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<document id="13DFA9314AEA82EA98F68BF76D6221C3" ID-CLB-Dataset="229105" ID-DOI="10.11646/zootaxa.5230.2.3" ID-GBIF-Dataset="997e16f6-a408-4411-93cc-53fa1b085c92" ID-ISSN="1175-5326" ID-Zenodo-Dep="7553485" ID-ZooBank="7D2F9E0B-9E32-40E4-8DED-9E6227BAEFD5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1674200759302" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2023" docId="1F070A1EC6603C4AA1DCFA049597FF69" docLanguage="en" docName="zootaxa.5230.2.3.pdf" docOrigin="Zootaxa 5230 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nesamblyops townsendi Sokolov 2023, sp. nov." docType="treatment" docVersion="7" lastPageNumber="198" masterDocId="E33E7266C6713C5EA14BFFB9901CFF90" masterDocTitle="Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island" masterLastPageNumber="201" masterPageNumber="179" pageNumber="196" updateTime="1699474157326" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="89F45CD835225ABEB9841C933A597762">Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island</mods:title>
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<mods:namePart id="D2B17ABD4BBAD97F2493F2C584CAED5B">Sokolov, Igor M.</mods:namePart>
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<mods:title id="49B52CA6D8F4E89FFD5A31D91A7BF1B7">Zootaxa</mods:title>
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<mods:date id="09D7B14AB411F97CB2D84500FC7CD893">2023</mods:date>
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<mods:number id="F25F1917CB61C7A14E692637ECB21E4A">2023-01-20</mods:number>
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<mods:number id="3396D7E93584111F9305A2E3105F2ED6">5230</mods:number>
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<treatment id="1F070A1EC6603C4AA1DCFA049597FF69" ID-DOI="http://doi.org/10.5281/zenodo.7555124" ID-GBIF-Taxon="209011118" ID-Zenodo-Dep="7555124" LSID="urn:lsid:plazi:treatment:1F070A1EC6603C4AA1DCFA049597FF69" httpUri="http://treatment.plazi.org/id/1F070A1EC6603C4AA1DCFA049597FF69" lastPageId="20" lastPageNumber="198" pageId="17" pageNumber="196">
<subSubSection id="DFB4E883C6603C4FA1DCFA049239FA47" box="[151,549,1469,1496]" pageId="17" pageNumber="196" type="nomenclature">
<paragraph id="9711BB08C6603C4FA1DCFA049239FA47" blockId="17.[151,549,1469,1532]" box="[151,549,1469,1496]" pageId="17" pageNumber="196">
<heading id="CC590C64C6603C4FA1DCFA049239FA47" bold="true" box="[151,549,1469,1496]" fontSize="11" level="1" pageId="17" pageNumber="196" reason="1">
<emphasis id="A5DA671AC6603C4FA1DCFA049239FA47" bold="true" box="[151,549,1469,1496]" pageId="17" pageNumber="196">
<taxonomicName id="50AEC08BC6603C4FA1DCFA0491A4FA48" authority="Sokolov, 2023" authorityName="Sokolov" authorityYear="2023" box="[151,440,1469,1496]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="196" phylum="Arthropoda" rank="species" species="townsendi" status="sp. nov.">
<emphasis id="A5DA671AC6603C4FA1DCFA0491A4FA48" bold="true" box="[151,440,1469,1496]" italics="true" pageId="17" pageNumber="196">Nesamblyops townsendi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BEE9DA61C6603C4FA08CFA049239FA47" box="[455,549,1469,1495]" pageId="17" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DFB4E883C6603C4FA1DCFA5891B3FA6B" box="[151,431,1505,1532]" pageId="17" pageNumber="196" type="description">
<paragraph id="9711BB08C6603C4FA1DCFA5891B3FA6B" blockId="17.[151,549,1469,1532]" box="[151,431,1505,1532]" pageId="17" pageNumber="196">
<figureCitation id="0F95A78DC6603C4FA1DCFA5890E4FA6B" box="[151,248,1505,1531]" captionStart="FIGURE 3" captionStartId="8.[151,250,957,981]" captionTargetBox="[151,1435,151,943]" captionTargetId="figure-392@8.[151,1435,151,943]" captionTargetPageId="8" captionText="FIGURE 3. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. parvulus (Mount Stokes, Marlborough Sounds, SO), B—N. tararua (Judd Ridge, Tararua Range, Wellington, NO), C—N. townsendi (Tennyson Inlet, Marlborough Sounds, SO). Scale bar 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7553495" httpUri="https://zenodo.org/record/7553495/files/figure.png" pageId="17" pageNumber="196">Figs. 3C</figureCitation>
,
<figureCitation id="0F95A78DC6603C4FA04FFA589102FA6B" box="[260,286,1505,1531]" captionStart="FIGURE 4" captionStartId="10.[151,250,1222,1246]" captionTargetBox="[151,1436,181,1197]" captionTargetId="figure-28@10.[151,1436,181,1198]" captionTargetPageId="10" captionText="FIGURE 4. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. brouni (Lewis Pass, North Canterbury, SO), B—N. carltoni (Dun Mountain, Nelson, SO), C—N. confusus (Mount Stokes, Marlborough Sounds, SO), D—N. distinctus (Fabians Valley, Marlborough, SO), E—N. lescheni (DUrville Island, Cook Strait, Marlborough Sounds, SO), F—N. oreobius (Mount Pirongia, Waikato, NO), G—N. parvulus (Mount Stokes, Marlborough Sounds, SO), H—N. tararua (Judd Ridge, Tararua Range, Wellington, NO), I—N. townsendi (Tennyson Inlet, Marlborough Sounds, SO). Scale bar 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7553497" httpUri="https://zenodo.org/record/7553497/files/figure.png" pageId="17" pageNumber="196">4I</figureCitation>
,
<figureCitation id="0F95A78DC6603C4FA062FA589179FA6B" box="[297,357,1505,1531]" captionStart="FIGURE 6" captionStartId="13.[151,250,993,1017]" captionTargetBox="[157,1426,159,968]" captionTargetId="figure-403@13.[151,1436,151,978]" captionTargetPageId="13" captionText="FIGURE 6. Line drawings of male genitalia of New Zealand Nesamblyops species. N. parvulus (Mount Stokes, Marlborough Sounds, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. tararua (Judd Ridge, Tararua Range, Wellington, NO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. townsendi (Tennyson Inlet, Marlborough Sounds, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. Scale bar 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.7553503" httpUri="https://zenodo.org/record/7553503/files/figure.png" pageId="17" pageNumber="196">6GI</figureCitation>
,
<figureCitation id="0F95A78DC6603C4FA03AFA5B9189FA6C" box="[369,405,1506,1532]" captionStart="FIGURE 7" captionStartId="15.[151,250,1470,1494]" captionTargetBox="[163,1424,161,1453]" captionTargetId="figure-210@15.[151,1436,151,1466]" captionTargetPageId="15" captionText="FIGURE 7. Line drawings of ring sclerite of New Zealand Nesamblyops species, male genitalia, dorsal aspect. A—N. brouni (Lewis Pass, North Canterbury, SO), B—N. carltoni (Dun Mountain, Nelson, SO), C—N. distinctus (Fabians Valley, Marlborough, SO), D—N. oreobius (Mount Pirongia, Waikato, NO), E—N. lescheni (DUrville Island, Cook Strait, Marlborough Sounds, SO), F—N. parvulus (Mount Stokes, Marlborough Sounds, SO), G—N. tararua (Judd Ridge, Tararua Range, Wellington, NO), H—N. townsendi (Tennyson Inlet, Marlborough Sounds, SO). Scale = 0.4mm." figureDoi="http://doi.org/10.5281/zenodo.7553507" httpUri="https://zenodo.org/record/7553507/files/figure.png" pageId="17" pageNumber="196">7H</figureCitation>
,
<figureCitation id="0F95A78DC6603C4FA0EAFA5891B3FA6B" box="[417,431,1505,1531]" captionStart="FIGURE 9" captionStartId="17.[151,250,1291,1315]" captionTargetBox="[151,1437,159,1280]" captionTargetId="figure-326@17.[151,1437,159,1280]" captionTargetPageId="17" captionText="FIGURE 9. Distributional records for the Nesamblyops species and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Black circles—N. tararua; yellow circles—N. townsendi; red circles—N. distinctus; green circles—N. brouni. Scale bar= 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7553513" httpUri="https://zenodo.org/record/7553513/files/figure.png" pageId="17" pageNumber="196">9</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="9711BB08C6603C4FA1DCF99D94A7F886" blockId="17.[151,1437,1572,2030]" pageId="17" pageNumber="196">
<emphasis id="A5DA671AC6603C4FA1DCF99D9120F9AE" bold="true" box="[151,316,1572,1598]" pageId="17" pageNumber="196">Type material.</emphasis>
<materialsCitation id="27C6B155C6603C4FA009F99D9109F9F2" ID-GBIF-Occurrence="4016386304" collectionCode="NZAC, DSIR" collectorName="J. I. Townsend" country="New Zealand" location="No" pageId="17" pageNumber="196" specimenCount="1" typeStatus="holotype">
<emphasis id="A5DA671AC6603C4FA009F99D91CEF9AE" bold="true" box="[322,466,1572,1598]" italics="true" pageId="17" pageNumber="196">
<typeStatus id="481505AAC6603C4FA009F99D91CEF9AE" box="[322,466,1572,1598]" pageId="17" pageNumber="196" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
, male, in
<collectionCode id="F1BF23CDC6603C4FA373F99D9298F9AE" box="[568,644,1572,1598]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="17" pageNumber="196" type="Other institutional type">NZAC</collectionCode>
, labeled: Tennyson Inlet Marlb. 22.5.64
<collectorName id="3A5BDEDEC6603C4FA567F99D94A4F9AE" box="[1068,1208,1572,1598]" pageId="17" pageNumber="196">J.I.Townsend</collectorName>
\ Forest litter \
<collectionCode id="F1BF23CDC6603C4FA41CF99D9588F9AE" box="[1367,1428,1572,1598]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="17" pageNumber="196">DSIR</collectionCode>
\
<location id="9271EDD3C6603C4FA1DCF9F190A4F9F2" LSID="urn:lsid:plazi:treatment:1F070A1EC6603C4AA1DCFA049597FF69:9271EDD3C6603C4FA1DCF9F190A4F9F2" box="[151,184,1608,1634]" country="New Zealand" name="No" pageId="17" pageNumber="196">No</location>
\
<collectingCountry id="EFB9FB98C6603C4FA181F9F190F2F9F2" box="[202,238,1608,1634]" name="New Zealand" pageId="17" pageNumber="196">NZ</collectingCountry>
PB
</materialsCitation>
\.
<materialsCitation id="27C6B155C6603C4FA06CF9F1939EF9F2" ID-GBIF-Occurrence="4016386399" box="[295,898,1608,1634]" collectionCode="NZAC" country="New Zealand" pageId="17" pageNumber="196" specimenCount="12" typeStatus="paratype">
<emphasis id="A5DA671AC6603C4FA06CF9F191A0F9F2" bold="true" box="[295,444,1608,1634]" italics="true" pageId="17" pageNumber="196">
<typeStatus id="481505AAC6603C4FA06CF9F191A0F9F2" box="[295,444,1608,1634]" pageId="17" pageNumber="196" type="paratype">PARATYPES</typeStatus>
</emphasis>
(
<specimenCount id="81A87081C6603C4FA082F9F19246F9F2" box="[457,602,1608,1634]" pageId="17" pageNumber="196" type="generic">12 specimens</specimenCount>
, in
<collectionCode id="F1BF23CDC6603C4FA336F9F192D5F9F2" box="[637,713,1608,1634]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="17" pageNumber="196" type="Other institutional type">NZAC</collectionCode>
, dissected 4 exx.)
</materialsCitation>
,
<specimenCount id="81A87081C6603C4FA2C0F9F193FAF9F2" box="[907,998,1608,1634]" pageId="17" pageNumber="196" type="female">1 female</specimenCount>
labeled same as
<typeStatus id="481505AAC6603C4FA5DFF9F194E8F9F2" box="[1172,1268,1608,1634]" pageId="17" pageNumber="196" type="holotype">holotype</typeStatus>
;
<materialsCitation id="27C6B155C6603C4FA5B6F9F193D7F916" ID-GBIF-Occurrence="4016386359" collectionCode="DSIR" collectorName="F. Alack Litter &amp; No" country="New Zealand" location="Opouri Sdle. Tennyson Inlet Rd." pageId="17" pageNumber="196" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="81A87081C6603C4FA5B6F9F19558F9F2" box="[1277,1348,1608,1634]" pageId="17" pageNumber="196" type="male">1 male</specimenCount>
labeled: \
<location id="9271EDD3C6603C4FA1EFF9D591E4F916" LSID="urn:lsid:plazi:treatment:1F070A1EC6603C4AA1DCFA049597FF69:9271EDD3C6603C4FA1EFF9D591E4F916" box="[164,504,1644,1670]" country="New Zealand" name="Opouri Sdle. Tennyson Inlet Rd." pageId="17" pageNumber="196">Opouri Sdle. Tennyson Inlet Rd.</location>
4.viii.67 \
<collectorName id="3A5BDEDEC6603C4FA32CF9D592E2F916" box="[615,766,1644,1670]" pageId="17" pageNumber="196">F.Alack Litter</collectorName>
67/225 \
<collectionCode id="F1BF23CDC6603C4FA22BF9D59380F916" box="[864,924,1644,1670]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="17" pageNumber="196">DSIR</collectionCode>
\
<collectorName id="3A5BDEDEC6603C4FA2E1F9D593D7F916" box="[938,971,1644,1670]" pageId="17" pageNumber="196">No</collectorName>
</materialsCitation>
\;
<materialsCitation id="27C6B155C6603C4FA2AFF9D591A2F93A" ID-GBIF-Occurrence="4016386338" collectorName="J. I. Townsend" country="New Zealand" location="Tennyson Inlet" pageId="17" pageNumber="196" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="81A87081C6603C4FA2AFF9D59423F916" box="[996,1087,1644,1670]" pageId="17" pageNumber="196" type="female">1 female</specimenCount>
labeled: \
<location id="9271EDD3C6603C4FA5E1F9D5955BF916" LSID="urn:lsid:plazi:treatment:1F070A1EC6603C4AA1DCFA049597FF69:9271EDD3C6603C4FA5E1F9D5955BF916" box="[1194,1351,1644,1670]" country="New Zealand" name="Tennyson Inlet" pageId="17" pageNumber="196">Tennyson Inlet</location>
22.5.64 litter
<collectorName id="3A5BDEDEC6603C4FA184F9299147F93A" box="[207,347,1680,1706]" pageId="17" pageNumber="196">J.I.Townsend</collectorName>
\
<collectingCountry id="EFB9FB98C6603C4FA03AF9299189F93A" box="[369,405,1680,1706]" name="New Zealand" pageId="17" pageNumber="196">NZ</collectingCountry>
PB
</materialsCitation>
\;
<materialsCitation id="27C6B155C6603C4FA099F9299314F95E" ID-GBIF-Occurrence="4016386328" collectingDate="1970-03-13" collectionCode="DSIR" collectorName="J. I. Townsend &amp; No" country="New Zealand" elevation="500" location="Robinson Ridge Kenepuru Sd." pageId="17" pageNumber="196" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Marlborough" typeStatus="paratype">
<specimenCount id="81A87081C6603C4FA099F9299239F93A" box="[466,549,1680,1706]" pageId="17" pageNumber="196" type="male">2 males</specimenCount>
and
<specimenCount id="81A87081C6603C4FA312F92992A3F93A" box="[601,703,1680,1706]" pageId="17" pageNumber="196" type="female">2 females</specimenCount>
labeled: \ M
<location id="9271EDD3C6603C4FA205F9299489F93A" LSID="urn:lsid:plazi:treatment:1F070A1EC6603C4AA1DCFA049597FF69:9271EDD3C6603C4FA205F9299489F93A" box="[846,1173,1680,1706]" country="New Zealand" name="Robinson Ridge Kenepuru Sd." pageId="17" pageNumber="196" stateProvince="Marlborough">Robinson Ridge Kenepuru Sd.</location>
<quantity id="505616EDC6603C4FA5D0F92994C4F93B" box="[1179,1240,1680,1707]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="196" unit="m" value="500.0">
<elevation id="1C835C3BC6603C4FA5D0F92994C4F93B" box="[1179,1240,1680,1707]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="196" unit="m" value="500.0">500m</elevation>
</quantity>
<collectingRegion id="556A75EAC6603C4FA5ABF9299570F93A" box="[1248,1388,1680,1706]" country="New Zealand" name="Marlborough" pageId="17" pageNumber="196">Marlborough</collectingRegion>
\
<date id="E3109DC8C6603C4FA4CAF92990E3F95E" pageId="17" pageNumber="196" value="1970-03-13">
<collectingDate id="F3546420C6603C4FA4CAF92990E3F95E" pageId="17" pageNumber="196" value="1970-03-13">13 Mar 1970</collectingDate>
</date>
<collectorName id="3A5BDEDEC6603C4FA04EF90D918BF95E" box="[261,407,1716,1742]" pageId="17" pageNumber="196">J.I. Townsend</collectorName>
\ litter 70/142 \
<collectionCode id="F1BF23CDC6603C4FA375F90D9266F95E" box="[574,634,1716,1742]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="17" pageNumber="196">DSIR</collectionCode>
\
<collectorName id="3A5BDEDEC6603C4FA3C3F90D92B5F95E" box="[648,681,1716,1742]" pageId="17" pageNumber="196">No</collectorName>
\
<collectingCountry id="EFB9FB98C6603C4FA3F7F90D92FCF95E" box="[700,736,1716,1742]" name="New Zealand" pageId="17" pageNumber="196">NZ</collectingCountry>
PB
</materialsCitation>
\;
<materialsCitation id="27C6B155C6603C4FA250F90D9227F962" ID-GBIF-Occurrence="4016386362" collectionCode="DSIR" collectorName="J. S. Dugdale" country="New Zealand" location="Nov." pageId="17" pageNumber="196" specimenCount="1" typeStatus="paratype">
<specimenCount id="81A87081C6603C4FA250F90D9370F95E" box="[795,876,1716,1742]" pageId="17" pageNumber="196" type="male">3 males</specimenCount>
labeled: \ Ship Cove
<specimenCount id="81A87081C6603C4FA506F90D948DF95F" box="[1101,1169,1716,1743]" pageId="17" pageNumber="196" type="generic">
<quantity id="505616EDC6603C4FA506F90D9492F95F" box="[1101,1166,1716,1743]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="17" pageNumber="196" unit="m" value="365.0">365m</quantity>
.
</specimenCount>
SD 30
<location id="9271EDD3C6603C4FA5ABF90D950FF95E" LSID="urn:lsid:plazi:treatment:1F070A1EC6603C4AA1DCFA049597FF69:9271EDD3C6603C4FA5ABF90D950FF95E" box="[1248,1299,1716,1742]" country="New Zealand" name="Nov." pageId="17" pageNumber="196">Nov.</location>
72
<collectorName id="3A5BDEDEC6603C4FA472F90D90DFF962" pageId="17" pageNumber="196">J.S. Dugdale</collectorName>
\
<collectorName id="3A5BDEDEC6603C4FA19DF961910CF962" box="[214,272,1752,1778]" pageId="17" pageNumber="196">Litter</collectorName>
72/273 \
<collectionCode id="F1BF23CDC6603C4FA039F96191B2F962" box="[370,430,1752,1778]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="17" pageNumber="196">DSIR</collectionCode>
\
<collectorName id="3A5BDEDEC6603C4FA0F7F96191C1F962" box="[444,477,1752,1778]" pageId="17" pageNumber="196">No</collectorName>
\
<collectingCountry id="EFB9FB98C6603C4FA0A4F961920FF962" box="[495,531,1752,1778]" name="New Zealand" pageId="17" pageNumber="196">NZ</collectingCountry>
PB
</materialsCitation>
\;
<materialsCitation id="27C6B155C6603C4FA305F9619581F962" ID-GBIF-Occurrence="4016386350" box="[590,1437,1752,1778]" collectionCode="DSIR" collectorName="J. McBurney" country="New Zealand" location="Shakespeare Bay" municipality="Picton" pageId="17" pageNumber="196" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="81A87081C6603C4FA305F96192B5F962" box="[590,681,1752,1778]" pageId="17" pageNumber="196" type="female">1 female</specimenCount>
labeled: \
<location id="9271EDD3C6603C4FA25FF96193D5F962" LSID="urn:lsid:plazi:treatment:1F070A1EC6603C4AA1DCFA049597FF69:9271EDD3C6603C4FA25FF96193D5F962" box="[788,969,1752,1778]" country="New Zealand" municipality="Picton" name="Shakespeare Bay" pageId="17" pageNumber="196">Shakespeare Bay</location>
,
<collectingMunicipality id="77752172C6603C4FA29FF961940BF962" box="[980,1047,1752,1778]" pageId="17" pageNumber="196">Picton</collectingMunicipality>
\ 11
<collectorName id="3A5BDEDEC6603C4FA50CF9619469F962" box="[1095,1141,1752,1778]" pageId="17" pageNumber="196">Aug</collectorName>
69
<collectorName id="3A5BDEDEC6603C4FA5D7F9619502F962" box="[1180,1310,1752,1778]" pageId="17" pageNumber="196">J.McBurney</collectorName>
\
<collectionCode id="F1BF23CDC6603C4FA47AF9619571F962" box="[1329,1389,1752,1778]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="17" pageNumber="196">DSIR</collectionCode>
\
<collectorName id="3A5BDEDEC6603C4FA430F9619581F962" box="[1403,1437,1752,1778]" pageId="17" pageNumber="196">No</collectorName>
</materialsCitation>
\;
<materialsCitation id="27C6B155C6603C4FA1E7F94594B2F886" ID-GBIF-Occurrence="4016386310" box="[172,1198,1788,1815]" collectorName="I. Mannering &amp; Leafmould" country="New Zealand" location="Mt. Robertson" pageId="17" pageNumber="196" specimenCount="1" specimenCount-female="1" stateProvince="Marlborough" typeStatus="paratype">
<specimenCount id="81A87081C6603C4FA1E7F945911BF886" box="[172,263,1788,1814]" pageId="17" pageNumber="196" type="female">1 female</specimenCount>
labeled: \
<collectingRegion id="556A75EAC6603C4FA03FF945921CF886" box="[372,512,1788,1814]" country="New Zealand" name="Marlborough" pageId="17" pageNumber="196">Marlborough</collectingRegion>
<location id="9271EDD3C6603C4FA34CF9459285F886" LSID="urn:lsid:plazi:treatment:1F070A1EC6603C4AA1DCFA049597FF69:9271EDD3C6603C4FA34CF9459285F886" box="[519,665,1788,1814]" country="New Zealand" name="Mt. Robertson" pageId="17" pageNumber="196" stateProvince="Marlborough">Mt.Robertson</location>
1,600 16-5-60
<collectorName id="3A5BDEDEC6603C4FA270F94593D9F886" box="[827,965,1788,1814]" pageId="17" pageNumber="196">I. Mannering</collectorName>
\
<collectorName id="3A5BDEDEC6603C4FA292F9459452F886" box="[985,1102,1788,1814]" pageId="17" pageNumber="196">Leafmould</collectorName>
\
<collectingCountry id="EFB9FB98C6603C4FA52AF9459499F886" box="[1121,1157,1788,1814]" name="New Zealand" pageId="17" pageNumber="196">NZ</collectingCountry>
PB
</materialsCitation>
\.
</paragraph>
</subSubSection>
<subSubSection id="DFB4E883C6603C4FA18CF89994D9F8CE" pageId="17" pageNumber="196" type="etymology">
<paragraph id="9711BB08C6603C4FA18CF89994D9F8CE" blockId="17.[151,1437,1572,2030]" pageId="17" pageNumber="196">
<emphasis id="A5DA671AC6603C4FA18CF8999152F8AA" bold="true" box="[199,334,1824,1850]" pageId="17" pageNumber="196">Etymology.</emphasis>
The specific epithet is a Latinized eponym in the genitive case, and is based on the surname of James Ian Townsend, a famous
<collectingCountry id="EFB9FB98C6603C4FA0B1F8FD928CF8CE" box="[506,656,1860,1886]" name="New Zealand" pageId="17" pageNumber="196">New Zealand</collectingCountry>
carabidologist and the collector of the
<typeStatus id="481505AAC6603C4FA50DF8FD946AF8CE" box="[1094,1142,1860,1886]" pageId="17" pageNumber="196">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="DFB4E883C6603C4FA18CF8D1958AF87E" pageId="17" pageNumber="196" type="materials_examined">
<paragraph id="9711BB08C6603C4FA18CF8D19448F812" blockId="17.[151,1437,1572,2030]" box="[199,1108,1896,1922]" pageId="17" pageNumber="196">
<materialsCitation id="27C6B155C6603C4FA18CF8D1944CF812" ID-GBIF-Occurrence="4016386376" box="[199,1104,1896,1922]" country="New Zealand" county="South Island" location="Tennyson Inlet" municipality="Sounds" pageId="17" pageNumber="196" specimenCount="1" stateProvince="Marlborough" typeStatus="holotype">
<emphasis id="A5DA671AC6603C4FA18CF8D1917AF812" bold="true" box="[199,358,1896,1922]" pageId="17" pageNumber="196">
<typeStatus id="481505AAC6603C4FA18CF8D1911EF812" box="[199,258,1896,1922]" pageId="17" pageNumber="196">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="EFB9FB98C6603C4FA027F8D19219F812" box="[364,517,1896,1922]" name="New Zealand" pageId="17" pageNumber="196">New Zealand</collectingCountry>
,
<collectingCounty id="7E70C384C6603C4FA35BF8D192BCF812" box="[528,672,1896,1922]" pageId="17" pageNumber="196">South Island</collectingCounty>
,
<collectingRegion id="556A75EAC6603C4FA3E0F8D19323F812" box="[683,831,1896,1922]" country="New Zealand" name="Marlborough" pageId="17" pageNumber="196">Marlborough</collectingRegion>
<collectingMunicipality id="77752172C6603C4FA20DF8D19387F812" box="[838,923,1896,1922]" pageId="17" pageNumber="196">Sounds</collectingMunicipality>
,
<location id="9271EDD3C6603C4FA2EDF8D1944CF812" LSID="urn:lsid:plazi:treatment:1F070A1EC6603C4AA1DCFA049597FF69:9271EDD3C6603C4FA2EDF8D1944CF812" box="[934,1104,1896,1922]" country="New Zealand" county="South Island" municipality="Sounds" name="Tennyson Inlet" pageId="17" pageNumber="196" stateProvince="Marlborough">Tennyson Inlet</location>
</materialsCitation>
.
</paragraph>
<paragraph id="9711BB08C6603C4FA18CF835958AF87E" blockId="17.[151,1437,1572,2030]" pageId="17" pageNumber="196">
<emphasis id="A5DA671AC6603C4FA18CF8359141F836" bold="true" box="[199,349,1932,1958]" pageId="17" pageNumber="196">Recognition.</emphasis>
Adults of this species (
<figureCitation id="0F95A78DC6603C4FA315F83592AEF836" box="[606,690,1932,1958]" captionStart="FIGURE 3" captionStartId="8.[151,250,957,981]" captionTargetBox="[151,1435,151,943]" captionTargetId="figure-392@8.[151,1435,151,943]" captionTargetPageId="8" captionText="FIGURE 3. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. parvulus (Mount Stokes, Marlborough Sounds, SO), B—N. tararua (Judd Ridge, Tararua Range, Wellington, NO), C—N. townsendi (Tennyson Inlet, Marlborough Sounds, SO). Scale bar 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7553495" httpUri="https://zenodo.org/record/7553495/files/figure.png" pageId="17" pageNumber="196">Fig. 3C</figureCitation>
) can be distinguished from the adults of majority species of
<taxonomicName id="50AEC08BC6603C4FA403F83490C3F85A" authorityName="Jeannel" authorityYear="1937" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="A5DA671AC6603C4FA403F83490C3F85A" italics="true" pageId="17" pageNumber="196">Nesamblyops</emphasis>
</taxonomicName>
by the distinctive shape of pronotum (
<figureCitation id="0F95A78DC6603C4FA337F80992DBF85A" box="[636,711,1968,1994]" captionStart="FIGURE 4" captionStartId="10.[151,250,1222,1246]" captionTargetBox="[151,1436,181,1197]" captionTargetId="figure-28@10.[151,1436,181,1198]" captionTargetPageId="10" captionText="FIGURE 4. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. brouni (Lewis Pass, North Canterbury, SO), B—N. carltoni (Dun Mountain, Nelson, SO), C—N. confusus (Mount Stokes, Marlborough Sounds, SO), D—N. distinctus (Fabians Valley, Marlborough, SO), E—N. lescheni (DUrville Island, Cook Strait, Marlborough Sounds, SO), F—N. oreobius (Mount Pirongia, Waikato, NO), G—N. parvulus (Mount Stokes, Marlborough Sounds, SO), H—N. tararua (Judd Ridge, Tararua Range, Wellington, NO), I—N. townsendi (Tennyson Inlet, Marlborough Sounds, SO). Scale bar 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7553497" httpUri="https://zenodo.org/record/7553497/files/figure.png" pageId="17" pageNumber="196">Fig. 4I</figureCitation>
), however are practically indistinguishable from the adults of some
<taxonomicName id="50AEC08BC6603C4FA1DCF86D9163F87E" box="[151,383,2004,2030]" pageId="17" pageNumber="196">
<emphasis id="A5DA671AC6603C4FA1DCF86D9136F87E" box="[151,298,2004,2030]" italics="true" pageId="17" pageNumber="196">Nesamblyops</emphasis>
species
</taxonomicName>
(cf. with
<figureCitation id="0F95A78DC6603C4FA0A2F86D9254F87E" box="[489,584,2004,2030]" captionStart="FIGURE 2" captionStartId="7.[151,250,1692,1716]" captionTargetBox="[151,1423,154,1661]" captionTargetId="figure-172@7.[151,1436,151,1669]" captionTargetPageId="7" captionText="FIGURE 2. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. brouni (Lewis Pass, North Canterbury, SO), B—N. carltoni (Dun Mountain, Nelson, SO), C—N. confusus (Mount Stokes, Marlborough Sounds, SO), D—N. distinctus (Fabians Valley, Marlborough, SO), E—N. lescheni (DUrville Island, Cook Strait, Marlborough Sounds, SO), F—N. oreobius (Mount Pirongia, Waikato, NO). Scale bar 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7553491" httpUri="https://zenodo.org/record/7553491/files/figure.png" pageId="17" pageNumber="196">Figs. 2D</figureCitation>
) and are distinguished from all of them by the structure of the male genitalia.
</paragraph>
</subSubSection>
<caption id="C3D1EB80C6633C4CA1DCF91B92C9F892" ID-DOI="http://doi.org/10.5281/zenodo.7553515" ID-Zenodo-Dep="7553515" httpUri="https://zenodo.org/record/7553515/files/figure.png" pageId="18" pageNumber="197" startId="18.[151,250,1698,1722]" targetBox="[195,1404,151,1693]" targetPageId="18">
<paragraph id="9711BB08C6633C4CA1DCF91B92C9F892" blockId="18.[151,1437,1698,1795]" pageId="18" pageNumber="197">
<emphasis id="A5DA671AC6633C4CA1DCF91B913DF92A" bold="true" box="[151,289,1698,1722]" pageId="18" pageNumber="197">FIGURE 10.</emphasis>
Distributional records for the
<taxonomicName id="50AEC08BC6633C4CA311F91A932DF92B" box="[602,817,1699,1723]" pageId="18" pageNumber="197">
<emphasis id="A5DA671AC6633C4CA311F91A92FDF92B" box="[602,737,1699,1723]" italics="true" pageId="18" pageNumber="197">Nesamblyops</emphasis>
species
</taxonomicName>
and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Black circles—
<taxonomicName id="50AEC08BC6633C4CA217F97E93D6F94F" authorityName="Sokolov" authorityYear="2023" box="[860,970,1735,1759]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="197" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="A5DA671AC6633C4CA217F97E93D6F94F" box="[860,970,1735,1759]" italics="true" pageId="18" pageNumber="197">N. carltoni</emphasis>
</taxonomicName>
; yellow circles—
<taxonomicName id="50AEC08BC6633C4CA534F97E94E9F94F" authorityName="Sokolov" authorityYear="2023" box="[1151,1269,1735,1759]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="197" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="A5DA671AC6633C4CA534F97E94E9F94F" box="[1151,1269,1735,1759]" italics="true" pageId="18" pageNumber="197">N. parvulus</emphasis>
</taxonomicName>
; red circles—
<taxonomicName id="50AEC08BC6633C4CA4CEF97E90F6F893" authorityName="Sokolov" authorityYear="2023" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="197" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="A5DA671AC6633C4CA4CEF97E90F6F893" italics="true" pageId="18" pageNumber="197">N. lescheni</emphasis>
</taxonomicName>
; green circles—
<taxonomicName id="50AEC08BC6633C4CA0D9F9529214F893" authorityName="Sokolov" authorityYear="2023" box="[402,520,1771,1795]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="197" phylum="Arthropoda" rank="species" species="confusus">
<emphasis id="A5DA671AC6633C4CA0D9F9529214F893" box="[402,520,1771,1795]" italics="true" pageId="18" pageNumber="197">N. confusus</emphasis>
</taxonomicName>
. Scale bar= 0.5mm.
</paragraph>
</caption>
<subSubSection id="DFB4E883C6633C4AA18CF8899597FF69" lastPageId="20" lastPageNumber="199" pageId="18" pageNumber="197" type="description">
<paragraph id="9711BB08C6633C4CA18CF889948CF8DB" blockId="18.[151,1436,1840,1975]" box="[199,1168,1840,1868]" pageId="18" pageNumber="197">
<emphasis id="A5DA671AC6633C4CA18CF8899145F8DA" bold="true" box="[199,345,1840,1866]" pageId="18" pageNumber="197">Description.</emphasis>
Large for genus (SBL range
<quantity id="505616EDC6633C4CA3D4F8889358F8DB" box="[671,836,1841,1868]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.05" metricValueMin="1.85" pageId="18" pageNumber="197" unit="mm" value="1.95" valueMax="2.05" valueMin="1.85">1.852.05 mm</quantity>
, mean 1.95±
<quantity id="505616EDC6633C4CA298F8889454F8DB" box="[979,1096,1841,1868]" metricMagnitude="-5" metricUnit="m" metricValue="7.1" pageId="18" pageNumber="197" unit="mm" value="0.071">0.071 mm</quantity>
, n=6).
</paragraph>
<paragraph id="9711BB08C6633C4CA18CF8ED9207F827" blockId="18.[151,1436,1840,1975]" pageId="18" pageNumber="197">
<emphasis id="A5DA671AC6633C4CA18CF8ED9136F8FF" bold="true" box="[199,298,1876,1903]" italics="true" pageId="18" pageNumber="197">Habitus.</emphasis>
Body form (
<figureCitation id="0F95A78DC6633C4CA0FBF8EC921FF8FF" box="[432,515,1877,1903]" captionStart="FIGURE 3" captionStartId="8.[151,250,957,981]" captionTargetBox="[151,1435,151,943]" captionTargetId="figure-392@8.[151,1435,151,943]" captionTargetPageId="8" captionText="FIGURE 3. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. parvulus (Mount Stokes, Marlborough Sounds, SO), B—N. tararua (Judd Ridge, Tararua Range, Wellington, NO), C—N. townsendi (Tennyson Inlet, Marlborough Sounds, SO). Scale bar 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7553495" httpUri="https://zenodo.org/record/7553495/files/figure.png" pageId="18" pageNumber="197">Fig.3C</figureCitation>
)markedly convex, elongate ovoid, general proportions wide (WE/SBL 0.41±0.010), head narrow relative to pronotum (WH/WPm 0.66±0.014), proportions of pronotum in comparison to elytra average for genus (WPm/WE 0.77±0.013).
</paragraph>
<caption id="C3D1EB80C6623C4DA1DCFBE59213FB2C" ID-DOI="http://doi.org/10.5281/zenodo.7553517" ID-Zenodo-Dep="7553517" httpUri="https://zenodo.org/record/7553517/files/figure.png" pageId="19" pageNumber="198" startId="19.[151,250,1116,1140]" targetBox="[151,1436,151,1105]" targetPageId="19">
<paragraph id="9711BB08C6623C4DA1DCFBE59213FB2C" blockId="19.[151,1437,1116,1212]" pageId="19" pageNumber="198">
<emphasis id="A5DA671AC6623C4DA1DCFBE59103FBE4" bold="true" box="[151,287,1116,1140]" pageId="19" pageNumber="198">FIGURE 11.</emphasis>
Distributional records for
<taxonomicName id="50AEC08BC6623C4DA364FBE59377FBE5" authority="(Broun)" authorityName="Det. I. Townsend" authorityYear="1973" baseAuthorityName="Broun" baseAuthorityYear="1893" box="[559,875,1116,1141]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="A5DA671AC6623C4DA364FBE59308FBE4" box="[559,788,1116,1140]" italics="true" pageId="19" pageNumber="198">Nesamblyops oreobius</emphasis>
(Broun)
</taxonomicName>
and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Red star—the type locality of
<taxonomicName id="50AEC08BC6623C4DA573FB39953EFB08" authority="Broun" authorityName="Broun, 1893:" authorityYear="1399" box="[1080,1314,1152,1176]" class="Insecta" family="Carabidae" genus="Tachys" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="oreobius">
<emphasis id="A5DA671AC6623C4DA573FB3994C0FB08" box="[1080,1244,1152,1176]" italics="true" pageId="19" pageNumber="198">Tachys oreobius</emphasis>
Broun
</taxonomicName>
and
<taxonomicName id="50AEC08BC6623C4DA41FFB39919AFB2C" authority="Broun. Scale" authorityName="Broun. Scale" authorityYear="1910" class="Insecta" family="Carabidae" genus="Anillus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="A5DA671AC6623C4DA41FFB3990E0FB2C" italics="true" pageId="19" pageNumber="198">Anillus monticola</emphasis>
Broun. Scale
</taxonomicName>
bar= 0.5mm.
</paragraph>
</caption>
<paragraph id="9711BB08C6623C4DA18CFAB79391FAB9" blockId="19.[151,1437,1294,2041]" box="[199,909,1294,1321]" pageId="19" pageNumber="198">
<emphasis id="A5DA671AC6623C4DA18CFAB79112FAB9" bold="true" box="[199,270,1294,1321]" italics="true" pageId="19" pageNumber="198">Color.</emphasis>
Body color brunneopiceous, appendages brunneorufuos.
</paragraph>
<paragraph id="9711BB08C6623C4DA18CFA8B950AFA29" blockId="19.[151,1437,1294,2041]" pageId="19" pageNumber="198">
<emphasis id="A5DA671AC6623C4DA18CFA8B9123FADD" bold="true" box="[199,319,1330,1357]" italics="true" pageId="19" pageNumber="198">Prothorax.</emphasis>
Pronotum (
<figureCitation id="0F95A78DC6623C4DA08BFA8A9215FADD" box="[448,521,1331,1357]" captionStart="FIGURE 4" captionStartId="10.[151,250,1222,1246]" captionTargetBox="[151,1436,181,1197]" captionTargetId="figure-28@10.[151,1436,181,1198]" captionTargetPageId="10" captionText="FIGURE 4. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. brouni (Lewis Pass, North Canterbury, SO), B—N. carltoni (Dun Mountain, Nelson, SO), C—N. confusus (Mount Stokes, Marlborough Sounds, SO), D—N. distinctus (Fabians Valley, Marlborough, SO), E—N. lescheni (DUrville Island, Cook Strait, Marlborough Sounds, SO), F—N. oreobius (Mount Pirongia, Waikato, NO), G—N. parvulus (Mount Stokes, Marlborough Sounds, SO), H—N. tararua (Judd Ridge, Tararua Range, Wellington, NO), I—N. townsendi (Tennyson Inlet, Marlborough Sounds, SO). Scale bar 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7553497" httpUri="https://zenodo.org/record/7553497/files/figure.png" pageId="19" pageNumber="198">Fig. 4I</figureCitation>
) moderately long in comparison to elytra (LP/LE 0.44±0.009) and moderately transverse (WPm/LP 1.22±0.016), with lateral margins rectilinear and slightly constricted posteriorly (WPm/WPp 1.18±0.019). Anterior angles indistinct, posterior angles obtuse (94100°), dented, and slightly protruding backward. Width between posterior angles much greater than between anterior angles (WPa/WPp 0.77±0.009). Basal margin concave.
</paragraph>
<paragraph id="9711BB08C6623C4DA18CFA7B92B4F9B5" blockId="19.[151,1437,1294,2041]" pageId="19" pageNumber="198">
<emphasis id="A5DA671AC6623C4DA18CFA7B910EFA4D" bold="true" box="[199,274,1474,1501]" italics="true" pageId="19" pageNumber="198">Elytra.</emphasis>
Ovoid, narrowly depressed along suture, comparatively long (LE/SBL 0.59±0.010) and moderately wide (WE/LE 0.69±0.008). Humeri completely rounded. Lateral margins slightly divergent at basal half, shortly subparallel at middle and evenly rounded to apex in apical half.
</paragraph>
<paragraph id="9711BB08C6623C4DA18CF997919DF8B1" blockId="19.[151,1437,1294,2041]" pageId="19" pageNumber="198">
<emphasis id="A5DA671AC6623C4DA18CF9979172F9D9" bold="true" box="[199,366,1582,1609]" italics="true" pageId="19" pageNumber="198">Male genitalia.</emphasis>
Median lobe of aedeagus (
<figureCitation id="0F95A78DC6623C4DA3DBF99692C4F9D8" box="[656,728,1582,1609]" captionStart="FIGURE 6" captionStartId="13.[151,250,993,1017]" captionTargetBox="[157,1426,159,968]" captionTargetId="figure-403@13.[151,1436,151,978]" captionTargetPageId="13" captionText="FIGURE 6. Line drawings of male genitalia of New Zealand Nesamblyops species. N. parvulus (Mount Stokes, Marlborough Sounds, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. tararua (Judd Ridge, Tararua Range, Wellington, NO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. townsendi (Tennyson Inlet, Marlborough Sounds, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. Scale bar 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.7553503" httpUri="https://zenodo.org/record/7553503/files/figure.png" pageId="19" pageNumber="198">Fig. 6I</figureCitation>
) strongly arcuate and moderately twisted. Shaft diverging basally, tapering in apical half. Apex moderately curved dorsally with narrowly rounded tip. Apical orifice long, occupies almost half of the shaft length. Ventral margin of median lobe straight. Walls of shaft with scattered poriferous canals. Dorsal copulatory sclerites in form of specific “birds head” plate, surrounded by small scaled membraneous field. Spines of internal sac absent. Left paramere (
<figureCitation id="0F95A78DC6623C4DA35EF9069276F948" box="[533,618,1726,1753]" captionStart="FIGURE 6" captionStartId="13.[151,250,993,1017]" captionTargetBox="[157,1426,159,968]" captionTargetId="figure-403@13.[151,1436,151,978]" captionTargetPageId="13" captionText="FIGURE 6. Line drawings of male genitalia of New Zealand Nesamblyops species. N. parvulus (Mount Stokes, Marlborough Sounds, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. tararua (Judd Ridge, Tararua Range, Wellington, NO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. townsendi (Tennyson Inlet, Marlborough Sounds, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. Scale bar 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.7553503" httpUri="https://zenodo.org/record/7553503/files/figure.png" pageId="19" pageNumber="198">Fig. 6G</figureCitation>
) of normal proportions, with moderately attenuate apex, bearing three long setae. Right paramere (
<figureCitation id="0F95A78DC6623C4DA0DBF95A91F9F96C" box="[400,485,1762,1789]" captionStart="FIGURE 6" captionStartId="13.[151,250,993,1017]" captionTargetBox="[157,1426,159,968]" captionTargetId="figure-403@13.[151,1436,151,978]" captionTargetPageId="13" captionText="FIGURE 6. Line drawings of male genitalia of New Zealand Nesamblyops species. N. parvulus (Mount Stokes, Marlborough Sounds, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. tararua (Judd Ridge, Tararua Range, Wellington, NO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. townsendi (Tennyson Inlet, Marlborough Sounds, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. Scale bar 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.7553503" httpUri="https://zenodo.org/record/7553503/files/figure.png" pageId="19" pageNumber="198">Fig. 6H</figureCitation>
) long and narrow, bearing three long setae, slightly shorter the length of paramere. Ring sclerite as on
<figureCitation id="0F95A78DC6623C4DA063F8BE9161F8B1" box="[296,381,1799,1825]" captionStart="FIGURE 7" captionStartId="15.[151,250,1470,1494]" captionTargetBox="[163,1424,161,1453]" captionTargetId="figure-210@15.[151,1436,151,1466]" captionTargetPageId="15" captionText="FIGURE 7. Line drawings of ring sclerite of New Zealand Nesamblyops species, male genitalia, dorsal aspect. A—N. brouni (Lewis Pass, North Canterbury, SO), B—N. carltoni (Dun Mountain, Nelson, SO), C—N. distinctus (Fabians Valley, Marlborough, SO), D—N. oreobius (Mount Pirongia, Waikato, NO), E—N. lescheni (DUrville Island, Cook Strait, Marlborough Sounds, SO), F—N. parvulus (Mount Stokes, Marlborough Sounds, SO), G—N. tararua (Judd Ridge, Tararua Range, Wellington, NO), H—N. townsendi (Tennyson Inlet, Marlborough Sounds, SO). Scale = 0.4mm." figureDoi="http://doi.org/10.5281/zenodo.7553507" httpUri="https://zenodo.org/record/7553507/files/figure.png" pageId="19" pageNumber="198">Fig. 7H</figureCitation>
.
</paragraph>
<paragraph id="9711BB08C6623C4DA18CF89392BDF8D5" blockId="19.[151,1437,1294,2041]" box="[199,673,1834,1861]" pageId="19" pageNumber="198">
<emphasis id="A5DA671AC6623C4DA18CF89391E8F8D5" bold="true" box="[199,500,1834,1861]" italics="true" pageId="19" pageNumber="198">Female internal genitalia.</emphasis>
Not examined.
</paragraph>
<paragraph id="9711BB08C6623C4DA18CF8F791F9F81D" blockId="19.[151,1437,1294,2041]" pageId="19" pageNumber="198">
<emphasis id="A5DA671AC6623C4DA18CF8F79214F8F8" bold="true" box="[199,520,1870,1896]" pageId="19" pageNumber="198">Geographical distribution.</emphasis>
This species is distributed within the
<collectingRegion id="556A75EAC6623C4DA283F8F69440F8F9" box="[968,1116,1871,1897]" country="New Zealand" name="Marlborough" pageId="19" pageNumber="198">Marlborough</collectingRegion>
Sounds area (
<bibRefCitation id="F33FC6F9C6623C4DA44FF8F690CCF81D" author="Crosby, T. K. &amp; Dugdale, J. S. &amp; Watt, J. C." pageId="19" pageNumber="198" pagination="175 - 183" refId="ref15708" refString="Crosby, T. K., Dugdale, J. S. &amp; Watt, J. C. (1998) Area codes for recording specimen localities in the New Zealand subregion. New Zealand Journal of Zoology, 25, 175 - 183. https: // doi. org / 10.1080 / 03014223.1998.9518148" type="journal article" year="1998">
Crosby
<emphasis id="A5DA671AC6623C4DA42AF8F69581F8F9" box="[1377,1437,1871,1897]" italics="true" pageId="19" pageNumber="198">et al.</emphasis>
1998
</bibRefCitation>
) (
<figureCitation id="0F95A78DC6623C4DA1A3F8CA9130F81D" box="[232,300,1907,1933]" captionStart="FIGURE 9" captionStartId="17.[151,250,1291,1315]" captionTargetBox="[151,1437,159,1280]" captionTargetId="figure-326@17.[151,1437,159,1280]" captionTargetPageId="17" captionText="FIGURE 9. Distributional records for the Nesamblyops species and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Black circles—N. tararua; yellow circles—N. townsendi; red circles—N. distinctus; green circles—N. brouni. Scale bar= 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7553513" httpUri="https://zenodo.org/record/7553513/files/figure.png" pageId="19" pageNumber="198">Fig. 9</figureCitation>
, yellow circles).
</paragraph>
<paragraph id="9711BB08C6623C4DA18CF82F91A8F845" blockId="19.[151,1437,1294,2041]" pageId="19" pageNumber="198">
<emphasis id="A5DA671AC6623C4DA18CF82F9136F820" bold="true" box="[199,298,1942,1968]" pageId="19" pageNumber="198">Habitat.</emphasis>
According to label data specimens were collected from leafmold and litter without more precise data about plant communities.
</paragraph>
<paragraph id="9711BB08C6623C4AA18CF8679597FF69" blockId="19.[151,1437,1294,2041]" lastBlockId="20.[151,1436,151,249]" lastPageId="20" lastPageNumber="199" pageId="19" pageNumber="198">
<emphasis id="A5DA671AC6623C4DA18CF867916CF868" bold="true" box="[199,368,2014,2040]" pageId="19" pageNumber="198">Relationships.</emphasis>
The structure of the male genitalia of
<taxonomicName id="50AEC08BC6623C4DA25DF86693B4F869" authorityName="Sokolov" authorityYear="2023" box="[790,936,2015,2041]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="townsendi">
<emphasis id="A5DA671AC6623C4DA25DF86693B4F869" box="[790,936,2015,2041]" italics="true" pageId="19" pageNumber="198">N. townsendi</emphasis>
</taxonomicName>
suggests relationship to other species having dorsal copulatory sclerite in a shape of “birds head,” such as
<taxonomicName id="50AEC08BC6653C4AA204FF2E93A6FF21" authorityName="Sokolov" authorityYear="2023" box="[847,954,151,177]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="199" phylum="Arthropoda" rank="species" species="brouni">
<emphasis id="A5DA671AC6653C4AA204FF2E93A6FF21" box="[847,954,151,177]" italics="true" pageId="20" pageNumber="199">N. brouni</emphasis>
</taxonomicName>
,
<taxonomicName id="50AEC08BC6653C4AA283FF2E9448FF21" authorityName="Sokolov" authorityYear="2023" box="[968,1108,151,177]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="199" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="A5DA671AC6653C4AA283FF2E9448FF21" box="[968,1108,151,177]" italics="true" pageId="20" pageNumber="199">N. distinctus</emphasis>
</taxonomicName>
, and
<taxonomicName id="50AEC08BC6653C4AA5D8FF2E9516FF20" authorityName="Sokolov" authorityYear="2023" box="[1171,1290,151,176]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="199" phylum="Arthropoda" rank="species" species="tararua">
<emphasis id="A5DA671AC6653C4AA5D8FF2E9516FF20" box="[1171,1290,151,176]" italics="true" pageId="20" pageNumber="199">N. tararua</emphasis>
</taxonomicName>
. The state of the scaled membraneous field and the presence of the area with poriferous canals suggest that
<taxonomicName id="50AEC08BC6653C4AA584FF029578FF45" authorityName="Sokolov" authorityYear="2023" box="[1231,1380,187,213]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="199" phylum="Arthropoda" rank="species" species="townsendi">
<emphasis id="A5DA671AC6653C4AA584FF029578FF45" box="[1231,1380,187,213]" italics="true" pageId="20" pageNumber="199">N. townsendi</emphasis>
</taxonomicName>
may share the common ancestry with
<taxonomicName id="50AEC08BC6653C4AA342FF669263FF68" authorityName="Sokolov" authorityYear="2023" box="[521,639,223,248]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="199" phylum="Arthropoda" rank="species" species="tararua">
<emphasis id="A5DA671AC6653C4AA342FF669263FF68" box="[521,639,223,248]" italics="true" pageId="20" pageNumber="199">N. tararua</emphasis>
</taxonomicName>
. Within-group relationships of
<taxonomicName id="50AEC08BC6653C4AA294FF66946EFF69" authorityName="Sokolov" authorityYear="2023" box="[991,1138,223,249]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="199" phylum="Arthropoda" rank="species" species="townsendi">
<emphasis id="A5DA671AC6653C4AA294FF66946EFF69" box="[991,1138,223,249]" italics="true" pageId="20" pageNumber="199">N. townsendi</emphasis>
</taxonomicName>
remains to be examined.
</paragraph>
</subSubSection>
</treatment>
</document>