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<mods:title id="B8003FEAE50F5EADAD644B91DC30C50D">Clarification of the status of the types of Australian Melolonthini (Coleoptera Scarabaeidae: Melolonthinae) described before 1950</mods:title>
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<treatment id="C264702F453C35192BE79C01FB8AFF52" ID-DOI="http://doi.org/10.5281/zenodo.4340938" ID-GBIF-Taxon="170719304" ID-Zenodo-Dep="4340938" LSID="urn:lsid:plazi:treatment:C264702F453C35192BE79C01FB8AFF52" httpUri="http://treatment.plazi.org/id/C264702F453C35192BE79C01FB8AFF52" lastPageId="7" lastPageNumber="457" pageId="6" pageNumber="457">
<subSubSection id="02D792B2453C35182BE79C01FDE5FA29" box="[151,634,1384,1411]" pageId="6" pageNumber="457" type="nomenclature">
<paragraph id="4A72C139453C35182BE79C01FDE5FA29" blockId="6.[151,634,1384,1411]" box="[151,634,1384,1411]" pageId="6" pageNumber="457">
<heading id="113A7655453C35182BE79C01FDE5FA29" bold="true" box="[151,634,1384,1411]" fontSize="11" level="1" pageId="6" pageNumber="457" reason="1">
<taxonomicName id="8DCDBABA453C35182BE79C01FDE5FA29" authority="(Blackburn, 1911)" baseAuthorityName="Blackburn" baseAuthorityYear="1911" box="[151,634,1384,1411]" class="Insecta" family="Melolonthidae" genus="Antitrogus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="planiceps">
<emphasis id="78B91D2B453C35182BE79C01FDE5FA29" bold="true" box="[151,634,1384,1411]" pageId="6" pageNumber="457">
<emphasis id="78B91D2B453C35182BE79C01FE0EFA28" bold="true" box="[151,401,1384,1411]" italics="true" pageId="6" pageNumber="457">Antitrogus planiceps</emphasis>
(
<bibRefCitation id="2E5CBCC8453C35182AD19C01FDEDFA29" author="Blackburn, T." box="[417,626,1384,1410]" pageId="6" pageNumber="457" pagination="173 - 203" refId="ref24884" refString="Blackburn, T. (1911) Further notes on Australian Coleoptera, with descriptions of new genera and species. No. XLI. Transactions of the Royal Society of South Australia, 35, 173 - 203." type="journal article" year="1911">Blackburn, 1911</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02D792B2453C35182BE79CC4FE49F91C" pageId="6" pageNumber="457" type="reference_group">
<paragraph id="4A72C139453C35182BE79CC4FDDEFA6E" blockId="6.[151,630,1453,1538]" box="[151,577,1453,1477]" pageId="6" pageNumber="457">
<treatmentCitationGroup id="6ADDE617453C35182BE79CC4FDDEFA6E" box="[151,577,1453,1477]" pageId="6" pageNumber="457">
<treatmentCitation id="CB6CE728453C35182BE79CC4FDA2FA6E" author="Blackburn, T." box="[151,573,1453,1477]" page="195" pageId="6" pageNumber="457" year="1911">
<taxonomicName id="8DCDBABA453C35182BE79CC4FDA2FA6E" authority="Blackburn, 1911: 195" authorityName="Blackburn" authorityPageNumber="195" authorityYear="1911" box="[151,573,1453,1477]" class="Insecta" family="Melolonthidae" genus="Rhopaea" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="planiceps">
<emphasis id="78B91D2B453C35182BE79CC4FEC6FA6E" box="[151,345,1453,1477]" italics="true" pageId="6" pageNumber="457">Rhopaea planiceps</emphasis>
<bibRefCitation id="2E5CBCC8453C35182A2F9CC4FDA2FA6E" author="Blackburn, T." box="[351,573,1453,1477]" pageId="6" pageNumber="457" pagination="173 - 203" refId="ref24884" refString="Blackburn, T. (1911) Further notes on Australian Coleoptera, with descriptions of new genera and species. No. XLI. Transactions of the Royal Society of South Australia, 35, 173 - 203." type="journal article" year="1911">Blackburn, 1911: 195</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4A72C139453C35182BE79CA5FDBAFA4F" blockId="6.[151,630,1453,1538]" box="[151,549,1484,1508]" pageId="6" pageNumber="457">
<treatmentCitationGroup id="6ADDE617453C35182BE79CA5FDBAFA4F" box="[151,549,1484,1508]" pageId="6" pageNumber="457">
<treatmentCitation id="CB6CE728453C35182BE79CA5FDBFFA4F" author="Britton, E. B." box="[151,544,1484,1508]" page="20" pageId="6" pageNumber="457" year="1978">
<taxonomicName id="8DCDBABA453C35182BE79CA5FDBFFA4F" authority=": Britton 1978: 20" authorityName="Britton" authorityPageNumber="20" authorityYear="1978" box="[151,544,1484,1508]" class="Insecta" family="Melolonthidae" genus="Antitrogus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="planiceps">
<emphasis id="78B91D2B453C35182BE79CA5FEF6FA4F" box="[151,361,1484,1508]" italics="true" pageId="6" pageNumber="457">Antitrogus planiceps</emphasis>
:
<bibRefCitation id="2E5CBCC8453C35182A069CA5FDBFFA4F" author="Britton, E. B." box="[374,544,1484,1508]" pageId="6" pageNumber="457" pagination="1 - 150" refId="ref25258" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton 1978: 20</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4A72C139453C35182BE79C83FDE9F9A9" blockId="6.[151,630,1453,1538]" box="[151,630,1514,1538]" pageId="6" pageNumber="457">
<emphasis id="78B91D2B453C35182BE79C83FF6EF9A9" box="[151,241,1514,1538]" italics="true" pageId="6" pageNumber="457">Rhodaea</emphasis>
<taxonomicName id="8DCDBABA453C35182B889C83FE57F9A9" authority=": Lea 1912" authorityName=": Lea" authorityYear="1912" box="[248,456,1514,1538]" class="Insecta" family="Melolonthidae" genus="Antitrogus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="planiceps">
<emphasis id="78B91D2B453C35182B889C83FEC6F9A9" box="[248,345,1514,1538]" italics="true" pageId="6" pageNumber="457">planiceps</emphasis>
:
<bibRefCitation id="2E5CBCC8453C35182A169C82FE57F9A9" author="Lea, A. M." box="[358,456,1514,1538]" pageId="6" pageNumber="457" refId="ref25947" refString="Lea, A. M. (1912) The late Rev. Canon Thomas Blackburn, B. A., and his entomological work. Transactions of the Royal Society of South Australia, 36, v-xl." type="book" year="1912">Lea 1912</bibRefCitation>
</taxonomicName>
: xxxvi (in error).
</paragraph>
<paragraph id="4A72C139453C35182BE79F58FE49F91C" blockId="6.[151,1437,1584,2043]" pageId="6" pageNumber="457">
<emphasis id="78B91D2B453C35182BE79F58FC4DF9E1" bold="true" box="[151,978,1584,1611]" pageId="6" pageNumber="457">
<typeStatus id="95767F9B453C35182BE79F58FE92F9E0" box="[151,269,1585,1611]" pageId="6" pageNumber="457" type="lectotype">Lectotype</typeStatus>
(designated by
<bibRefCitation id="2E5CBCC8453C35182AB59F58FC9FF9E1" author="Houston, W. W. K. &amp; Weir, T. A." box="[453,768,1584,1611]" pageId="6" pageNumber="457" refId="ref25821" refString="Houston, W. W. K. &amp; Weir, T. A. (1992) Melolonthinae. In: Houston, W. W. K. (Ed.), Zoological Catalogue of Australia. Coleoptera: Scarabaeoidea. Vol. 9. Australian Government Publishing Service, Canberra, pp. 174 - 358." type="book" year="1992">Houston &amp; Weir 1992: 306</bibRefCitation>
) male (
<figureCitation id="D2F6DDBC453C351828259F59FC5CF9E1" box="[853,963,1584,1610]" captionStart="FIGURES 17. 13" captionStartId="7.[151,264,1584,1608]" captionTargetBox="[161,1437,731,1560]" captionTargetId="figure-176@7.[151,1437,730,1560]" captionTargetPageId="7" captionText="FIGURES 17. 13, Lectotype male of Rhopaea planiceps Blackburn, 1911 (now Antitrogus planiceps). 1, dorsal; 2, lateral; 3, labels. 47, Holotype male of Rhopaea polita Lea, 1920 (now Antitrogus politus). 4, dorsal; 5, lateral; 6, labels; 7, mesotarsi. Photographs courtesy of the South Australian Museum Board." figureDoi="http://doi.org/10.5281/zenodo.4296850" httpUri="https://zenodo.org/record/4296850/files/figure.png" pageId="6" pageNumber="457">Figs. 13</figureCitation>
):
</emphasis>
Museum Balhannah 12-85 (handwritten) | TYPE
<taxonomicName id="8DCDBABA453C35182B809F3DFDB7F9C5" authority=", Blackb." authorityName="Blackb." box="[240,552,1620,1646]" class="Insecta" family="Melolonthidae" genus="Rhopaea" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="planiceps">Rhopaea planiceps, Blackb.</taxonomicName>
(handwritten by Blackburn) | I.4526
<taxonomicName id="8DCDBABA453C351828B89F3DFB46F9C4" authority="Bl S." authorityName="Bl S." box="[968,1241,1620,1647]" class="Insecta" family="Melolonthidae" genus="Rhopaea" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="planiceps">Rhopaea planiceps Bl S.</taxonomicName>
<collectingCountry id="32DA81A9453C35182FAF9F3DFAD6F9C5" box="[1247,1353,1620,1646]" name="Australia" pageId="6" pageNumber="457">Australia</collectingCountry>
. (handwritten with S.
<collectingCountry id="32DA81A9453C35182A339F11FE35F939" box="[323,426,1656,1682]" name="Australia" pageId="6" pageNumber="457">Australia</collectingCountry>
crossed out) TYPE (handwritten in red, at right angle) | SAMA Database No. 25-034468 | my
<typeStatus id="95767F9B453C35182BB29FF5FEB6F91D" box="[194,297,1692,1718]" pageId="6" pageNumber="457" type="lectotype">lectotype</typeStatus>
label; in SAM.
</paragraph>
</subSubSection>
<subSubSection id="02D792B2453C35192BB79FA9FB8AFF52" lastPageId="7" lastPageNumber="458" pageId="6" pageNumber="457" type="description">
<paragraph id="4A72C139453C35182BB79FA9FC33F87C" blockId="6.[151,1437,1584,2043]" pageId="6" pageNumber="457">
<bibRefCitation id="2E5CBCC8453C35182BB79FA9FE10F970" author="Blackburn, T." box="[199,399,1728,1755]" pageId="6" pageNumber="457" pagination="173 - 203" refId="ref24884" refString="Blackburn, T. (1911) Further notes on Australian Coleoptera, with descriptions of new genera and species. No. XLI. Transactions of the Royal Society of South Australia, 35, 173 - 203." type="journal article" year="1911">Blackburn (1911)</bibRefCitation>
did not define the type series, seeing an unknown number of males (vide Articles 73.1.2, 72.1.1), but he said that the “Type” was in SAM and it was listed by
<bibRefCitation id="2E5CBCC8453C351828E29F8CFB92F954" author="Lea, A. M." box="[914,1037,1765,1791]" pageId="6" pageNumber="457" refId="ref25947" refString="Lea, A. M. (1912) The late Rev. Canon Thomas Blackburn, B. A., and his entomological work. Transactions of the Royal Society of South Australia, 36, v-xl." type="book" year="1912">Lea (1912)</bibRefCitation>
as such.
<bibRefCitation id="2E5CBCC8453C35182F039F8DFA8AF954" author="Britton, E. B." box="[1139,1301,1764,1791]" pageId="6" pageNumber="457" pagination="1 - 150" refId="ref25258" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton (1978)</bibRefCitation>
,
<bibRefCitation id="2E5CBCC8453C35182E519F8CFEBEF888" author="Houston, W. W. K. &amp; Weir, T. A." pageId="6" pageNumber="457" refId="ref25821" refString="Houston, W. W. K. &amp; Weir, T. A. (1992) Melolonthinae. In: Houston, W. W. K. (Ed.), Zoological Catalogue of Australia. Coleoptera: Scarabaeoidea. Vol. 9. Australian Government Publishing Service, Canberra, pp. 174 - 358." type="book" year="1992">Houston &amp; Weir (1992)</bibRefCitation>
, and
<bibRefCitation id="2E5CBCC8453C35182A2B9E61FDF2F888" author="Miller, L. J. &amp; Allsopp, P. G." box="[347,621,1800,1827]" pageId="6" pageNumber="457" pagination="377 - 409" refId="ref26403" refString="Miller, L. J. &amp; Allsopp, P. G. (2000) Identification of Australian canegrubs (Coleoptera: Scarabaeidae: Melolonthini). Invertebrate Taxonomy, 14, 377 - 409. https: // doi. org / 10.1071 / IT 98019" type="journal article" year="2000">Miller &amp; Allsopp (2000)</bibRefCitation>
all referred to a male “
<typeStatus id="95767F9B453C3518281D9E61FC4BF889" box="[877,980,1800,1826]" pageId="6" pageNumber="457" type="holotype">holotype</typeStatus>
” in SAM, but none provided any discussion as to why they considered that
<bibRefCitation id="2E5CBCC8453C351829429E45FD67F8EC" author="Blackburn, T." box="[562,760,1836,1863]" pageId="6" pageNumber="457" pagination="173 - 203" refId="ref24884" refString="Blackburn, T. (1911) Further notes on Australian Coleoptera, with descriptions of new genera and species. No. XLI. Transactions of the Royal Society of South Australia, 35, 173 - 203." type="journal article" year="1911">Blackburn (1911)</bibRefCitation>
might have had only one specimen. As
<bibRefCitation id="2E5CBCC8453C35182FB39E45FAFAF8EC" author="Britton, E. B." box="[1219,1381,1836,1863]" pageId="6" pageNumber="457" pagination="1 - 150" refId="ref25258" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton (1978)</bibRefCitation>
provided no discussion, did not label any specimen in ZMB and did not correctly state the collection data on the specimen label, I considered that his mention of a
<typeStatus id="95767F9B453C351829E89E1DFD65F825" box="[664,762,1908,1934]" pageId="6" pageNumber="457" type="holotype">holotype</typeStatus>
does not constitute a valid
<typeStatus id="95767F9B453C35182F419E1DFB07F825" box="[1073,1176,1908,1934]" pageId="6" pageNumber="457" type="lectotype">lectotype</typeStatus>
designation. However,
<bibRefCitation id="2E5CBCC8453C35182BE79EF0FE3AF818" author="Houston, W. W. K. &amp; Weir, T. A." box="[151,421,1944,1971]" pageId="6" pageNumber="457" refId="ref25821" refString="Houston, W. W. K. &amp; Weir, T. A. (1992) Melolonthinae. In: Houston, W. W. K. (Ed.), Zoological Catalogue of Australia. Coleoptera: Scarabaeoidea. Vol. 9. Australian Government Publishing Service, Canberra, pp. 174 - 358." type="book" year="1992">Houston &amp; Weir (1992)</bibRefCitation>
cited the SAM register number I.4526 that fixes a particular specimen and is a
<typeStatus id="95767F9B453C35182E469EF1FA02F819" box="[1334,1437,1944,1970]" pageId="6" pageNumber="457" type="lectotype">lectotype</typeStatus>
designation by an incorrect reference to a
<typeStatus id="95767F9B453C3518291E9ED5FD4FF87D" box="[622,720,1980,2006]" pageId="6" pageNumber="457" type="holotype">holotype</typeStatus>
(vide Article 74.5).
</paragraph>
<paragraph id="4A72C139453C35192BB79E89FB8AFF52" blockId="6.[151,1437,1584,2043]" lastBlockId="7.[151,1437,150,249]" lastPageId="7" lastPageNumber="458" pageId="6" pageNumber="457">
The type locality was given by
<bibRefCitation id="2E5CBCC8453C3518296D9E89FD7FF850" author="Blackburn, T." box="[541,736,2016,2043]" pageId="6" pageNumber="457" pagination="173 - 203" refId="ref24884" refString="Blackburn, T. (1911) Further notes on Australian Coleoptera, with descriptions of new genera and species. No. XLI. Transactions of the Royal Society of South Australia, 35, 173 - 203." type="journal article" year="1911">Blackburn (1911)</bibRefCitation>
as “
<collectingRegion id="88090FDB453C3518287C9E89FC21F851" box="[780,958,2016,2042]" country="Australia" name="South Australia" pageId="6" pageNumber="457">South Australia</collectingRegion>
” and on the label as “Balhannah” [
<geoCoordinate id="2FF9A7FE453C35182E499E88FA07F850" box="[1337,1432,2017,2043]" degrees="34.98" direction="south" orientation="latitude" pageId="6" pageNumber="457" precision="555" value="-34.98">34.98°S</geoCoordinate>
,
<geoCoordinate id="2FF9A7FE453D35192BE799FFFE99FF1A" box="[151,262,150,177]" degrees="138.82" direction="east" orientation="longitude" pageId="7" pageNumber="458" precision="555" value="138.82">138.82°E</geoCoordinate>
], about
<quantity id="8D356CDC453D35192A1299FEFE34FF1A" box="[354,427,151,177]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="458" unit="km" value="30.0">30 km</quantity>
south-east of Adelaide; it does not accord with the known distribution of the species in northeastern
<collectingRegion id="88090FDB453D35192A5899D2FE71FF7E" box="[296,494,187,213]" country="Australia" name="New South Wales" pageId="7" pageNumber="458">New South Wales</collectingRegion>
and southeastern
<collectingRegion id="88090FDB453D351929C699D2FCA5FF7E" box="[694,826,187,213]" country="Australia" name="Queensland" pageId="7" pageNumber="458">Queensland</collectingRegion>
(
<bibRefCitation id="2E5CBCC8453D3519283899D2FC45FF7E" author="Britton, E. B." box="[840,986,187,213]" pageId="7" pageNumber="458" pagination="1 - 150" refId="ref25258" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton 1978</bibRefCitation>
;
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). Perhaps, this is why South Australia is crossed out on one of the labels.
</paragraph>
</subSubSection>
</treatment>
</document>