDF6018FF4B0568F7D1" box="[283,311,187,214]" captionStart="FIGURES 57–58" captionStartId="81.[151,264,1928,1952]" captionTargetBox="[151,1435,181,1905]" captionTargetId="figure-15@81.[151,1435,181,1905]" captionTargetPageId="81" captionText="FIGURES 57–58. Geographical distributions. 57. Endemics and near endemics. 58. H. bituberculatus." figureDoi="http://doi.org/10.5281/zenodo.5043356" httpUri="https://zenodo.org/record/5043356/files/figure.png" pageId="32" pageNumber="33">57</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFDCFFDF6194FEF3017EF61A" box="[151,1313,259,286]" pageId="32" pageNumber="33" type="materials_examined">
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<paragraph id="8BB736C8FFDCFFDF6194FEF3017EF61A" blockId="32.[151,1437,259,1365]" box="[151,1313,259,286]" pageId="32" pageNumber="33">
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<emphasis id="B97CEADAFFDCFFDF6194FEF30517F61A" bold="true" box="[151,328,259,285]" pageId="32" pageNumber="33">Type Material.</emphasis>
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<materialsCitation id="3B603C95FFDCFFDF604CFEF3017EF61A" box="[335,1313,259,286]" collectingDate="1969-07-09" collectingDateMax="1969-07-10" collectingDateMin="1969-07-09" collectionCode="NMNH" collectorName="P. & P. Spangler" country="Brazil" location="Para" municipality="Belem" pageId="32" pageNumber="33" specimenCount="1" typeStatus="holotype">
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<emphasis id="B97CEADAFFDCFFDF604CFEF30647F61A" bold="true" box="[335,536,259,285]" pageId="32" pageNumber="33">
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<typeStatus id="54B3886AFFDCFFDF604CFEF305E3F61A" box="[335,444,259,285]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
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(male):
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</emphasis>
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“
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<collectingCountry id="F31F7658FFDCFFDF632AFEF306CAF61A" box="[553,661,259,285]" name="Brazil" pageId="32" pageNumber="33">BRAZIL</collectingCountry>
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:
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<location id="8ED76013FFDCFFDF63A3FEF4068AF619" LSID="urn:lsid:plazi:treatment:03A187DEFFDCFFDF6194FF670571F252:8ED76013FFDCFFDF63A3FEF4068AF619" box="[672,725,260,286]" country="Brazil" municipality="Belem" name="Para" pageId="32" pageNumber="33">Para</location>
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,
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<collectingMunicipality id="6BD3ACB2FFDCFFDF63DCFEF30774F61A" box="[735,811,259,285]" pageId="32" pageNumber="33">Belem</collectingMunicipality>
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,
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<date id="FFB61008FFDCFFDF6235FEF30781F61A" box="[822,990,259,285]" pageId="32" pageNumber="33" value="1969-07-09" valueMax="1969-07-10" valueMin="1969-07-09">
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<collectingDate id="EFF2E9E0FFDCFFDF6235FEF30781F61A" box="[822,990,259,285]" pageId="32" pageNumber="33" value="1969-07-09" valueMax="1969-07-10" valueMin="1969-07-09">VII-9-10-1969</collectingDate>
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</date>
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,
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<collectorName id="26FD531EFFDCFFDF62EAFEF400FEF61A" box="[1001,1185,259,286]" pageId="32" pageNumber="33">P. & P. Spangler</collectorName>
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” (
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<collectionCode id="ED19AE0DFFDCFFDF65BAFEF3014AF61A" box="[1209,1301,259,285]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="32" pageNumber="33" type="Museum">NMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFDCFFDF61C4FED7013EF6AA" pageId="32" pageNumber="33" type="diagnosis">
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<paragraph id="8BB736C8FFDCFFDF61C4FED7013EF6AA" blockId="32.[151,1437,259,1365]" pageId="32" pageNumber="33">
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<emphasis id="B97CEADAFFDCFFDF61C4FED7058CF646" bold="true" box="[199,467,295,321]" pageId="32" pageNumber="33">Differential Diagnosis.</emphasis>
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Among Brazilian species, recognized by the combination of small size (ca.
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<quantity id="4CF09B2DFFDCFFDF642BFED701CFF645" box="[1320,1424,295,322]" metricMagnitude="-3" metricUnit="m" metricValue="2.41" pageId="32" pageNumber="33" unit="mm" value="2.41">2.41 mm</quantity>
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), the pronotum with very shallow impressions, the pronotum with dense and rather uniformly distributed punctuation, the pronotal shape, the large black spots on the elytra, and the male genitalia (
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<figureCitation id="13332A4DFFDCFFDF62F3FE9F001DF68E" box="[1008,1090,367,393]" captionStart="FIGURES 32–33" captionStartId="68.[151,264,1944,1968]" captionTargetBox="[163,1427,196,1890]" captionTargetId="figure-28@68.[151,1435,181,1920]" captionTargetPageId="68" captionText="FIGURES 32–33. Habitus and male genitalia of holotypes. 32. Hydrochus quinarius n. sp. 33. Hydrochus baloghi Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043316" httpUri="https://zenodo.org/record/5043316/files/figure.png" pageId="32" pageNumber="33">Fig. 32</figureCitation>
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; described below). Differentiation from other similarly small sized species with similarly shaped pronotum will require dissection of males.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFDCFFDF61C4FE470719F20A" pageId="32" pageNumber="33" type="description">
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<paragraph id="8BB736C8FFDCFFDF61C4FE470565F582" blockId="32.[151,1437,259,1365]" pageId="32" pageNumber="33">
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<emphasis id="B97CEADAFFDCFFDF61C4FE47050EF6D6" bold="true" box="[199,337,439,465]" pageId="32" pageNumber="33">Description</emphasis>
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. Size:
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<typeStatus id="54B3886AFFDCFFDF60A7FE470659F6D6" box="[420,518,439,465]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 2.41/0.84; head width 0.60; pronotum l/w 0.54/0.54; PA 0.54; PB 0.45; elytra 1.53/0.84. Habitus as illustrated (
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<figureCitation id="13332A4DFFDCFFDF6549FE2B00FFF6F2" box="[1098,1184,475,501]" captionStart="FIGURES 32–33" captionStartId="68.[151,264,1944,1968]" captionTargetBox="[163,1427,196,1890]" captionTargetId="figure-28@68.[151,1435,181,1920]" captionTargetPageId="68" captionText="FIGURES 32–33. Habitus and male genitalia of holotypes. 32. Hydrochus quinarius n. sp. 33. Hydrochus baloghi Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043316" httpUri="https://zenodo.org/record/5043316/files/figure.png" pageId="32" pageNumber="33">Fig. 32</figureCitation>
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). Body size small (ca.
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<quantity id="4CF09B2DFFDCFFDF6194FE0F055DF51D" box="[151,258,511,538]" metricMagnitude="-3" metricUnit="m" metricValue="2.41" pageId="32" pageNumber="33" unit="mm" value="2.41">2.41 mm</quantity>
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). Dorsum brown (
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<typeStatus id="54B3886AFFDCFFDF60D6FE0F0666F51E" box="[469,569,511,537]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
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is teneral), elytra with several large black spots. Legs testaceous, with tibiofemoral joints darker. Punctation on elytra ca. 2–3x that of pronotum. Elytra interstriae ca. 0.5x strial puncture diameter, or less. Interstria 9
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<superScript id="7C7D9B80FFDCFFDF60D4FDB705BBF552" attach="left" box="[471,484,583,597]" fontSize="6" pageId="32" pageNumber="33">th</superScript>
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more convex than others, overhanging 10
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<superScript id="7C7D9B80FFDCFFDF62BFFDB70796F552" attach="left" box="[956,969,583,597]" fontSize="6" pageId="32" pageNumber="33">th</superScript>
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interstria. Usual area of callus on 5
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<superScript id="7C7D9B80FFDCFFDF6454FDB7013BF552" attach="left" box="[1367,1380,583,597]" fontSize="6" pageId="32" pageNumber="33">th</superScript>
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very weakly raised.
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</paragraph>
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<paragraph id="8BB736C8FFDCFFDF61C4FD600525F5F5" blockId="32.[151,1437,259,1365]" pageId="32" pageNumber="33">Pronotum slightly wider than long (ratio ca. 31/30), widest at anterior margin, narrowed at base, sides straight, smooth; anterior margin arcuate or very slightly bisinuate; depressions very shallow; punctation dense, quite uniform over pronotum</paragraph>
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<paragraph id="8BB736C8FFDCFFDF61C4FD0B0724F43E" blockId="32.[151,1437,259,1365]" pageId="32" pageNumber="33">Elytra with spaces between strial punctures ca. 0.5–1x their diameter; apices rather sharply separately rounded in dorsal view, in lateral view outer margin with slight angulation.</paragraph>
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<paragraph id="8BB736C8FFDCFFDF61C4FCB30505F486" blockId="32.[151,1437,259,1365]" pageId="32" pageNumber="33">Ventral characters: Mentum punctate, with deep central depression; submentum with two wide, deep, nearly confluent foveae.</paragraph>
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<paragraph id="8BB736C8FFDCFFDF61C4FC7B0569F3A6" blockId="32.[151,1437,259,1365]" pageId="32" pageNumber="33">
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Male genitalia (
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<figureCitation id="13332A4DFFDCFFDF6074FC7B0597F4A2" box="[375,456,907,933]" captionStart="FIGURES 32–33" captionStartId="68.[151,264,1944,1968]" captionTargetBox="[163,1427,196,1890]" captionTargetId="figure-28@68.[151,1435,181,1920]" captionTargetPageId="68" captionText="FIGURES 32–33. Habitus and male genitalia of holotypes. 32. Hydrochus quinarius n. sp. 33. Hydrochus baloghi Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043316" httpUri="https://zenodo.org/record/5043316/files/figure.png" pageId="32" pageNumber="33">Fig. 32</figureCitation>
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) general characters: genitalia narrow, elongate, length ca. 5x greatest width, basal piece much shorter than parameres (ratio ca. 1/3); paramere tips extending well beyond aedeagus tip, tips lobate and widely separated from aedeagus, plsa absent; in lateral view parameres very narrow, widened slightly subapically, sharply pointed apically; pseudoparameres large, long, ca. ¾ length of parameres, rather wide, with very sparse reticulation; aedeagus narrow, in dorsal/ventral views decreasing in width from agw, at apex, to adbl; basal piece in dorsal/ventral views decreasing in width from distal end. to orifice, in lateral view very slightly arcuate, dorsal surface more arcuate than ventral, maximum width in dorsal/ventral views slightly greater than lateral width, orifice asymmetrical.
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</paragraph>
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<paragraph id="8BB736C8FFDCFFDF61C4FB5B0698F3ED" blockId="32.[151,1437,259,1365]" pageId="32" pageNumber="33">Dorsal surface: adtl subapical, very faint; pdmm widely separated from alm, except underneath lateral margin of adbl; adbl small, width much smaller than agw.</paragraph>
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<paragraph id="8BB736C8FFDCFFDF61C4FB030719F20A" blockId="32.[151,1437,259,1365]" box="[199,838,1267,1294]" pageId="32" pageNumber="33">Ventral surface: avtl not apparent; pvmm same as pdmm.</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFDCFFDF61C4FAE70571F252" pageId="32" pageNumber="33" type="etymology">
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<paragraph id="8BB736C8FFDCFFDF61C4FAE70571F252" blockId="32.[151,1437,259,1365]" pageId="32" pageNumber="33">
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<emphasis id="B97CEADAFFDCFFDF61C4FAE70517F236" bold="true" box="[199,328,1303,1329]" pageId="32" pageNumber="33">Etymology</emphasis>
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. Named in reference to the five distal pieces of the male genitalia: aedeagus, 2 parameres, 2 pseudoparameres.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |