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<document id="3FFDC9BEFCCD49505C5160F0B4A48681" ID-DOI="10.5281/zenodo.186868" ID-GBIF-Dataset="eb1f2a4c-93fc-40a4-ab13-c9728b6c7730" ID-ISSN="1175-5326" ID-Zenodo-Dep="186868" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461058779892" checkinUser="plazi" docAuthor="Mendes, Humberto Fonseca &amp; Andersen, Trond" docDate="2009" docId="9F680F0F461AFF8BFF4DFA14FC574E11" docLanguage="en" docName="zt02063p045.pdf" docOrigin="Zootaxa 2063" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Nilothauma strebulosum Adam et Saether, comb. n." docType="treatment" docVersion="9" lastPageNumber="37" masterDocId="635177774639FFAFFFDAFFD1FFC74F37" masterDocTitle="Neotropical Nilothauma Kieffer, 1921, with the description of thirteen new species (Diptera: Chironomidae)" masterLastPageNumber="45" masterPageNumber="1" pageNumber="36" updateTime="1698581706666" updateUser="plazi">
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<mods:title id="BAA171C3E7F9AAE221827DECF7165894">Neotropical Nilothauma Kieffer, 1921, with the description of thirteen new species (Diptera: Chironomidae)</mods:title>
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<mods:namePart id="C25BA98CFD2B50D407898EB1F6A657D8">Mendes, Humberto Fonseca</mods:namePart>
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<mods:namePart id="990CA28A3756DEED5EA4928E80DC403E">Andersen, Trond</mods:namePart>
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<mods:typeOfResource id="7ED4095887EB40A424617ABE4B05BD35">text</mods:typeOfResource>
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<mods:date id="602282B4B4505819D7171C9CAE116CB0">2009</mods:date>
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<mods:number id="B8A55568DAEE397F27BF1DF7A94D0751">2063</mods:number>
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<mods:identifier id="B6FBB2F5909CBD2367A5F284312779CD" type="DOI">10.5281/zenodo.186868</mods:identifier>
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<treatment id="9F680F0F461AFF8BFF4DFA14FC574E11" ID-DOI="http://doi.org/10.5281/zenodo.6224202" ID-GBIF-Taxon="119613661" ID-Zenodo-Dep="6224202" LSID="urn:lsid:plazi:treatment:9F680F0F461AFF8BFF4DFA14FC574E11" httpUri="http://treatment.plazi.org/id/9F680F0F461AFF8BFF4DFA14FC574E11" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<subSubSection id="5FDBED92461AFF8CFF4DFA14FEEA4935" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="177EBE19461AFF8CFF4DFA14FCDD4AE8" blockId="35.[151,794,1477,1538]" box="[151,794,1477,1503]" pageId="35" pageNumber="36">
<heading id="4C360975461AFF8CFF4DFA14FCDD4AE8" bold="true" box="[151,794,1477,1503]" fontSize="11" level="1" pageId="35" pageNumber="36" reason="1">
<emphasis id="25B5620B461AFF8CFF4DFA14FCDD4AE8" bold="true" box="[151,794,1477,1503]" pageId="35" pageNumber="36">
<taxonomicName id="D0C1C59A461AFF8CFF4DFA14FD604AE8" authority="Adam et Saether" authorityName="Adam et Saether" box="[151,679,1477,1503]" class="Insecta" family="Chironomidae" genus="Nilothauma" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="strebulosum" status="comb. nov.">
<emphasis id="25B5620B461AFF8CFF4DFA14FE044AE8" bold="true" box="[151,451,1477,1503]" italics="true" pageId="35" pageNumber="36">Nilothauma strebulosum</emphasis>
(Adam
<emphasis id="25B5620B461AFF8CFDF2FA14FDFA4AE8" bold="true" box="[552,573,1477,1503]" italics="true" pageId="35" pageNumber="36">et</emphasis>
Saether)
</taxonomicName>
<taxonomicNameLabel id="3E86DF70461AFF8CFD75FA14FCDD4AE8" box="[687,794,1477,1503]" pageId="35" pageNumber="36">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="177EBE19461AFF8CFF4DFA39FEEA4935" blockId="35.[151,794,1477,1538]" box="[151,301,1512,1538]" pageId="35" pageNumber="36">
(
<figureCitation id="8FFAA29C461AFF8CFF45FA39FEE24935" box="[159,293,1512,1538]" captionStart="FIGURES 61 63" captionStartId="36.[151,274,1263,1287]" captionTargetBox="[188,1383,356,1262]" captionTargetId="figure@36.[151,1436,323,1265]" captionTargetPageId="36" captionText="FIGURES 61 63. Nilothauma strebulosum (Adam et Saether) comb. n., male. 61 — dorsal aspect of hypopygium. 62 — hypopygium with tergite IX and anal point removed, dorsal view left, ventral view right. 63 — median and superior volsella, dorsal view." httpUri="https://zenodo.org/record/186887/files/figure.png" pageId="35" pageNumber="36">Figs 6163</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5FDBED92461AFF8CFF4DF9E4FCED497B" box="[151,810,1589,1612]" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="177EBE19461AFF8CFF4DF9E4FCED497B" blockId="35.[151,810,1589,1612]" box="[151,810,1589,1612]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="37D19937461AFF8CFF4DF9E4FCED497B" box="[151,810,1589,1612]" pageId="35" pageNumber="36">
<taxonomicName id="D0C1C59A461AFF8CFF4DF9E4FD1E497B" ID-CoL="4D65N" authority="Adam et Saether, 2000: 21" authorityName="Adam et Saether" authorityPageNumber="21" authorityYear="2000" box="[151,729,1589,1612]" class="Insecta" family="Chironomidae" genus="Paranilothauma" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="strebulosa">
<emphasis id="25B5620B461AFF8CFF4DF9E4FE7D497B" box="[151,442,1589,1612]" italics="true" pageId="35" pageNumber="36">Paranilothauma strebulosa</emphasis>
<treatmentCitation id="96609808461AFF8CFE1BF9E4FD1E497B" author="Adam" box="[449,729,1589,1612]" page="21" pageId="35" pageNumber="36" year="2000">
<bibRefCitation id="7350C3E8461AFF8CFE1BF9E4FD75497B" author="Adam" box="[449,690,1589,1612]" pageId="35" pageNumber="42" refString="Adam, J. I. &amp; Saether, O. A. (2000) Paranilothauma strebulosa sp. n. from Costa Rica (Diptera: Chironomidae). In: Hoffrichter, O. (Ed.), Late 20 th Century Research on Chironomidae: An Anthology from the 13 th International Symposium on Chironomidae. Shaker Verlag, Aachen, pp. 19 - 23." type="book chapter" year="2000">
Adam
<emphasis id="25B5620B461AFF8CFDD3F9E4FDDB497B" box="[521,540,1589,1612]" italics="true" pageId="35" pageNumber="36">et</emphasis>
Saether, 2000
</bibRefCitation>
: 21
</treatmentCitation>
</taxonomicName>
,
<figureCitation id="8FFAA29C461AFF8CFD3EF9E4FCE3497B" box="[740,804,1589,1612]" captionStart="FIGURES 1 2" captionStartId="4.[151,271,1941,1965]" captionTargetBox="[186,1311,291,1940]" captionTargetId="figure@4.[151,1436,291,1978]" captionTargetPageId="4" captionText="FIGURES 1 2. Parsimony analysis of Nilothauma Kieffer species with Paratendipes Kieffer and Pseudochironomus Malloch fixed as outgroup. Characters and character states as in text, matrix of characters as in Appendix 1. strict consensus of shortest tree. 2 — strict consensus of shortest tree after reweighting according to rescaled consistency index." httpUri="https://zenodo.org/record/186869/files/figure.png" pageId="35" pageNumber="36">Fig. 1</figureCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="5FDBED92461AFF8CFF4DF9AAFBCB4807" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="177EBE19461AFF8CFF4DF9AAFA51498B" blockId="35.[151,1437,1659,1956]" pageId="35" pageNumber="36">
<emphasis id="25B5620B461AFF8CFF4DF9AAFDAD49A2" bold="true" box="[151,618,1659,1685]" pageId="35" pageNumber="36">
Material examined.
<collectingCountry id="6FD6FE89461AFF8CFE78F9AAFDA249A2" box="[418,613,1659,1685]" name="Costa Rica" pageId="35" pageNumber="36">COSTA RICA</collectingCountry>
:
</emphasis>
La Selva,
<typeStatus id="C87A00BB461AFF8CFD28F9AAFC9849A2" box="[754,863,1659,1685]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
male,
<date id="637F98D9461AFF8CFC6EF9AAFBF949A2" box="[948,1086,1659,1685]" pageId="35" pageNumber="36" value="1993-03-16">16.iii.1993</date>
, Malaise trap, O. A. Saether (
<collectionCode id="71D026DC461AFF8CFF45F973FF31498B" box="[159,246,1698,1724]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="35" pageNumber="36">ZMBN</collectionCode>
<typeStatus id="C87A00BB461AFF8CFF27F973FEF0498B" box="[253,311,1698,1724]" pageId="35" pageNumber="36">Type</typeStatus>
no. 267).
<emphasis id="25B5620B461AFF8CFE74F973FDAB498B" bold="true" box="[430,620,1698,1724]" pageId="35" pageNumber="36">
<collectingCountry id="6FD6FE89461AFF8CFE74F973FDA0498B" box="[430,615,1698,1724]" name="Costa Rica" pageId="35" pageNumber="36">COSTA RICA</collectingCountry>
:
</emphasis>
Rancho Quemado,
<specimenCount id="01C77590461AFF8CFC8FF973FC63498B" box="[853,932,1698,1724]" pageId="35" pageNumber="36" type="male">2 male</specimenCount>
<typeStatus id="C87A00BB461AFF8CFC71F973FBD9498B" box="[939,1054,1698,1724]" pageId="35" pageNumber="36" type="paratype">paratypes</typeStatus>
, ix &amp; xii, 1990, W. Riss (
<collectionCode id="71D026DC461AFF8CFA9DF973FA4E498B" box="[1351,1417,1698,1724]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="35" pageNumber="36">ZSM</collectionCode>
).
</paragraph>
<paragraph id="177EBE19461AFF8CFF1FF919FBCB4807" blockId="35.[151,1437,1659,1956]" pageId="35" pageNumber="36">
<emphasis id="25B5620B461AFF8CFF1FF919FE1C49D5" bold="true" box="[197,475,1736,1762]" pageId="35" pageNumber="36">Diagnostic characters.</emphasis>
The two dorsal setose lobes on tergite IX and lack of anal point combined with a strongly curved inferior volsella with some apically split setae, and median volsella with strong apical seta will separate the male of
<taxonomicName id="D0C1C59A461AFF8CFE64F8C6FDA94807" box="[446,622,1815,1840]" class="Insecta" family="Chironomidae" genus="Nilothauma" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="strebulosum">
<emphasis id="25B5620B461AFF8CFE64F8C6FDA94807" box="[446,622,1815,1840]" italics="true" pageId="35" pageNumber="36">N. strebulosum</emphasis>
</taxonomicName>
from all other
<taxonomicName id="D0C1C59A461AFF8CFCC4F8C6FC6F4807" box="[798,936,1815,1840]" class="Insecta" family="Chironomidae" genus="Nilothauma" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="25B5620B461AFF8CFCC4F8C6FC6F4807" box="[798,936,1815,1840]" italics="true" pageId="35" pageNumber="36">Nilothauma</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="5FDBED92461AFF8CFF1FF8ECFDA94893" pageId="35" pageNumber="36" type="description">
<paragraph id="177EBE19461AFF8CFF1FF8ECFE33484A" blockId="35.[151,1437,1659,1956]" pageId="35" pageNumber="36">
<emphasis id="25B5620B461AFF8CFF1FF8ECFECC4860" bold="true" box="[197,267,1853,1879]" pageId="35" pageNumber="36">Male.</emphasis>
The male is described in detail by
<bibRefCitation id="7350C3E8461AFF8CFD7EF8ECFC084860" author="Adam" box="[676,975,1853,1879]" pageId="35" pageNumber="42" refString="Adam, J. I. &amp; Saether, O. A. (2000) Paranilothauma strebulosa sp. n. from Costa Rica (Diptera: Chironomidae). In: Hoffrichter, O. (Ed.), Late 20 th Century Research on Chironomidae: An Anthology from the 13 th International Symposium on Chironomidae. Shaker Verlag, Aachen, pp. 19 - 23." type="book chapter" year="2000">Adam and Saether (2000)</bibRefCitation>
. Hypopygium and superior and median volsellae as in
<figureCitation id="8FFAA29C461AFF8CFE9EF8B2FE37484A" box="[324,496,1891,1917]" captionStart="FIGURES 61 63" captionStartId="36.[151,274,1263,1287]" captionTargetBox="[188,1383,356,1262]" captionTargetId="figure@36.[151,1436,323,1265]" captionTargetPageId="36" captionText="FIGURES 61 63. Nilothauma strebulosum (Adam et Saether) comb. n., male. 61 — dorsal aspect of hypopygium. 62 — hypopygium with tergite IX and anal point removed, dorsal view left, ventral view right. 63 — median and superior volsella, dorsal view." httpUri="https://zenodo.org/record/186887/files/figure.png" pageId="35" pageNumber="36">Figures 6163</figureCitation>
.
</paragraph>
<paragraph id="177EBE19461AFF8CFF1FF85BFDA94893" blockId="35.[151,1437,1659,1956]" box="[197,622,1930,1956]" pageId="35" pageNumber="36">
<emphasis id="25B5620B461AFF8CFF1FF85BFE2B4893" bold="true" box="[197,492,1930,1956]" pageId="35" pageNumber="36">Female and immatures.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="5FDBED92461DFF8BFF1FFF46FC574E11" pageId="36" pageNumber="37" type="discussion">
<paragraph id="177EBE19461DFF8BFF1FFF46FC574E11" blockId="36.[151,1436,151,294]" pageId="36" pageNumber="37">
<emphasis id="25B5620B461DFF8BFF1FFF46FE864F86" bold="true" box="[197,321,151,177]" pageId="36" pageNumber="37">Remarks.</emphasis>
This species was originally described in
<taxonomicName id="D0C1C59A461DFF8BFCE6FF49FBC54F86" box="[828,1026,152,177]" class="Insecta" family="Chironomidae" genus="Paranilothauma" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="25B5620B461DFF8BFCE6FF49FBC54F86" box="[828,1026,152,177]" italics="true" pageId="36" pageNumber="37">Paranilothauma</emphasis>
</taxonomicName>
as the two dorsal setose lobes on tergite IX were regarded as belonging to laterosternite IX (
<bibRefCitation id="7350C3E8461DFF8BFCE5FF6FFB824FEF" author="Adam" box="[831,1093,190,216]" pageId="36" pageNumber="42" refString="Adam, J. I. &amp; Saether, O. A. (2000) Paranilothauma strebulosa sp. n. from Costa Rica (Diptera: Chironomidae). In: Hoffrichter, O. (Ed.), Late 20 th Century Research on Chironomidae: An Anthology from the 13 th International Symposium on Chironomidae. Shaker Verlag, Aachen, pp. 19 - 23." type="book chapter" year="2000">Adam &amp; Saether 2000</bibRefCitation>
). The addition of several new Neotropical species with similar lobes clearly demonstrates that these lobes originate on tergite IX.
<emphasis id="25B5620B461DFF8BFF1FFEDAFEA24E12" bold="true" box="[197,357,267,293]" pageId="36" pageNumber="37">Distribution.</emphasis>
This species is recorded only from
<collectingCountry id="6FD6FE89461DFF8BFCD2FEDDFC4C4E11" box="[776,907,268,294]" name="Costa Rica" pageId="36" pageNumber="37">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>