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<document id="C85E3F6B848848C1854C0612D1C6B1FC" ID-CLB-Dataset="29192" ID-DOI="10.11646/zootaxa.4482.2.2" ID-GBIF-Dataset="7f45fc71-fc58-4ca9-a320-5052c4aeba9d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1453225" ID-ZooBank="E0375F43-9552-4FC7-B79B-DA8A9B1B3D42" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539088841754" checkinUser="plazi" docAuthor="Recuero, Ernesto &amp; Sánchez-Vialas, Alberto" docDate="2018" docId="03A187BECD06FFF5FF35FB147AA6FE0D" docLanguage="en" docName="zootaxa.4482.2.2.pdf" docOrigin="Zootaxa 4482 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Barydesmus nangaritza Recuero &amp; Sánchez-Vialas 2018, new species" docType="treatment" docUuid="1AECF043-2201-4B6E-91D0-C0B177F9CE7D" docUuidSource="ZooBank" docVersion="9" lastPageNumber="250" masterDocId="FF98FFC6CD04FFF0FFA2FF987B41FFF8" masterDocTitle="A new distinctive species of Barydesmus (Diplopoda, Polydesmida, Platyrhacidae) from Ecuador, with an annotated bibliographical checklist of the American Platyrhacidae" masterLastPageNumber="273" masterPageNumber="245" pageNumber="247" updateTime="1698570743588" updateUser="ExternalLinkService">
<mods:mods id="82EC620F26A4D6FB4B70D4ECBB2FF303" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E5F11F36599A344472D975721F516305">A new distinctive species of Barydesmus (Diplopoda, Polydesmida, Platyrhacidae) from Ecuador, with an annotated bibliographical checklist of the American Platyrhacidae</mods:title>
</mods:titleInfo>
<mods:name id="C0D95B6D4888FE00942AE8DE3235A82A" type="personal">
<mods:role id="A137D0E95C89213B2756ED958EC159E1">
<mods:roleTerm id="71A19D506C376BA0C79FCB640EB1E14F">Author</mods:roleTerm>
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<mods:namePart id="DD36417FBDD7DC303E6E6A57D5DC35DA">Recuero, Ernesto</mods:namePart>
</mods:name>
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<mods:namePart id="B728D2B41EEDB7160BFAFAF9EEBF4007">Sánchez-Vialas, Alberto</mods:namePart>
</mods:name>
<mods:typeOfResource id="44E38D40CDD1ED74338B3957A807652D">text</mods:typeOfResource>
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<mods:title id="DF08FA233962F63EAE3678FF3530A6DD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A71F4CF8FD9E108A0C84AED39CA212D8">
<mods:date id="66FB721CF56BE0D38FB7B07021409BCC">2018</mods:date>
<mods:detail id="1BC46BB7D436CA615CB7CFF6B91B0CFE" type="pubDate">
<mods:number id="852AF900DEA4DD571E6E269F7F017E09">2018-09-18</mods:number>
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<mods:detail id="44A1DC8ED416AE2665D6315A96A5EAB5" type="volume">
<mods:number id="14065215D3297A51D6A52F0C1D25A4EC">4482</mods:number>
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<mods:number id="CCA057C653C333A39ACB982AFCF6D9E7">2</mods:number>
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<mods:start id="5FA85BF2E762CB215763ED309FC5390D">245</mods:start>
<mods:end id="042F460AF1C283DB8E089EFB73D55E6F">273</mods:end>
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<mods:classification id="15E56C97952D362DB19CFE84C02172D1">journal article</mods:classification>
<mods:identifier id="49F77B8A4848ED02F7CEDE6172A523C6" type="CLB-Dataset">29192</mods:identifier>
<mods:identifier id="BD1F5DE242266956D5A0A1CD13EC4816" type="DOI">10.11646/zootaxa.4482.2.2</mods:identifier>
<mods:identifier id="4A9D5D85C2D1BCFDDF145A0559F415BE" type="GBIF-Dataset">7f45fc71-fc58-4ca9-a320-5052c4aeba9d</mods:identifier>
<mods:identifier id="E682BAA4FF638AFAB99FDAACF43C5975" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="83F5A711634DB5774AA2D59F646B8925" type="Zenodo-Dep">1453225</mods:identifier>
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<treatment id="03A187BECD06FFF5FF35FB147AA6FE0D" ID-DOI="http://doi.org/10.5281/zenodo.5963364" ID-GBIF-Taxon="148663878" ID-Zenodo-Dep="5963364" LSID="urn:lsid:zoobank.org:act:1AECF043-2201-4B6E-91D0-C0B177F9CE7D" httpUri="http://treatment.plazi.org/id/03A187BECD06FFF5FF35FB147AA6FE0D" lastPageId="5" lastPageNumber="250" pageId="2" pageNumber="247">
<subSubSection id="C3126523CD06FFF2FF35FB147A6FFB3F" pageId="2" pageNumber="247" type="nomenclature">
<paragraph id="8BB736A8CD06FFF2FF35FB147912FB5E" blockId="2.[151,595,1164,1224]" box="[151,595,1164,1190]" pageId="2" pageNumber="247">
<heading id="D0FF81C4CD06FFF2FF35FB147912FB5E" bold="true" box="[151,595,1164,1190]" fontSize="11" level="1" pageId="2" pageNumber="247" reason="1">
<emphasis id="B97CEABACD06FFF2FF35FB147912FB5E" bold="true" box="[151,595,1164,1190]" pageId="2" pageNumber="247">
<taxonomicName id="4C084D2BCD06FFF2FF35FB147AF5FB5E" ID-CoL="68DJB" authority="Recuero &amp; Sánchez-Vialas, 2018" authorityName="Recuero &amp; Sánchez-Vialas" authorityYear="2018" box="[151,436,1164,1190]" class="Diplopoda" family="Platyrhacidae" genus="Barydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="247" phylum="Arthropoda" rank="species" species="nangaritza" status="sp. nov.">
<emphasis id="B97CEABACD06FFF2FF35FB147AF5FB5E" bold="true" box="[151,436,1164,1190]" italics="true" pageId="2" pageNumber="247">Barydesmus nangaritza</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24F57C1CD06FFF2FE61FB147912FB5E" box="[451,595,1164,1190]" pageId="2" pageNumber="247" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB736A8CD06FFF2FF35FB377A6FFB3F" blockId="2.[151,595,1164,1224]" box="[151,302,1199,1224]" pageId="2" pageNumber="247">
(
<figureCitation id="13332A2DCD06FFF2FF3CFB377A4AFB3F" box="[158,267,1199,1224]" captionStart="FIGURE 2" captionStartId="3.[151,250,1790,1812]" captionTargetBox="[157,1426,734,1762]" captionTargetId="figure@3.[151,1436,733,1766]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Barydesmus nangaritza sp. nov. A: Head and collum; B: 5 th to 7 th segments; C: 10 th to 12 th segments; D: 18 th and 19 th segments and epiproct; E: Hypoproct and paraprocts; F: Gonopods in situ. Scales = 1 mm." httpUri="https://zenodo.org/record/1453229/files/figure.png" pageId="2" pageNumber="247">Figures 2</figureCitation>
,
<figureCitation id="13332A2DCD06FFF2FEB5FB377A64FB3F" box="[279,293,1199,1223]" captionStart="FIGURE 3" captionStartId="4.[151,250,1557,1579]" captionTargetBox="[151,1436,481,1535]" captionTargetId="figure@4.[151,1436,481,1535]" captionTargetPageId="4" captionText="FIGURE 3. Left gonopod of Barydesmus nangaritza sp. nov. A: ectal view; B: mesal view; C: tip of telopodite, dorsal view." httpUri="https://zenodo.org/record/1453231/files/figure.png" pageId="2" pageNumber="247">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3126523CD06FFF2FF35FB6F78C4FAA0" pageId="2" pageNumber="247" type="materials_examined">
<paragraph id="8BB736A8CD06FFF2FF35FB6F78C4FAA0" blockId="2.[151,1437,1271,2016]" pageId="2" pageNumber="247">
<materialsCitation id="3B603CF5CD06FFF2FF35FB6F78C4FAA0" ID-GBIF-Occurrence="1915274020" collectingDate="05-10-25" collectingDateMax="2016-12-08" collectingDateMin="05-10-25" collectionCode="QCAZI" collectorName="A. Sanchez" country="Ecuador" county="Museo de Zoologia de la Pontificia Universidad Catolica" elevation="1040" latitude="-4.2491665" location="Las Orquideas" longLatPrecision="21" longitude="-78.66389" municipality="Canton Nangaritza" pageId="2" pageNumber="247" specimenCount="1" stateProvince="Zamora-Chinchipe" typeStatus="holotype">
<emphasis id="B97CEABACD06FFF2FF35FB6F7A47FAE8" bold="true" box="[151,262,1271,1296]" pageId="2" pageNumber="247">Material.</emphasis>
Male
<typeStatus id="54B3880ACD06FFF2FEEDFB6F7AF0FAE8" box="[335,433,1271,1296]" pageId="2" pageNumber="247" type="holotype">holotype</typeStatus>
(
<collectingDate id="EFF2E980CD06FFF2FE63FB6F7956FAE8" box="[449,535,1271,1296]" pageId="2" pageNumber="247" value="05-10-25">
<date id="FFB61068CD06FFF2FE63FB6F7956FAE8" box="[449,535,1271,1296]" pageId="2" pageNumber="247" value="05-10-25">251005</date>
</collectingDate>
<collectionCode id="ED19AE6DCD06FFF2FDBCFB6F7937FAF7" box="[542,630,1271,1295]" pageId="2" pageNumber="247">QCAZI</collectionCode>
,
<collectingCounty id="62D64E24CD06FFF2FD21FB607E46FAE8" box="[643,1287,1271,1296]" pageId="2" pageNumber="247">Museo de Zoología de la Pontificia Universidad Católica</collectingCounty>
del
<collectingCountry id="F31F7638CD06FFF2FA9BFB6F7ED9FAE8" box="[1337,1432,1271,1296]" name="Ecuador" pageId="2" pageNumber="247">Ecuador</collectingCountry>
, sección de invertebrados) from
<collectingCountry id="F31F7638CD06FFF2FE5BFA837919FACC" box="[505,600,1307,1332]" name="Ecuador" pageId="2" pageNumber="247">Ecuador</collectingCountry>
, Provincia
<collectingRegion id="49CCF84ACD06FFF2FD77FA8378EDFACC" box="[725,940,1307,1332]" country="Ecuador" name="Zamora-Chinchipe" pageId="2" pageNumber="247">Zamora-Chinchipe</collectingRegion>
,
<collectingMunicipality id="6BD3ACD2CD06FFF2FC15FA837FCCFACC" box="[951,1165,1307,1332]" pageId="2" pageNumber="247">Cantón Nangaritza</collectingMunicipality>
,
<location id="8ED76073CD06FFF2FB3AFA847E7BFACC" LSID="urn:lsid:plazi:treatment:03A187BECD06FFF5FF35FB147AA6FE0D:8ED76073CD06FFF2FB3AFA847E7BFACC" box="[1176,1338,1307,1332]" country="Ecuador" county="Museo de Zoologia de la Pontificia Universidad Catolica" latitude="-4.2491665" longLatPrecision="21" longitude="-78.66389" municipality="Canton Nangaritza" name="Las Orquideas" pageId="2" pageNumber="247" stateProvince="Zamora-Chinchipe">Las Orquideas</location>
(
<figureCitation id="13332A2DCD06FFF2FAEBFA837ECAFACB" box="[1353,1419,1307,1332]" captionStart="FIGURE 1" captionStartId="2.[151,250,992,1014]" captionTargetBox="[156,1436,195,968]" captionTargetId="figure@2.[151,1436,195,968]" captionTargetPageId="2" captionText="FIGURE 1. Location of the type locality of Barydesmus nangaritza sp. nov. and view of the habitat." httpUri="https://zenodo.org/record/1453227/files/figure.png" pageId="2" pageNumber="247">Fig. 1</figureCitation>
),
<elevation id="0025D19BCD06FFF2FF35FAA77BAFFAAF" box="[151,238,1343,1367]" metricMagnitude="3" metricUnit="m" metricValue="1.04" pageId="2" pageNumber="247" unit="m" value="1040.0">
<quantity id="4CF09B4DCD06FFF2FF35FAA77BAFFAAF" box="[151,238,1343,1367]" metricMagnitude="3" metricUnit="m" metricValue="1.04" pageId="2" pageNumber="247" unit="m" value="1040.0">1040 m</quantity>
</elevation>
,
<geoCoordinate id="EE3C506FCD06FFF2FF58FAA77A26FAA0" box="[250,359,1343,1368]" direction="south" orientation="latitude" pageId="2" pageNumber="247" precision="15" value="-4.2491665">4°14'57''S</geoCoordinate>
<geoCoordinate id="EE3C506FCD06FFF2FECCFAA77AB4FAA0" box="[366,501,1343,1368]" direction="west" orientation="longitude" pageId="2" pageNumber="247" precision="15" value="-78.66389">78°39'50''W</geoCoordinate>
,
<collectingDate id="EFF2E980CD06FFF2FDA2FAA7798BFAA0" box="[512,714,1343,1368]" pageId="2" pageNumber="247" value="2016-12-08">
<date id="FFB61068CD06FFF2FDA2FAA7798BFAA0" box="[512,714,1343,1368]" pageId="2" pageNumber="247" value="2016-12-08">8 December 2016</date>
</collectingDate>
,
<collectorName id="26FD537ECD06FFF2FD77FAD87815FAA0" box="[725,852,1343,1368]" pageId="2" pageNumber="247">A. Sánchez</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3126523CD06FFF2FF65FAFB7E0FFA83" box="[199,1358,1379,1404]" pageId="2" pageNumber="247" type="etymology">
<paragraph id="8BB736A8CD06FFF2FF65FAFB7E0FFA83" blockId="2.[151,1437,1271,2016]" box="[199,1358,1379,1404]" pageId="2" pageNumber="247">
<emphasis id="B97CEABACD06FFF2FF65FAFB7A0FFA84" bold="true" box="[199,334,1379,1404]" pageId="2" pageNumber="247">Etymology.</emphasis>
The species name, a noun in apposition, refers to the region where the
<typeStatus id="54B3880ACD06FFF2FBC9FAFC7FDAFA84" box="[1131,1179,1380,1404]" pageId="2" pageNumber="247">type</typeStatus>
locality occurs.
</paragraph>
</subSubSection>
<subSubSection id="C3126523CD06FFF2FF65FA1F7F01FA3C" pageId="2" pageNumber="247" type="diagnosis">
<paragraph id="8BB736A8CD06FFF2FF65FA1F7F01FA3C" blockId="2.[151,1437,1271,2016]" pageId="2" pageNumber="247">
<emphasis id="B97CEABACD06FFF2FF65FA1F7A00FA58" bold="true" box="[199,321,1415,1440]" pageId="2" pageNumber="247">Diagnosis.</emphasis>
This new species differs from any other species in the genus by the deeply notched paraterga (
<figureCitation id="13332A2DCD06FFF2FACDFA1F7BE6FA3B" captionStart="FIGURE 2" captionStartId="3.[151,250,1790,1812]" captionTargetBox="[157,1426,734,1762]" captionTargetId="figure@3.[151,1436,733,1766]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Barydesmus nangaritza sp. nov. A: Head and collum; B: 5 th to 7 th segments; C: 10 th to 12 th segments; D: 18 th and 19 th segments and epiproct; E: Hypoproct and paraprocts; F: Gonopods in situ. Scales = 1 mm." httpUri="https://zenodo.org/record/1453229/files/figure.png" pageId="2" pageNumber="247">Fig. 2</figureCitation>
), giving the appearance of having two pairs of lateral teeth in each trunk segment.
</paragraph>
</subSubSection>
<subSubSection id="C3126523CD06FFF4FF65FA577F52FE55" lastPageId="4" lastPageNumber="249" pageId="2" pageNumber="247" type="description">
<paragraph id="8BB736A8CD06FFF2FF65FA5778CEF9D7" blockId="2.[151,1437,1271,2016]" pageId="2" pageNumber="247">
<emphasis id="B97CEABACD06FFF2FF65FA577A18FA10" bold="true" box="[199,345,1487,1512]" pageId="2" pageNumber="247">Description.</emphasis>
Adult male
<typeStatus id="54B3880ACD06FFF2FE45FA57790DFA10" box="[487,588,1487,1512]" pageId="2" pageNumber="247" type="holotype">holotype</typeStatus>
, ca.
<quantity id="4CF09B4DCD06FFF2FD22FA577991FA1F" box="[640,720,1487,1512]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="247" unit="mm" value="60.0">60 mm</quantity>
long, width, at collum =
<quantity id="4CF09B4DCD06FFF2FC53FA577F0BFA1F" box="[1009,1098,1487,1511]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="2" pageNumber="247" unit="mm" value="4.8">4.8 mm</quantity>
, at 2nd segment =
<quantity id="4CF09B4DCD06FFF2FA80FA577E3AFA1F" box="[1314,1403,1487,1512]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="247" unit="mm" value="7.5">7.5 mm</quantity>
, at 10th segment =
<quantity id="4CF09B4DCD06FFF2FEF4FA6B7AF3F9F3" box="[342,434,1523,1547]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="2" pageNumber="247" unit="mm" value="8.7">8.7 mm</quantity>
, at 19th segment =
<quantity id="4CF09B4DCD06FFF2FD02FA6B79BCF9F3" box="[672,765,1523,1547]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="247" unit="mm" value="3.8">3.8 mm</quantity>
. General habitus slender, with W/L ratio ca. 14.5%, with convex dorsum and paranota set high and slightly upturned (
<figureCitation id="13332A2DCD06FFF2FC9FF98F78C3F9D7" box="[829,898,1559,1584]" captionStart="FIGURE 2" captionStartId="3.[151,250,1790,1812]" captionTargetBox="[157,1426,734,1762]" captionTargetId="figure@3.[151,1436,733,1766]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Barydesmus nangaritza sp. nov. A: Head and collum; B: 5 th to 7 th segments; C: 10 th to 12 th segments; D: 18 th and 19 th segments and epiproct; E: Hypoproct and paraprocts; F: Gonopods in situ. Scales = 1 mm." httpUri="https://zenodo.org/record/1453229/files/figure.png" pageId="2" pageNumber="247">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BB736A8CD06FFF2FF65F9A37A4AF964" blockId="2.[151,1437,1271,2016]" pageId="2" pageNumber="247">Color: Head, body, legs and antennae black; collum and metaterga decorated with transverse rows of large, rounded, grey tubercles. Paraterga covered with smaller, greyish tubercles. Margins of paranota greyish. Labrum yellowish.</paragraph>
<paragraph id="8BB736A8CD06FFF2FF65F93F7BBFF86F" blockId="2.[151,1437,1271,2016]" pageId="2" pageNumber="247">
Head (
<figureCitation id="13332A2DCD06FFF2FEB3F93F7A21F947" box="[273,352,1703,1728]" captionStart="FIGURE 2" captionStartId="3.[151,250,1790,1812]" captionTargetBox="[157,1426,734,1762]" captionTargetId="figure@3.[151,1436,733,1766]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Barydesmus nangaritza sp. nov. A: Head and collum; B: 5 th to 7 th segments; C: 10 th to 12 th segments; D: 18 th and 19 th segments and epiproct; E: Hypoproct and paraprocts; F: Gonopods in situ. Scales = 1 mm." httpUri="https://zenodo.org/record/1453229/files/figure.png" pageId="2" pageNumber="247">Fig. 2a</figureCitation>
) granulated, except for labrum and a semicircular area of the clypeus with coriaceous texture just above labrum, and glabrous, except for a few large setae on gnathal lobes (18+
<quantity id="4CF09B4DCD06FFF2FBBFF9537F16F91C" box="[1053,1111,1739,1764]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="2" pageNumber="247" unit="in" value="18.0">18 in</quantity>
frontal area and a few more irregularly dispersed elsewhere), and a few compound setae (7+7 labral, 4+4 prelabral, 2+2 clypeal). Labrum tridentate, deeply offset from level of clypeus. Epicranial sulcus deep. Antennae robust, extending up to the third segment when stretched backwards. Antennomeres covered with long setae. Relative lengths of antennomeres 6&gt;3&gt;5=4=2&gt;1&gt;7. Four apical sensory cones. Antennomere 4 of left antenna with a teratological extra antennomere (
<figureCitation id="13332A2DCD06FFF2FF3DF8E77BB0F86F" box="[159,241,1919,1944]" captionStart="FIGURE 2" captionStartId="3.[151,250,1790,1812]" captionTargetBox="[157,1426,734,1762]" captionTargetId="figure@3.[151,1436,733,1766]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Barydesmus nangaritza sp. nov. A: Head and collum; B: 5 th to 7 th segments; C: 10 th to 12 th segments; D: 18 th and 19 th segments and epiproct; E: Hypoproct and paraprocts; F: Gonopods in situ. Scales = 1 mm." httpUri="https://zenodo.org/record/1453229/files/figure.png" pageId="2" pageNumber="247">Fig. 2a</figureCitation>
).
</paragraph>
<paragraph id="8BB736A8CD06FFF3FF65F83B7E1FFF2D" blockId="2.[151,1437,1271,2016]" lastBlockId="3.[151,1437,151,680]" lastPageId="3" lastPageNumber="248" pageId="2" pageNumber="247">Collum glabrous, elevated caudad, with frontal and posterior margins almost straight; paranota progressively narrowed laterad, broadly rounded, but margin with 7 small indentations. Surface irregularly granulated, less marked in the middle, and with one anterior and one posterior transverse row of medium to large grey tubercles, the former with 9 tubercles, the latter with 10. Paraterga also covered with granulations and small grey tubercles.</paragraph>
<paragraph id="8BB736A8CD07FFF3FF65FF477977FE55" blockId="3.[151,1437,151,680]" pageId="3" pageNumber="248">
Second segment also clearly elevated caudad, following segments (
<figureCitation id="13332A2DCD07FFF3FC1AFF477F62FF00" box="[952,1059,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1790,1812]" captionTargetBox="[157,1426,734,1762]" captionTargetId="figure@3.[151,1436,733,1766]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Barydesmus nangaritza sp. nov. A: Head and collum; B: 5 th to 7 th segments; C: 10 th to 12 th segments; D: 18 th and 19 th segments and epiproct; E: Hypoproct and paraprocts; F: Gonopods in situ. Scales = 1 mm." httpUri="https://zenodo.org/record/1453229/files/figure.png" pageId="3" pageNumber="248">Fig. 2b, c</figureCitation>
) increasingly levelled. Surface of postcollum metaterga without setae, granulated in posterior third and next to paranota, the rest being coriaceous. A conspicuous transverse row of 8 large grey tubercles, rarely with 9 or 10, near caudal margin. These tubercles present in every segment, but increasingly reduced towards terminal segments. There are also one (segments 2–9) or two (segments 10–19) additional irregular transverse rows of much lower, weaker, dark tubercles. Prozonae uniformly and very finely punctured.
</paragraph>
<paragraph id="8BB736A8CD07FFF3FF65FE207A75FD50" blockId="3.[151,1437,151,680]" pageId="3" pageNumber="248">
Paranota (
<figureCitation id="13332A2DCD07FFF3FE9AFE2F7AFBFE28" box="[312,442,439,464]" captionStart="FIGURE 2" captionStartId="3.[151,250,1790,1812]" captionTargetBox="[157,1426,734,1762]" captionTargetId="figure@3.[151,1436,733,1766]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Barydesmus nangaritza sp. nov. A: Head and collum; B: 5 th to 7 th segments; C: 10 th to 12 th segments; D: 18 th and 19 th segments and epiproct; E: Hypoproct and paraprocts; F: Gonopods in situ. Scales = 1 mm." httpUri="https://zenodo.org/record/1453229/files/figure.png" pageId="3" pageNumber="248">Fig 2a, b, d</figureCitation>
) well-developed, profusely covered dorsally with small to medium-sized grey tubercles. Posterior corners of paranota 2 projected laterad; both anterior and posterior corners of following paranota projected laterad, thus looking deeply notched laterally and forming two large teeth in each paratergum, posterior tooth being more slender than anterior one. Notch smaller in segment 18 and vestigial in segment 19. Anterior and posterior margins finely serrulated, the former with a defined whitish rim. Lateral margins also covered with irregularly sized grey tubercles. Ventral surface granulated. Pore formula normal, ozopores opening dorsally at base of notch.
</paragraph>
<caption id="DF776620CD07FFF3FF35F9667FEDF8CD" httpUri="https://zenodo.org/record/1453229/files/figure.png" pageId="3" pageNumber="248" startId="3.[151,250,1790,1812]" targetBox="[157,1426,734,1762]" targetPageId="3">
<paragraph id="8BB736A8CD07FFF3FF35F9667FEDF8CD" blockId="3.[151,1436,1787,1845]" pageId="3" pageNumber="248">
<emphasis id="B97CEABACD07FFF3FF35F9667A53F8EB" bold="true" box="[151,274,1790,1812]" pageId="3" pageNumber="248">FIGURE 2.</emphasis>
Habitus of
<taxonomicName id="4C084D2BCD07FFF3FE2AF9667939F8EC" authority="Recuero &amp; Sánchez-Vialas, 2018" authorityName="Recuero &amp; Sánchez-Vialas" authorityYear="2018" box="[392,632,1790,1812]" class="Diplopoda" family="Platyrhacidae" genus="Barydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="nangaritza" status="sp. nov.">
<emphasis id="B97CEABACD07FFF3FE2AF9667939F8EC" box="[392,632,1790,1812]" italics="true" pageId="3" pageNumber="248">Barydesmus nangaritza</emphasis>
</taxonomicName>
<emphasis id="B97CEABACD07FFF3FDDFF96679A7F8EB" bold="true" box="[637,742,1790,1811]" pageId="3" pageNumber="248">
<taxonomicNameLabel id="A24F57C1CD07FFF3FDDFF966798CF8EB" box="[637,717,1790,1811]" pageId="3" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
: Head and collum;
<emphasis id="B97CEABACD07FFF3FC0EF96678FCF8EB" bold="true" box="[940,957,1790,1811]" pageId="3" pageNumber="248">B</emphasis>
: 5th to 7th segments;
<emphasis id="B97CEABACD07FFF3FB28F9667FDDF8EC" bold="true" box="[1162,1180,1790,1812]" pageId="3" pageNumber="248">C</emphasis>
: 10th to 12th segments;
<emphasis id="B97CEABACD07FFF3FA21F9667ED4F8EB" bold="true" box="[1411,1429,1790,1811]" pageId="3" pageNumber="248">D</emphasis>
: 18th and 19th segments and epiproct;
<emphasis id="B97CEABACD07FFF3FDA4F8877956F8CC" bold="true" box="[518,535,1823,1844]" pageId="3" pageNumber="248">E</emphasis>
: Hypoproct and paraprocts;
<emphasis id="B97CEABACD07FFF3FC9BF8877809F8CC" bold="true" box="[825,840,1823,1844]" pageId="3" pageNumber="248">F</emphasis>
: Gonopods in situ. Scales = 1 mm.
</paragraph>
</caption>
<paragraph id="8BB736A8CD07FFF3FF65F8C47984F819" blockId="3.[151,1436,1884,2017]" pageId="3" pageNumber="248">
Epiproct (
<figureCitation id="13332A2DCD07FFF3FE9AF8C47ACBF88D" box="[312,394,1884,1909]" captionStart="FIGURE 2" captionStartId="3.[151,250,1790,1812]" captionTargetBox="[157,1426,734,1762]" captionTargetId="figure@3.[151,1436,733,1766]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Barydesmus nangaritza sp. nov. A: Head and collum; B: 5 th to 7 th segments; C: 10 th to 12 th segments; D: 18 th and 19 th segments and epiproct; E: Hypoproct and paraprocts; F: Gonopods in situ. Scales = 1 mm." httpUri="https://zenodo.org/record/1453229/files/figure.png" pageId="3" pageNumber="248">Fig. 2d</figureCitation>
) granulated, flattened and rounded, fringed by four long setae (three of them missing in the preserved
<typeStatus id="54B3880ACD07FFF3FEB2F8187A37F861" box="[272,374,1920,1945]" pageId="3" pageNumber="248" type="holotype">holotype</typeStatus>
), with two additional dorsal setae (one missing). Paraprocts (
<figureCitation id="13332A2DCD07FFF3FBECF8187FE2F861" box="[1102,1187,1920,1945]" captionStart="FIGURE 2" captionStartId="3.[151,250,1790,1812]" captionTargetBox="[157,1426,734,1762]" captionTargetId="figure@3.[151,1436,733,1766]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Barydesmus nangaritza sp. nov. A: Head and collum; B: 5 th to 7 th segments; C: 10 th to 12 th segments; D: 18 th and 19 th segments and epiproct; E: Hypoproct and paraprocts; F: Gonopods in situ. Scales = 1 mm." httpUri="https://zenodo.org/record/1453229/files/figure.png" pageId="3" pageNumber="248">Fig. 2e</figureCitation>
) slightly convex and granulated, with two pairs of setigerous tubercules. Hypoproct (
<figureCitation id="13332A2DCD07FFF3FCD1F83C7884F845" box="[883,965,1956,1981]" captionStart="FIGURE 2" captionStartId="3.[151,250,1790,1812]" captionTargetBox="[157,1426,734,1762]" captionTargetId="figure@3.[151,1436,733,1766]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Barydesmus nangaritza sp. nov. A: Head and collum; B: 5 th to 7 th segments; C: 10 th to 12 th segments; D: 18 th and 19 th segments and epiproct; E: Hypoproct and paraprocts; F: Gonopods in situ. Scales = 1 mm." httpUri="https://zenodo.org/record/1453229/files/figure.png" pageId="3" pageNumber="248">Fig. 2e</figureCitation>
) small, longer than wide, granulated and with two large paramedian processes at rear edge.
</paragraph>
<paragraph id="8BB736A8CD00FFF4FF65FF0F7F79FF00" blockId="4.[151,1436,151,429]" pageId="4" pageNumber="249">
Sternites elevated, without processes, with rugose surface and sparse long setae, more abundant at front margins of segments 6 and 8.
<collectingRegion id="49CCF84ACD00FFF4FE4AFF247964FF2D" box="[488,549,188,213]" country="Haiti" name="Sud-Est" pageId="4" pageNumber="249">Sides</collectingRegion>
of metazonae finely granulated. Legs uniformly covered with erect setae, with no traces of spines or processes. Gonapophyses opening on swollen, blunt coxae 2.
</paragraph>
<paragraph id="8BB736A8CD00FFF4FF65FE9C7F52FE55" blockId="4.[151,1436,151,429]" pageId="4" pageNumber="249">
Gonopod aperture (
<figureCitation id="13332A2DCD00FFF4FE04FE9C7AB7FEE5" box="[422,502,260,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1790,1812]" captionTargetBox="[157,1426,734,1762]" captionTargetId="figure@3.[151,1436,733,1766]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Barydesmus nangaritza sp. nov. A: Head and collum; B: 5 th to 7 th segments; C: 10 th to 12 th segments; D: 18 th and 19 th segments and epiproct; E: Hypoproct and paraprocts; F: Gonopods in situ. Scales = 1 mm." httpUri="https://zenodo.org/record/1453229/files/figure.png" pageId="4" pageNumber="249">Fig. 2f</figureCitation>
) transversely oval, fitting closely with width of gonopodal coxae. Posterior edge elevated. Gonopods (
<figureCitation id="13332A2DCD00FFF4FE28FEBF7A8EFEB8" box="[394,463,295,320]" captionStart="FIGURE 3" captionStartId="4.[151,250,1557,1579]" captionTargetBox="[151,1436,481,1535]" captionTargetId="figure@4.[151,1436,481,1535]" captionTargetPageId="4" captionText="FIGURE 3. Left gonopod of Barydesmus nangaritza sp. nov. A: ectal view; B: mesal view; C: tip of telopodite, dorsal view." httpUri="https://zenodo.org/record/1453231/files/figure.png" pageId="4" pageNumber="249">Fig. 3</figureCitation>
) slender, extended forward between legs of segment 6, coxa brown in external half, white inside and around articulation, with no trace of setae; telopodite brown over most of its lower half, yellow in distal part, except at tip which is brown again. Base of telopodite with dense setation, this part shorter than glabrous one. Both solenomere and apical lamina (tibiotarsus) acute and directed dorsad.
</paragraph>
</subSubSection>
<caption id="DF776620CD00FFF4FF35F98D7ECDF9D2" box="[151,1420,1556,1579]" httpUri="https://zenodo.org/record/1453231/files/figure.png" pageId="4" pageNumber="249" startId="4.[151,250,1557,1579]" targetBox="[151,1436,481,1535]" targetPageId="4">
<paragraph id="8BB736A8CD00FFF4FF35F98D7ECDF9D2" blockId="4.[151,1420,1556,1579]" box="[151,1420,1556,1579]" pageId="4" pageNumber="249">
<emphasis id="B97CEABACD00FFF4FF35F98D7A53F9D2" bold="true" box="[151,274,1557,1579]" pageId="4" pageNumber="249">FIGURE 3.</emphasis>
Left gonopod of
<taxonomicName id="4C084D2BCD00FFF4FE66F98C79F5F9D3" authority="Recuero &amp; Sánchez-Vialas, 2018" authorityName="Recuero &amp; Sánchez-Vialas" authorityYear="2018" box="[452,692,1556,1579]" class="Diplopoda" family="Platyrhacidae" genus="Barydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="nangaritza" status="sp. nov.">
<emphasis id="B97CEABACD00FFF4FE66F98C79F5F9D3" box="[452,692,1556,1579]" italics="true" pageId="4" pageNumber="249">Barydesmus nangaritza</emphasis>
</taxonomicName>
<emphasis id="B97CEABACD00FFF4FD18F98D7862F9D2" bold="true" box="[698,803,1557,1578]" pageId="4" pageNumber="249">
<taxonomicNameLabel id="A24F57C1CD00FFF4FD18F98D784BF9D2" box="[698,778,1557,1578]" pageId="4" pageNumber="249" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
: ectal view;
<emphasis id="B97CEABACD00FFF4FC07F98D78F7F9D2" bold="true" box="[933,950,1557,1578]" pageId="4" pageNumber="249">B</emphasis>
: mesal view;
<emphasis id="B97CEABACD00FFF4FBE6F98D7F17F9D3" bold="true" box="[1092,1110,1557,1579]" pageId="4" pageNumber="249">C</emphasis>
: tip of telopodite, dorsal view.
</paragraph>
</caption>
<subSubSection id="C3126523CD00FFF5FF65F9CF7AA6FE0D" lastPageId="5" lastPageNumber="250" pageId="4" pageNumber="249" type="discussion">
<paragraph id="8BB736A8CD00FFF4FF65F9CF7F44F820" blockId="4.[151,1436,1623,2008]" pageId="4" pageNumber="249">
<emphasis id="B97CEABACD00FFF4FF65F9CF7A0EF988" bold="true" box="[199,335,1623,1648]" pageId="4" pageNumber="249">Comments.</emphasis>
The form of the paranota in
<taxonomicName id="4C084D2BCD00FFF4FD12F9CF78FFF997" authority="Recuero &amp; Sánchez-Vialas, 2018" authorityName="Recuero &amp; Sánchez-Vialas" authorityYear="2018" box="[688,958,1623,1648]" class="Diplopoda" family="Platyrhacidae" genus="Barydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="nangaritza" status="sp. nov.">
<emphasis id="B97CEABACD00FFF4FD12F9CF78FFF997" box="[688,958,1623,1648]" italics="true" pageId="4" pageNumber="249">Barydesmus nangaritza</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57C1CD00FFF4FC6BF9CF7F66F988" box="[969,1063,1623,1648]" pageId="4" pageNumber="249" rank="species">
<emphasis id="B97CEABACD00FFF4FC6BF9CF7F66F988" bold="true" box="[969,1063,1623,1648]" pageId="4" pageNumber="249">sp. nov.</emphasis>
</taxonomicNameLabel>
is unique among the American
<taxonomicName id="4C084D2BCD00FFF4FF35F9E37A72F96C" authorityName="Pocock" authorityYear="1895" box="[151,307,1659,1684]" class="Diplopoda" family="Platyrhacidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="249" phylum="Arthropoda" rank="family">Platyrhacidae</taxonomicName>
. Checking all descriptions and published figures, only
<taxonomicName id="4C084D2BCD00FFF4FC38F9E37E4CF96C" authority="(Cook, 1896 a)" authorityName="Recuero &amp; Sánchez-Vialas" authorityYear="2018" baseAuthorityName="Cook" baseAuthorityYear="1896" box="[922,1293,1659,1684]" class="Diplopoda" family="Platyrhacidae" genus="Barydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="exsul">
<emphasis id="B97CEABACD00FFF4FC38F9E37F22F96C" box="[922,1123,1659,1684]" italics="true" pageId="4" pageNumber="249">Barydesmus exsul</emphasis>
(
<bibRefCitation id="EF994B59CD00FFF4FBD0F9E37E45F96C" box="[1138,1284,1659,1684]" pageId="4" pageNumber="249" refId="ref15722" refString="Cook, O. F. (1896 a) New american Platyrhacidae. Brandtia, 12, 51 - 54." type="journal article">Cook, 1896a</bibRefCitation>
)
</taxonomicName>
leaves some doubt about its possible similarity to the paranota of
<taxonomicName id="4C084D2BCD00FFF4FD45F93878C1F94F" authority="Recuero &amp; Sánchez-Vialas, 2018" authorityName="Recuero &amp; Sánchez-Vialas" authorityYear="2018" box="[743,896,1696,1719]" class="Diplopoda" family="Platyrhacidae" genus="Barydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="nangaritza" status="sp. nov.">
<emphasis id="B97CEABACD00FFF4FD45F93878C1F94F" box="[743,896,1696,1719]" italics="true" pageId="4" pageNumber="249">B. nangaritza</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57C1CD00FFF4FC25F90778A0F940" box="[903,993,1695,1720]" pageId="4" pageNumber="249" rank="species">
<emphasis id="B97CEABACD00FFF4FC25F90778A0F940" bold="true" box="[903,993,1695,1720]" pageId="4" pageNumber="249">sp. nov.</emphasis>
</taxonomicNameLabel>
, as in the original description it is said: &quot;lateral carinae usually deeply excised opposite to the pore&quot; (
<bibRefCitation id="EF994B59CD00FFF4FCEAF95B7F43F924" box="[840,1026,1731,1756]" pageId="4" pageNumber="249" refId="ref15722" refString="Cook, O. F. (1896 a) New american Platyrhacidae. Brandtia, 12, 51 - 54." type="journal article">Cook 1896a: 54</bibRefCitation>
). We have examined photographs of the
<typeStatus id="54B3880ACD00FFF4FF62F9707BB1F8F8" box="[192,240,1768,1792]" pageId="4" pageNumber="249">type</typeStatus>
specimen of
<taxonomicName id="4C084D2BCD00FFF4FE27F9707A9FF8F8" authorityName="Recuero &amp; Sánchez-Vialas" authorityYear="2018" baseAuthorityName="Cook" baseAuthorityYear="1896" box="[389,478,1767,1792]" class="Diplopoda" family="Platyrhacidae" genus="Barydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="exsul">
<emphasis id="B97CEABACD00FFF4FE27F9707A9FF8F8" box="[389,478,1767,1792]" italics="true" pageId="4" pageNumber="249">B. exsul</emphasis>
</taxonomicName>
and the form of the paranota is highly variable among segments. Only paranota 5 and 12 appear clearly excised opposite to the pore (even if not as deeply as in
<taxonomicName id="4C084D2BCD00FFF4FC65F8947F1EF8DB" authority="Recuero &amp; Sánchez-Vialas, 2018" authorityName="Recuero &amp; Sánchez-Vialas" authorityYear="2018" box="[967,1119,1804,1827]" class="Diplopoda" family="Platyrhacidae" genus="Barydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="nangaritza" status="sp. nov.">
<emphasis id="B97CEABACD00FFF4FC65F8947F1EF8DB" box="[967,1119,1804,1827]" italics="true" pageId="4" pageNumber="249">B. nangaritza</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57C1CD00FFF4FBC7F8937FFFF8DC" box="[1125,1214,1803,1828]" pageId="4" pageNumber="249" rank="species">
<emphasis id="B97CEABACD00FFF4FBC7F8937FFFF8DC" bold="true" box="[1125,1214,1803,1828]" pageId="4" pageNumber="249">sp. nov.</emphasis>
</taxonomicNameLabel>
), but most paranota of
<taxonomicName id="4C084D2BCD00FFF4FF14F8A87A4EF8B0" authorityName="Recuero &amp; Sánchez-Vialas" authorityYear="2018" baseAuthorityName="Cook" baseAuthorityYear="1896" box="[182,271,1839,1864]" class="Diplopoda" family="Platyrhacidae" genus="Barydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="exsul">
<emphasis id="B97CEABACD00FFF4FF14F8A87A4EF8B0" box="[182,271,1839,1864]" italics="true" pageId="4" pageNumber="249">B. exsul</emphasis>
</taxonomicName>
have 3–4 blunt teeth providing the rough, irregular external margins typical of several other species of the family. Gonopod structure among
<taxonomicName id="4C084D2BCD00FFF4FDEBF8CB7993F894" authorityName="Cook" authorityYear="1896" box="[585,722,1875,1900]" class="Diplopoda" family="Platyrhacidae" genus="Barydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEABACD00FFF4FDEBF8CB7993F894" box="[585,722,1875,1900]" italics="true" pageId="4" pageNumber="249">Barydesmus</emphasis>
</taxonomicName>
species, as in other
<taxonomicName id="4C084D2BCD00FFF4FC1EF8CB7F16F894" authorityName="Pocock" authorityYear="1895" box="[956,1111,1875,1900]" class="Diplopoda" family="Platyrhacidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="249" phylum="Arthropoda" rank="family">Platyrhacidae</taxonomicName>
, is highly homogeneous and peripheral characters should be considered as major diagnostic features in this group (
<bibRefCitation id="EF994B59CD00FFF4FB3DF8EF7ED6F868" box="[1183,1431,1911,1936]" pageId="4" pageNumber="249" refId="ref16434" refString="Hoffman, R. L., Martinez-Torres, D. &amp; Florez D., E. (2011) A new Colombian species in the milliped genus Psammodesmus, symbiotic host for bryophytes (Polydesmida: Platyrhacidae). Zootaxa, 3015, 52 - 60." type="journal article">
Hoffman
<emphasis id="B97CEABACD00FFF4FAB0F8E07E08F868" box="[1298,1353,1911,1936]" italics="true" pageId="4" pageNumber="249">et al</emphasis>
. 2011
</bibRefCitation>
;
<bibRefCitation id="EF994B59CD00FFF4FF35F8047AD3F84C" box="[151,402,1947,1972]" pageId="4" pageNumber="249" refId="ref17075" refString="Martinez-Torres, D. (2016) Revision taxonomica y distribucion geografica de los milpies del genero Psammodesmus Cook, 1896 (Diplopoda: Polydesmida: Platyrhacidae). MSc Thesis in Biology, Universidad Nacional de Colombia, Facultad de Ciencias, Departamento de Biologia, Bogota D. C., Colombia, pp. 1 - 182." type="book chapter">Martínez-Torres 2016</bibRefCitation>
). Given the available information, it is not feasible to postulate solid hypotheses regarding the affinities among species of
<taxonomicName id="4C084D2BCD00FFF4FE51F827793DF820" authorityName="Cook" authorityYear="1896" box="[499,636,1983,2008]" class="Diplopoda" family="Platyrhacidae" genus="Barydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEABACD00FFF4FE51F827793DF820" box="[499,636,1983,2008]" italics="true" pageId="4" pageNumber="249">Barydesmus</emphasis>
</taxonomicName>
without a full review of the genus.
</paragraph>
<paragraph id="8BB736A8CD01FFF5FF65FF0F7E7BFEB8" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="250">
<taxonomicName id="4C084D2BCD01FFF5FF65FF0F7A91FF48" authority="Recuero &amp; Sánchez-Vialas, 2018" authorityName="Recuero &amp; Sánchez-Vialas" authorityYear="2018" box="[199,464,151,176]" class="Diplopoda" family="Platyrhacidae" genus="Barydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="nangaritza" status="sp. nov.">
<emphasis id="B97CEABACD01FFF5FF65FF0F7A91FF48" box="[199,464,151,176]" italics="true" pageId="5" pageNumber="250">Barydesmus nangaritza</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57C1CD01FFF5FE74FF0F796EFF48" box="[470,559,151,176]" pageId="5" pageNumber="250" rank="species">
<emphasis id="B97CEABACD01FFF5FE74FF0F796EFF48" bold="true" box="[470,559,151,176]" pageId="5" pageNumber="250">sp. nov.</emphasis>
</taxonomicNameLabel>
comes from the Nangaritza River Basin, an area characterized by the presence of isolated sandstone mountains named &quot;tepuyes&quot;, and a humid subtropical climate (
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). The habitat has a sandy soil covered by a thin layer of leaf litter, with dispersed stones and rotten logs. Most trees are covered with dense epiphytic vegetation. The
<typeStatus id="54B3880ACD01FFF5FD54FE9C7817FEE5" box="[758,854,260,285]" pageId="5" pageNumber="250" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C084D2BCD01FFF5FCD9FE9D7F50FEE4" authorityName="Recuero &amp; Sánchez-Vialas" authorityYear="2018" box="[891,1041,261,284]" class="Diplopoda" family="Platyrhacidae" genus="Barydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="nangaritza">
<emphasis id="B97CEABACD01FFF5FCD9FE9D7F50FEE4" box="[891,1041,261,284]" italics="true" pageId="5" pageNumber="250">B. nangaritza</emphasis>
</taxonomicName>
was found active by night, walking on a vertical trunk of a living tree, in a dense piedmont forest located on the eastern slope of a tepuy (
<figureCitation id="13332A2DCD01FFF5FB4BFEBF7E68FEB8" box="[1257,1321,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,992,1014]" captionTargetBox="[156,1436,195,968]" captionTargetId="figure@2.[151,1436,195,968]" captionTargetPageId="2" captionText="FIGURE 1. Location of the type locality of Barydesmus nangaritza sp. nov. and view of the habitat." httpUri="https://zenodo.org/record/1453227/files/figure.png" pageId="5" pageNumber="250">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BB736A8CD01FFF5FF65FED47AA6FE0D" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="250">
The biodiversity in this region has remained largely unexplored for a long time. Recent investigations, focused on regional diversity, have resulted in the description of at least four species of Amphibia, one of Reptilia, 10 of Phasmida and 13 of
<taxonomicName id="4C084D2BCD01FFF5FE2DFE0C7896FE54" authority="(Guayasamin and Bonaccorso 2011)" baseAuthorityName="Guayasamin and Bonaccorso" baseAuthorityYear="2011" box="[399,983,404,429]" class="Insecta" family="Tettigoniidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="family">
Tettigoniidae (
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)
</taxonomicName>
. This suggests that more effort is still needed to attain a better knowledge of the biodiversity and biological history of this remarkable region and its unique geological formations.
</paragraph>
</subSubSection>
</treatment>
</document>
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