468 lines
62 KiB
XML
468 lines
62 KiB
XML
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<mods:title id="D55F6BBF7BCDD317C11696DAEA1BE1ED">The genus Asiagomphus in Vietnam, with descriptions of three new species, and first descriptions of the male of Asiagomphus auricolor (Fraser, 1926) and of the female of Asiagomphus reinhardti Kosterin & Yokoi, 2016 (Odonata: Gomphidae)</mods:title>
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<mods:namePart id="70C768038AA4309C2F68CE10B8C8E119">Kompier, Tom</mods:namePart>
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<mods:title id="4D9BB1723469EDB0C8991CC840F37C7B">Zootaxa</mods:title>
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<mods:date id="B4B445E379EBD5157F468E809B8E63C9">2018</mods:date>
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<treatment id="03C887DB607C8C1BFF67D0BC18E2F86D" ID-DOI="http://doi.org/10.5281/zenodo.6495794" ID-GBIF-Taxon="148424956" ID-Zenodo-Dep="6495794" LSID="urn:lsid:plazi:treatment:03C887DB607C8C1BFF67D0BC18E2F86D" httpUri="http://treatment.plazi.org/id/03C887DB607C8C1BFF67D0BC18E2F86D" lastPageId="26" lastPageNumber="327" pageId="25" pageNumber="326">
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<paragraph id="8BDE36CD607C8C18FF67D0BC1B9DFE22" blockId="25.[151,715,390,450]" box="[151,715,390,416]" pageId="25" pageNumber="326">
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<taxonomicName id="4C614D4E607C8C18FF67D0BC1B9DFE22" ID-CoL="H9SX" authority="(Williamson, 1907)" baseAuthorityName="Williamson" baseAuthorityYear="1907" box="[151,715,390,416]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
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<emphasis id="B915EADF607C8C18FF67D0BC1B9DFE22" bold="true" box="[151,715,390,416]" pageId="25" pageNumber="326">
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<emphasis id="B915EADF607C8C18FF67D0BC188FFE22" bold="true" box="[151,473,390,416]" italics="true" pageId="25" pageNumber="326">Asiagomphus xanthenatus</emphasis>
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(
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<bibRefCitation id="EFF04B3C607C8C18FE17D0BC1B95FE22" box="[487,707,390,416]" pageId="25" pageNumber="326" refId="ref17719" refString="Williamson, E. B. (1907) The dragonflies (Odonata) of Burma and Lower Siam. II. Subfamilies Cordulegasterinae, Chlorogomphinae, and Gomphinae. Proceedings U. S. National Museum, 33, 267 - 317. https: // doi. org / 10.5479 / si. 00963801.33 - 1571.267" type="journal article">Williamson, 1907</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BDE36CD607C8C18FF67D0931B54FE43" blockId="25.[151,715,390,450]" box="[151,514,425,450]" pageId="25" pageNumber="326">
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(
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<figureCitation id="135A2A48607C8C18FF6FD0931845FE40" box="[159,275,425,450]" captionStart="FIGURE 3" captionStartId="8.[151,250,1943,1965]" captionTargetBox="[182,1417,193,1925]" captionTargetId="figure@8.[182,1417,193,1925]" captionTargetPageId="8" captionText="FIGURE 3. Anterior and posterior hamuli [a,c,e,g,h] lateral view, [b,d,f] antero-ventral lateral view. (a–b) Asiagomphus acco, Tam Dao NP, 18-V-2014; (c–d) A. kosterini sp. nov. holotype; (e–f) A. xanthenatus Quang Nam Prov., 15-V-2016; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. kosterini paratype." httpUri="https://zenodo.org/record/1441617/files/figure.png" pageId="25" pageNumber="326">Figs. 3e–f</figureCitation>
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,
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||
<figureCitation id="135A2A48607C8C18FEEED093186CFE40" box="[286,314,425,450]" captionStart="FIGURE 5" captionStartId="10.[151,250,1939,1961]" captionTargetBox="[191,1400,193,1919]" captionTargetId="figure@10.[191,1400,193,1920]" captionTargetPageId="10" captionText="FIGURE 5. Male appendages in dorsal view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441621/files/figure.png" pageId="25" pageNumber="326">5e</figureCitation>
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,
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<figureCitation id="135A2A48607C8C18FEB6D0931835FE40" box="[326,355,425,450]" captionStart="FIGURE 6" captionStartId="11.[151,250,1945,1967]" captionTargetBox="[201,1402,193,1927]" captionTargetId="figure@11.[188,1403,193,1927]" captionTargetPageId="11" captionText="FIGURE 6. Male appendages in lateral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441623/files/figure.png" pageId="25" pageNumber="326">6e</figureCitation>
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,
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<figureCitation id="135A2A48607C8C18FE9FD09018DAFE43" box="[367,396,426,449]" captionStart="FIGURE 7" captionStartId="12.[151,250,1946,1968]" captionTargetBox="[173,1418,193,1934]" captionTargetId="figure@12.[173,1418,193,1934]" captionTargetPageId="12" captionText="FIGURE 7. Male appendages in ventral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. superciliaris paratype Huu Lien NR, 24-V-2014." httpUri="https://zenodo.org/record/1441625/files/figure.png" pageId="25" pageNumber="326">7e</figureCitation>
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,
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<figureCitation id="135A2A48607C8C18FE67D0931895FE40" box="[407,451,425,450]" captionStart="FIGURE 10" captionStartId="19.[151,250,1872,1894]" captionTargetBox="[150,1436,481,1851]" captionTargetId="figure@19.[150,1436,481,1851]" captionTargetPageId="19" captionText="FIGURE 10. Asiagomphus spp. alive in hand [a–d, g] male, [e–f, h] female. (a) A. auricolor, Bac Kan Prov., 26-VI-2014; (b) A. auricolor, Huu Lien NR, 1-V-2014; (c) A. auricolor, Huu Lien NR, 24-V-2014; (d) A. xanthenatus, Quang Nam Prov., 15-V- 2016; (e) A. auricolor, used for description; (f) A. auricolor, Huu Lien NR, 26-IV-2014; (g) A. pacificus, Cao Bang Prov., 19-V- 2015; (h) A. pacificus, Bac Kan Prov., 25-VI-2014." httpUri="https://zenodo.org/record/1441631/files/figure.png" pageId="25" pageNumber="326">10d</figureCitation>
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,
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<figureCitation id="135A2A48607C8C18FE3FD09318ACFE43" box="[463,506,425,449]" captionStart="FIGURE 12" captionStartId="21.[151,250,1586,1608]" captionTargetBox="[150,1436,193,1565]" captionTargetId="figure@21.[150,1436,193,1565]" captionTargetPageId="21" captionText="FIGURE 12. Asiagomphus spp. alive in hand, [a–e] A. reinhardti habitus, [a–b] male, [c–e] female, [f–h] male abdomen tip in dorsal view. (a) Gia Lai Prov., 19-VI-2016; (b) Lam Dong Prov., 17-V-2016; (c) Gia Lai Prov., 20-VI-2016; (d) Gia Lai Prov., 19-VI-2016; (e) female used for description; (f) A. reinhardti, Lam Dong Prov., 17-V-2016; (g) A. xanthenatus, Quang Nam Prov., 15-V-2016; (h) A. kosterini holotype." httpUri="https://zenodo.org/record/1441635/files/figure.png" pageId="25" pageNumber="326">
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</figureCitation>
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)
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</paragraph>
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<subSubSection id="C37B6546607C8C18FF67D0D01A6DFD82" box="[151,827,490,512]" pageId="25" pageNumber="326" type="description">
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<paragraph id="8BDE36CD607C8C18FF67D0D01A6DFD82" blockId="25.[151,1436,490,758]" box="[151,827,490,512]" pageId="25" pageNumber="326">
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<taxonomicName id="4C614D4E607C8C18FF67D0D01876FD82" authorityName="Asahina" authorityYear="1985" box="[151,288,490,512]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="genus">
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<emphasis id="B915EADF607C8C18FF67D0D01876FD82" box="[151,288,490,512]" italics="true" pageId="25" pageNumber="326">Asiagomphus</emphasis>
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</taxonomicName>
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||
sp.:
|
||
<bibRefCitation id="EFF04B3C607C8C18FEA0D0D018ACFD82" box="[336,506,490,512]" pageId="25" pageNumber="326" refId="ref16938" refString="Asahina, S. (1986 a) A list of the Odonata from Thailand Part XIII. Gomphidae - 1. Cho Cho, 9, 29 - 43." type="journal article">Asahina (1986a)</bibRefCitation>
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||
, pp. 29–31, figs. 1–6 [females].
|
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</paragraph>
|
||
</subSubSection>
|
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<subSubSection id="C37B6546607C8C18FF67D3331A07FD74" pageId="25" pageNumber="326" type="reference_group">
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<paragraph id="8BDE36CD607C8C18FF67D3331D97FDBC" blockId="25.[151,1436,490,758]" pageId="25" pageNumber="326">
|
||
<taxonomicName id="4C614D4E607C8C18FF67D3331B09FD9D" ID-CoL="H9SX" authority=": Asahina (1986 b)" authorityName=": Asahina" authorityYear="1986" box="[151,607,521,543]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
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<emphasis id="B915EADF607C8C18FF67D33318F4FD9D" box="[151,418,521,543]" italics="true" pageId="25" pageNumber="326">Asiagomphus xanthenatus</emphasis>
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||
:
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||
<bibRefCitation id="EFF04B3C607C8C18FE41D3331B09FD9D" box="[433,607,521,543]" pageId="25" pageNumber="326" refId="ref16969" refString="Asahina, S. (1986 b) Descriptions of two Asiagomphus species from Assam-Burma and Burma-Thailand areas (Odonata, Gomphidae). Gekkan-Mushi, 186, 9 - 14." type="journal article">Asahina (1986b)</bibRefCitation>
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||
</taxonomicName>
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||
, pp. 9–14, figs. 13–25 [description of Thai specimens];
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||
<bibRefCitation id="EFF04B3C607C8C18FB58D333185AFDBC" pageId="25" pageNumber="326" refId="ref17961" refString="Yokoi, N. & Souphanthong, V. (2014) A list of Lao dragonflies. Privately published by the first author, Koriyama, 110 pp." type="book">Yokoi & Souphanthong (2014)</bibRefCitation>
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||
, p. 37 [scan of male], p. 57 fig. 61 [anal appendages in lateral view], p. 84 [distribution map].
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||
</paragraph>
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||
<paragraph id="8BDE36CD607C8C18FF67D37C1865FDF9" blockId="25.[151,1436,490,758]" pageId="25" pageNumber="326">
|
||
<taxonomicName id="4C614D4E607C8C18FF67D37C1BEBFDDE" ID-CoL="H9SH" authority=": Karube (1990)" authorityName=": Karube" authorityYear="1990" box="[151,701,582,604]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="subSpecies" species="xanthenatus" subSpecies="malayanus">
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||
<emphasis id="B915EADF607C8C18FF67D37C1B40FDDE" box="[151,534,582,604]" italics="true" pageId="25" pageNumber="326">Asiagomphus xanthenatus malayanus</emphasis>
|
||
:
|
||
<bibRefCitation id="EFF04B3C607C8C18FDD5D37C1BEBFDDE" box="[549,701,582,604]" pageId="25" pageNumber="326" refId="ref17337" refString="Karube, H. (1990) Description of a new subspecies of Asiagomphus xanthenatus (Williamson) from West Malaysia. Tombo, 33, 21 - 24." type="journal article">Karube (1990)</bibRefCitation>
|
||
</taxonomicName>
|
||
, pp. 21–24, figs. 1–10 [description of dark subspecies and female from
|
||
<collectingCountry id="F376765D607C8C18FF37D35F187CFDF9" box="[199,298,613,635]" name="Malaysia" pageId="25" pageNumber="326">Malaysia</collectingCountry>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="8BDE36CD607C8C18FF67D3BE1D20FD18" blockId="25.[151,1436,490,758]" box="[151,1142,644,666]" pageId="25" pageNumber="326">
|
||
<taxonomicName id="4C614D4E607C8C18FF67D3BE1B50FD18" ID-CoL="9X2SZ" authority=": Fraser (1925)" authorityName="Fraser" authorityYear="1925" box="[151,518,644,666]" class="Insecta" family="Gomphidae" genus="Gomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="personatus">
|
||
<emphasis id="B915EADF607C8C18FF67D3BE183BFD18" box="[151,365,644,666]" italics="true" pageId="25" pageNumber="326">Gomphus personatus</emphasis>
|
||
:
|
||
<bibRefCitation id="EFF04B3C607C8C18FE8BD3BF1B50FD18" box="[379,518,644,666]" pageId="25" pageNumber="326" refId="ref17215" refString="Fraser, F. C. (1925) Indian Dragonflies, Part XXI. Journal of the Bombay Natural History Society, 30, 657 - 663, 1 pl." type="journal article">Fraser (1925)</bibRefCitation>
|
||
</taxonomicName>
|
||
, p. 658–660 [inclusion of
|
||
<taxonomicName id="4C614D4E607C8C18FCFFD3BE1AF3FD18" ID-CoL="H9SX" authorityName=": Asahina" authorityYear="1986" box="[783,933,644,666]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607C8C18FCFFD3BE1AF3FD18" box="[783,933,644,666]" italics="true" pageId="25" pageNumber="326">G. xanthenatus</emphasis>
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||
</taxonomicName>
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||
as junior synonym].
|
||
</paragraph>
|
||
<paragraph id="8BDE36CD607C8C18FF67D3981B9CFD3A" blockId="25.[151,1436,490,758]" box="[151,714,674,696]" pageId="25" pageNumber="326">
|
||
<taxonomicName id="4C614D4E607C8C18FF67D39819AEFD3A" ID-CoL="9CHZC" authorityName="Leach" authorityYear="1815" box="[151,248,674,696]" class="Insecta" family="Gomphidae" genus="Gomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B915EADF607C8C18FF67D39819AEFD3A" box="[151,248,674,696]" italics="true" pageId="25" pageNumber="326">Gomphus</emphasis>
|
||
</taxonomicName>
|
||
sp.:
|
||
<bibRefCitation id="EFF04B3C607C8C18FED8D398189EFD3A" box="[296,456,674,696]" pageId="25" pageNumber="326" refId="ref16880" refString="Asahina, S. (1981) Records of little or unknown Odonata from Thailand. Tombo, 23, 3 - 16." type="journal article">Asahina (1981)</bibRefCitation>
|
||
, p. 10 [record of female].
|
||
</paragraph>
|
||
<paragraph id="8BDE36CD607C8C18FF67D3FB1A07FD74" blockId="25.[151,1436,490,758]" pageId="25" pageNumber="326">
|
||
<taxonomicName id="4C614D4E607C8C18FF67D3FB1B0FFD55" ID-CoL="9X2TY" authority=": Williamson (1907)" authorityName=": Williamson" authorityYear="1907" box="[151,601,705,727]" class="Insecta" family="Gomphidae" genus="Gomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607C8C18FF67D3FB1828FD55" box="[151,382,705,727]" italics="true" pageId="25" pageNumber="326">Gomphus xanthenatus</emphasis>
|
||
:
|
||
<bibRefCitation id="EFF04B3C607C8C18FE62D3FB1B0FFD55" box="[402,601,705,727]" pageId="25" pageNumber="326" refId="ref17719" refString="Williamson, E. B. (1907) The dragonflies (Odonata) of Burma and Lower Siam. II. Subfamilies Cordulegasterinae, Chlorogomphinae, and Gomphinae. Proceedings U. S. National Museum, 33, 267 - 317. https: // doi. org / 10.5479 / si. 00963801.33 - 1571.267" type="journal article">Williamson (1907)</bibRefCitation>
|
||
</taxonomicName>
|
||
, p. 303, fig. 11 [thorax pattern], pp. 305–308, fig. 33 [anal appendages and secondary genitalia];
|
||
<bibRefCitation id="EFF04B3C607C8C18FE51D3DA1B16FD74" box="[417,576,736,758]" pageId="25" pageNumber="326" refId="ref17499" refString="Laidlaw, F. F. (1922) A list of the dragonflies recorded from the Indian Empire. Part V. The subfamily Gomphinae. Records of the Indian Museum, 24, 367 - 426, 1 pl." type="journal article">Laidlaw (1922)</bibRefCitation>
|
||
, p. 398 [short description].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B6546607C8C18FF67D2261B41FCFE" pageId="25" pageNumber="326" type="materials_examined">
|
||
<paragraph id="8BDE36CD607C8C18FF67D2261CD0FCB7" blockId="25.[151,1437,796,2036]" box="[151,1414,796,821]" pageId="25" pageNumber="326">
|
||
<materialsCitation id="3B093C90607C8C18FF67D2261CD0FCB7" ID-GBIF-Occurrence="1913893024" box="[151,1414,796,821]" collectingDate="2016-05-15" latitude="15.583" location="Material" longLatPrecision="76" longitude="107.833" pageId="25" pageNumber="326" specimenCount="1" specimenCount-male="1" stateProvince="Quang Nam">
|
||
<emphasis id="B915EADF607C8C18FF67D22618D4FCB7" bold="true" box="[151,386,796,821]" pageId="25" pageNumber="326">
|
||
<location id="8EBE6016607C8C18FF67D22619A9FCB7" LSID="urn:lsid:plazi:treatment:03C887DB607C8C1BFF67D0BC18E2F86D:8EBE6016607C8C18FF67D22619A9FCB7" box="[151,255,796,821]" latitude="15.583" longLatPrecision="76" longitude="107.833" name="Material" pageId="25" pageNumber="326" stateProvince="Quang Nam">Material</location>
|
||
examined:
|
||
</emphasis>
|
||
<specimenCount id="9D67FD44607C8C18FE79D22618E3FCB6" box="[393,437,796,820]" pageId="25" pageNumber="326" type="male">1 ♂</specimenCount>
|
||
, along road AH17,
|
||
<collectingRegion id="49A5F82F607C8C18FD62D2261A4CFCB6" box="[658,794,796,820]" country="Vietnam" name="Quang Nam" pageId="25" pageNumber="326">Quang Nam</collectingRegion>
|
||
Prov. (appr.
|
||
<geoCoordinate id="EE55500A607C8C18FC59D2261D43FCB6" box="[937,1045,796,821]" direction="north" orientation="latitude" pageId="25" pageNumber="326" precision="55" value="15.583">15.583 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE55500A607C8C18FBD0D2261DC0FCB6" box="[1056,1174,796,820]" direction="east" orientation="longitude" pageId="25" pageNumber="326" precision="55" value="107.833">107.833 E</geoCoordinate>
|
||
),
|
||
<collectingDate id="EF9BE9E5607C8C18FB5AD2261C73FCB7" box="[1194,1317,796,821]" pageId="25" pageNumber="326" value="2016-05-15">
|
||
<date id="FFDF100D607C8C18FB5AD2261C73FCB7" box="[1194,1317,796,821]" pageId="25" pageNumber="326" value="2016-05-15">15-V-2016</date>
|
||
</collectingDate>
|
||
, TK leg.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BDE36CD607C8C18FF37D2051B41FCFE" blockId="25.[151,1437,796,2036]" pageId="25" pageNumber="326">
|
||
<materialsCitation id="3B093C90607C8C18FF37D2051B41FCFE" ID-GBIF-Occurrence="1913893017" collectingDate="2016-05-15" latitude="15.583" location="Quang Nam" longLatPrecision="76" longitude="107.833" pageId="25" pageNumber="326" specimenCount="2" specimenCount-male="2" stateProvince="Quang Nam">
|
||
<emphasis id="B915EADF607C8C18FF37D2051A0EFCDA" bold="true" box="[199,856,831,856]" pageId="25" pageNumber="326">Additional records based on field observations by TK:</emphasis>
|
||
<specimenCount id="9D67FD44607C8C18FC92D27A1AF2FCD5" box="[866,932,832,856]" pageId="25" pageNumber="326" type="male">2 ♂♂</specimenCount>
|
||
, along road AH17,
|
||
<collectingRegion id="49A5F82F607C8C18FB77D27A1C46FCD5" box="[1159,1296,832,856]" country="Vietnam" name="Quang Nam" pageId="25" pageNumber="326">Quang Nam</collectingRegion>
|
||
Prov. (appr.
|
||
<geoCoordinate id="EE55500A607C8C18FF67D2591854FCF9" box="[151,258,867,892]" direction="north" orientation="latitude" pageId="25" pageNumber="326" precision="55" value="15.583">15.583 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE55500A607C8C18FEFDD25E18D5FCFE" box="[269,387,868,892]" direction="east" orientation="longitude" pageId="25" pageNumber="326" precision="55" value="107.833">107.833 E</geoCoordinate>
|
||
),
|
||
<collectingDate id="EF9BE9E5607C8C18FE67D2591B45FCFE" box="[407,531,867,892]" pageId="25" pageNumber="326" value="2016-05-15">
|
||
<date id="FFDF100D607C8C18FE67D2591B45FCFE" box="[407,531,867,892]" pageId="25" pageNumber="326" value="2016-05-15">15-V-2016</date>
|
||
</collectingDate>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B6546607C8C18FF37D2BD1A50FBAD" pageId="25" pageNumber="326" type="diagnosis">
|
||
<paragraph id="8BDE36CD607C8C18FF37D2BD1CCAFC46" blockId="25.[151,1437,796,2036]" pageId="25" pageNumber="326">
|
||
<emphasis id="B915EADF607C8C18FF37D2BD189BFC22" bold="true" box="[199,461,903,928]" pageId="25" pageNumber="326">Differential diagnosis.</emphasis>
|
||
The details of the shape of the secondary genitalia and anal appendages (
|
||
<figureCitation id="135A2A48607C8C18FB0DD2BD1C26FC22" box="[1277,1392,903,928]" captionStart="FIGURE 3" captionStartId="8.[151,250,1943,1965]" captionTargetBox="[182,1417,193,1925]" captionTargetId="figure@8.[182,1417,193,1925]" captionTargetPageId="8" captionText="FIGURE 3. Anterior and posterior hamuli [a,c,e,g,h] lateral view, [b,d,f] antero-ventral lateral view. (a–b) Asiagomphus acco, Tam Dao NP, 18-V-2014; (c–d) A. kosterini sp. nov. holotype; (e–f) A. xanthenatus Quang Nam Prov., 15-V-2016; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. kosterini paratype." httpUri="https://zenodo.org/record/1441617/files/figure.png" pageId="25" pageNumber="326">Figs. 3e–f</figureCitation>
|
||
,
|
||
<figureCitation id="135A2A48607C8C18FA8BD2BD1CC1FC22" box="[1403,1431,903,928]" captionStart="FIGURE 5" captionStartId="10.[151,250,1939,1961]" captionTargetBox="[191,1400,193,1919]" captionTargetId="figure@10.[191,1400,193,1920]" captionTargetPageId="10" captionText="FIGURE 5. Male appendages in dorsal view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441621/files/figure.png" pageId="25" pageNumber="326">5e</figureCitation>
|
||
,
|
||
<figureCitation id="135A2A48607C8C18FF67D29119E2FC46" box="[151,180,939,964]" captionStart="FIGURE 6" captionStartId="11.[151,250,1945,1967]" captionTargetBox="[201,1402,193,1927]" captionTargetId="figure@11.[188,1403,193,1927]" captionTargetPageId="11" captionText="FIGURE 6. Male appendages in lateral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441623/files/figure.png" pageId="25" pageNumber="326">6e</figureCitation>
|
||
,
|
||
<figureCitation id="135A2A48607C8C18FF4FD291198CFC41" box="[191,218,939,963]" captionStart="FIGURE 7" captionStartId="12.[151,250,1946,1968]" captionTargetBox="[173,1418,193,1934]" captionTargetId="figure@12.[173,1418,193,1934]" captionTargetPageId="12" captionText="FIGURE 7. Male appendages in ventral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. superciliaris paratype Huu Lien NR, 24-V-2014." httpUri="https://zenodo.org/record/1441625/files/figure.png" pageId="25" pageNumber="326">7e</figureCitation>
|
||
) in combination with the black metepisternum help to separate this taxon from all other Vietnamese species.
|
||
</paragraph>
|
||
<paragraph id="8BDE36CD607C8C18FF37D2F41A50FBAD" blockId="25.[151,1437,796,2036]" pageId="25" pageNumber="326">
|
||
<emphasis id="B915EADF607C8C18FF37D2F418EDFC65" bold="true" box="[199,443,974,999]" pageId="25" pageNumber="326">Habitat and ecology.</emphasis>
|
||
Three males were found in close proximity of each other perched on rocks in a mediumsized open stream, or low on bushes and shrubs by the side of the stream, the stream coming down from a forested valley with heavily logged forest, crossing a main road.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B6546607C8C1BFF37D50318E2F86D" lastPageId="26" lastPageNumber="327" pageId="25" pageNumber="326" type="discussion">
|
||
<paragraph id="8BDE36CD607C8C18FF37D503188DF97A" blockId="25.[151,1437,796,2036]" pageId="25" pageNumber="326">
|
||
<emphasis id="B915EADF607C8C18FF37D503181AFBD3" bold="true" box="[199,332,1081,1106]" pageId="25" pageNumber="326">Discussion.</emphasis>
|
||
<taxonomicName id="4C614D4E607C8C18FEA6D5031B28FBD0" authorityName=": Asahina" authorityYear="1986" box="[342,638,1081,1106]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607C8C18FEA6D5031B28FBD0" box="[342,638,1081,1106]" italics="true" pageId="25" pageNumber="326">Asiagomphus xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
was described as
|
||
<taxonomicName id="4C614D4E607C8C18FCA6D5031D04FBD0" authorityName=": Williamson" authorityYear="1907" box="[854,1106,1081,1106]" class="Insecta" family="Gomphidae" genus="Gomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607C8C18FCA6D5031D04FBD0" box="[854,1106,1081,1106]" italics="true" pageId="25" pageNumber="326">Gomphus xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
by Williamson from
|
||
<collectingCountry id="F376765D607C8C18FAA0D5001CCAFBD3" box="[1360,1436,1082,1105]" name="Myanmar" pageId="25" pageNumber="326">Burma</collectingCountry>
|
||
(present day
|
||
<collectingCountry id="F376765D607C8C18FEDFD56418F7FBF4" box="[303,417,1118,1142]" name="Myanmar" pageId="25" pageNumber="326">Myanmar</collectingCountry>
|
||
).
|
||
<bibRefCitation id="EFF04B3C607C8C18FE47D5641B04FBF4" box="[439,594,1117,1142]" pageId="25" pageNumber="326" refId="ref17215" refString="Fraser, F. C. (1925) Indian Dragonflies, Part XXI. Journal of the Bombay Natural History Society, 30, 657 - 663, 1 pl." type="journal article">Fraser (1925)</bibRefCitation>
|
||
included
|
||
<taxonomicName id="4C614D4E607C8C18FD38D5641A24FBF4" authorityName=": Williamson" authorityYear="1907" box="[712,882,1117,1142]" class="Insecta" family="Gomphidae" genus="Gomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607C8C18FD38D5641A24FBF4" box="[712,882,1117,1142]" italics="true" pageId="25" pageNumber="326">G. xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C614D4E607C8C18FC51D5671D9CFBF4" authorityName=": Asahina" authorityYear="1986" box="[929,1226,1117,1142]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607C8C18FC51D5671D9CFBF4" box="[929,1226,1117,1142]" italics="true" pageId="25" pageNumber="326">Asiagomphus xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
) in
|
||
<taxonomicName id="4C614D4E607C8C18FB0ED564184FFB18" authority="Selys, 1973" authorityName="Selys" authorityYear="1973" class="Insecta" family="Gomphidae" genus="Gomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="personatus">
|
||
<emphasis id="B915EADF607C8C18FB0ED5641CCAFBF4" box="[1278,1436,1118,1142]" italics="true" pageId="25" pageNumber="326">G. personatus</emphasis>
|
||
Selys, 1973
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C614D4E607C8C18FEC9D5BB1B07FB1B" baseAuthorityName="Selys" baseAuthorityYear="1873" box="[313,593,1153,1178]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="personatus">
|
||
<emphasis id="B915EADF607C8C18FEC9D5BB1B07FB1B" box="[313,593,1153,1178]" italics="true" pageId="25" pageNumber="326">Asiagomphus personatus</emphasis>
|
||
</taxonomicName>
|
||
), because he perceived the differences in thorax pattern to be part of a cline and therefore no basis for separation. Possibly the similarity in structure of appendages of these species, at a time when few representatives of the current genus
|
||
<taxonomicName id="4C614D4E607C8C18FD5DD5F21A12FB63" authorityName="Asahina" authorityYear="1985" box="[685,836,1224,1249]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EADF607C8C18FD5DD5F21A12FB63" box="[685,836,1224,1249]" italics="true" pageId="25" pageNumber="326">Asiagomphus</emphasis>
|
||
</taxonomicName>
|
||
and the difficulty to separate them were known, led Fraser to this conclusion. But Williamson had given a detailed description of the abdomen pattern of his
|
||
<taxonomicName id="4C614D4E607C8C18FA74D5D71848FAAB" authorityName=": Asahina" authorityYear="1986" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607C8C18FA74D5D71848FAAB" italics="true" pageId="25" pageNumber="326">G. xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
, which was clearly a ‘ringed’ species, rather than a species characterised by a longitudinal pattern on the abdomen (as is
|
||
<taxonomicName id="4C614D4E607C8C18FE75D40E1BF5FACE" baseAuthorityName="Selys" baseAuthorityYear="1873" box="[389,675,1332,1357]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="personatus">
|
||
<emphasis id="B915EADF607C8C18FE75D40E1BF5FACE" box="[389,675,1332,1357]" italics="true" pageId="25" pageNumber="326">Asiagomphus personatus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<bibRefCitation id="EFF04B3C607C8C18FD30D40E1A3AFACE" box="[704,876,1332,1357]" pageId="25" pageNumber="326" refId="ref16880" refString="Asahina, S. (1981) Records of little or unknown Odonata from Thailand. Tombo, 23, 3 - 16." type="journal article">Asahina (1981</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF04B3C607C8C18FC8DD40E1A94FACF" box="[893,962,1332,1357]" pageId="25" pageNumber="326" refId="ref16938" refString="Asahina, S. (1986 a) A list of the Odonata from Thailand Part XIII. Gomphidae - 1. Cho Cho, 9, 29 - 43." type="journal article">1986a</bibRefCitation>
|
||
) published several female
|
||
<taxonomicName id="4C614D4E607C8C18FAF5D40E1CCAFACF" authorityName="Asahina" authorityYear="1985" box="[1285,1436,1332,1357]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EADF607C8C18FAF5D40E1CCAFACF" box="[1285,1436,1332,1357]" italics="true" pageId="25" pageNumber="326">Asiagomphus</emphasis>
|
||
</taxonomicName>
|
||
specimens from
|
||
<collectingCountry id="F376765D607C8C18FEA3D46D18E1FAF2" box="[339,439,1367,1392]" name="Thailand" pageId="25" pageNumber="326">Thailand</collectingCountry>
|
||
as
|
||
<taxonomicName id="4C614D4E607C8C18FE2ED46D1B23FAF2" authorityName="Asahina" authorityYear="1985" box="[478,629,1367,1392]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EADF607C8C18FE2ED46D1B23FAF2" box="[478,629,1367,1392]" italics="true" pageId="25" pageNumber="326">Asiagomphus</emphasis>
|
||
</taxonomicName>
|
||
sp.. Subsequently he received a male specimen, which he identified as
|
||
<emphasis id="B915EADF607C8C18FF67D4461810FA16" box="[151,326,1403,1428]" italics="true" pageId="25" pageNumber="326">
|
||
<taxonomicName id="4C614D4E607C8C18FF67D4461814FA16" authorityName=": Asahina" authorityYear="1986" box="[151,322,1403,1428]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">A. xanthenatus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
He then published as
|
||
<taxonomicName id="4C614D4E607C8C18FDB3D4461BBCFA16" authorityName=": Asahina" authorityYear="1986" box="[579,746,1403,1428]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607C8C18FDB3D4461BBCFA16" box="[579,746,1403,1428]" italics="true" pageId="25" pageNumber="326">A. xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
both this male, additional females, and the females formerly only identified as
|
||
<taxonomicName id="4C614D4E607C8C18FE92D4A518AFFA3A" authorityName="Asahina" authorityYear="1985" box="[354,505,1439,1464]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EADF607C8C18FE92D4A518AFFA3A" box="[354,505,1439,1464]" italics="true" pageId="25" pageNumber="326">Asiagomphus</emphasis>
|
||
</taxonomicName>
|
||
sp. Although all these specimens were collected from
|
||
<collectingCountry id="F376765D607C8C18FB95D4A51D9DFA3A" box="[1125,1227,1439,1464]" name="Thailand" pageId="25" pageNumber="326">Thailand</collectingCountry>
|
||
, they came from a variety of locations, with only the male collected from Thung Yai in
|
||
<collectingRegion id="49A5F82F607C8C18FC64D4F91DCAFA5E" box="[916,1180,1475,1500]" country="Thailand" name="Kanchanaburi" pageId="25" pageNumber="326">Kanchanaburi Province</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFF04B3C607C8C18FB59D4F91C0EFA5E" box="[1193,1368,1475,1500]" pageId="25" pageNumber="326" refId="ref16969" refString="Asahina, S. (1986 b) Descriptions of two Asiagomphus species from Assam-Burma and Burma-Thailand areas (Odonata, Gomphidae). Gekkan-Mushi, 186, 9 - 14." type="journal article">Asahina 1986b</bibRefCitation>
|
||
). This raises some doubt whether these specimens all belong to the same species, even if they share similar abdomen pattern. In
|
||
<collectingCountry id="F376765D607C8C18FEE8D730182CF9A1" box="[280,378,1546,1571]" name="Vietnam" pageId="25" pageNumber="326">Vietnam</collectingCountry>
|
||
, at least five other species occur with a ringed abdomen.
|
||
<bibRefCitation id="EFF04B3C607C8C18FBD3D7301DB4F9A1" box="[1059,1250,1546,1571]" pageId="25" pageNumber="326" refId="ref16969" refString="Asahina, S. (1986 b) Descriptions of two Asiagomphus species from Assam-Burma and Burma-Thailand areas (Odonata, Gomphidae). Gekkan-Mushi, 186, 9 - 14." type="journal article">Asahina (1986b)</bibRefCitation>
|
||
pointed out the differences between
|
||
<taxonomicName id="4C614D4E607C8C18FE8ED7151B4EF9C4" baseAuthorityName="Selys" baseAuthorityYear="1873" box="[382,536,1583,1606]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="personatus">
|
||
<emphasis id="B915EADF607C8C18FE8ED7151B4EF9C4" box="[382,536,1583,1606]" italics="true" pageId="25" pageNumber="326">A. personatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C614D4E607C8C18FDBFD7151BA3F9C5" authorityName=": Asahina" authorityYear="1986" box="[591,757,1582,1607]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607C8C18FDBFD7151BA3F9C5" box="[591,757,1582,1607]" italics="true" pageId="25" pageNumber="326">A. xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
, supporting the point of view that both are
|
||
<emphasis id="B915EADF607C8C18FB2CD7141C42F9C5" box="[1244,1300,1582,1607]" italics="true" pageId="25" pageNumber="326">bona</emphasis>
|
||
species. On the other hand, Asahina did not compare his Thai male to
|
||
<typeStatus id="54DA886F607C8C18FCEFD7681A19F9E8" box="[799,847,1618,1642]" pageId="25" pageNumber="326">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="4C614D4E607C8C18FC27D7681D2BF9E8" authorityName=": Asahina" authorityYear="1986" box="[983,1149,1617,1642]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607C8C18FC27D7681D2BF9E8" box="[983,1149,1617,1642]" italics="true" pageId="25" pageNumber="326">A. xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
, but based himself on the similarity between both the description and the illustrations provided by
|
||
<bibRefCitation id="EFF04B3C607C8C18FBE2D74F1DA6F90C" box="[1042,1264,1653,1678]" pageId="25" pageNumber="326" refId="ref17719" refString="Williamson, E. B. (1907) The dragonflies (Odonata) of Burma and Lower Siam. II. Subfamilies Cordulegasterinae, Chlorogomphinae, and Gomphinae. Proceedings U. S. National Museum, 33, 267 - 317. https: // doi. org / 10.5479 / si. 00963801.33 - 1571.267" type="journal article">Williamson (1907)</bibRefCitation>
|
||
. Although his conclusion seems warranted, it remains possible that eventually
|
||
<taxonomicName id="4C614D4E607C8C18FC9CD7A01DEBF930" authority="sensu Asahina" authorityName="Asahina" box="[876,1213,1689,1714]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607C8C18FC9CD7A01D0FF933" box="[876,1113,1689,1714]" italics="true" pageId="25" pageNumber="326">A. xanthenatus sensu</emphasis>
|
||
Asahina
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C614D4E607C8C18FB05D7A01808F954" authority="sensu Williamson" authorityName="Williamson" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607C8C18FB05D7A01983F957" italics="true" pageId="25" pageNumber="326">A. xanthenatus sensu</emphasis>
|
||
Williamson
|
||
</taxonomicName>
|
||
turn out to be different. Here, I follow the interpretation by Asahina that his male specimen from Thung Yai is
|
||
<taxonomicName id="4C614D4E607C8C18FEDDD7DB1885F97B" box="[301,467,1760,1785]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607C8C18FEDDD7DB1885F97B" box="[301,467,1760,1785]" italics="true" pageId="25" pageNumber="326">A. xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDE36CD607C8C18FF37D63E1BAFF876" blockId="25.[151,1437,796,2036]" pageId="25" pageNumber="326">
|
||
The Vietnamese male from
|
||
<collectingRegion id="49A5F82F607C8C18FE0FD63F1BD1F89E" box="[511,647,1797,1821]" country="Vietnam" name="Quang Nam" pageId="25" pageNumber="326">Quang Nam</collectingRegion>
|
||
shares with
|
||
<taxonomicName id="4C614D4E607C8C18FCE5D63F1AEAF89F" box="[789,956,1796,1821]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607C8C18FCE5D63F1AEAF89F" box="[789,956,1796,1821]" italics="true" pageId="25" pageNumber="326">A. xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
the widely divaricate superior appendages and epiproct branches (
|
||
<figureCitation id="135A2A48607C8C18FE57D6121B50F8C3" box="[423,518,1832,1857]" captionStart="FIGURE 5" captionStartId="10.[151,250,1939,1961]" captionTargetBox="[191,1400,193,1919]" captionTargetId="figure@10.[191,1400,193,1920]" captionTargetPageId="10" captionText="FIGURE 5. Male appendages in dorsal view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441621/files/figure.png" pageId="25" pageNumber="326">Figs. 5e</figureCitation>
|
||
,
|
||
<figureCitation id="135A2A48607C8C18FDE6D6131B65F8C2" box="[534,563,1833,1856]" captionStart="FIGURE 7" captionStartId="12.[151,250,1946,1968]" captionTargetBox="[173,1418,193,1934]" captionTargetId="figure@12.[173,1418,193,1934]" captionTargetPageId="12" captionText="FIGURE 7. Male appendages in ventral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. superciliaris paratype Huu Lien NR, 24-V-2014." httpUri="https://zenodo.org/record/1441625/files/figure.png" pageId="25" pageNumber="326">7e</figureCitation>
|
||
,
|
||
<figureCitation id="135A2A48607C8C18FDB2D6121B39F8C2" box="[578,623,1832,1856]" captionStart="FIGURE 12" captionStartId="21.[151,250,1586,1608]" captionTargetBox="[150,1436,193,1565]" captionTargetId="figure@21.[150,1436,193,1565]" captionTargetPageId="21" captionText="FIGURE 12. Asiagomphus spp. alive in hand, [a–e] A. reinhardti habitus, [a–b] male, [c–e] female, [f–h] male abdomen tip in dorsal view. (a) Gia Lai Prov., 19-VI-2016; (b) Lam Dong Prov., 17-V-2016; (c) Gia Lai Prov., 20-VI-2016; (d) Gia Lai Prov., 19-VI-2016; (e) female used for description; (f) A. reinhardti, Lam Dong Prov., 17-V-2016; (g) A. xanthenatus, Quang Nam Prov., 15-V-2016; (h) A. kosterini holotype." httpUri="https://zenodo.org/record/1441635/files/figure.png" pageId="25" pageNumber="326">
|
||
<quantity id="4C999B28607C8C18FDB2D6121B39F8C2" box="[578,623,1832,1856]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="25" pageNumber="326" unit="g" value="12.0">12g</quantity>
|
||
</figureCitation>
|
||
), the serrated anterior edge of the posterior hamulus (
|
||
<figureCitation id="135A2A48607C8C18FB18D6121C6BF8C2" box="[1256,1341,1832,1857]" captionStart="FIGURE 3" captionStartId="8.[151,250,1943,1965]" captionTargetBox="[182,1417,193,1925]" captionTargetId="figure@8.[182,1417,193,1925]" captionTargetPageId="8" captionText="FIGURE 3. Anterior and posterior hamuli [a,c,e,g,h] lateral view, [b,d,f] antero-ventral lateral view. (a–b) Asiagomphus acco, Tam Dao NP, 18-V-2014; (c–d) A. kosterini sp. nov. holotype; (e–f) A. xanthenatus Quang Nam Prov., 15-V-2016; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. kosterini paratype." httpUri="https://zenodo.org/record/1441617/files/figure.png" pageId="25" pageNumber="326">Fig. 3e</figureCitation>
|
||
), paired yellow spots on the labrum, large yellow basal ring on S3 and basal half-rings on S4–7, large yellow postero-dorsal mark on S9 and small yellow lateral spots at posterior margin of S8 (
|
||
<figureCitation id="135A2A48607C8C18FC31D6551D70F80A" box="[961,1062,1903,1928]" captionStart="FIGURE 12" captionStartId="21.[151,250,1586,1608]" captionTargetBox="[150,1436,193,1565]" captionTargetId="figure@21.[150,1436,193,1565]" captionTargetPageId="21" captionText="FIGURE 12. Asiagomphus spp. alive in hand, [a–e] A. reinhardti habitus, [a–b] male, [c–e] female, [f–h] male abdomen tip in dorsal view. (a) Gia Lai Prov., 19-VI-2016; (b) Lam Dong Prov., 17-V-2016; (c) Gia Lai Prov., 20-VI-2016; (d) Gia Lai Prov., 19-VI-2016; (e) female used for description; (f) A. reinhardti, Lam Dong Prov., 17-V-2016; (g) A. xanthenatus, Quang Nam Prov., 15-V-2016; (h) A. kosterini holotype." httpUri="https://zenodo.org/record/1441635/files/figure.png" pageId="25" pageNumber="326">
|
||
Fig.
|
||
<quantity id="4C999B28607C8C18FC09D64A1D70F80A" box="[1017,1062,1904,1928]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="25" pageNumber="326" unit="g" value="12.0">12g</quantity>
|
||
</figureCitation>
|
||
). It differs by the absence of an antehumeral spot.
|
||
<bibRefCitation id="EFF04B3C607C8C18FE96D6A91B5DF82E" box="[358,523,1939,1964]" pageId="25" pageNumber="326" refId="ref17337" refString="Karube, H. (1990) Description of a new subspecies of Asiagomphus xanthenatus (Williamson) from West Malaysia. Tombo, 33, 21 - 24." type="journal article">Karube (1990)</bibRefCitation>
|
||
described almost identical specimens from
|
||
<collectingCountry id="F376765D607C8C18FC09D6A91D09F82E" box="[1017,1119,1939,1964]" name="Malaysia" pageId="25" pageNumber="326">Malaysia</collectingCountry>
|
||
as a separate subspecies,
|
||
<taxonomicName id="4C614D4E607C8C18FA74D6AE187CF852" authorityName=": Karube" authorityYear="1990" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="subSpecies" species="xanthenatus" subSpecies="malayanus">
|
||
<emphasis id="B915EADF607C8C18FA74D6AE187CF852" italics="true" pageId="25" pageNumber="326">A. x. malayanus</emphasis>
|
||
</taxonomicName>
|
||
. His material also lacks antehumeral spots or stripes and has largely black (sometimes completely so) metepisternum. However, labrum and S8 are all black.
|
||
</paragraph>
|
||
<paragraph id="8BDE36CD607F8C1BFF37D1AD1DD0FE9F" blockId="26.[151,1436,151,285]" pageId="26" pageNumber="327">
|
||
<bibRefCitation id="EFF04B3C607F8C1BFF37D1AD1B7CFF32" box="[199,554,151,176]" pageId="26" pageNumber="327" refId="ref17961" refString="Yokoi, N. & Souphanthong, V. (2014) A list of Lao dragonflies. Privately published by the first author, Koriyama, 110 pp." type="book">Yokoi & Souphanthong (2014)</bibRefCitation>
|
||
recorded
|
||
<taxonomicName id="4C614D4E607F8C1BFD6FD1A21A1EFF32" box="[671,840,151,176]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607F8C1BFD6FD1A21A1EFF32" box="[671,840,151,176]" italics="true" pageId="26" pageNumber="327">A. xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
widely from
|
||
<collectingCountry id="F376765D607F8C1BFC18D1A21D77FF2D" box="[1000,1057,152,175]" name="Laos" pageId="26" pageNumber="327">Laos</collectingCountry>
|
||
, although not as far south as the latitude of
|
||
<collectingRegion id="49A5F82F607F8C1BFEE2D18618C8FF56" box="[274,414,188,212]" country="Vietnam" name="Quang Nam" pageId="26" pageNumber="327">Quang Nam</collectingRegion>
|
||
. The scan of a male included in their book (p. 37) appears to have a tiny antehumeral spot and small yellow markings on the metepisternum in line with
|
||
<bibRefCitation id="EFF04B3C607F8C1BFCA3D1E51D7DFF7A" box="[851,1067,223,248]" pageId="26" pageNumber="327" refId="ref17719" refString="Williamson, E. B. (1907) The dragonflies (Odonata) of Burma and Lower Siam. II. Subfamilies Cordulegasterinae, Chlorogomphinae, and Gomphinae. Proceedings U. S. National Museum, 33, 267 - 317. https: // doi. org / 10.5479 / si. 00963801.33 - 1571.267" type="journal article">Williamson (1907)</bibRefCitation>
|
||
. It seems reasonable to conclude that the Vietnamese male is well within the range of variability of this rather dark species.
|
||
</paragraph>
|
||
<caption id="DF1E6645607F8C1BFF67D7FC1AC6F979" httpUri="https://zenodo.org/record/1441639/files/figure.png" pageId="26" pageNumber="327" startId="26.[151,250,1734,1756]" targetBox="[169,1417,396,1666]" targetPageId="26">
|
||
<paragraph id="8BDE36CD607F8C1BFF67D7FC1AC6F979" blockId="26.[151,1437,1734,1787]" pageId="26" pageNumber="327">
|
||
<emphasis id="B915EADF607F8C1BFF67D7FC1849F95E" bold="true" box="[151,287,1734,1756]" pageId="26" pageNumber="327">FIGURE 14.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="4C614D4E607F8C1BFE31D7FC1B1FF95E" authorityName="Asahina" authorityYear="1985" box="[449,585,1734,1756]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EADF607F8C1BFE31D7FC1B1FF95E" box="[449,585,1734,1756]" italics="true" pageId="26" pageNumber="327">Asiagomphus</emphasis>
|
||
</taxonomicName>
|
||
spp. (A)
|
||
<taxonomicName id="4C614D4E607F8C1BFD59D7FD1BA3F95F" box="[681,757,1735,1757]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="acco">
|
||
<emphasis id="B915EADF607F8C1BFD59D7FD1BA3F95F" box="[681,757,1735,1757]" italics="true" pageId="26" pageNumber="327">A. acco</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B915EADF607F8C1BFCF4D7FD1A41F95E" bold="true" box="[772,791,1735,1756]" pageId="26" pageNumber="327">•</emphasis>
|
||
),
|
||
<taxonomicName id="4C614D4E607F8C1BFCC2D7FD1AFCF95E" authorityName=": Wilson" authorityYear="2005" box="[818,938,1734,1756]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="pacificus">
|
||
<emphasis id="B915EADF607F8C1BFCC2D7FD1AFCF95E" box="[818,938,1734,1756]" italics="true" pageId="26" pageNumber="327">A. pacificus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B915EADF607F8C1BFC48D7FD1A9DF95E" bold="true" box="[952,971,1735,1756]" pageId="26" pageNumber="327">•</emphasis>
|
||
),
|
||
<taxonomicName id="4C614D4E607F8C1BFC17D7FD1D3EF95E" authorityName=": Kosterin & Yokoi" authorityYear="2016" box="[999,1128,1734,1756]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="reinhardti">
|
||
<emphasis id="B915EADF607F8C1BFC17D7FD1D3EF95E" box="[999,1128,1734,1756]" italics="true" pageId="26" pageNumber="327">A. reinhardti</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B915EADF607F8C1BFB86D7FD1DDFF95E" bold="true" box="[1142,1161,1735,1756]" pageId="26" pageNumber="327">•</emphasis>
|
||
); (B)
|
||
<taxonomicName id="4C614D4E607F8C1BFB3DD7FD1C1CF95E" box="[1229,1354,1734,1756]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="auricolor">
|
||
<emphasis id="B915EADF607F8C1BFB3DD7FD1C1CF95E" box="[1229,1354,1734,1756]" italics="true" pageId="26" pageNumber="327">A. auricolor</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B915EADF607F8C1BFAA8D7FD1C3DF95E" bold="true" box="[1368,1387,1735,1756]" pageId="26" pageNumber="327">•</emphasis>
|
||
),
|
||
<taxonomicName id="4C614D4E607F8C1BFA76D7FD19AAF979" authorityName="Kompier" authorityYear="2018" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="monticola">
|
||
<emphasis id="B915EADF607F8C1BFA76D7FD19AAF979" italics="true" pageId="26" pageNumber="327">A. monticola</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B915EADF607F8C1BFEFBD7DC1848F979" bold="true" box="[267,286,1766,1787]" pageId="26" pageNumber="327">•</emphasis>
|
||
),
|
||
<taxonomicName id="4C614D4E607F8C1BFEC9D7DC1882F979" authorityName="Kompier" authorityYear="2018" box="[313,468,1765,1787]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="superciliaris">
|
||
<emphasis id="B915EADF607F8C1BFEC9D7DC1882F979" box="[313,468,1765,1787]" italics="true" pageId="26" pageNumber="327">A. superciliaris</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B915EADF607F8C1BFE13D7DC18A0F979" bold="true" box="[483,502,1766,1787]" pageId="26" pageNumber="327">•</emphasis>
|
||
),
|
||
<taxonomicName id="4C614D4E607F8C1BFDE1D7DC1BFEF979" box="[529,680,1765,1787]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607F8C1BFDE1D7DC1BFEF979" box="[529,680,1765,1787]" italics="true" pageId="26" pageNumber="327">A. xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B915EADF607F8C1BFD46D7DC1B9FF979" bold="true" box="[694,713,1766,1787]" pageId="26" pageNumber="327">•</emphasis>
|
||
),
|
||
<taxonomicName id="4C614D4E607F8C1BFD15D7DC1A0CF979" authorityName="Kompier" authorityYear="2018" box="[741,858,1765,1787]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="kosterini">
|
||
<emphasis id="B915EADF607F8C1BFD15D7DC1A0CF979" box="[741,858,1765,1787]" italics="true" pageId="26" pageNumber="327">A. kosterini</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B915EADF607F8C1BFC98D7DC1A2DF979" bold="true" box="[872,891,1766,1787]" pageId="26" pageNumber="327">•</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDE36CD607F8C1BFF37D61818E2F86D" blockId="26.[151,1436,1826,2031]" pageId="26" pageNumber="327">
|
||
<bibRefCitation id="EFF04B3C607F8C1BFF37D6181B36F8B9" box="[199,608,1826,1851]" pageId="26" pageNumber="327" refId="ref17573" refString="Reels, G. & Zhang, H. (2015) A Field Guide to the Dragonflies of Hainan. China Forestry Publishing House, Beijing, 463 pp." type="book">Reels & Zhang (2015, pp. 274–275)</bibRefCitation>
|
||
illustrate an
|
||
<taxonomicName id="4C614D4E607F8C1BFD03D6181ADCF8B9" authorityName="Asahina" authorityYear="1985" box="[755,906,1826,1851]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EADF607F8C1BFD03D6181ADCF8B9" box="[755,906,1826,1851]" italics="true" pageId="26" pageNumber="327">Asiagomphus</emphasis>
|
||
</taxonomicName>
|
||
species from
|
||
<collectingRegion id="49A5F82F607F8C1BFBD8D6181D2FF8B9" box="[1064,1145,1826,1851]" country="China" name="Hainan" pageId="26" pageNumber="327">Hainan</collectingRegion>
|
||
as
|
||
<taxonomicName id="4C614D4E607F8C1BFB50D6181C61F8B9" authorityName="Asahina" authorityYear="1985" box="[1184,1335,1826,1851]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EADF607F8C1BFB50D6181C61F8B9" box="[1184,1335,1826,1851]" italics="true" pageId="26" pageNumber="327">Asiagomphus</emphasis>
|
||
</taxonomicName>
|
||
sp. cf
|
||
<taxonomicName id="4C614D4E607F8C1BFA74D6191848F8DD" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607F8C1BFA74D6191848F8DD" italics="true" pageId="26" pageNumber="327">A. xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
. It is very similar in outward appearance to both
|
||
<taxonomicName id="4C614D4E607F8C1BFC81D67D1D4AF8DD" box="[881,1052,1862,1887]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607F8C1BFC81D67D1D4AF8DD" box="[881,1052,1862,1887]" italics="true" pageId="26" pageNumber="327">A. xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B915EADF607F8C1BFBACD67D1C45F8DD" box="[1116,1299,1862,1887]" italics="true" pageId="26" pageNumber="327">
|
||
<taxonomicName id="4C614D4E607F8C1BFBACD67D1C59F8DD" authorityName="Kompier" authorityYear="2018" box="[1116,1295,1862,1887]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="superciliaris">A. superciliaris</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Abdominal markings are particularly close to the individual of
|
||
<taxonomicName id="4C614D4E607F8C1BFD2ED6511ADDF801" authorityName="Kompier" authorityYear="2018" box="[734,907,1898,1923]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="superciliaris">
|
||
<emphasis id="B915EADF607F8C1BFD2ED6511ADDF801" box="[734,907,1898,1923]" italics="true" pageId="26" pageNumber="327">A. superciliaris</emphasis>
|
||
</taxonomicName>
|
||
illustrated in
|
||
<figureCitation id="135A2A48607F8C1BFBD9D6501DDDF801" box="[1065,1163,1898,1923]" captionStart="FIGURE 11" captionStartId="20.[151,246,1215,1237]" captionTargetBox="[159,1426,193,1203]" captionTargetId="figure@20.[159,1427,193,1203]" captionTargetPageId="20" captionText="FIGURE 11. Asiagomphus spp. males alive in hand. (a) A. monticola holotype; (b) A. monticola paratype, Yen Bai Prov., 1-VI-2014; (c) A. superciliaris holotype; (d) A. superciliaris paratype, Bac Kan Prov., 5-IV-2015; (e) A. superciliaris paratype, Huu Lien NR, 24- V-2014; (f) A. kosterini holotype." httpUri="https://zenodo.org/record/1441633/files/figure.png" pageId="26" pageNumber="327">Fig. 11d</figureCitation>
|
||
., but the frontal view of the head (p. 275) shows yellow spots on the labrum, absent in
|
||
<taxonomicName id="4C614D4E607F8C1BFC98D6B51D43F825" authorityName="Kompier" authorityYear="2018" box="[872,1045,1934,1959]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="superciliaris">
|
||
<emphasis id="B915EADF607F8C1BFC98D6B51D43F825" box="[872,1045,1934,1959]" italics="true" pageId="26" pageNumber="327">A. superciliaris</emphasis>
|
||
</taxonomicName>
|
||
. In view of the otherwise tropical distribution of
|
||
<taxonomicName id="4C614D4E607F8C1BFEB4D68918BBF849" box="[324,493,1970,1995]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607F8C1BFEB4D68918BBF849" box="[324,493,1970,1995]" italics="true" pageId="26" pageNumber="327">A. xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
, it is worth investigating the possibility their
|
||
<taxonomicName id="4C614D4E607F8C1BFBF6D6881DCBF849" authorityName="Asahina" authorityYear="1985" box="[1030,1181,1970,1995]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EADF607F8C1BFBF6D6881DCBF849" box="[1030,1181,1970,1995]" italics="true" pageId="26" pageNumber="327">Asiagomphus</emphasis>
|
||
</taxonomicName>
|
||
sp. cf
|
||
<taxonomicName id="4C614D4E607F8C1BFB02D6891CCDF849" box="[1266,1435,1970,1995]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="xanthenatus">
|
||
<emphasis id="B915EADF607F8C1BFB02D6891CCDF849" box="[1266,1435,1970,1995]" italics="true" pageId="26" pageNumber="327">A. xanthenatus</emphasis>
|
||
</taxonomicName>
|
||
concerns
|
||
<taxonomicName id="4C614D4E607F8C1BFEF1D6ED18FAF86D" authorityName="Kompier" authorityYear="2018" box="[257,428,2006,2031]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="superciliaris">
|
||
<emphasis id="B915EADF607F8C1BFEF1D6ED18FAF86D" box="[257,428,2006,2031]" italics="true" pageId="26" pageNumber="327">A. superciliaris</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |