treatments-xml/data/5C/33/87/5C338787FE714B0F82FD0FC1FC14C5DE.xml
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<document id="6D7664C453A1ECF09574109224EE212F" ID-DOI="10.11646/zootaxa.5437.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10984997" ID-ZooBank="79196FA8-AC75-4768-AC43-9C9C1B515FB9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1713344825674" checkinUser="plazi" docAuthor="Lopes, Rogério B. &amp; Noll, Fernando B." docDate="2024" docId="5C338787FE714B0F82FD0FC1FC14C5DE" docLanguage="en" docName="zootaxa.5437.3.3.pdf" docOrigin="Zootaxa 5437 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5437.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zethus (Zethoides) peruvicus Bohart &amp; Stange 1965" docType="treatment" docVersion="3" lastPageNumber="377" masterDocId="A00AFFFFFE7F4B01826A0F56FFBEC354" masterDocTitle="Taxonomic notes on the Zethus olmecus de Saussure, 1875 species group (Hymenoptera, Vespidae, Eumeninae), with the description of five new species" masterLastPageNumber="383" masterPageNumber="363" pageNumber="377" updateTime="1713815074103" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="26EE82B2F812EAAB5FC9D4B168B34FD4">Taxonomic notes on the Zethus olmecus de Saussure, 1875 species group (Hymenoptera, Vespidae, Eumeninae), with the description of five new species</mods:title>
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<mods:namePart id="0A8C7EAFBE76E9F53879A6A292158808">Lopes, Rogério B.</mods:namePart>
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<mods:affiliation id="4C12D94E9B5921E612B8D61ACD904541">Universidade Estadual Paulista “ Júlio de Mesquita Filho ”, Instituto de Biociências, Letras e Ciências Exatas, Departamento de Ciências Biológicas</mods:affiliation>
<mods:nameIdentifier id="86E989CCEE1F6446A03462F6E7D99C36" type="email">rbotlopes@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B5778E47B264B3E98B721BB7E9F1491C">Noll, Fernando B.</mods:namePart>
<mods:affiliation id="98373CA3E27281C044C6D01AF79B0BF3">Universidade Estadual Paulista “ Júlio de Mesquita Filho ”, Instituto de Biociências, Letras e Ciências Exatas, Departamento de Ciências Biológicas</mods:affiliation>
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<mods:title id="71587F8D68610A9BF53E7EC25498CF5D">Zootaxa</mods:title>
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<mods:date id="BB3D4C941195767D86108F29316CCDF5">2024</mods:date>
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<treatment id="5C338787FE714B0F82FD0FC1FC14C5DE" LSID="urn:lsid:plazi:treatment:5C338787FE714B0F82FD0FC1FC14C5DE" httpUri="http://treatment.plazi.org/id/5C338787FE714B0F82FD0FC1FC14C5DE" lastPageNumber="377" pageId="14" pageNumber="377">
<subSubSection id="9C80651AFE714B0F82FD0FC1FCA2C3E5" box="[151,796,151,178]" pageId="14" pageNumber="377" type="nomenclature">
<paragraph id="D4253691FE714B0F82FD0FC1FCA2C3E5" blockId="14.[151,796,151,178]" box="[151,796,151,178]" pageId="14" pageNumber="377">
<heading id="8F6D81FDFE714B0F82FD0FC1FCA2C3E5" bold="true" box="[151,796,151,178]" fontSize="11" level="1" pageId="14" pageNumber="377" reason="1">
<taxonomicName id="139A4D12FE714B0F82FD0FC1FCA2C3E5" ID-CoL="7GFLW" authority="Bohart &amp; Stange, 1965" authorityName="Bohart &amp; Stange" authorityYear="1965" box="[151,796,151,178]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="377" phylum="Arthropoda" rank="species" species="peruvicus" subGenus="Zethoides">
<emphasis id="E6EEEA83FE714B0F82FD0FC1FCA2C3E5" bold="true" box="[151,796,151,178]" pageId="14" pageNumber="377">
<emphasis id="E6EEEA83FE714B0F82FD0FC1FF54C3E6" bold="true" box="[151,234,151,178]" italics="true" pageId="14" pageNumber="377">Zethus</emphasis>
(
<emphasis id="E6EEEA83FE714B0F82910FC1FECEC3E6" bold="true" box="[251,368,151,178]" italics="true" pageId="14" pageNumber="377">Zethoides</emphasis>
)
<emphasis id="E6EEEA83FE714B0F83E80FC1FE49C3E5" bold="true" box="[386,503,151,177]" italics="true" pageId="14" pageNumber="377">peruvicus</emphasis>
<bibRefCitation id="B00B4B60FE714B0F83950FC1FCA2C3E5" author="Bohart, R. M. &amp; Stange, L. A." box="[511,796,151,177]" pageId="14" pageNumber="377" pagination="1 - 208" refId="ref14458" refString="Bohart, R. M. &amp; Stange, L. A. (1965) A Revision of the genus Zethus Fabricius in the Western Hemisphere (Hymenoptera: Eumenidae). University of California Publications in Entomology, 40, 1 - 208." type="journal article" year="1965">Bohart &amp; Stange, 1965</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9C80651AFE714B0F82FD0F8AFEF1C265" pageId="14" pageNumber="377" type="reference_group">
<paragraph id="D4253691FE714B0F82FD0F8AFEF1C265" blockId="14.[151,1436,219,305]" pageId="14" pageNumber="377">
<treatmentCitationGroup id="F48A11BFFE714B0F82FD0F8AFEF1C265" pageId="14" pageNumber="377">
<treatmentCitation id="553B1080FE714B0F82FD0F8AFD17C3A7" author="Bohart, R. M. &amp; Stange, L. A." box="[151,681,219,244]" page="160" pageId="14" pageNumber="377" year="1965">
<taxonomicName id="139A4D12FE714B0F82FD0F8AFD17C3A7" ID-CoL="7GFLW" authority="Bohart &amp; Stange, 1965: 159 - 160" authorityName="Bohart &amp; Stange" authorityPageNumber="159 - 160" authorityYear="1965" box="[151,681,219,244]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="377" phylum="Arthropoda" rank="species" species="peruvicus">
<emphasis id="E6EEEA83FE714B0F82FD0F8AFEF8C3A7" box="[151,326,220,244]" italics="true" pageId="14" pageNumber="377">Zethus peruvicus</emphasis>
<bibRefCitation id="B00B4B60FE714B0F83250F8AFD17C3A7" author="Bohart, R. M. &amp; Stange, L. A." box="[335,681,219,244]" pageId="14" pageNumber="377" pagination="1 - 208" refId="ref14458" refString="Bohart, R. M. &amp; Stange, L. A. (1965) A Revision of the genus Zethus Fabricius in the Western Hemisphere (Hymenoptera: Eumenidae). University of California Publications in Entomology, 40, 1 - 208." type="journal article" year="1965">Bohart &amp; Stange, 1965: 159160</bibRefCitation>
</taxonomicName>
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(key), 167, 171 (Figs 165167) (in subgenus
<taxonomicName id="139A4D12FE714B0F86E40F8AFB4FC3A0" box="[1166,1265,220,244]" pageId="14" pageNumber="377" rank="subGenus" subGenus="Zethoides">
<emphasis id="E6EEEA83FE714B0F86E40F8AFB4FC3A0" box="[1166,1265,220,244]" italics="true" pageId="14" pageNumber="377">Zethoides</emphasis>
</taxonomicName>
) 204 (aedeagus, fig. 343).
<treatmentCitation id="553B1080FE714B0F83460FADFDD8C246" author="Rasmussen, C. &amp; Asenjo, A." box="[300,614,250,274]" page="43" pageId="14" pageNumber="377" year="2009">
<bibRefCitation id="B00B4B60FE714B0F83460FADFDD8C246" author="Rasmussen, C. &amp; Asenjo, A." box="[300,614,250,274]" pageId="14" pageNumber="377" pagination="1 - 78" refId="ref15654" refString="Rasmussen, C. &amp; Asenjo, A. (2009) A checklist to the wasps of Peru (Hymenoptera, Aculeata). ZooKeys, 15, 1 - 78. https: // doi. org / 10.3897 / zookeys. 15.196" type="journal article" year="2009">Rasmussen &amp; Asenjo 2009: 43</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="AC8D7601FE714B0F801E0FACFD16C246" box="[628,680,250,274]" name="Peru" pageId="14" pageNumber="377">Peru</collectingCountry>
). Type data:
<typeStatus id="0B218833FE714B0F81410FACFC37C246" box="[811,905,250,274]" pageId="14" pageNumber="377" type="holotype">Holotype</typeStatus>
male, Belém [MPEG]. Type locality: “Iquitos,
<collectingCountry id="AC8D7601FE714B0F87040FADFA22C246" box="[1390,1436,251,274]" name="Peru" pageId="14" pageNumber="377">Peru</collectingCountry>
Amazonico”.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9C80651AFE714B0F82FD0E09FB86C22E" box="[151,1080,351,378]" pageId="14" pageNumber="377" type="diagnosis">
<paragraph id="D4253691FE714B0F82FD0E09FB86C22E" blockId="14.[151,1437,351,1674]" box="[151,1080,351,378]" pageId="14" pageNumber="377">
<emphasis id="E6EEEA83FE714B0F82FD0E09FEB4C22D" bold="true" box="[151,266,351,377]" pageId="14" pageNumber="377">Diagnosis</emphasis>
. The tegula posteriorly bowed out and wide is diagnostic for both sexes.
</paragraph>
</subSubSection>
<subSubSection id="9C80651AFE714B0F82AD0ED5FB2DC295" pageId="14" pageNumber="377" type="materials_examined">
<paragraph id="D4253691FE714B0F82AD0ED5FB2DC295" blockId="14.[151,1437,351,1674]" pageId="14" pageNumber="377">
<emphasis id="E6EEEA83FE714B0F82AD0ED5FE15C2C9" bold="true" box="[199,427,387,413]" pageId="14" pageNumber="377">Material examined</emphasis>
.
<materialsCitation id="64F23CCCFE714B0F83D60ED5FA27C2C9" box="[444,1433,387,416]" collectionCode="MPEG" country="Peru" location="Iquitos" pageId="14" pageNumber="377" specimenCount="2" specimenCount-female="2" stateProvince="Loreto" typeStatus="holotype">
<specimenCount id="C29CFD18FE714B0F83D60ED5FE5EC2C9" box="[444,480,387,413]" count="2" pageId="14" pageNumber="377" type="female">2♀</specimenCount>
, 1♁ (see Supplementary Material 1).
<collectingCountry id="AC8D7601FE714B0F81F90ED2FC61C2CA" box="[915,991,388,414]" name="Peru" pageId="14" pageNumber="377">PERU</collectingCountry>
:
<typeStatus id="0B218833FE714B0F81860ED5FBEFC2C9" box="[1004,1105,387,413]" pageId="14" pageNumber="377" type="holotype">holotype</typeStatus>
, ♁,
<collectingRegion id="165EF873FE714B0F86E90ED2FB6FC2CA" box="[1155,1233,388,414]" country="Peru" name="Loreto" pageId="14" pageNumber="377">Loreto</collectingRegion>
,
<location id="D145604AFE714B0F86B40ED5FA92C2C9" LSID="urn:lsid:plazi:treatment:5C338787FE714B0F82FD0FC1FC14C5DE:D145604AFE714B0F86B40ED5FA92C2C9" box="[1246,1324,387,413]" country="Peru" name="Iquitos" pageId="14" pageNumber="377" stateProvince="Loreto">Iquitos</location>
[
<collectionCode id="B28BAE54FE714B0F87510ED5FA2CC2C9" box="[1339,1426,387,413]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="14" pageNumber="377">MPEG</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="64F23CCCFE714B0F82FD0EF1FD25C296" box="[151,667,423,450]" collectionCode="DZUP" country="Brazil" location="Caceres" pageId="14" pageNumber="377" specimenCount="1" specimenCount-female="1" stateProvince="Mato Grosso" typeStatus="holotype">
<collectingCountry id="AC8D7601FE714B0F82FD0EF1FEBFC295" box="[151,257,423,449]" name="Brazil" pageId="14" pageNumber="377">BRAZIL</collectingCountry>
:
<specimenCount id="C29CFD18FE714B0F83610EF1FE90C295" box="[267,302,423,449]" count="1" pageId="14" pageNumber="377" type="female">1♀</specimenCount>
,
<collectingRegion id="165EF873FE714B0F83560EFEFE6EC295" box="[316,464,423,450]" country="Brazil" name="Mato Grosso" pageId="14" pageNumber="377">Mato Grosso</collectingRegion>
,
<location id="D145604AFE714B0F83B60EF1FD8BC295" LSID="urn:lsid:plazi:treatment:5C338787FE714B0F82FD0FC1FC14C5DE:D145604AFE714B0F83B60EF1FD8BC295" box="[476,565,423,449]" country="Brazil" name="Caceres" pageId="14" pageNumber="377" stateProvince="Mato Grosso">Cáceres</location>
[
<collectionCode id="B28BAE54FE714B0F80280EFEFD2AC296" box="[578,660,424,450]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="14" pageNumber="377" type="University or college">DZUP</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="64F23CCCFE714B0F80CC0EF1FB2EC295" box="[678,1168,423,450]" collectionCode="UFGD" country="Brazil" location="Pantanal" pageId="14" pageNumber="377" specimenCount="1" specimenCount-female="1" stateProvince="Mato Grosso do Sul" typeStatus="holotype">
<specimenCount id="C29CFD18FE714B0F80CC0EF1FD77C295" box="[678,713,423,449]" count="1" pageId="14" pageNumber="377" type="female">1♀</specimenCount>
,
<collectingRegion id="165EF873FE714B0F80BD0EFEFC04C295" box="[727,954,423,450]" country="Brazil" name="Mato Grosso do Sul" pageId="14" pageNumber="377">Mato Grosso do Sul</collectingRegion>
,
<location id="D145604AFE714B0F81AF0EF1FB9BC295" LSID="urn:lsid:plazi:treatment:5C338787FE714B0F82FD0FC1FC14C5DE:D145604AFE714B0F81AF0EF1FB9BC295" box="[965,1061,423,449]" country="Brazil" name="Pantanal" pageId="14" pageNumber="377" stateProvince="Mato Grosso do Sul">Pantanal</location>
[
<collectionCode id="B28BAE54FE714B0F865E0EF1FB37C295" box="[1076,1161,423,449]" pageId="14" pageNumber="377">UFGD</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9C80651AFE714B0F82AD0E9DFF4AC6AE" pageId="14" pageNumber="377" type="description">
<paragraph id="D4253691FE714B0F82AD0E9DFF4AC6AE" blockId="14.[151,1437,351,1674]" pageId="14" pageNumber="377">
<emphasis id="E6EEEA83FE714B0F82AD0E9DFEEFC2B1" bold="true" box="[199,337,459,485]" pageId="14" pageNumber="377">Description</emphasis>
.
<emphasis id="E6EEEA83FE714B0F830A0E9DFE0AC2B1" box="[352,436,459,485]" italics="true" pageId="14" pageNumber="377">Female</emphasis>
(hitherto unknown).
<emphasis id="E6EEEA83FE714B0F80C80E9DFC94C2B1" bold="true" box="[674,810,459,485]" pageId="14" pageNumber="377">Coloration.</emphasis>
Black, with the following markings orange: mandibles; clypeus; scape and pedicel entirely; inner-ventral surface of flagellum most of gena, except narrow stripes on dorsal half along eye margin and genal margin; spots wide band from submedian portion of pronotum extending towards, but not reaching, pronotal lobe; large scrobal spot; apical half of parategula; very small pair of spots on scutellum and metanotum; fore and midlegs. Pale yellow: Hypostoma; gena inferiorly; proepisternum; anterior surface for forelegs apical bands of T2 and 3. Yellow: Outer surface of mid-tibia; apical band of T1. Propodeal apical lamella dark with hyaline tip and valvula amber. Wings hyaline with light brown costa and dark brown venation.
<emphasis id="E6EEEA83FE714B0F87480DF5FA22C1E9" bold="true" box="[1314,1436,675,701]" pageId="14" pageNumber="377">Structure.</emphasis>
Mandible 5-toothed. Clypeus with straight apex. Dorsum of clypeus and interantennal carina flat. Gena in dorsal view angulate. Pronotal carina with constant height. Humeral angles not projected. Pronotum short, humeral-tegular distance shorter than inter-humeral distance. Parategula digitiform. Tegula with outer margin posteriorly bowed out. Scutellum flat. Metanotum convex. Mid-tibia with two spurs. Dorsum of propodeum lower than metanotum and scutellum. Apical propodeal lamella triangular with wide rounded tip. T
<quantity id="13629B74FE714B0F81A30C01FC4FC025" box="[969,1009,855,881]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="377" unit="in" value="1.0">1 in</quantity>
dorsal view with angulate expansion. S2 angulate in profile with a long straight posterior portion.
<emphasis id="E6EEEA83FE714B0F81070C2DFC57C0C1" bold="true" box="[877,1001,891,917]" pageId="14" pageNumber="377">Sculpture.</emphasis>
Clypeus moderately striato-punctate. Dense macro and micropunctation on frons and vertex, sparser on gena where a small dorso-lateral portion is rather smooth, but not shiny. Pronotum and mesoscutum shiny with moderate macropunctation intercalated my very dense micropunctation. Scutellum with microstriae between sparse macropunctation, Mesepisternum with moderate micro and macropunctation. Metanotum densely micro and macropunctate. Propodeum with a few large sparse crater-like punctures, smaller and coalescent laterally with weak stria medially. Lateral carina complete. Submedian carina on basal 2/3 of propodeum, with base curved in lateral view. Submarginal carina slightly convex on dorsal portion, with obtuse dorso-lateral angles. T1 with basal transverse carina, and dense macropunctation with smooth shiny interspaces. T23 with very dense micropunctation obscuring macropunctures stopping close to apex forming a narrow smooth band, wider on T3. S23 moderately macropunctate with smooth shiny interspaces. Very dense micropunctation on T45 and S46, becoming moderate to sparse on T6.
<emphasis id="E6EEEA83FE714B0F86EA0BB5FB5EC7A9" bold="true" box="[1152,1248,1251,1277]" pageId="14" pageNumber="377">Pilosity.</emphasis>
Clypeus mostly bare, except for dense patch along dorso-lateral margins. Short, thin, moderate to sparse setae along frons, most of vertex, dorsum of pronotum, scutellum and inferior half of mesepisternum. Moderate to dense on mesoscutum. Very dense, tomentose, on vertex behind ocellar region, lateral surface of pronotum, most of mesopleura, metapleural and propodeum (becomes longer on apex). T1 with sparse short setae. T2 tomentose. S2 with moderate, thick, and long pilosity. T35 and S35 with apical row of long setae, longer on sterna T6 and S6 with erect thin setae, short towards middle of sclerite and longer (not as long as preceding sclerites) apically.
<emphasis id="E6EEEA83FE714B0F865B0AEDFB63C681" bold="true" box="[1073,1245,1467,1493]" pageId="14" pageNumber="377">Measurement.</emphasis>
Forewing length
<quantity id="13629B74FE714B0F82FD0A89FF4EC6AE" box="[151,240,1503,1530]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="14" pageNumber="377" unit="mm" value="6.8">6,8 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="9C80651AFE714B0F82AD0955FC14C5DE" pageId="14" pageNumber="377" type="distribution">
<paragraph id="D4253691FE714B0F82AD0955FAEAC549" blockId="14.[151,1437,351,1674]" box="[199,1364,1539,1566]" pageId="14" pageNumber="377">
<emphasis id="E6EEEA83FE714B0F82AD0955FEE7C549" bold="true" box="[199,345,1539,1565]" pageId="14" pageNumber="377">Distribution</emphasis>
.
<collectingCountry id="AC8D7601FE714B0F830D0952FE23C54A" box="[359,413,1540,1566]" name="Peru" pageId="14" pageNumber="377">Peru</collectingCountry>
, *
<collectingCountry id="AC8D7601FE714B0F83DE0955FE45C549" box="[436,507,1539,1565]" name="Brazil" pageId="14" pageNumber="377">Brazil</collectingCountry>
(
<collectingRegion id="165EF873FE714B0F80600955FD1EC549" box="[522,672,1539,1565]" country="Brazil" name="Mato Grosso" pageId="14" pageNumber="377">Mato Grosso</collectingRegion>
,
<collectingRegion id="165EF873FE714B0F80C10952FC33C549" box="[683,909,1539,1566]" country="Brazil" name="Mato Grosso do Sul" pageId="14" pageNumber="377">Mato Grosso do Sul</collectingRegion>
) (
<bibRefCitation id="B00B4B60FE714B0F81CE0955FB1AC54A" author="Bohart, R. M. &amp; Stange, L. A." box="[932,1188,1539,1566]" pageId="14" pageNumber="377" pagination="1 - 208" refId="ref14458" refString="Bohart, R. M. &amp; Stange, L. A. (1965) A Revision of the genus Zethus Fabricius in the Western Hemisphere (Hymenoptera: Eumenidae). University of California Publications in Entomology, 40, 1 - 208." type="journal article" year="1965">Bohart &amp; Stange 1965</bibRefCitation>
; and new data).
</paragraph>
<paragraph id="D4253691FE714B0F82AD0971FC14C5DE" blockId="14.[151,1437,351,1674]" pageId="14" pageNumber="377">
<emphasis id="E6EEEA83FE714B0F82AD0971FED8C515" bold="true" box="[199,358,1575,1601]" pageId="14" pageNumber="377">Observations</emphasis>
. The examined females, although collected very far South from the
<typeStatus id="0B218833FE714B0F8616097EFB12C516" box="[1148,1196,1576,1602]" pageId="14" pageNumber="377">type</typeStatus>
location, present the bowed tegula unique to this species regarding the group. Some pale markings are smaller or missing in the female from Pantanal, which is also smaller, with forewing length of
<quantity id="13629B74FE714B0F81270939FC18C5DE" box="[845,934,1647,1674]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="14" pageNumber="377" unit="mm" value="6.4">6,4 mm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>