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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.544.6074" ID-GBIF-Dataset="0e5e47a0-f8f3-4039-b0e1-c159fef0b9f6" ID-PMC="PMC4714384" ID-Pensoft-Pub="1313-2970-544-1" ID-PubMed="26798289" ID-ZBK="C5978BD0145B40F8ACDEB27371B7B9A4" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-544-1" ModsDocOrigin="ZooKeys 544" ModsDocTitle="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation" checkinTime="1451243640953" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2015" docId="2C761CCB2203FD18C61C47BEEE420E94" docLanguage="en" docName="ZooKeys 544: 1-407" docOrigin="ZooKeys 544" docSource="http://dx.doi.org/10.3897/zookeys.544.6074" docTitle="Mecyclothorax giffardi Liebherr" docType="treatment" docVersion="4" lastPageNumber="105" masterDocId="FF92FFC2FF9FFFC7191EFFB8FF91AF5D" masterDocTitle="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation" masterLastPageNumber="407" masterPageNumber="1" pageNumber="104" updateTime="1668162300918" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liebherr, James K.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>544</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>407</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.544.6074</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.544.6074</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-544-1</mods:identifier>
<mods:identifier type="ZBK">C5978BD0145B40F8ACDEB27371B7B9A4</mods:identifier>
<mods:identifier type="ZooBank">C5978BD0145B40F8ACDEB27371B7B9A4</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127853613" LSID="urn:lsid:plazi:treatment:2C761CCB2203FD18C61C47BEEE420E94" httpUri="http://treatment.plazi.org/id/2C761CCB2203FD18C61C47BEEE420E94" lastPageId="104" lastPageNumber="105" pageId="103" pageNumber="104">
<subSubSection pageId="103" pageNumber="104" type="multiple">
<paragraph pageId="103" pageNumber="104">Taxon classification Animalia Coleoptera Carabidae</paragraph>
</subSubSection>
<subSubSection pageId="103" pageNumber="104" type="nomenclature">
<paragraph pageId="103" pageNumber="104">
(039)
<taxonomicName authority="Liebherr" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax giffardi" order="Coleoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="giffardi">Mecyclothorax giffardi Liebherr</taxonomicName>
Figs 52D, 53G, 56
</paragraph>
</subSubSection>
<subSubSection pageId="104" pageNumber="105" type="reference_group">
<paragraph pageId="104" pageNumber="105">
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax giffardi" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="giffardi">
<pageBreakToken pageId="104" pageNumber="105" start="start">Mecyclothorax</pageBreakToken>
giffardi
</taxonomicName>
<bibRefCitation author="Liebherr, JK" journalOrPublisher="Journal of the New York Entomological Society" pageId="309" pageNumber="310" pagination="97 - 128" title="New species of Mecyclothorax (Coleoptera: Carabidae, Psydrini) from Polipoli, Maui define an area of endemism on Haleakala Volcano, Hawaii." url="10.1664/0028-7199(2005)113" volume="113" year="2005 b">Liebherr 2005b</bibRefCitation>
: 108.
</paragraph>
</subSubSection>
<subSubSection pageId="104" pageNumber="105" type="diagnosis">
<paragraph pageId="104" pageNumber="105">Diagnosis.</paragraph>
<paragraph pageId="104" pageNumber="105">
Of the beetles comprising this taxonomic group, only individuals of this species simultaneously exhibit: 1, two dorsal elytral setae on interval 3 but no additional setae on intervals 5 or 7; 2, laterally convex elytra with the greatest width behind midlength in combination with slightly narrowed humeri, MEW/HuW = 1.98; and 3, moderately smaller body size, standardized body length 4.5-4.9 mm. The dorsal body surface is reflective due to largely transverse microsculpture: 1, vertex and pronotal disc covered with a distinct transverse mesh, sculpticell breadth subequal to twice sculpticell length; and 2, elytra with a transverse mesh, sculpticell breadth 2
<normalizedToken originalValue="3×">-3x</normalizedToken>
length. Only the pronotal median base exhibits the upraised isodiametric
<normalizedToken originalValue="sculpticellsirregularly">sculpticells-irregularly</normalizedToken>
swirling based on the orientation of the cuticular
<normalizedToken originalValue="surfacecharacteristic">surface-characteristic</normalizedToken>
of other species in the group. Setal formula 2 2 2 2.
</paragraph>
</subSubSection>
<subSubSection pageId="104" pageNumber="105" type="identification">
<paragraph pageId="104" pageNumber="105">Identification</paragraph>
<paragraph pageId="104" pageNumber="105">
(n = 2). The eyes are smaller, covering only
<normalizedToken originalValue="¾">3/4</normalizedToken>
of the protruded ocular lobe, and narrowly convex, ocular ratio = 1.41-1.42. The forebody is narrower relative to the elytra than in the other species with only the 2 dorsal elytral setae; MEW/MWH = 2.14, MEW/MPW = 1.46. The discal elytral striae are minutely punctate, and the dorsal elytral setae are in broad, foveate depressions that span interval 3 plus the adjacent halves of intervals 2 and 4. The forebody is distinctly darker than the elytra, with frons and vertex plus pronotal disc rufopiceous, contrasted to the rufobrunneous elytra. The apex of the elytral sutural stria and the adjoining elytral apex are paler, rufoflavous. Like related species, the femora are flavous with a basal piceous cloud, and the tibiae are rufobrunneous with a more pronounced piceous cast apically.
</paragraph>
<paragraph pageId="104" pageNumber="105">
Male genitalia (n = 1). Aedeagal median lobe gracile, distance from parameral articulation to tip 4.1
<normalizedToken originalValue="×">x</normalizedToken>
depth at midlength (Fig. 53G); apex broadly expanded with flat apical face, the ventral tip rightly rounded; internal sac short and broad, with broad dorsal ostial microtrichial patch at base, and ventral surface broadly spiculate as a ventral patch; flagellar plate short, length 0.30
<normalizedToken originalValue="×">x</normalizedToken>
parameral articulation-tip distance.
</paragraph>
</subSubSection>
<subSubSection pageId="104" pageNumber="105" type="holotype">
<paragraph pageId="104" pageNumber="105">Holotype.</paragraph>
<paragraph pageId="104" pageNumber="105">
Male (BPBM) dissected and designated by
<bibRefCitation author="Liebherr, JK" journalOrPublisher="Journal of the New York Entomological Society" pageId="309" pageNumber="310" pagination="97 - 128" title="New species of Mecyclothorax (Coleoptera: Carabidae, Psydrini) from Polipoli, Maui define an area of endemism on Haleakala Volcano, Hawaii." url="10.1664/0028-7199(2005)113" volume="113" year="2005 b">Liebherr (2005b</bibRefCitation>
: 109). Type locality is: HI: Maui,
<normalizedToken originalValue="Haleakalā">Haleakala</normalizedToken>
, Polipoli Springs area, 5000 ft. el.
</paragraph>
</subSubSection>
<subSubSection pageId="104" pageNumber="105" type="distribution and habitat">
<paragraph pageId="104" pageNumber="105">Distribution and habitat.</paragraph>
<paragraph pageId="104" pageNumber="105">
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax giffardi" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="giffardi">Mecyclothorax giffardi</taxonomicName>
is known from only two specimens, the holotype collected by W.M. Giffard at 1525 m elevation in the Kula Forest Reserve below Polipoli Springs, and a second male specimen collected by P.D. Krushelnycky in the Kahikinui Forest Reserve on
<normalizedToken originalValue="Haleakalās">Haleakala's</normalizedToken>
south slope (Fig. 56). The Kahikinui specimen was collected in
<taxonomicName lsidName="koa" pageId="104" pageNumber="105" rank="species" species="koa">koa</taxonomicName>
-
<normalizedToken originalValue="ōhia">'ōhi'a</normalizedToken>
leaf litter.
</paragraph>
</subSubSection>
</treatment>
</document>