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<treatment id="C23987DDFFFF2935FF44F9FDEBF4BC6F" ID-DOI="http://doi.org/10.5281/zenodo.11505085" ID-GBIF-Taxon="232155315" ID-Zenodo-Dep="11505085" LSID="urn:lsid:plazi:treatment:C23987DDFFFF2935FF44F9FDEBF4BC6F" httpUri="http://treatment.plazi.org/id/C23987DDFFFF2935FF44F9FDEBF4BC6F" lastPageId="9" lastPageNumber="8" pageId="7" pageNumber="6">
<subSubSection id="028A6540FFFF293BFF44F9FDECB1BB1A" box="[219,693,1625,1653]" pageId="7" pageNumber="6" type="nomenclature">
<paragraph id="4A2F36CBFFFF293BFF44F9FDECB1BB1A" blockId="7.[219,693,1625,1685]" box="[219,693,1625,1653]" pageId="7" pageNumber="6">
<heading id="116781A7FFFF293BFF44F9FDECB1BB1A" allCaps="true" box="[219,693,1625,1653]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="6" reason="2">
<taxonomicName id="8D904D48FFFF293BFF44F9FDEC75BB1A" authority="WEAVER" authorityName="Weaver, Vokes" authorityYear="1912" box="[219,625,1625,1653]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFFF293BFF44F9FDEC0FBB1A" box="[219,523,1625,1653]" italics="true" pageId="7" pageNumber="6">CRENELLA PORTERENSIS</emphasis>
WEAVER
</taxonomicName>
, 1912
</heading>
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFFF293BFEF6F9D9EF02BA17" pageId="7" pageNumber="6" type="description">
<paragraph id="4A2F36CBFFFF293BFEF6F9D9EC23BBFA" blockId="7.[219,693,1625,1685]" box="[361,551,1661,1685]" pageId="7" pageNumber="6">
<figureCitation id="D2AB2A4EFFFF293BFEF6F9D9EFF9BBFB" box="[361,509,1661,1685]" captionStart="Figure 3" captionStartId="8.[112,187,1396,1420]" captionTargetBox="[115,1515,197,1370]" captionTargetId="figure-220@8.[115,1515,195,1374]" captionTargetPageId="8" captionText="Figure 3. Crenella porterensis Weaver (1912) (AC, E) and C. washingtonensis Weaver (1916) (D, F). A. Articulated, left valve, hypotype UCMP 32410, Lincoln Creek Formation, height 18.2 mm. B. articulated, hypotype USNM 561803, Keasey Formation, height 13.5 mm. C. Exterior mold in matrix, hypotype UCMP 110725, Keasey Formation, height 10.8 mm. D. Right valve, hypotype UCMP 32250, Blakeley Formation, height 12.5 mm. E. Partial shell exterior in matrix, hypotype UCMP 110726, Keasey Formation, height 10.3 mm. F. Partial shell exterior in matrix, hypotype 110727, Keasey Formation, height 5.4 mm." figureDoi="http://doi.org/10.5281/zenodo.10913483" httpUri="https://zenodo.org/record/10913483/files/figure.png" pageId="7" pageNumber="6">FIGS. 3AC, E</figureCitation>
;
<figureCitation id="D2AB2A4EFFFF293BFD98F9D9EC23BBFA" box="[519,551,1661,1685]" captionStart="Figure 4" captionStartId="9.[112,187,1863,1887]" captionTargetBox="[115,1515,1136,1835]" captionTargetId="figure-396@9.[115,1515,1136,1835]" captionTargetPageId="9" captionText="Figure 4. A. Crenella porterensis detail of coarser ribs and rib increase by splitting (yellow arrow) and by intercalation (black arrows) (enlarged from Fig. 3A). B. Crenella washingtonensis detail of finer ribs and rib increase by intercalation (black arrows) (comparable enlargement from Fig. 3D)." figureDoi="http://doi.org/10.5281/zenodo.10913485" httpUri="https://zenodo.org/record/10913485/files/figure.png" pageId="7" pageNumber="6">4A</figureCitation>
</paragraph>
<paragraph id="4A2F36CBFFFF293BFFEFF914EED5BB87" blockId="7.[112,799,1711,1956]" pageId="7" pageNumber="6">
<taxonomicName id="8D904D48FFFF293BFFEFF914EC23BBA4" authority="Weaver (1912)" authorityName="Weaver, Vokes" authorityYear="1912" box="[112,551,1711,1740]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFFF293BFFEFF914EF60BBA4" box="[112,356,1711,1740]" italics="true" pageId="7" pageNumber="6">Crenella porterensis</emphasis>
Weaver (1912)
</taxonomicName>
. p. 15; pl. 14, fig. 115.
</paragraph>
<paragraph id="4A2F36CBFFFF293BFFEFF95CEF23BA5F" blockId="7.[112,799,1711,1956]" pageId="7" pageNumber="6">
<taxonomicName id="8D904D48FFFF293BFFEFF95CEC8ABA7C" authority="Weaver, Weaver (1916)" authorityName="Weaver, Weaver" authorityYear="1916" box="[112,654,1783,1812]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFFF293BFFEFF95CEF60BA7C" box="[112,356,1783,1812]" italics="true" pageId="7" pageNumber="6">Crenella porterensis</emphasis>
Weaver, Weaver (1916)
</taxonomicName>
. pp. 3637; figs. 41, 42.
</paragraph>
<paragraph id="4A2F36CBFFFF293BFFEFF8E4EF02BA17" blockId="7.[112,799,1711,1956]" pageId="7" pageNumber="6">
<taxonomicName id="8D904D48FFFF293BFFEFF8E4EC8ABA34" authority="Weaver, Weaver (1943)" authorityName="Weaver, Weaver" authorityYear="1943" box="[112,654,1855,1884]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFFF293BFFEFF8E4EF60BA34" box="[112,356,1855,1884]" italics="true" pageId="7" pageNumber="6">Crenella porterensis</emphasis>
Weaver, Weaver (1943)
</taxonomicName>
. p. 115; figs. 3, 4.
</paragraph>
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<subSubSection id="028A6540FFFF293BFFEFF82CEB72BC4D" pageId="7" pageNumber="6" type="reference_group">
<paragraph id="4A2F36CBFFFF293BFFEFF82CEB2CBDB2" blockId="7.[112,799,1711,1956]" box="[112,1320,194,1956]" lastBlockId="7.[832,1514,194,781]" pageId="7" pageNumber="6">
<taxonomicName id="8D904D48FFFF293BFFEFF82CEC93BACC" authority="Weaver, Durham (1944)" authorityName="Weaver, Durham" authorityYear="1944" box="[112,663,1927,1956]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFFF293BFFEFF82CEF60BACC" box="[112,356,1927,1956]" italics="true" pageId="7" pageNumber="6">Crenella porterensis</emphasis>
Weaver,
<bibRefCitation id="2E014B3AFFFF293BFE4DF82CEC93BACC" author="Durham, J. W." box="[466,663,1928,1955]" pageId="7" pageNumber="6" pagination="101 - 212" refId="ref29671" refString="Durham, J. W. 1944. Megafaunal zones of the Oligocene of northwestern Washington. University of California Publications, Department of Geological Sciences, Bulletin 27 (5): 101 - 212. pls. 13 - 18, figs. 1 - 7." type="journal article" year="1944">Durham (1944)</bibRefCitation>
</taxonomicName>
. p. 140. (unfigured hypotype
<collectionCode id="2C81AE0EFFFF293BFBECFF66EABBBDB3" box="[1139,1215,194,220]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="7" pageNumber="6" type="University or college">UCMP</collectionCode>
35163).
</paragraph>
<paragraph id="4A2F36CBFFFF293BFCDFFF4FEB72BC4D" blockId="7.[832,1514,194,781]" pageId="7" pageNumber="6">
<taxonomicName id="8D904D48FFFF293BFCDFFF4FEAEFBC6B" authority="Weaver, Vokes" authorityName="Weaver, Vokes" authorityYear="1912" box="[832,1259,233,263]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFFF293BFCDFFF4FEA30BC6A" box="[832,1076,233,263]" italics="true" pageId="7" pageNumber="6">Crenella porterensis</emphasis>
Weaver, Vokes
</taxonomicName>
in Warren, Norbisrath and Grivetti (1945).(checklist)
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFFF293BFCDFFE94EA26BC08" pageId="7" pageNumber="6" type="description">
<paragraph id="4A2F36CBFFFF293BFCDFFE94EA26BC08" blockId="7.[832,1514,194,781]" pageId="7" pageNumber="6">
<taxonomicName id="8D904D48FFFF293BFCDFFE94EDC7BC08" authority="Weaver, R. C. Moore and Vokes (1953)" authorityName="Weaver, R. C. Moore and Vokes" authorityYear="1953" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFFF293BFCDFFE94EA30BC25" box="[832,1076,302,332]" italics="true" pageId="7" pageNumber="6">Crenella porterensis</emphasis>
Weaver, R.C. Moore and Vokes (1953)
</taxonomicName>
. p. 119.
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFFF293BFCDFFED7EA0ABF55" pageId="7" pageNumber="6" type="reference_group">
<paragraph id="4A2F36CBFFFF293BFCDFFED7EDB9BCC5" blockId="7.[832,1514,194,781]" pageId="7" pageNumber="6">
<taxonomicName id="8D904D48FFFF293BFCDFFED7EB56BCE2" authority="Weaver, Kanno (1971)" authorityName="Weaver, Kanno" authorityYear="1971" box="[832,1362,369,399]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFFF293BFCDFFED7EA30BCE2" box="[832,1076,369,399]" italics="true" pageId="7" pageNumber="6">Crenella porterensis</emphasis>
Weaver,
<bibRefCitation id="2E014B3AFFFF293BFB3DFED7EB56BCE2" author="Kanno, S." box="[1186,1362,370,397]" pageId="7" pageNumber="6" pagination="1 - 154" refId="ref31982" refString="Kanno, S. 1971. Tertiary molluscan fauna from the Yakataga district and adjacent areas of southern Alaska. Palaeontological Society of Japan Special Papers 16: 1 - 154, 11 pls." type="book chapter" year="1971">Kanno (1971)</bibRefCitation>
</taxonomicName>
. p. 47. pl. 1, fig. 27.
</paragraph>
<paragraph id="4A2F36CBFFFF293BFCDFFE1FEA79BC9D" blockId="7.[832,1514,194,781]" pageId="7" pageNumber="6">
<taxonomicName id="8D904D48FFFF293BFCDFFE1FEB91BCBA" authority="Weaver, Armentrout (1973)" authorityName="Weaver, Armentrout" authorityYear="1973" box="[832,1429,441,471]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFFF293BFCDFFE1FEA30BCBA" box="[832,1076,441,471]" italics="true" pageId="7" pageNumber="6">Crenella porterensis</emphasis>
Weaver,
<bibRefCitation id="2E014B3AFFFF293BFB3DFE1EEB91BCBA" author="Armentrout, J. M." box="[1186,1429,442,469]" pageId="7" pageNumber="6" refId="ref27107" refString="Armentrout, J. M. 1973. Molluscan biostratigraphy and paleontology of the Lincoln Creek Formation (late Eocene - Oligocene) Southwestern Washington. Ph. D. diss. University of Washington, Seattle, WA. 479 pp." type="book" year="1973">Armentrout (1973)</bibRefCitation>
</taxonomicName>
. pp. 387389; pl. 8, fig. 23.
</paragraph>
<paragraph id="4A2F36CBFFFF293BFCDFFDA7EA0ABF55" blockId="7.[832,1514,194,781]" pageId="7" pageNumber="6">
<taxonomicName id="8D904D48FFFF293BFCDFFDA7EB7ABF72" authority="Weaver, E. J. Moore (1976)" authorityName="Weaver, E. J. Moore" authorityYear="1976" box="[832,1406,513,543]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFFF293BFCDFFDA7EA30BF72" box="[832,1076,513,543]" italics="true" pageId="7" pageNumber="6">Crenella porterensis</emphasis>
Weaver, E.J. Moore (1976)
</taxonomicName>
. p. 45; pl. 12, fig. 12.
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFFF293BFCDFFDEFEA26BFED" pageId="7" pageNumber="6" type="description">
<paragraph id="4A2F36CBFFFF293BFCDFFDEFEA26BFED" blockId="7.[832,1514,194,781]" pageId="7" pageNumber="6">
<taxonomicName id="8D904D48FFFF293BFCDFFDEFEAB0BF0B" authority="(Weaver)" baseAuthorityName="Weaver" box="[832,1204,585,615]" class="Bivalvia" family="Mytilidae" genus="Solamen" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFFF293BFCDFFDEFEA3CBF0A" box="[832,1080,585,615]" italics="true" pageId="7" pageNumber="6">Solamen porterensis</emphasis>
(Weaver)
</taxonomicName>
,
<bibRefCitation id="2E014B3AFFFF293BFB20FDEEEB9DBF0A" author="Addicott, W. O." box="[1215,1433,586,613]" pageId="7" pageNumber="6" pagination="1 - 44" refId="ref26918" refString="Addicott, W. O. 1976 b. Molluscan paleontology of the lower Miocene Clallum Formation Northwestern Washington. U. S. Geological Survey Professional Paper 976: 1 - 44, 9 pls., 10 figs., 7 tables." type="book chapter" year="1976">Addicott (1976b)</bibRefCitation>
. p. 29; pl. 4, figs. 3,11.
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFFF293BFCDFFD29EAA9BEF3" pageId="7" pageNumber="6" type="reference_group">
<paragraph id="4A2F36CBFFFF293BFCDFFD29EAE0BFAB" blockId="7.[832,1514,194,781]" pageId="7" pageNumber="6">
<taxonomicName id="8D904D48FFFF293BFCDFFD29EDC7BFAA" authority="Weaver, Allison and Marincovich (1981)" authorityName="Weaver, Allison and Marincovich" authorityYear="1981" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFFF293BFCDFFD29EA30BFC7" box="[832,1076,652,681]" italics="true" pageId="7" pageNumber="6">Crenella porterensis</emphasis>
Weaver,
<bibRefCitation id="2E014B3AFFFF293BFB3DFD29EDC7BFAA" author="Allison, R. C. &amp; L. Marincovich, Jr." pageId="7" pageNumber="6" pagination="1 - 11" refId="ref26963" refString="Allison, R. C., and L. Marincovich, Jr. 1981. A late Oligoene or earliest Miocene molluscan fauna from Sitkinak Island, Alaska. United States Geological Survey Professionnal Paper 1233: 1 - 11, 3 pls., 3 figs." type="book chapter" year="1981">Allison and Marincovich (1981)</bibRefCitation>
</taxonomicName>
. pp. 4, 8; pl. 1, figs. 5, 6.
</paragraph>
<paragraph id="4A2F36CBFFFF293BFCDFFD71EA2ABE63" blockId="7.[832,1514,194,781]" pageId="7" pageNumber="6">
?
<taxonomicName id="8D904D48FFFF293BFCD3FD71EDB5BF9E" ID-CoL="7NTRR" authorityName="Hickman" authorityYear="2023" box="[844,945,725,753]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="genus">
<emphasis id="78E4EAD9FFFF293BFCD3FD71EDB5BF9E" box="[844,945,725,753]" italics="true" pageId="7" pageNumber="6">Crenella</emphasis>
</taxonomicName>
cf.
<taxonomicName id="8D904D48FFFF293BFC45FD71EBCDBF9F" authority="Weaver, E. J. Moore (1963)" authorityName="Weaver, E. J. Moore" authorityYear="1963" box="[986,1481,724,753]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFFF293BFC45FD71EA7BBF9F" box="[986,1151,724,753]" italics="true" pageId="7" pageNumber="6">C. porterensis</emphasis>
Weaver, E.J. Moore (1963)
</taxonomicName>
. p. 63; pl. 25, fig. 6.
</paragraph>
<paragraph id="4A2F36CBFFFF293BFCDFFCB9EB41BE58" blockId="7.[832,1481,797,925]" box="[832,1349,797,825]" pageId="7" pageNumber="6">
?
<taxonomicName id="8D904D48FFFF293BFCD3FCB9EDDEBE58" box="[844,986,797,825]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="78E4EAD9FFFF293BFCD3FCB9EDB5BE56" box="[844,945,797,825]" italics="true" pageId="7" pageNumber="6">Crenella</emphasis>
sp.
</taxonomicName>
?
<bibRefCitation id="2E014B3AFFFF293BFC72FCB9EAB8BE57" author="Hickman, C. J. S." box="[1005,1212,797,824]" pageId="7" pageNumber="6" refId="ref30984" refString="Hickman, C. J. S. 1969. Oligocene marine molluscan fauna of the Eugene Formation in Oregon. University of Oregon Museum of Natural History Bulletin 16. 112 pp., 14 pls." type="book" year="1969">Hickman (1969)</bibRefCitation>
. pp. 33-34.
</paragraph>
<paragraph id="4A2F36CBFFFF293BFCDFFCECEAA9BEF3" blockId="7.[832,1481,797,925]" pageId="7" pageNumber="6">
?
<taxonomicName id="8D904D48FFFF293BFCD3FCECEAFABE0D" authority="Addicott (1976 b)" authorityName="Addicott" authorityYear="1976" box="[844,1278,840,868]" class="Bivalvia" family="Mytilidae" genus="Solamen" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="snavelyi">
<emphasis id="78E4EAD9FFFF293BFCD3FCECEA1ABE0B" box="[844,1054,840,868]" italics="true" pageId="7" pageNumber="6">Solamen snavelyi</emphasis>
<bibRefCitation id="2E014B3AFFFF293BFBBBFCECEAFABE0D" author="Addicott, W. O." box="[1060,1278,840,866]" pageId="7" pageNumber="6" pagination="1 - 44" refId="ref26918" refString="Addicott, W. O. 1976 b. Molluscan paleontology of the lower Miocene Clallum Formation Northwestern Washington. U. S. Geological Survey Professional Paper 976: 1 - 44, 9 pls., 10 figs., 7 tables." type="book chapter" year="1976">Addicott (1976b)</bibRefCitation>
</taxonomicName>
. p. 29; pl. 4, figs. 6,9. not
<taxonomicName id="8D904D48FFFF293BFC51FCC1EB21BEEF" authority="WEAVER" authorityName="Weaver, Vokes" authorityYear="1912" box="[974,1317,868,897]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFFF293BFC51FCC1EAC6BEEF" box="[974,1218,868,897]" italics="true" pageId="7" pageNumber="6">Crenella porterensis</emphasis>
Weaver
</taxonomicName>
, Tegland (1933). p. 112; pl. 6, fig. 2.
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFFF2935FCFFFC14EC04BCBC" lastPageId="9" lastPageNumber="8" pageId="7" pageNumber="6" type="discussion">
<paragraph id="4A2F36CBFFFF293BFCFFFC14EA8FB8B5" blockId="7.[832,1521,943,1955]" pageId="7" pageNumber="6">
<emphasis id="78E4EAD9FFFF293BFCFFFC14EDF5BEA5" bold="true" box="[864,1009,944,970]" pageId="7" pageNumber="6">Discussion</emphasis>
<taxonomicName id="8D904D48FFFF293BFB91FC15EB06BEA4" authorityName="Weaver, E. J. Moore" authorityYear="1963" box="[1038,1282,943,973]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFFF293BFB91FC15EB06BEA4" box="[1038,1282,943,973]" italics="true" pageId="7" pageNumber="6">Crenella porterensis</emphasis>
</taxonomicName>
is distinguished by its thin and strongly inflated shell, ovate-elongate shape, prominent prosogyrous umbones, external elongate posterior ligament groove, and sculpture of numerous (&gt;100) very fine radial ribs with distinct narrow interspaces and weaker commarginal ribs. Ribs terminate at the ventral margin to produce fine interlocking crenulations. Increase in the number of radial ribs is primarily by intercalation (
<figureCitation id="D2AB2A4EFFFF293BFB89FB6EEA76B98B" box="[1046,1138,1226,1252]" captionStart="Figure 4" captionStartId="9.[112,187,1863,1887]" captionTargetBox="[115,1515,1136,1835]" captionTargetId="figure-396@9.[115,1515,1136,1835]" captionTargetPageId="9" captionText="Figure 4. A. Crenella porterensis detail of coarser ribs and rib increase by splitting (yellow arrow) and by intercalation (black arrows) (enlarged from Fig. 3A). B. Crenella washingtonensis detail of finer ribs and rib increase by intercalation (black arrows) (comparable enlargement from Fig. 3D)." figureDoi="http://doi.org/10.5281/zenodo.10913485" httpUri="https://zenodo.org/record/10913485/files/figure.png" pageId="7" pageNumber="6">Fig. 4A</figureCitation>
, black arrows) of new ribs, although bifurcation also occurs (
<figureCitation id="D2AB2A4EFFFF293BFB4FFB49EB28B868" box="[1232,1324,1261,1287]" captionStart="Figure 4" captionStartId="9.[112,187,1863,1887]" captionTargetBox="[115,1515,1136,1835]" captionTargetId="figure-396@9.[115,1515,1136,1835]" captionTargetPageId="9" captionText="Figure 4. A. Crenella porterensis detail of coarser ribs and rib increase by splitting (yellow arrow) and by intercalation (black arrows) (enlarged from Fig. 3A). B. Crenella washingtonensis detail of finer ribs and rib increase by intercalation (black arrows) (comparable enlargement from Fig. 3D)." figureDoi="http://doi.org/10.5281/zenodo.10913485" httpUri="https://zenodo.org/record/10913485/files/figure.png" pageId="7" pageNumber="6">Fig. 4A</figureCitation>
, yellow arrow). Both modes of rib numerical increase may occur on a single specimen. Rib increase is most prominent late in growth on the medial portion of the shell. Shell growth is strongly allometric with an initially ovate shape increasing to elongate and clearly visible at growth checks on a single adult shell (
<figureCitation id="D2AB2A4EFFFF293BFBBEFA64EA7AB8B5" box="[1057,1150,1472,1498]" captionStart="Figure 2" captionStartId="6.[111,186,1868,1892]" captionTargetBox="[121,1508,1369,1846]" captionTargetId="figure-518@6.[121,1508,1369,1846]" captionTargetPageId="6" captionText="Figure 2. Alternative reconstructions of the life habit of Crenella decussata. A. Completely encased in crypt of cemented sedimentary grains, living below sediment-water interface in North Atlantic. B, C. Byssus attached to shell fragments, living at sedimentwater interface in North Pacific." figureDoi="http://doi.org/10.5281/zenodo.10913481" httpUri="https://zenodo.org/record/10913481/files/figure.png" pageId="7" pageNumber="6">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="4A2F36CBFFFF293BFCFFFA47EB72BACC" blockId="7.[832,1521,943,1955]" pageId="7" pageNumber="6">
The finely decussate shell surface layer is translucent, and the thin underlying (middle) nacreous layer is visible as a pattern that is enhanced as light passes through fine interruptions of crossing radial and concentric elements of surface sculpture. A third non-nacreous layer is present interior to the middle nacreous layer in some of the Keasey shells. This is consistent with observations of complex crossed lamellar aragonite and other aragonitic microstructures in the interior layer of crenellids (
<bibRefCitation id="2E014B3AFFFF293BFA3DF959ED85BA55" author="Carter, J. G." pageId="7" pageNumber="6" refId="ref27944" refString="Carter, J. G. 1990. Evolutionary significance of shell microstructure in the Palaeotaxodota, Pteriomorpha and Isofilibranchia (Bival- via: Mollusca). Pp. 336 - 296 in Carter J. G., ed., Skeletal Biomineralization: Patterns, Processes and Evolutionary Trends Vol. I. Van Nostrand Reinhold, New York. 832 pp." type="book" year="1990">Carter 1990</bibRefCitation>
). The interior of well-preserved dry shells of living
<taxonomicName id="8D904D48FFFF293BFCDFF8E7EA1BBA32" baseAuthorityName="Motagu" baseAuthorityYear="1808" box="[832,1055,1857,1887]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="decussata">
<emphasis id="78E4EAD9FFFF293BFCDFF8E7EA1BBA32" box="[832,1055,1857,1887]" italics="true" pageId="7" pageNumber="6">Crenella decussata</emphasis>
</taxonomicName>
(Hickman, personal observation) suggests a patchy, discontinuous distribution of mineralogy and microstructure that includes some nacre.
</paragraph>
<caption id="1EEF6643FFF02934FFEFFAD0EA7EB899" ID-DOI="http://doi.org/10.5281/zenodo.10913483" ID-Zenodo-Dep="10913483" httpUri="https://zenodo.org/record/10913483/files/figure.png" pageId="8" pageNumber="7" startId="8.[112,187,1396,1420]" targetBox="[115,1515,197,1370]" targetPageId="8" targetType="figure">
<paragraph id="4A2F36CBFFF02934FFEFFAD0EA7EB899" blockId="8.[112,1507,1396,1527]" pageId="8" pageNumber="7">
<emphasis id="78E4EAD9FFF02934FFEFFAD0EED2B8E3" bold="true" box="[112,214,1396,1420]" pageId="8" pageNumber="7">Figure 3.</emphasis>
<taxonomicName id="8D904D48FFF02934FF44FAD1EC52B8E3" authority="Weaver (1912)" authorityName="Weaver, Vokes" authorityYear="1912" box="[219,598,1396,1421]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFF02934FF44FAD1EFA9B8E2" box="[219,429,1397,1421]" italics="true" pageId="8" pageNumber="7">Crenella porterensis</emphasis>
Weaver (1912)
</taxonomicName>
(
<emphasis id="78E4EAD9FFF02934FDFAFAD0ECAEB8E3" bold="true" box="[613,682,1396,1420]" pageId="8" pageNumber="7">AC, E</emphasis>
) and
<taxonomicName id="8D904D48FFF02934FD78FAD1EA52B8E3" authority="Weaver (1916)" authorityName="Weaver" authorityYear="1916" box="[743,1110,1396,1421]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="washingtonensis">
<emphasis id="78E4EAD9FFF02934FD78FAD1EDABB8E2" box="[743,943,1397,1421]" italics="true" pageId="8" pageNumber="7">C. washingtonensis</emphasis>
Weaver (1916)
</taxonomicName>
(
<emphasis id="78E4EAD9FFF02934FBFAFAD0EA94B8E3" bold="true" box="[1125,1168,1396,1420]" pageId="8" pageNumber="7">D, F</emphasis>
).
<emphasis id="78E4EAD9FFF02934FB3BFAD0EAB1B8E3" bold="true" box="[1188,1205,1396,1420]" pageId="8" pageNumber="7">A</emphasis>
. Articulated, left valve, hypotype UCMP 32410, Lincoln Creek Formation, height 18.2 mm.
<emphasis id="78E4EAD9FFF02934FCBCFA2BED37B8C8" bold="true" box="[803,819,1423,1447]" pageId="8" pageNumber="7">B</emphasis>
. articulated, hypotype USNM 561803, Keasey Formation, height 13.5 mm.
<emphasis id="78E4EAD9FFF02934FF45FA0DEEEDB8AE" bold="true" box="[218,233,1449,1473]" pageId="8" pageNumber="7">C</emphasis>
. Exterior mold in matrix, hypotype UCMP 110725, Keasey Formation, height 10.8 mm.
<emphasis id="78E4EAD9FFF02934FB11FA0EEAA4B8AD" bold="true" box="[1166,1184,1450,1474]" pageId="8" pageNumber="7">D</emphasis>
. Right valve, hypotype UCMP 32250, Blakeley Formation, height 12.5 mm.
<emphasis id="78E4EAD9FFF02934FDCDFA60EC65B8B3" bold="true" box="[594,609,1476,1500]" pageId="8" pageNumber="7">E</emphasis>
. Partial shell exterior in matrix, hypotype UCMP 110726, Keasey Formation, height 10.3 mm.
<emphasis id="78E4EAD9FFF02934FF45FA7BEEECB898" bold="true" box="[218,232,1503,1527]" pageId="8" pageNumber="7">F</emphasis>
. Partial shell exterior in matrix, hypotype 110727, Keasey Formation, height 5.4 mm.
</paragraph>
</caption>
<paragraph id="4A2F36CBFFF02934FF0FF987EBC8BBC8" blockId="8.[112,800,1571,1949]" lastBlockId="8.[833,1522,1571,1949]" pageId="8" pageNumber="7">
Although none of the specimens examined are sufficiently well preserved for excavation of the hinge, offset of the valves below the beaks reveals part of a thickened denticulate plate (
<figureCitation id="D2AB2A4EFFF02934FECBF928EFB6BBC8" box="[340,434,1676,1703]" captionStart="Figure 3" captionStartId="8.[112,187,1396,1420]" captionTargetBox="[115,1515,197,1370]" captionTargetId="figure-220@8.[115,1515,195,1374]" captionTargetPageId="8" captionText="Figure 3. Crenella porterensis Weaver (1912) (AC, E) and C. washingtonensis Weaver (1916) (D, F). A. Articulated, left valve, hypotype UCMP 32410, Lincoln Creek Formation, height 18.2 mm. B. articulated, hypotype USNM 561803, Keasey Formation, height 13.5 mm. C. Exterior mold in matrix, hypotype UCMP 110725, Keasey Formation, height 10.8 mm. D. Right valve, hypotype UCMP 32250, Blakeley Formation, height 12.5 mm. E. Partial shell exterior in matrix, hypotype UCMP 110726, Keasey Formation, height 10.3 mm. F. Partial shell exterior in matrix, hypotype 110727, Keasey Formation, height 5.4 mm." figureDoi="http://doi.org/10.5281/zenodo.10913483" httpUri="https://zenodo.org/record/10913483/files/figure.png" pageId="8" pageNumber="7">Fig. 3A</figureCitation>
). A denticulate hinge plate is characteristic of
<typeStatus id="952B8869FFF02934FED4F914EF87BBA5" box="[331,387,1712,1738]" pageId="8" pageNumber="7">type</typeStatus>
<taxonomicName id="8D904D48FFF02934FE11F914EFF1BBA3" authorityName="Hickman" authorityYear="2023" box="[398,501,1712,1740]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="8" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="78E4EAD9FFF02934FE11F914EFF1BBA3" box="[398,501,1712,1740]" italics="true" pageId="8" pageNumber="7">Crenella</emphasis>
</taxonomicName>
, distinguishing it from edentulous living species assigned to the genus
<taxonomicName id="8D904D48FFF02934FD2DF977ED1EBB80" authorityName="Iredale" authorityYear="1924" box="[690,794,1747,1775]" class="Bivalvia" family="Mytilidae" genus="Solamen" kingdom="Animalia" order="Mytilida" pageId="8" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="78E4EAD9FFF02934FD2DF977ED1EBB80" box="[690,794,1747,1775]" italics="true" pageId="8" pageNumber="7">Solamen</emphasis>
</taxonomicName>
. This is probably a long-ranging species complex that includes the Miocene crenellid described by
<bibRefCitation id="2E014B3AFFF02934FD2BF8BDEED3BA38" author="Addicott, W. O." pageId="8" pageNumber="7" pagination="1 - 44" refId="ref26918" refString="Addicott, W. O. 1976 b. Molluscan paleontology of the lower Miocene Clallum Formation Northwestern Washington. U. S. Geological Survey Professional Paper 976: 1 - 44, 9 pls., 10 figs., 7 tables." type="book chapter" year="1976">Addicott (1976b)</bibRefCitation>
as
<taxonomicName id="8D904D48FFF02934FF64F899EFCCBA36" authorityName="Addicott" authorityYear="1976" box="[251,456,1853,1881]" class="Bivalvia" family="Mytilidae" genus="Solamen" kingdom="Animalia" order="Mytilida" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="snavelyi">
<emphasis id="78E4EAD9FFF02934FF64F899EFCCBA36" box="[251,456,1853,1881]" italics="true" pageId="8" pageNumber="7">Solamen snavelyi</emphasis>
</taxonomicName>
.
<bibRefCitation id="2E014B3AFFF02934FE4CF899ECC6BA38" author="Armentrout, J. M." box="[467,706,1853,1879]" pageId="8" pageNumber="7" refId="ref27107" refString="Armentrout, J. M. 1973. Molluscan biostratigraphy and paleontology of the Lincoln Creek Formation (late Eocene - Oligocene) Southwestern Washington. Ph. D. diss. University of Washington, Seattle, WA. 479 pp." type="book" year="1973">Armentrout (1973)</bibRefCitation>
reported it from six molluscan zones extending from the late Eocene into the early Miocene. The Miocene specimen from the Astoria Formation, illustrated by E.J. Moore (1963, pl. 25, fig. 6) has a denticulate hinge, and the mode of duplication of radial sculpture is consistent with the two modes documented here in
<taxonomicName id="8D904D48FFF02934FB4DF929EBC2BBC8" authorityName="Weaver, Vokes" authorityYear="1912" box="[1234,1478,1675,1705]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFF02934FB4DF929EBC2BBC8" box="[1234,1478,1675,1705]" italics="true" pageId="8" pageNumber="7">Crenella porterensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="4A2F36CBFFF02935FCFEF914EF67BDB2" blockId="8.[833,1522,1571,1949]" lastBlockId="9.[112,800,195,1098]" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
Geographically, this is a high-latitude, cool-water species complex, occurring from the Keasey Formation in
<collectingRegion id="8854F829FFF02934FCDEF952ED99BA7F" box="[833,925,1782,1808]" country="United States of America" name="Oregon" pageId="8" pageNumber="7">Oregon</collectingRegion>
to the Poul Creek and Narrow Cape Formations in the Gulf of
<collectingRegion id="8854F829FFF02934FC6CF8BDEA42BA5C" box="[1011,1094,1817,1843]" country="United States of America" name="Alaska" pageId="8" pageNumber="7">Alaska</collectingRegion>
(
<bibRefCitation id="2E014B3AFFF02934FBC5F8BEEAF1BA5C" author="Kanno, S." box="[1114,1269,1817,1844]" pageId="8" pageNumber="7" pagination="1 - 154" refId="ref31982" refString="Kanno, S. 1971. Tertiary molluscan fauna from the Yakataga district and adjacent areas of southern Alaska. Palaeontological Society of Japan Special Papers 16: 1 - 154, 11 pls." type="book chapter" year="1971">Kanno 1971</bibRefCitation>
,
<bibRefCitation id="2E014B3AFFF02934FA9BF8BDEDDDBA38" author="Allison, R. C. &amp; L. Marincovich, Jr." pageId="8" pageNumber="7" pagination="1 - 11" refId="ref26963" refString="Allison, R. C., and L. Marincovich, Jr. 1981. A late Oligoene or earliest Miocene molluscan fauna from Sitkinak Island, Alaska. United States Geological Survey Professionnal Paper 1233: 1 - 11, 3 pls., 3 figs." type="book chapter" year="1981">Allison and Marincovich 1981</bibRefCitation>
). The fauna of the Narrow Cape Formation is noted for its mixture of Asiatic, North American, and endemic high-latitude North Pacific taxa (
<bibRefCitation id="2E014B3AFFF02934FAFFF827EBF5BAF2" author="Allison, R. C. &amp; L. Marincovich, Jr." box="[1376,1521,1923,1949]" pageId="8" pageNumber="7" pagination="1 - 11" refId="ref26963" refString="Allison, R. C., and L. Marincovich, Jr. 1981. A late Oligoene or earliest Miocene molluscan fauna from Sitkinak Island, Alaska. United States Geological Survey Professionnal Paper 1233: 1 - 11, 3 pls., 3 figs." type="book chapter" year="1981">Allison and</bibRefCitation>
Marincovich 1981).
</paragraph>
<paragraph id="4A2F36CBFFF12935FF0FFF42EC04BCBC" blockId="9.[112,800,195,1098]" pageId="9" pageNumber="8">
The Danian
<taxonomicName id="8D904D48FFF12935FEBCFF42ECDABC6E" authority="Marincovich (1993)" authorityName="Marincovich" authorityYear="1993" box="[291,734,229,258]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="kannoi">
<emphasis id="78E4EAD9FFF12935FEBCFF42EFE5BC6E" box="[291,481,229,258]" italics="true" pageId="9" pageNumber="8">Crenella kannoi</emphasis>
Marincovich (1993)
</taxonomicName>
from the Prince Creek Formation on the Arctic coastal plain of
<collectingRegion id="8854F829FFF12935FF10FE88EEE6BC29" box="[143,226,300,326]" country="United States of America" name="Alaska" pageId="9" pageNumber="8">Alaska</collectingRegion>
is strikingly similar in size and ovate-elongate shape to
<taxonomicName id="8D904D48FFF12935FF7AFEF4EF89BC05" authorityName="Weaver, Vokes" authorityYear="1912" box="[229,397,334,364]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="porterensis">
<emphasis id="78E4EAD9FFF12935FF7AFEF4EF89BC05" box="[229,397,334,364]" italics="true" pageId="9" pageNumber="8">C. porterensis</emphasis>
</taxonomicName>
, but differs in a distinctive pattern of concentric undulations and wavy radial threads. As noted above, Arctic and Northeastern Pacific faunas were not connected at this time.
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFF12935FF0FFE78EBF4BC6F" pageId="9" pageNumber="8" type="materials_examined">
<paragraph id="4A2F36CBFFF12935FF0FFE78EC5FBC99" blockId="9.[112,800,195,1098]" box="[144,603,476,503]" pageId="9" pageNumber="8">
<emphasis id="78E4EAD9FFF12935FF0FFE78EF88BC99" bold="true" box="[144,396,476,502]" pageId="9" pageNumber="8">Material examined</emphasis>
<specimenCount id="5C96FD42FFF12935FE31FE79EC52BC99" box="[430,598,476,503]" count="13" pageId="9" pageNumber="8" type="generic">13 specimens</specimenCount>
.
</paragraph>
<paragraph id="4A2F36CBFFF12935FF0FFE5BEEC3BFA5" blockId="9.[112,800,195,1098]" pageId="9" pageNumber="8">
<emphasis id="78E4EAD9FFF12935FF0FFE5BEFEABF75" bold="true" box="[144,494,511,538]" pageId="9" pageNumber="8">Figured Keasey hypotypes</emphasis>
<collectionCode id="2C81AE0EFFF12935FD95FDA4EC5CBF75" box="[522,600,512,538]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="9" pageNumber="8" type="University or college">UCMP</collectionCode>
110725 (in matrix), length 7.9 mm, height 10.8 mm, Loc.
<collectionCode id="2C81AE0EFFF12935FDECFD87ECBBBF52" box="[627,703,547,573]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="9" pageNumber="8" type="University or college">UCMP</collectionCode>
IP7984 (=
<collectionCode id="2C81AE0EFFF12935FF19FDE2EEC9BF0F" box="[134,205,582,608]" country="USA" httpUri="http://grbio.org/cool/3n47-ewde" name="U.S. Geological Survey" pageId="9" pageNumber="8">USGS</collectionCode>
25031);
<collectionCode id="2C81AE0EFFF12935FEDAFDE2EF95BF0F" box="[325,401,582,608]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="9" pageNumber="8" type="University or college">UCMP</collectionCode>
110726 (in matrix), length 7.9 mm, height 9.8 mm, Loc.
<collectionCode id="2C81AE0EFFF12935FE3BFDCDEFF4BFEC" box="[420,496,617,643]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="9" pageNumber="8" type="University or college">UCMP</collectionCode>
IP2286 (=
<collectionCode id="2C81AE0EFFF12935FDF1FDCDECB1BFEC" box="[622,693,617,643]" country="USA" httpUri="http://grbio.org/cool/3n47-ewde" name="U.S. Geological Survey" pageId="9" pageNumber="8">USGS</collectionCode>
15265);
<collectionCode id="2C81AE0EFFF12935FFEFFD29EEB9BFC8" box="[112,189,653,679]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="8" type="Museum">USNM</collectionCode>
561803, length 8.9 mm, height 9.5 mm. Loc.
<collectionCode id="2C81AE0EFFF12935FD41FD29ED24BFC8" box="[734,800,653,679]" country="USA" httpUri="http://grbio.org/cool/3n47-ewde" name="U.S. Geological Survey" pageId="9" pageNumber="8">USGS</collectionCode>
15282.
</paragraph>
<paragraph id="4A2F36CBFFF12935FF0FFD76EC66BE39" blockId="9.[112,800,195,1098]" pageId="9" pageNumber="8">
<emphasis id="78E4EAD9FFF12935FF0FFD76EC51BF82" bold="true" box="[144,597,722,749]" pageId="9" pageNumber="8">Figured Lincoln Creek hypotypes</emphasis>
<collectionCode id="2C81AE0EFFF12935FDEEFD77ECBBBF82" box="[625,703,723,749]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="9" pageNumber="8" type="University or college">UCMP</collectionCode>
32410, length 10.18 mm, height 18.2 mm, Loc.
<collectionCode id="2C81AE0EFFF12935FDC4FD52ECA3BE7F" box="[603,679,758,784]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="9" pageNumber="8" type="University or college">UCMP</collectionCode>
A9 (=
<collectionCode id="2C81AE0EFFF12935FD71FD52ED24BE7F" box="[750,800,758,784]" country="USA" name="University of Washington Fish Collection" pageId="9" pageNumber="8" type="Museum">UW</collectionCode>
256, Weavers
<typeStatus id="952B8869FFF12935FEB7FCBDEF5BBE5C" box="[296,351,793,819]" pageId="9" pageNumber="8">type</typeStatus>
locality);
<collectionCode id="2C81AE0EFFF12935FE41FCBDEC2EBE5C" box="[478,554,793,819]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="9" pageNumber="8" type="University or college">UCMP</collectionCode>
35163, length 11.7 mm, height 18.0 mm, Loc.
<collectionCode id="2C81AE0EFFF12935FE29FC98EC06BE39" box="[438,514,828,854]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="9" pageNumber="8" type="University or college">UCMP</collectionCode>
A1811.
</paragraph>
<paragraph id="4A2F36CBFFF12935FF0FFCFAEC13BE8D" blockId="9.[112,800,195,1098]" pageId="9" pageNumber="8">
<emphasis id="78E4EAD9FFF12935FF0FFCFAEC2CBE17" bold="true" box="[144,552,862,888]" pageId="9" pageNumber="8">Keasey Formation localities</emphasis>
—Lower member:
<collectionCode id="2C81AE0EFFF12935FFEFFC25EEBABEF4" box="[112,190,897,923]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="9" pageNumber="8" type="University or college">UCMP</collectionCode>
IP2286,
<collectionCode id="2C81AE0EFFF12935FEA6FC25EF79BEF4" box="[313,381,897,923]" country="USA" httpUri="http://grbio.org/cool/3n47-ewde" name="U.S. Geological Survey" pageId="9" pageNumber="8">USGS</collectionCode>
15265. Middle member:
<collectionCode id="2C81AE0EFFF12935FD4DFC25ED1BBEF4" box="[722,799,897,923]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="9" pageNumber="8" type="University or college">UCMP</collectionCode>
IP7984,
<collectionCode id="2C81AE0EFFF12935FF48FC00EF1DBED1" box="[215,281,932,958]" country="USA" httpUri="http://grbio.org/cool/3n47-ewde" name="U.S. Geological Survey" pageId="9" pageNumber="8">USGS</collectionCode>
15282, 15276, 15277, 15279, 15525. Upper member:
<collectionCode id="2C81AE0EFFF12935FE86FC63EF5FBE8E" box="[281,347,967,993]" country="USA" httpUri="http://grbio.org/cool/3n47-ewde" name="U.S. Geological Survey" pageId="9" pageNumber="8">USGS</collectionCode>
13515, 15518.
</paragraph>
<paragraph id="4A2F36CBFFF12935FF0FFC4EEBF4BC6F" blockId="9.[112,800,195,1098]" lastBlockId="9.[832,1520,195,256]" pageId="9" pageNumber="8">
<emphasis id="78E4EAD9FFF12935FF0FFC4EEF8CB96B" bold="true" box="[144,392,1002,1028]" pageId="9" pageNumber="8">Other occurrences</emphasis>
—In
<collectingRegion id="8854F829FFF12935FE59FC4EEC26B96B" box="[454,546,1002,1028]" country="United States of America" name="Oregon" pageId="9" pageNumber="8">Oregon</collectingRegion>
this species also occurs stratigraphically higher in the overlying Pittsburg Bluff Formation and Miocene Astoria Formation. In
<materialsCitation id="FAF83C96FFF12935FCDFFF67EBE8BC6F" ID-GBIF-Occurrence="4873742322" location="Eo-Oligocene Lincoln Creek" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Washington">
<collectingRegion id="8854F829FFF12935FCDFFF67EDD1BDB2" box="[832,981,195,221]" country="United States of America" name="Washington" pageId="9" pageNumber="8">Washington</collectingRegion>
it occurs in the
<location id="4F4F6010FFF12935FB06FF67EBF4BDB2" LSID="urn:lsid:plazi:treatment:C23987DDFFFF2935FF44F9FDEBF4BC6F:4F4F6010FFF12935FB06FF67EBF4BDB2" box="[1177,1520,195,221]" name="Eo-Oligocene Lincoln Creek" pageId="9" pageNumber="8" stateProvince="Washington">Eo-Oligocene Lincoln Creek</location>
Formation, Quimper Sandstone, and Blakeley Formation
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>