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<document id="296BD5E59887EFD450129BA485514219" ID-DOI="10.5281/zenodo.191513" ID-GBIF-Dataset="2228c9ca-2836-42aa-8ba6-bb75baf534b5" ID-ISSN="1175-5326" ID-Zenodo-Dep="191513" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461129078534" checkinUser="plazi" docAuthor="Sonnenberg, Rainer &amp; Busch, Eckhard" docDate="2009" docId="039C5022FFAFD75A2CDB1CB0A451FA62" docLanguage="en" docName="zt02294p022.pdf" docOrigin="Zootaxa 2294" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Epiplatys maeseni Poll 1941, new combination" docType="treatment" docVersion="7" lastPageNumber="11" masterDocId="FFA5285AFFA6D7502C4C1A05A51EFFE5" masterDocTitle="Description of a new genus and two new species of killifish (Cyprinodontiformes: Nothobranchiidae) from West Africa, with a discussion of the taxonomic status of Aphyosemion maeseni Poll, 1941" masterLastPageNumber="22" masterPageNumber="1" pageNumber="10" updateTime="1698597310946" updateUser="plazi">
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<mods:title id="9CEBCE8F1FA3E65ECB5DFA114B86B648">Description of a new genus and two new species of killifish (Cyprinodontiformes: Nothobranchiidae) from West Africa, with a discussion of the taxonomic status of Aphyosemion maeseni Poll, 1941</mods:title>
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<mods:namePart id="362210134BB89365B207BF76023BC770">Sonnenberg, Rainer</mods:namePart>
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<mods:namePart id="A85717C48AA2F54C4D1667534AF343CA">Busch, Eckhard</mods:namePart>
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<mods:date id="9A5AC5FD3F5E92922001F931777D7809">2009</mods:date>
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<mods:number id="653EC7ADD90AF85BCAF4D07D35243E1E">2294</mods:number>
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<mods:identifier id="982325E7EE7E80DEC7EC7B9C7E5518CF" type="DOI">10.5281/zenodo.191513</mods:identifier>
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<treatment id="039C5022FFAFD75A2CDB1CB0A451FA62" ID-DOI="http://doi.org/10.5281/zenodo.6216136" ID-GBIF-Taxon="119626767" ID-Zenodo-Dep="6216136" LSID="urn:lsid:plazi:treatment:039C5022FFAFD75A2CDB1CB0A451FA62" httpUri="http://treatment.plazi.org/id/039C5022FFAFD75A2CDB1CB0A451FA62" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<subSubSection id="C32FB2BFFFAFD7592CDB1CB0A427F913" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B8AE134FFAFD7592CDB1CB0A7F5F92A" blockId="9.[151,747,1717,1782]" box="[151,747,1717,1743]" pageId="9" pageNumber="10">
<heading id="D0C25658FFAFD7592CDB1CB0A7F5F92A" bold="true" box="[151,747,1717,1743]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="B9413D26FFAFD7592CDB1CB0A7F5F92A" bold="true" box="[151,747,1717,1743]" pageId="9" pageNumber="10">
<taxonomicName id="4C359AB7FFAFD7592CDB1CB0A719F92A" ID-CoL="6G6PK" authority="Poll, 1941" authorityName="Poll" authorityYear="1941" box="[151,519,1717,1743]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="maeseni" status="comb. nov.">
<emphasis id="B9413D26FFAFD7592CDB1CB0A46FF92A" bold="true" box="[151,369,1717,1743]" italics="true" pageId="9" pageNumber="10">Epiplatys maeseni</emphasis>
(
<bibRefCitation id="EFA49CC5FFAFD7592DCD1CB0A4E1F92A" author="Poll" box="[385,511,1717,1743]" pageId="9" pageNumber="22" refString="Poll, M. (1941) Poissons nouveaux de la Cote d'Ivoire. Revue de Zoologie et de Botanique Africaines, 44, 133 - 143." type="journal article" year="1941">Poll, 1941</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A272805DFFAFD7592E591CB0A7F5F92A" box="[533,747,1717,1743]" pageId="9" pageNumber="10">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8AE134FFAFD7592CDB1CD9A427F913" blockId="9.[151,747,1717,1782]" box="[151,313,1756,1782]" pageId="9" pageNumber="10">
(
<figureCitation id="130EFDB1FFAFD7592CD31CD9A42FF913" box="[159,305,1756,1782]" captionStart="FIGURE 4. A &amp; B" captionStartId="11.[151,255,954,978]" captionTargetBox="[153,1433,194,930]" captionTargetId="figure@11.[153,1433,194,930]" captionTargetPageId="11" captionText="FIGURE 4. A &amp; B. Ventral and lateral view of the holotype of Epiplatys maeseni (MRAC P- 66392), 37 mm TL; C &amp; D. Ventral and lateral view of a paratype of N. leucopterygius (MRAC 73 - 39 - P- 1897); 58 mm TL; and E &amp; F. Ventral and lateral view of a syntype of E. olbrechtsi (MRAC P- 66388), ca. 55 mm TL (Photos by T. Musschoot, MRAC)." httpUri="https://zenodo.org/record/191517/files/figure.png" pageId="9" pageNumber="10">Figs. 4A&amp;B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32FB2BFFFAFD7592CDB1D21A7D3F8D9" box="[151,717,1828,1852]" pageId="9" pageNumber="10" type="description">
<paragraph id="8B8AE134FFAFD7592CDB1D21A7D3F8D9" blockId="9.[151,717,1828,1852]" box="[151,717,1828,1852]" pageId="9" pageNumber="10">
<emphasis id="B9413D26FFAFD7592CDB1D21A40AF8D9" bold="true" box="[151,276,1828,1852]" pageId="9" pageNumber="10">Synonymy.</emphasis>
<taxonomicName id="4C359AB7FFAFD7592D571D20A79EF8D9" authority="Poll, 1941" authorityName="Poll" authorityYear="1941" box="[283,640,1829,1852]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="maeseni">
<emphasis id="B9413D26FFAFD7592D571D20A716F8D9" box="[283,520,1829,1852]" italics="true" pageId="9" pageNumber="10">Aphyosemion maeseni</emphasis>
<bibRefCitation id="EFA49CC5FFAFD7592E431D20A79EF8D9" author="Poll" box="[527,640,1829,1852]" pageId="9" pageNumber="22" refString="Poll, M. (1941) Poissons nouveaux de la Cote d'Ivoire. Revue de Zoologie et de Botanique Africaines, 44, 133 - 143." type="journal article" year="1941">Poll, 1941</bibRefCitation>
</taxonomicName>
: p. 141
</paragraph>
</subSubSection>
<subSubSection id="C32FB2BFFFAFD7592CDB1D62A04EF813" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8B8AE134FFAFD7592CDB1D62A04EF813" blockId="9.[151,1437,1895,2038]" pageId="9" pageNumber="10">
<emphasis id="B9413D26FFAFD7592CDB1D62A494F864" bold="true" box="[151,394,1895,1921]" pageId="9" pageNumber="10">Material examined.</emphasis>
<taxonomicName id="4C359AB7FFAFD7592DDE1D6DA788F864" box="[402,662,1896,1921]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="maeseni">
<emphasis id="B9413D26FFAFD7592DDE1D6DA788F864" box="[402,662,1896,1921]" italics="true" pageId="9" pageNumber="10">Aphyosemion maeseni</emphasis>
</taxonomicName>
:
<typeStatus id="548E5F96FFAFD7592EEA1D6DA611F867" box="[678,783,1896,1922]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
,
<collectionCode id="ED2479F1FFAFD7592F561D6DA66FF867" box="[794,881,1896,1922]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="9" pageNumber="10">MRAC</collectionCode>
<collectionCode id="ED2479F1FFAFD7592F341D6DA68CF867" box="[888,914,1896,1922]" pageId="9" pageNumber="10">P-</collectionCode>
66392, 37 mm TL; collected at the confluent of the small Masei River and the Bon de Boan River, which is a tributary of the Nuon (at the border
<collectingCountry id="F322A1A4FFAFD759297D1D8BA09BF84D" box="[1329,1413,1934,1960]" name="Liberia" pageId="9" pageNumber="10">Liberia</collectingCountry>
–
<collectingCountry id="F322A1A4FFAFD7592CDB1DB0A430F82A" box="[151,302,1973,1999]" name="Ivory Coast" pageId="9" pageNumber="10">Ivory Coast</collectingCountry>
); A. Maesen, F. M. Olbrechts &amp; P. J. Vandenhoute,
<date id="FF8BC7F4FFAFD7592FFC1DB0A18EF82A" box="[944,1168,1973,1999]" pageId="9" pageNumber="10" value="1939-04-12" valueMax="1939-04-13" valueMin="1939-04-12">12–13 April 1939</date>
.
<taxonomicName id="4C359AB7FFAFD75928D01DB0A08AF82B" box="[1180,1428,1973,1998]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="olbrechtsi">
<emphasis id="B9413D26FFAFD75928D01DB0A08AF82B" box="[1180,1428,1973,1998]" italics="true" pageId="9" pageNumber="10">Epiplatys olbrechtsi</emphasis>
</taxonomicName>
:
<typeStatus id="548E5F96FFAFD7592CDB1DD9A41EF813" box="[151,256,2012,2038]" pageId="9" pageNumber="10" type="syntype">syntypes</typeStatus>
,
<collectionCode id="ED2479F1FFAFD7592D401DD9A47DF813" box="[268,355,2012,2038]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="9" pageNumber="10">MRAC</collectionCode>
<collectionCode id="ED2479F1FFAFD7592D261DD9A49AF813" box="[362,388,2012,2038]" pageId="9" pageNumber="10">P-</collectionCode>
66388–66391, same data as
<taxonomicName id="4C359AB7FFAFD7592E9A1DD9A6C5F810" box="[726,987,2012,2037]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="maeseni">
<emphasis id="B9413D26FFAFD7592E9A1DD9A6C5F810" box="[726,987,2012,2037]" italics="true" pageId="9" pageNumber="10">Aphyosemion maeseni</emphasis>
</taxonomicName>
. Additional material see below.
</paragraph>
</subSubSection>
<subSubSection id="C32FB2BFFFACD75A2C891A92A4D5FD67" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="8B8AE134FFACD75A2C891A92A075FDEB" blockId="10.[151,1437,151,1415]" pageId="10" pageNumber="11">
<emphasis id="B9413D26FFACD75A2C891A92A459FF54" bold="true" box="[197,327,151,177]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
The
<typeStatus id="548E5F96FFACD75A2DCA1A9DA4F0FF57" box="[390,494,152,178]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
is, as far as it can be assessed by its present state of conservation, by external morphology similar to the
<typeStatus id="548E5F96FFACD75A2D911ABBA759FF3D" box="[477,583,190,216]" pageId="10" pageNumber="11" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C359AB7FFACD75A2E3D1ABAA6F2FF3D" authority="Poll, 1941" authorityName="Poll" authorityYear="1941" box="[625,1004,190,216]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="olbrechtsi">
<emphasis id="B9413D26FFACD75A2E3D1ABAA67AFF3D" box="[625,868,191,216]" italics="true" pageId="10" pageNumber="11">Epiplatys olbrechtsi</emphasis>
<bibRefCitation id="EFA49CC5FFACD75A2F201ABBA6F2FF3D" author="Poll" box="[876,1004,190,216]" pageId="10" pageNumber="22" refString="Poll, M. (1941) Poissons nouveaux de la Cote d'Ivoire. Revue de Zoologie et de Botanique Africaines, 44, 133 - 143." type="journal article" year="1941">Poll, 1941</bibRefCitation>
</taxonomicName>
, having a robust and more rounded body versus more slender and laterally slightly compressed in
<emphasis id="B9413D26FFACD75A2FE51AE0A172FF1B" box="[937,1132,229,254]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C359AB7FFACD75A2FE51AE0A176FF1B" box="[937,1128,229,254]" class="Actinopterygii" family="Nothobranchiidae" genus="Nimbapanchax" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Nimbapanchax</taxonomicName>
.
</emphasis>
It shares with
<taxonomicName id="4C359AB7FFACD75A296B1AE0A40DFEC0" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="olbrechtsi">
<emphasis id="B9413D26FFACD75A296B1AE0A40DFEC0" italics="true" pageId="10" pageNumber="11">Epiplatys olbrechtsi</emphasis>
</taxonomicName>
the distinct morphology of the throat and lower jaw, diagnostic for
<emphasis id="B9413D26FFACD75A28231B09A1F3FEC0" box="[1135,1261,268,293]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C359AB7FFACD75A28231B09A1F6FEC0" box="[1135,1256,268,293]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Epiplatys</taxonomicName>
,
</emphasis>
which differs considerably from the head morphology of
<taxonomicName id="4C359AB7FFACD75A2EFD1B37A68BFEAE" box="[689,917,306,331]" class="Actinopterygii" family="Nothobranchiidae" genus="Archiaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B9413D26FFACD75A2EFD1B37A68BFEAE" box="[689,917,306,331]" italics="true" pageId="10" pageNumber="11">Archiaphyosemion</emphasis>
</taxonomicName>
,
<taxonomicName id="4C359AB7FFACD75A2FE91B37A140FEAE" box="[933,1118,306,331]" class="Actinopterygii" family="Nothobranchiidae" genus="Nimbapanchax" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B9413D26FFACD75A2FE91B37A140FEAE" box="[933,1118,306,331]" italics="true" pageId="10" pageNumber="11">Nimbapanchax</emphasis>
</taxonomicName>
and other nothobranchiid genera (
<figureCitation id="130EFDB1FFACD75A2CB61B5CA44FFE96" box="[250,337,345,371]" captionStart="FIGURE 4. A &amp; B" captionStartId="11.[151,255,954,978]" captionTargetBox="[153,1433,194,930]" captionTargetId="figure@11.[153,1433,194,930]" captionTargetPageId="11" captionText="FIGURE 4. A &amp; B. Ventral and lateral view of the holotype of Epiplatys maeseni (MRAC P- 66392), 37 mm TL; C &amp; D. Ventral and lateral view of a paratype of N. leucopterygius (MRAC 73 - 39 - P- 1897); 58 mm TL; and E &amp; F. Ventral and lateral view of a syntype of E. olbrechtsi (MRAC P- 66388), ca. 55 mm TL (Photos by T. Musschoot, MRAC)." httpUri="https://zenodo.org/record/191517/files/figure.png" pageId="10" pageNumber="11">Figs. 4</figureCitation>
C&amp;D) (see Van der Zee 2002; Wildekamp &amp; Van der
<bibRefCitation id="EFA49CC5FFACD75A2FAB1B5CA17EFE96" author="Zee" box="[999,1120,345,371]" pageId="10" pageNumber="22" refString="Wildekamp, R. H. &amp; Van der Zee, J. R. (2003) Cyprinodontiformes. In: Paugy, D., Leveque, C. &amp; Teugels, G. G. (Eds) The Fresh and Brackish Water Fishes of West Africa. Volume II. IRD Editions, Paris, pp. 298 - 442." type="book chapter" year="2003">Zee 2003</bibRefCitation>
). In
<taxonomicName id="4C359AB7FFACD75A28DB1B5CA015FE97" box="[1175,1291,345,370]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B9413D26FFACD75A28DB1B5CA015FE97" box="[1175,1291,345,370]" italics="true" pageId="10" pageNumber="11">Epiplatys</emphasis>
</taxonomicName>
the opercle borders on the ventral side have their smallest distance at or slightly posterior of the eye level and then widen anteriorly on the lower jaw (
<figureCitation id="130EFDB1FFACD75A2DB61BA3A79AFE25" box="[506,644,422,448]" captionStart="FIGURE 4. A &amp; B" captionStartId="11.[151,255,954,978]" captionTargetBox="[153,1433,194,930]" captionTargetId="figure@11.[153,1433,194,930]" captionTargetPageId="11" captionText="FIGURE 4. A &amp; B. Ventral and lateral view of the holotype of Epiplatys maeseni (MRAC P- 66392), 37 mm TL; C &amp; D. Ventral and lateral view of a paratype of N. leucopterygius (MRAC 73 - 39 - P- 1897); 58 mm TL; and E &amp; F. Ventral and lateral view of a syntype of E. olbrechtsi (MRAC P- 66388), ca. 55 mm TL (Photos by T. Musschoot, MRAC)." httpUri="https://zenodo.org/record/191517/files/figure.png" pageId="10" pageNumber="11">Figs. 4A,B</figureCitation>
,E&amp;F). In the remaining
<taxonomicName id="4C359AB7FFACD75A2FFA1BA3A14EFE25" box="[950,1104,422,448]" class="Insecta" family="Richardidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subFamily" subFamily="Epiplateinae">Epiplateinae</taxonomicName>
the opercle borders on the ventral side run almost parallely towards the anterior part of the lower jaw (
<figureCitation id="130EFDB1FFACD75A28551BC8A173FE02" box="[1049,1133,461,487]" captionStart="FIGURE 4. A &amp; B" captionStartId="11.[151,255,954,978]" captionTargetBox="[153,1433,194,930]" captionTargetId="figure@11.[153,1433,194,930]" captionTargetPageId="11" captionText="FIGURE 4. A &amp; B. Ventral and lateral view of the holotype of Epiplatys maeseni (MRAC P- 66392), 37 mm TL; C &amp; D. Ventral and lateral view of a paratype of N. leucopterygius (MRAC 73 - 39 - P- 1897); 58 mm TL; and E &amp; F. Ventral and lateral view of a syntype of E. olbrechtsi (MRAC P- 66388), ca. 55 mm TL (Photos by T. Musschoot, MRAC)." httpUri="https://zenodo.org/record/191517/files/figure.png" pageId="10" pageNumber="11">Figs. 4</figureCitation>
C&amp;D).
<taxonomicName id="4C359AB7FFACD75A28881BC8A083FE03" box="[1220,1437,461,486]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="maeseni">
<emphasis id="B9413D26FFACD75A28881BC8A083FE03" box="[1220,1437,461,486]" italics="true" pageId="10" pageNumber="11">Epiplatys maeseni</emphasis>
</taxonomicName>
has a short and more rounded snout versus longer and more pointed in
<taxonomicName id="4C359AB7FFACD75A2F991BF1A1DEFDE8" box="[981,1216,500,525]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="olbrechtsi">
<emphasis id="B9413D26FFACD75A2F991BF1A1DEFDE8" box="[981,1216,500,525]" italics="true" pageId="10" pageNumber="11">Epiplatys olbrechtsi</emphasis>
</taxonomicName>
(
<figureCitation id="130EFDB1FFACD75A289C1BF1A03FFDEB" box="[1232,1313,500,526]" captionStart="FIGURE 4. A &amp; B" captionStartId="11.[151,255,954,978]" captionTargetBox="[153,1433,194,930]" captionTargetId="figure@11.[153,1433,194,930]" captionTargetPageId="11" captionText="FIGURE 4. A &amp; B. Ventral and lateral view of the holotype of Epiplatys maeseni (MRAC P- 66392), 37 mm TL; C &amp; D. Ventral and lateral view of a paratype of N. leucopterygius (MRAC 73 - 39 - P- 1897); 58 mm TL; and E &amp; F. Ventral and lateral view of a syntype of E. olbrechtsi (MRAC P- 66388), ca. 55 mm TL (Photos by T. Musschoot, MRAC)." httpUri="https://zenodo.org/record/191517/files/figure.png" pageId="10" pageNumber="11">Figs. 4</figureCitation>
E&amp;F).
</paragraph>
<paragraph id="8B8AE134FFACD75A2C89181FA4D5FD67" blockId="10.[151,1437,151,1415]" pageId="10" pageNumber="11">
<emphasis id="B9413D26FFACD75A2C89181FA44DFDD1" bold="true" box="[197,339,538,564]" pageId="10" pageNumber="11">Coloration.</emphasis>
Preserved in ethanol. Body nearly completely uniform brownish, unpaired fins darker brown. No traces of color pattern visible, no whitish or light border on unpaired fins.
<bibRefCitation id="EFA49CC5FFACD75A28731844A1D9FDBE" author="Poll" box="[1087,1223,577,603]" pageId="10" pageNumber="22" refString="Poll, M. (1941) Poissons nouveaux de la Cote d'Ivoire. Revue de Zoologie et de Botanique Africaines, 44, 133 - 143." type="journal article" year="1941">Poll (1941)</bibRefCitation>
reported black or dark dots in the dorsal fin.
</paragraph>
</subSubSection>
<subSubSection id="C32FB2BFFFACD75A2C89188BA7FFFCFE" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8B8AE134FFACD75A2C89188BA7FFFCFE" blockId="10.[151,1437,151,1415]" pageId="10" pageNumber="11">
<emphasis id="B9413D26FFACD75A2C89188BA475FD4D" bold="true" box="[197,363,654,680]" pageId="10" pageNumber="11">Distribution.</emphasis>
Currently only known from the
<typeStatus id="548E5F96FFACD75A2EB7188BA631FD4D" box="[763,815,654,680]" pageId="10" pageNumber="11">type</typeStatus>
locality in northwestern
<collectingCountry id="F322A1A4FFACD75A2828188BA1E9FD4D" box="[1124,1271,654,680]" name="Ivory Coast" pageId="10" pageNumber="11">Ivory Coast</collectingCountry>
. All previous records in the literature, which mention
<taxonomicName id="4C359AB7FFACD75A2EEC18B0A616FD2B" box="[672,776,693,718]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="maeseni">
<emphasis id="B9413D26FFACD75A2EEC18B0A616FD2B" box="[672,776,693,718]" italics="true" pageId="10" pageNumber="11">maeseni</emphasis>
</taxonomicName>
under the generic names
<taxonomicName id="4C359AB7FFACD75A281918B0A01EFD2B" box="[1109,1280,693,718]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B9413D26FFACD75A281918B0A01EFD2B" box="[1109,1280,693,718]" italics="true" pageId="10" pageNumber="11">Aphyosemion</emphasis>
</taxonomicName>
,
<taxonomicName id="4C359AB7FFACD75A295D18B0A067FD2B" box="[1297,1401,693,718]" family="Nothobranchiidae" genus="Roloffia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B9413D26FFACD75A295D18B0A067FD2B" box="[1297,1401,693,718]" italics="true" pageId="10" pageNumber="11">Roloffia</emphasis>
</taxonomicName>
or
<taxonomicName id="4C359AB7FFACD75A2CDB18D9A46CFD10" box="[151,370,732,757]" class="Actinopterygii" family="Nothobranchiidae" genus="Archiaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B9413D26FFACD75A2CDB18D9A46CFD10" box="[151,370,732,757]" italics="true" pageId="10" pageNumber="11">Archiaphyosemion</emphasis>
</taxonomicName>
most probably are dealing with
<emphasis id="B9413D26FFACD75A2EBD18D9A6D6FD10" box="[753,968,732,757]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C359AB7FFACD75A2EBD18D9A6DDFD10" box="[753,963,732,757]" class="Actinopterygii" family="Nothobranchiidae" genus="Nimbapanchax" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="leucopterygius" status="sp. nov.">N. leucopterygius</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A272805DFFACD75A2F9C18D9A17FFD13" box="[976,1121,732,758]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
, and at least were a record for a
<taxonomicName id="4C359AB7FFACD75A2C981907A498FCFE" box="[212,390,770,795]" class="Actinopterygii" family="Nothobranchiidae" genus="Nimbapanchax" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B9413D26FFACD75A2C981907A498FCFE" box="[212,390,770,795]" italics="true" pageId="10" pageNumber="11">Nimbapanchax</emphasis>
</taxonomicName>
species and not an
<emphasis id="B9413D26FFACD75A2E271907A7FFFCFE" box="[619,737,770,795]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C359AB7FFACD75A2E271907A7C0FCFE" box="[619,734,770,795]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Epiplatys</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32FB2BFFFACD75A2C89192DA12BFCA6" box="[197,1077,808,835]" pageId="10" pageNumber="11" type="etymology">
<paragraph id="8B8AE134FFACD75A2C89192DA12BFCA6" blockId="10.[151,1437,151,1415]" box="[197,1077,808,835]" pageId="10" pageNumber="11">
<emphasis id="B9413D26FFACD75A2C89192DA44FFCA7" bold="true" box="[197,337,808,834]" pageId="10" pageNumber="11">Etymology.</emphasis>
The species was named after one of the collectors, A. Maesen.
</paragraph>
</subSubSection>
<subSubSection id="C32FB2BFFFACD75A2C8A194AA451FA62" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8B8AE134FFACD75A2C8A194AA451FA62" blockId="10.[151,1437,151,1415]" pageId="10" pageNumber="11">
<emphasis id="B9413D26FFACD75A2C8A194AA45EFC8C" bold="true" box="[198,320,847,873]" pageId="10" pageNumber="11">Remarks.</emphasis>
The single
<typeStatus id="548E5F96FFACD75A2D9B1955A714FC8F" box="[471,522,848,874]" pageId="10" pageNumber="11">type</typeStatus>
specimen of
<taxonomicName id="4C359AB7FFACD75A2EF81955A6A0FC8C" box="[692,958,848,873]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="maeseni">
<emphasis id="B9413D26FFACD75A2EF81955A6A0FC8C" box="[692,958,848,873]" italics="true" pageId="10" pageNumber="11">Aphyosemion maeseni</emphasis>
</taxonomicName>
(MRAC P-66392) was compared with specimens from other localities from
<collectingCountry id="F322A1A4FFACD75A2E201973A7D7FC75" box="[620,713,886,912]" name="Guinea-Bissau" pageId="10" pageNumber="11">Guinea</collectingCountry>
,
<collectingCountry id="F322A1A4FFACD75A2E9A1973A62EFC75" box="[726,816,886,912]" name="Liberia" pageId="10" pageNumber="11">Liberia</collectingCountry>
and
<collectingCountry id="F322A1A4FFACD75A2F221973A11AFC75" box="[878,1028,886,912]" name="Ivory Coast" pageId="10" pageNumber="11">Ivory Coast</collectingCountry>
, which are currently regarded as conspecific (see list of material under the respective descriptions below). All characters that distinguish the
<typeStatus id="548E5F96FFACD75A2CDB19C1A5D4FC3B" box="[151,202,964,990]" pageId="10" pageNumber="11">type</typeStatus>
of
<taxonomicName id="4C359AB7FFACD75A2CBF19C1A468FC38" box="[243,374,964,989]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="maeseni">
<emphasis id="B9413D26FFACD75A2CBF19C1A468FC38" box="[243,374,964,989]" italics="true" pageId="10" pageNumber="11">A. maeseni</emphasis>
</taxonomicName>
from
<taxonomicName id="4C359AB7FFACD75A2D8C19C1A76DFC38" box="[448,627,964,989]" class="Actinopterygii" family="Nothobranchiidae" genus="Nimbapanchax" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B9413D26FFACD75A2D8C19C1A76DFC38" box="[448,627,964,989]" italics="true" pageId="10" pageNumber="11">Nimbapanchax</emphasis>
</taxonomicName>
are shared with the
<typeStatus id="548E5F96FFACD75A2F2A19C1A6D0FC3B" box="[870,974,964,990]" pageId="10" pageNumber="11" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C359AB7FFACD75A2FBB19C1A190FC38" box="[1015,1166,964,989]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="olbrechtsi">
<emphasis id="B9413D26FFACD75A2FBB19C1A190FC38" box="[1015,1166,964,989]" italics="true" pageId="10" pageNumber="11">E. olbrechtsi</emphasis>
</taxonomicName>
, with the exception of the snout shape, which is slightly shorter and more rounded in
<taxonomicName id="4C359AB7FFACD75A2FE819EFA134FBE6" box="[932,1066,1002,1027]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="maeseni">
<emphasis id="B9413D26FFACD75A2FE819EFA134FBE6" box="[932,1066,1002,1027]" italics="true" pageId="10" pageNumber="11">E. maeseni</emphasis>
</taxonomicName>
(
<figureCitation id="130EFDB1FFACD75A287119EFA1C7FBE1" box="[1085,1241,1002,1028]" captionStart="FIGURE 4. A &amp; B" captionStartId="11.[151,255,954,978]" captionTargetBox="[153,1433,194,930]" captionTargetId="figure@11.[153,1433,194,930]" captionTargetPageId="11" captionText="FIGURE 4. A &amp; B. Ventral and lateral view of the holotype of Epiplatys maeseni (MRAC P- 66392), 37 mm TL; C &amp; D. Ventral and lateral view of a paratype of N. leucopterygius (MRAC 73 - 39 - P- 1897); 58 mm TL; and E &amp; F. Ventral and lateral view of a syntype of E. olbrechtsi (MRAC P- 66388), ca. 55 mm TL (Photos by T. Musschoot, MRAC)." httpUri="https://zenodo.org/record/191517/files/figure.png" pageId="10" pageNumber="11">Figs. 4A&amp;B</figureCitation>
). However, it is possible that the
<typeStatus id="548E5F96FFACD75A2D201E14A4BFFBCE" box="[364,417,1041,1067]" pageId="10" pageNumber="11">type</typeStatus>
is a damaged or deformed specimen of
<taxonomicName id="4C359AB7FFACD75A2FD51E14A127FBCF" box="[921,1081,1041,1066]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="olbrechtsi">
<emphasis id="B9413D26FFACD75A2FD51E14A127FBCF" box="[921,1081,1041,1066]" italics="true" pageId="10" pageNumber="11">E. olbrechtsi</emphasis>
</taxonomicName>
or might represent a second
<taxonomicName id="4C359AB7FFACD75A2CDB1E3DA418FBB4" box="[151,262,1080,1105]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B9413D26FFACD75A2CDB1E3DA418FBB4" box="[151,262,1080,1105]" italics="true" pageId="10" pageNumber="11">Epiplatys</emphasis>
</taxonomicName>
species within the same collection. Unfortunately no useful traces of color patterns are visible. The current assignment of a
<taxonomicName id="4C359AB7FFACD75A2DFA1E5AA777FB9D" box="[438,617,1119,1144]" class="Actinopterygii" family="Nothobranchiidae" genus="Nimbapanchax" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B9413D26FFACD75A2DFA1E5AA777FB9D" box="[438,617,1119,1144]" italics="true" pageId="10" pageNumber="11">Nimbapanchax</emphasis>
</taxonomicName>
species to the name
<taxonomicName id="4C359AB7FFACD75A2F2F1E5AA175FB9D" box="[867,1131,1119,1144]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="maeseni">
<emphasis id="B9413D26FFACD75A2F2F1E5AA175FB9D" box="[867,1131,1119,1144]" italics="true" pageId="10" pageNumber="11">Aphyosemion maeseni</emphasis>
</taxonomicName>
seems to be based on the dark coloration of the unpaired fins mentioned by
<bibRefCitation id="EFA49CC5FFACD75A2E921E80A678FB7A" author="Poll" box="[734,870,1157,1183]" pageId="10" pageNumber="22" refString="Poll, M. (1941) Poissons nouveaux de la Cote d'Ivoire. Revue de Zoologie et de Botanique Africaines, 44, 133 - 143." type="journal article" year="1941">Poll (1941)</bibRefCitation>
. However,
<bibRefCitation id="EFA49CC5FFACD75A2FA51E80A173FB7A" author="Poll" box="[1001,1133,1157,1183]" pageId="10" pageNumber="22" refString="Poll, M. (1941) Poissons nouveaux de la Cote d'Ivoire. Revue de Zoologie et de Botanique Africaines, 44, 133 - 143." type="journal article" year="1941">Poll (1941)</bibRefCitation>
and later
<bibRefCitation id="EFA49CC5FFACD75A28931E80A5D2FB23" author="Scheel" pageId="10" pageNumber="22" refString="Scheel, J. J. (1968) Rivulins of the Old World. TFH Publications, New Jersey, 480 pp." type="book" year="1968" yearSuffix="p">Scheel (1968, p. 289)</bibRefCitation>
did not mention the white border of all unpaired fins, typical for the
<taxonomicName id="4C359AB7FFACD75A2FA11EA9A181FB20" box="[1005,1183,1196,1221]" class="Actinopterygii" family="Nothobranchiidae" genus="Nimbapanchax" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B9413D26FFACD75A2FA11EA9A181FB20" box="[1005,1183,1196,1221]" italics="true" pageId="10" pageNumber="11">Nimbapanchax</emphasis>
</taxonomicName>
species for which the name is currently used. With regard to the morphological characters, the
<typeStatus id="548E5F96FFACD75A28641ED7A143FB09" box="[1064,1117,1234,1260]" pageId="10" pageNumber="11">type</typeStatus>
of
<taxonomicName id="4C359AB7FFACD75A28C51ED6A083FB09" box="[1161,1437,1235,1260]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="maeseni">
<emphasis id="B9413D26FFACD75A28C51ED6A083FB09" box="[1161,1437,1235,1260]" italics="true" pageId="10" pageNumber="11">Aphyosemion maeseni</emphasis>
</taxonomicName>
represents an
<taxonomicName id="4C359AB7FFACD75A2D081EFCA4A7FAF7" box="[324,441,1273,1298]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B9413D26FFACD75A2D081EFCA4A7FAF7" box="[324,441,1273,1298]" italics="true" pageId="10" pageNumber="11">Epiplatys</emphasis>
</taxonomicName>
species, and the
<taxonomicName id="4C359AB7FFACD75A2EDE1EFCA653FAF7" box="[658,845,1273,1298]" class="Actinopterygii" family="Nothobranchiidae" genus="Nimbapanchax" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B9413D26FFACD75A2EDE1EFCA653FAF7" box="[658,845,1273,1298]" italics="true" pageId="10" pageNumber="11">Nimbapanchax</emphasis>
</taxonomicName>
species for which this name is currently in use represents a scientifically unnamed species, which is described in the following. Until a revision of
<taxonomicName id="4C359AB7FFACD75A29611F25A082FADC" box="[1325,1436,1312,1337]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B9413D26FFACD75A29611F25A082FADC" box="[1325,1436,1312,1337]" italics="true" pageId="10" pageNumber="11">Epiplatys</emphasis>
</taxonomicName>
might show its synonymy with another species, the new combination
<taxonomicName id="4C359AB7FFACD75A2FA91F42A045FA85" authority="Poll, 1941" authorityName="Poll" authorityYear="1941" box="[997,1371,1350,1376]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="maeseni">
<emphasis id="B9413D26FFACD75A2FA91F42A1DCFA85" box="[997,1218,1351,1376]" italics="true" pageId="10" pageNumber="11">Epiplatys maeseni</emphasis>
(
<bibRefCitation id="EFA49CC5FFACD75A289F1F43A04CFA85" author="Poll" box="[1235,1362,1350,1376]" pageId="10" pageNumber="22" refString="Poll, M. (1941) Poissons nouveaux de la Cote d'Ivoire. Revue de Zoologie et de Botanique Africaines, 44, 133 - 143." type="journal article" year="1941">Poll, 1941</bibRefCitation>
)
</taxonomicName>
is an available name.
</paragraph>
</subSubSection>
</treatment>
</document>
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