treatments-xml/data/C2/39/87/C23987DDFFDA2919FCC7FEE1EDC4B9E6.xml
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<document id="2F765FDE8685F37DA8081C0C6E00D2E6" ID-DOI="10.5070/P940561331" ID-ISSN="0031-0298" ID-Zenodo-Dep="10913477" ID-ZooBank="1756B24A-813B-423F-896F-91B21FF58A79" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712169731283" checkinUser="felipe" docAuthor="Hickman, Carole S." docDate="2023" docId="C23987DDFFDA2919FCC7FEE1EDC4B9E6" docLanguage="en" docName="PaleoBios.40.5.1-51.pdf" docOrigin="PaleoBios 40 (5)" docSource="http://dx.doi.org/10.5070/p940561331" docStyle="DocumentStyle:76F55B8832C4952BA7DD61B33E1F0E81.1:PaleoBios.2023-.journal_article" docStyleId="76F55B8832C4952BA7DD61B33E1F0E81" docStyleName="PaleoBios.2023-.journal_article" docStyleVersion="1" docTitle="Propeamussium (Parvamussium) mistensis Hickman 2023, N. SP." docType="treatment" docVersion="2" lastPageNumber="36" masterDocId="3E00FFA5FFF8293CFF9FFFA4EE04BD6F" masterDocTitle="Paleogene marine bivalves of the deep-water Keasey Formation in Oregon, Part II: The pteriomorphs" masterLastPageNumber="51" masterPageNumber="1" pageNumber="33" updateTime="1712170163329" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="AE9557D0E0092318445C2232048EE617">Paleogene marine bivalves of the deep-water Keasey Formation in Oregon, Part II: The pteriomorphs</mods:title>
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<treatment id="C23987DDFFDA2919FCC7FEE1EDC4B9E6" LSID="urn:lsid:plazi:treatment:C23987DDFFDA2919FCC7FEE1EDC4B9E6" httpUri="http://treatment.plazi.org/id/C23987DDFFDA2919FCC7FEE1EDC4B9E6" lastPageId="37" lastPageNumber="36" pageId="34" pageNumber="33">
<subSubSection id="028A6540FFDA291EFCC7FEE1EBDEBC0E" box="[856,1498,325,353]" pageId="34" pageNumber="33" type="nomenclature">
<paragraph id="4A2F36CBFFDA291EFCC7FEE1EBDEBC0E" blockId="34.[856,1498,325,385]" box="[856,1498,325,353]" pageId="34" pageNumber="33">
<heading id="116781A7FFDA291EFCC7FEE1EBDEBC0E" allCaps="true" box="[856,1498,325,353]" centered="true" fontSize="10" level="2" pageId="34" pageNumber="33" reason="2">
<taxonomicName id="8D904D48FFDA291EFCC7FEE1EB9FBC0E" authority="Hickman, 2023" authorityName="Hickman" authorityYear="2023" box="[856,1435,325,353]" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" kingdom="Animalia" order="Pectinida" pageId="34" pageNumber="1" phylum="Mollusca" rank="species" species="mistensis" status="n. sp." subGenus="Parvamussium">
<emphasis id="78E4EAD9FFDA291EFCC7FEE1EB9FBC0E" box="[856,1435,325,353]" italics="true" pageId="34" pageNumber="33">PROPEAMUSSIUM (PARVAMUSSIUM) MISTENSIS</emphasis>
</taxonomicName>
<taxonomicNameLabel id="63D757A2FFDA291EFA3EFEEDEBDEBC0E" box="[1441,1498,329,353]" pageId="34" pageNumber="33" rank="species">N. SP.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFDA291EFC60FECDEAF6BF3F" pageId="34" pageNumber="33" type="description">
<paragraph id="4A2F36CBFFDA291EFC60FECDEB36BCEE" blockId="34.[856,1498,325,385]" box="[1023,1330,361,385]" pageId="34" pageNumber="33">
<figureCitation id="D2AB2A4EFFDA291EFC60FECDEA83BCEE" box="[1023,1159,361,385]" captionStart="Figure 21" captionStartId="35.[115,190,1833,1857]" captionTargetBox="[131,1504,192,1811]" captionTargetId="figure-13@35.[131,1504,192,1811]" captionTargetPageId="35" captionText="Figure 21. Propeamussium (Parvamussium) mistensis n. sp., paired color images of molds in matrix and corresponding ammonium chloride-coated casts. A. Left valve exterior, holotype USNM 561849, length 8 mm. B. Left valve exterior, paratype USNM 561856, length 8.1 mm. C. Right valve exterior, paratype USNM 561850, length 5 mm. D. Right valve exterior on top of left valve, paratype USNM 561854, length 7 mm. E. Left valve interior, Paratype USNM 561852, length 6 mm." figureDoi="http://doi.org/10.5281/zenodo.10913529" httpUri="https://zenodo.org/record/10913529/files/figure.png" pageId="34" pageNumber="33">FIGS. 21AE</figureCitation>
.
<figureCitation id="D2AB2A4EFFDA291EFB0FFECDEADFBCEE" box="[1168,1243,361,385]" captionStart="Figure 22" captionStartId="36.[114,189,1847,1871]" captionTargetBox="[144,764,195,1827]" captionTargetId="figure-438@36.[144,764,195,1827]" captionTargetPageId="36" captionText="Figure 22. Propeamussium (Parvamussium) mistensis n. sp., uncoated latex casts of three paratypes. A. Left valve exterior, USNM 561853. B. Left valve exterior, USNM 561856. Right valve exterior, USNM 561850. Scale bars=3 mm." figureDoi="http://doi.org/10.5281/zenodo.10913533" httpUri="https://zenodo.org/record/10913533/files/figure.png" pageId="34" pageNumber="33">22AC</figureCitation>
,
<figureCitation id="D2AB2A4EFFDA291EFB7AFECDEB36BCEE" box="[1253,1330,361,385]" captionStart="Figure 23" captionStartId="37.[112,187,1860,1884]" captionTargetBox="[112,1521,1352,1834]" captionTargetId="figure-546@37.[112,1521,1352,1834]" captionTargetPageId="37" captionText="Figure 23. Propeamussium (Parvamussium) mistensis n. sp., reconstruction of major features of shell morphology based on all material examined. A. Left valve exterior. B. Left valve interior. C. Right valve exterior. D. Right valve interior. Dashed lines in C, D indicate extent of the larger left valve and uncalcified flexible apron of right valve." figureDoi="http://doi.org/10.5281/zenodo.10913535" httpUri="https://zenodo.org/record/10913535/files/figure.png" pageId="34" pageNumber="33">23AD</figureCitation>
</paragraph>
<paragraph id="4A2F36CBFFDA291EFCDFFE3FEDA5BCBC" blockId="34.[832,1484,411,467]" pageId="34" pageNumber="33">
<emphasis id="78E4EAD9FFDA291EFCDFFE3FEDF0BCD8" box="[832,1012,411,439]" italics="true" pageId="34" pageNumber="33">Propeamssium</emphasis>
<taxonomicNameLabel id="63D757A2FFDA291EFC65FE38EA3EBCD9" box="[1018,1082,412,438]" pageId="34" pageNumber="33" rank="species">n. sp.</taxonomicNameLabel>
<collectionCode id="2C81AE0EFFDA291EFBDEFE38EA51BCD9" box="[1089,1109,412,438]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="34" pageNumber="33">R</collectionCode>
.C. Moore and Vokes (1953). p. 119.
</paragraph>
<paragraph id="4A2F36CBFFDA291EFCDFFE47EB75BC91" blockId="34.[832,1393,483,511]" box="[832,1393,483,511]" pageId="34" pageNumber="33">
<taxonomicName id="8D904D48FFDA291EFCDFFE47EA29BC91" box="[832,1069,483,511]" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" kingdom="Animalia" order="Pectinida" pageId="34" pageNumber="33" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="78E4EAD9FFDA291EFCDFFE47EA00BC90" box="[832,1028,483,511]" italics="true" pageId="34" pageNumber="33">Propeamussium</emphasis>
sp.
</taxonomicName>
<bibRefCitation id="2E014B3AFFDA291EFBABFE40EB06BC91" author="Hickman, C. S." box="[1076,1282,484,510]" pageId="34" pageNumber="33" pagination="1215 - 1234" refId="ref31128" refString="Hickman, C. S. 1984. Composition, structure, ecology, and evolution of six Cenozoic deep-water mollusk communities. Journal of Paleontology 58: 1215 - 1234." type="journal article" year="1984">Hickman (1984)</bibRefCitation>
. p. 1224.
</paragraph>
<paragraph id="4A2F36CBFFDA291EFCFFFDB6EAF6BF3F" blockId="34.[832,1521,529,1927]" pageId="34" pageNumber="33">
<emphasis id="78E4EAD9FFDA291EFCFFFDB6EA12BF44" bold="true" box="[864,1046,529,556]" pageId="34" pageNumber="33">
ZooBank
<collectionCode id="2C81AE0EFFDA291EFC46FDB5EA12BF44" box="[985,1046,529,555]" pageId="34" pageNumber="33">LSID</collectionCode>
</emphasis>
<uri id="3E013AC9FFDA291EFBA9FDB6EAF6BF3F" pageId="34" pageNumber="33">
urn:lsid:zoobank.org:act:
<uuid id="3E360C1EFFDA291EFAC6FDB6EAF6BF3F" pageId="34" pageNumber="33">3BEBE80B- 2C47-4694-A13C-DB5A3B9FAB0D</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFDA291EFCFFFDFCEDB9B9EF" pageId="34" pageNumber="33" type="diagnosis">
<paragraph id="4A2F36CBFFDA291EFCFFFDFCEDB9B9EF" blockId="34.[832,1521,529,1927]" pageId="34" pageNumber="33">
<emphasis id="78E4EAD9FFDA291EFCFFFDFCEDE5BF1D" bold="true" box="[864,993,600,626]" pageId="34" pageNumber="33">Diagnosis</emphasis>
—Shell small (&lt;
<quantity id="8D689B2EFFDA291EFB3FFDFCEAE3BF1C" box="[1184,1255,600,627]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="33" unit="mm" value="5.0">5 mm</quantity>
),thin, outline circular, equilateral with length slightly greater than or equal to height; inequivalve: larger, right valve with smaller cal- cified portion and missing uncalcified maginal flexible apron (not preserved) that fits adpressed to the ventral margin of larger left valve; hingeline narrow with small auricles distinctly set off from disc, right anterior auricle with byssal notch but no ctenolium; exterior surface sculpture discrepant; right valve with numerous sharply defined commarginal riblets and no radial riblets, left valve scabrous, with prominent primary, secondary and tertiary radial riblets crossed by weaker commarginal lirae with semi-erect spines or nodules forming at intersections; interior of both valves smooth with prominent radial slats of lathic calcite imbedded in crossed-lamellar aragonite.
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFDA2918FCFFFB2DEBF4BC4C" lastPageId="36" lastPageNumber="35" pageId="34" pageNumber="33" type="description">
<paragraph id="4A2F36CBFFDA291EFCFFFB2DEA40BA72" blockId="34.[832,1521,529,1927]" pageId="34" pageNumber="33">
<emphasis id="78E4EAD9FFDA291EFCFFFB2DEA05B9CC" bold="true" box="[864,1025,1161,1187]" pageId="34" pageNumber="33">Description</emphasis>
—The squamose or cancellate surface texture of the larger left valves is well-preserved in exterior molds (
<figureCitation id="D2AB2A4EFFDA291EFC70FB74EA7AB985" box="[1007,1150,1232,1258]" captionStart="Figure 21" captionStartId="35.[115,190,1833,1857]" captionTargetBox="[131,1504,192,1811]" captionTargetId="figure-13@35.[131,1504,192,1811]" captionTargetPageId="35" captionText="Figure 21. Propeamussium (Parvamussium) mistensis n. sp., paired color images of molds in matrix and corresponding ammonium chloride-coated casts. A. Left valve exterior, holotype USNM 561849, length 8 mm. B. Left valve exterior, paratype USNM 561856, length 8.1 mm. C. Right valve exterior, paratype USNM 561850, length 5 mm. D. Right valve exterior on top of left valve, paratype USNM 561854, length 7 mm. E. Left valve interior, Paratype USNM 561852, length 6 mm." figureDoi="http://doi.org/10.5281/zenodo.10913529" httpUri="https://zenodo.org/record/10913529/files/figure.png" pageId="34" pageNumber="33">Fig. 21A, B</figureCitation>
). Although the surface of the smaller right valves is more subdued in the absence of radial riblets, the commarginal riblets are well defined in exterior molds (
<figureCitation id="D2AB2A4EFFDA291EFBBBFA9DEAABB83B" box="[1060,1199,1337,1364]" captionStart="Figure 21" captionStartId="35.[115,190,1833,1857]" captionTargetBox="[131,1504,192,1811]" captionTargetId="figure-13@35.[131,1504,192,1811]" captionTargetPageId="35" captionText="Figure 21. Propeamussium (Parvamussium) mistensis n. sp., paired color images of molds in matrix and corresponding ammonium chloride-coated casts. A. Left valve exterior, holotype USNM 561849, length 8 mm. B. Left valve exterior, paratype USNM 561856, length 8.1 mm. C. Right valve exterior, paratype USNM 561850, length 5 mm. D. Right valve exterior on top of left valve, paratype USNM 561854, length 7 mm. E. Left valve interior, Paratype USNM 561852, length 6 mm." figureDoi="http://doi.org/10.5281/zenodo.10913529" httpUri="https://zenodo.org/record/10913529/files/figure.png" pageId="34" pageNumber="33">Fig. 21C, D</figureCitation>
). Radial riblets are absent in the umbonal region of the left valve, with inception occurring at approximately 1/3 of the distance from the umbones and continuing to the ventral margin of the disc. Secondary and tertiary riblets are added by intercalation. The total number of riblets is variable, with 55 on the
<typeStatus id="952B8869FFDA291EFCDFF9A8EDB5BB49" box="[832,945,1548,1574]" pageId="34" pageNumber="33" type="holotype">holotype</typeStatus>
. The total number of commarginal riblets on the right valve is approximately 2530. Sculpture of the anterior and posterior auricles of the right valve includes both radial and commarginal sculpture while those of the left valve are predominantly concentric on the posterior ear, with strongly developed radials on the anterior ear. Interior molds preserve deep impressions of 12 to 14 narrow slat-like ribs.
</paragraph>
<paragraph id="4A2F36CBFFDA2918FCFFF882EBF4BC4C" blockId="34.[832,1521,529,1927]" lastBlockId="36.[832,1521,195,1946]" lastPageId="36" lastPageNumber="35" pageId="34" pageNumber="33">
An incomplete interior mold of a valve from Mist (
<collectionCode id="2C81AE0EFFDA291EFCD8F8EDED93BA0C" box="[839,919,1865,1891]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="34" pageNumber="33" type="University or college">UCMP</collectionCode>
110764) lacks auricles (it may be either a right or a left valve), but the disc preserves deep impressions of 14 ribs that extend to the ventral margin of the valve. A small fragment of adhering shell, removed from the mold, preserves portions of five of the imbedded slat-like ribs.
</paragraph>
</subSubSection>
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<paragraph id="4A2F36CBFFDB291FFFECF88DEA2EBAFE" blockId="35.[115,1514,1833,1938]" pageId="35" pageNumber="34">
<emphasis id="78E4EAD9FFDB291FFFECF88DEEE5BA2E" bold="true" box="[115,225,1833,1857]" pageId="35" pageNumber="34">Figure 21</emphasis>
.
<taxonomicName id="8D904D48FFDB291FFF73F88FECB4BA2C" authority="Hickman, 2023" authorityName="Hickman" authorityYear="2023" box="[236,688,1835,1859]" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" kingdom="Animalia" order="Pectinida" pageId="35" pageNumber="34" phylum="Mollusca" rank="species" species="mistensis" status="sp. nov." subGenus="Parvamussium">
<emphasis id="78E4EAD9FFDB291FFF73F88FECB4BA2C" box="[236,688,1835,1859]" italics="true" pageId="35" pageNumber="34">Propeamussium (Parvamussium) mistensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="63D757A2FFDB291FFD29F88EECEABA2E" box="[694,750,1834,1857]" pageId="35" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
, paired color images of molds in matrix and corresponding ammonium chloride-coated casts.
<emphasis id="78E4EAD9FFDB291FFEFDF8E0EF77BA33" bold="true" box="[354,371,1860,1884]" pageId="35" pageNumber="34">A</emphasis>
. Left valve exterior, holotype USNM 561849, length 8 mm.
<emphasis id="78E4EAD9FFDB291FFC77F8E0EDFCBA33" bold="true" box="[1000,1016,1860,1884]" pageId="35" pageNumber="34">B</emphasis>
. Left valve exterior, paratype USNM 561856, length 8.1 mm.
<emphasis id="78E4EAD9FFDB291FFE87F8FBEF23BA18" bold="true" box="[280,295,1887,1911]" pageId="35" pageNumber="34">C</emphasis>
. Right valve exterior, paratype USNM 561850, length 5 mm.
<emphasis id="78E4EAD9FFDB291FFC32F8FBEDBBBA18" bold="true" box="[941,959,1887,1911]" pageId="35" pageNumber="34">D</emphasis>
. Right valve exterior on top of left valve, paratype USNM 561854, length 7 mm.
<emphasis id="78E4EAD9FFDB291FFE32F8DEEFB8BAFD" bold="true" box="[429,444,1914,1938]" pageId="35" pageNumber="34">E</emphasis>
. Left valve interior, Paratype USNM 561852, length 6 mm.
</paragraph>
</caption>
<subSubSection id="028A6540FFDC2919FCFFFE88EF7EB992" lastPageId="37" lastPageNumber="36" pageId="36" pageNumber="35" type="discussion">
<paragraph id="4A2F36CBFFDC2918FCFFFE88EB20BFEB" blockId="36.[832,1521,195,1946]" pageId="36" pageNumber="35">
<emphasis id="78E4EAD9FFDC2918FCFFFE88EDF7BC29" bold="true" box="[864,1011,300,326]" pageId="36" pageNumber="35">Discussion</emphasis>
—This is the first population of an un- equivocal propeamussiid named and described from the Paleogene of the Cascadia Margin. It is also the earliest documentation of Eastern Pacific propeamussiids from a methane seep hardground. However,
<bibRefCitation id="2E014B3AFFDC2918FAFDFE1DEBF4BCBB" author="Kiel, S." box="[1378,1520,441,468]" pageId="36" pageNumber="35" pagination="121 - 137" refId="ref32175" refString="Kiel, S. 2006. New Records and species of molluscs from Tertiary cold-seep carbonates in Washington State, USA. Journal of Paleontology 80 (1): 121 - 137." type="journal article" year="2006">Kiel (2006)</bibRefCitation>
extracted a suspect propeamussiid shell with well-preserved foliated calcite microstructure from an Oligocene limestone float block in
<collectingRegion id="8854F829FFDC2918FBF3FD87EB43BF52" box="[1132,1351,547,573]" country="United States of America" name="Washington" pageId="36" pageNumber="35">Washington State</collectingRegion>
yielding δ
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C values indicating methane seep origin, describing it in open nomenclature as
<taxonomicName id="8D904D48FFDC2918FBC4FDCEEB20BFEB" box="[1115,1316,618,646]" class="Bivalvia" family="Propeamussiidae" genus="Catillopecten" kingdom="Animalia" order="Pectinida" pageId="36" pageNumber="35" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="78E4EAD9FFDC2918FBC4FDCEEAFFBFE9" box="[1115,1275,618,646]" italics="true" pageId="36" pageNumber="35">Catillopecten</emphasis>
sp.
</taxonomicName>
</paragraph>
<paragraph id="4A2F36CBFFDC2918FCFFFD28EA26B9FC" blockId="36.[832,1521,195,1946]" pageId="36" pageNumber="35">
Byssally- attached “living fossil” propeamussiids were discovered earlier on the periphery of hydrothermal vent systems at 13
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and 9
<superScript id="BDE59B83FFDC2918FB0FFD77EA9FBF89" attach="left" box="[1168,1179,723,742]" fontSize="8" pageId="36" pageNumber="35">o</superScript>
N on the East Pacific Rise (Schein-Fatton 1985). Described as a new genus and species,
<taxonomicName id="8D904D48FFDC2918FC37FCBEEBB7BE5B" authority="Schein-Fatton (1988)" authorityName="Schein-Fatton" authorityYear="1988" box="[936,1459,792,822]" class="Bivalvia" family="Propeamussiidae" genus="Bathypecten" kingdom="Animalia" order="Pectinida" pageId="36" pageNumber="35" phylum="Mollusca" rank="species" species="vulcani">
<emphasis id="78E4EAD9FFDC2918FC37FCBEEAA5BE5B" box="[936,1185,792,822]" italics="true" pageId="36" pageNumber="35">Bathypecten vulcani</emphasis>
Schein-Fatton (1988)
</taxonomicName>
, it is now assigned to
<taxonomicName id="8D904D48FFDC2918FB8CFC99EB77BE38" authority="Iredale (1939)" authorityName="Iredale" authorityYear="1939" box="[1043,1395,828,857]" class="Bivalvia" family="Propeamussiidae" genus="Catillopecten" kingdom="Animalia" order="Pectinida" pageId="36" pageNumber="35" phylum="Mollusca" rank="genus">
<emphasis id="78E4EAD9FFDC2918FB8CFC99EAB7BE36" box="[1043,1203,829,857]" italics="true" pageId="36" pageNumber="35">Catillopecten</emphasis>
<bibRefCitation id="2E014B3AFFDC2918FB24FC98EB77BE38" author="Iredale, T." box="[1211,1395,828,855]" pageId="36" pageNumber="35" pagination="209 - 425" refId="ref31545" refString="Iredale, T. 1939. Mollusca. Part 1. Great Barrier Reef Expedition 1928 - 29 Scientific Reports 5 (6): 209 - 425, pls, 1 - 7." type="journal article" year="1939">Iredale (1939)</bibRefCitation>
</taxonomicName>
. Detailed anatomical study of this peri-hydrothermal vent species (
<bibRefCitation id="2E014B3AFFDC2918FCD4FC27EA54BEF2" author="Beninger, P. G. &amp; S. C. Dufour &amp; P. Decottignies &amp; M. Le Pennec" box="[843,1104,899,925]" pageId="36" pageNumber="35" pagination="183 - 195" refId="ref27364" refString="Beninger, P. G., S. C. Dufour, P. Decottignies, and M. Le Pennec. 2003. Particle processing mechanisms in the archaic peri-hydrothermal vent bivalve Bathypecten vulcani, inferred from cilia and mucocyte distributions on the gill. Marine Ecology Progress Series 246: 183 - 195." type="book chapter" year="2003">Beninger et al. 2003</bibRefCitation>
, LePennec et al. 2003) reveals a unique and putatively archaic particle processing mechanism. This reinforces previous evidence for a peripheral zone of seep mollusks uniquely adapted to hypoxic and chemically hostile environments as well as opportunistic suspension feeding on an unconventional carbon source—in this instance chemosynthetic microbes in the water column.
</paragraph>
<paragraph id="4A2F36CBFFDC2918FCFFFB38EA68B8C2" blockId="36.[832,1521,195,1946]" pageId="36" pageNumber="35">
The new Keasey propeamussiid is easily distinguished from all previously described Eastern Pacific species. Details of the discrepant sculpture on the discs and auricles of the right and left valves, delayed ontogenetic appearance of the slat-like internal ribs, the presence of a byssal notch, and squamose sculpture on the right valve support its assignment to
<taxonomicName id="8D904D48FFDC2918FB7FFACAEB93B8E5" authorityName="SACCO" authorityYear="1897" box="[1248,1431,1390,1418]" class="Bivalvia" family="Propeamussiidae" genus="Parvamussium" kingdom="Animalia" order="Pectinida" pageId="36" pageNumber="35" phylum="Mollusca" rank="genus">
<emphasis id="78E4EAD9FFDC2918FB7FFACAEB93B8E5" box="[1248,1431,1390,1418]" italics="true" pageId="36" pageNumber="35">Parvamussium</emphasis>
</taxonomicName>
rather than
<taxonomicName id="8D904D48FFDC2918FCE0FA35EA47B8C2" authorityName="DE GREGORIO" authorityYear="1884" box="[895,1091,1425,1453]" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" kingdom="Animalia" order="Pectinida" pageId="36" pageNumber="35" phylum="Mollusca" rank="genus" sensu="stricto">
<emphasis id="78E4EAD9FFDC2918FCE0FA35EA47B8C2" box="[895,1091,1425,1453]" italics="true" pageId="36" pageNumber="35">Propeamussium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="63D757A2FFDC2918FBD6FA35EA68B8C2" box="[1097,1132,1425,1453]" pageId="36" pageNumber="35" sensu="stricto">
<emphasis id="78E4EAD9FFDC2918FBD6FA35EA61B8C2" box="[1097,1125,1425,1453]" italics="true" pageId="36" pageNumber="35">s.s</emphasis>
.
</taxonomicNameLabel>
</paragraph>
<paragraph id="4A2F36CBFFDC2919FCFFFA12EC2FBC6F" blockId="36.[832,1521,195,1946]" lastBlockId="37.[112,800,195,1312]" lastPageId="37" lastPageNumber="36" pageId="36" pageNumber="35">
Although Paleogene propeamussiids have a long history of recognition in
<collectingRegion id="8854F829FFDC2918FBC1FA7DEADFB89C" box="[1118,1243,1497,1523]" country="United States of America" name="California" pageId="36" pageNumber="35">California</collectingRegion>
, most of the material is preserved in mudstone facies as poorly-preserved interior casts recognized primarily by sharply-incised impressions of the radial ribs.
<taxonomicName id="8D904D48FFDC2918FB48F9E5EA5CBBEE" authority="(Gabb, 1869)" baseAuthorityName="Gabb" baseAuthorityYear="1869" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" kingdom="Animalia" order="Pectinida" pageId="36" pageNumber="35" phylum="Mollusca" rank="species" species="interradiatus">
<emphasis id="78E4EAD9FFDC2918FB48F9E5EDA3BBEE" italics="true" pageId="36" pageNumber="35">Propeamussium interradiatus</emphasis>
(
<bibRefCitation id="2E014B3AFFDC2918FC23F9C2EA49BBEF" author="Gabb, W. M." box="[956,1101,1638,1664]" pageId="36" pageNumber="35" refId="ref29855" refString="Gabb, W. M. 1869. Paleontology of California, Cretaceous and Tertiary fossils: Descriptions of new species. Geological Survey of California, Paleontology 2, 299 pp." type="book" year="1869">Gabb, 1869</bibRefCitation>
)
</taxonomicName>
has been used for much of this material, and difficulties with the typification and handdrawn illustrations of Gabbs species have been discussed by many authors (e.g., Stewart 1930,
<bibRefCitation id="2E014B3AFFDC2918FAB2F974ED85BA62" author="Grant, U. S. &amp; H. R. Gale" pageId="36" pageNumber="35" refId="ref30199" refString="Grant, U. S., IV, and H. R. Gale. 1931. Catalogue of the marine Pliocene and Pleistocene Mollusca of California and adjacent regions. San Diego Society of Natural History, Memoir I. 1036 pp., 32 pls." type="book" year="1931">Grant and Gale 1931</bibRefCitation>
,
<bibRefCitation id="2E014B3AFFDC2918FC12F957EA3BBA62" author="Addicott, W. O." box="[909,1087,1779,1805]" pageId="36" pageNumber="35" refId="ref26829" refString="Addicott, W. O. 1971. Some Paleogene mud pectens of the genus Propeamussium from Alaska and California. The Veliger 13 (3): 226 - 130, 13 figs." type="book" year="1971">Addicott 1971</bibRefCitation>
). Five
<collectingRegion id="8854F829FFDC2918FB10F957EB0DBA62" box="[1167,1289,1779,1805]" country="United States of America" name="California" pageId="36" pageNumber="35">California</collectingRegion>
Paleogene species of
<taxonomicName id="8D904D48FFDC2918FCFEF8B1EA12BA5E" authorityName="SACCO" authorityYear="1897" box="[865,1046,1813,1841]" class="Bivalvia" family="Propeamussiidae" genus="Parvamussium" kingdom="Animalia" order="Pectinida" pageId="36" pageNumber="35" phylum="Mollusca" rank="genus">
<emphasis id="78E4EAD9FFDC2918FCFEF8B1EA12BA5E" box="[865,1046,1813,1841]" italics="true" pageId="36" pageNumber="35">Parvamussium</emphasis>
</taxonomicName>
are recognized and illustrated by E.J. Moore (1984b). Propeamussiids in
<collectingRegion id="8854F829FFDC2918FB63F89DEB4BBA3C" box="[1276,1359,1849,1875]" country="United States of America" name="Alaska" pageId="36" pageNumber="35">Alaska</collectingRegion>
(Scholl et al. 1970), in rocks originally considered Paleozoic, were identified, and described by
<bibRefCitation id="2E014B3AFFDC2918FB00F824EB63BAF5" author="Addicott, W. O." box="[1183,1383,1920,1946]" pageId="36" pageNumber="35" refId="ref26829" refString="Addicott, W. O. 1971. Some Paleogene mud pectens of the genus Propeamussium from Alaska and California. The Veliger 13 (3): 226 - 130, 13 figs." type="book" year="1971">Addicott (1971)</bibRefCitation>
along with Eocene Foraminifera, extending the distribution to high latitude in the Northeastern Pacific.
</paragraph>
<caption id="1EEF6643FFDC2918FFEDF893EC33BAF1" ID-DOI="http://doi.org/10.5281/zenodo.10913533" ID-Zenodo-Dep="10913533" httpUri="https://zenodo.org/record/10913533/files/figure.png" pageId="36" pageNumber="35" startId="36.[114,189,1847,1871]" targetBox="[144,764,195,1827]" targetPageId="36" targetType="figure">
<paragraph id="4A2F36CBFFDC2918FFEDF893EC33BAF1" blockId="36.[114,800,1847,1951]" pageId="36" pageNumber="35">
<emphasis id="78E4EAD9FFDC2918FFEDF893EEE5BA20" bold="true" box="[114,225,1847,1871]" pageId="36" pageNumber="35">Figure 22</emphasis>
.
<taxonomicName id="8D904D48FFDC2918FF74F89CECB4BA3F" authority="Hickman, 2023" authorityName="Hickman" authorityYear="2023" box="[235,688,1848,1872]" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" kingdom="Animalia" order="Pectinida" pageId="36" pageNumber="35" phylum="Mollusca" rank="species" species="mistensis" status="sp. nov." subGenus="Parvamussium">
<emphasis id="78E4EAD9FFDC2918FF74F89CECB4BA3F" box="[235,688,1848,1872]" italics="true" pageId="36" pageNumber="35">Propeamussium (Parvamussium) mistensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="63D757A2FFDC2918FD2AF893ECE9BA21" box="[693,749,1847,1870]" pageId="36" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
, uncoated latex casts of three paratypes.
<emphasis id="78E4EAD9FFDC2918FDBEF8F5EC36BA06" bold="true" box="[545,562,1873,1897]" pageId="36" pageNumber="35">A</emphasis>
. Left valve exterior, USNM 561853.
<emphasis id="78E4EAD9FFDC2918FE85F8C8EF2EBAEB" bold="true" box="[282,298,1900,1924]" pageId="36" pageNumber="35">B</emphasis>
. Left valve exterior, USNM 561856. Right valve exterior, USNM 561850. Scale bars=3 mm.
</paragraph>
</caption>
<paragraph id="4A2F36CBFFDD2919FF0FFEADEEC5BFC9" blockId="37.[112,800,195,1312]" pageId="37" pageNumber="36">
Unfortunately, the exterior and interior molds of the new Keasey species are difficult to illuminate and photograph, appearing in a common optical illusion that reverses positive and negative features (
<figureCitation id="D2AB2A4EFFDD2919FD2AFED7ED15BCE2" box="[693,785,371,397]" captionStart="Figure 21" captionStartId="35.[115,190,1833,1857]" captionTargetBox="[131,1504,192,1811]" captionTargetId="figure-13@35.[131,1504,192,1811]" captionTargetPageId="35" captionText="Figure 21. Propeamussium (Parvamussium) mistensis n. sp., paired color images of molds in matrix and corresponding ammonium chloride-coated casts. A. Left valve exterior, holotype USNM 561849, length 8 mm. B. Left valve exterior, paratype USNM 561856, length 8.1 mm. C. Right valve exterior, paratype USNM 561850, length 5 mm. D. Right valve exterior on top of left valve, paratype USNM 561854, length 7 mm. E. Left valve interior, Paratype USNM 561852, length 6 mm." figureDoi="http://doi.org/10.5281/zenodo.10913529" httpUri="https://zenodo.org/record/10913529/files/figure.png" pageId="37" pageNumber="36">Fig. 21</figureCitation>
). Latex casts of these specimens, coated with ammonium chloride (
<figureCitation id="D2AB2A4EFFDD2919FF77FE1DEF44BCBC" box="[232,320,441,467]" captionStart="Figure 21" captionStartId="35.[115,190,1833,1857]" captionTargetBox="[131,1504,192,1811]" captionTargetId="figure-13@35.[131,1504,192,1811]" captionTargetPageId="35" captionText="Figure 21. Propeamussium (Parvamussium) mistensis n. sp., paired color images of molds in matrix and corresponding ammonium chloride-coated casts. A. Left valve exterior, holotype USNM 561849, length 8 mm. B. Left valve exterior, paratype USNM 561856, length 8.1 mm. C. Right valve exterior, paratype USNM 561850, length 5 mm. D. Right valve exterior on top of left valve, paratype USNM 561854, length 7 mm. E. Left valve interior, Paratype USNM 561852, length 6 mm." figureDoi="http://doi.org/10.5281/zenodo.10913529" httpUri="https://zenodo.org/record/10913529/files/figure.png" pageId="37" pageNumber="36">Fig. 21</figureCitation>
), provide a true indication of negative and positive but fail to capture the natural appearance of specimens as they appear in the field and in museum collections. The diagnosis and description of
<taxonomicName id="8D904D48FFDD2919FD07FD85ED24BF52" authorityName="Hickman" authorityYear="2023" box="[664,800,545,573]" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" kingdom="Animalia" order="Pectinida" pageId="37" pageNumber="36" phylum="Mollusca" rank="species" species="mistensis">
<emphasis id="78E4EAD9FFDD2919FD07FD85ED24BF52" box="[664,800,545,573]" italics="true" pageId="37" pageNumber="36">P. mistensis</emphasis>
</taxonomicName>
is based microscopic study and drawings combined with data extracted from three different image sets described below.
</paragraph>
<paragraph id="4A2F36CBFFDD2919FF0FFD14EF54BE5B" blockId="37.[112,800,195,1312]" pageId="37" pageNumber="36">In spite of the difficulties of specimen illustration, this is the best-preserved fossil propeamussiid species from the Northeastern Pacific, prompting the following note on the image sets.</paragraph>
<paragraph id="4A2F36CBFFDD2919FF0FFC99EF7EB992" blockId="37.[112,800,195,1312]" pageId="37" pageNumber="36">
<emphasis id="78E4EAD9FFDD2919FF0FFC99EF1BBE38" bold="true" box="[144,287,829,855]" pageId="37" pageNumber="36">Image sets</emphasis>
—Three kinds of images were assembled for the
<typeStatus id="952B8869FFDD2919FF5CFCC4EF2BBE15" box="[195,303,864,890]" pageId="37" pageNumber="36" type="holotype">holotype</typeStatus>
and each of
<specimenCount id="5C96FD42FFDD2919FE20FCC4EC7FBE15" box="[447,635,864,890]" count="8" pageId="37" pageNumber="36" type="generic" typeStatus="paratype">eight paratypes</specimenCount>
from the Mist locality.
<figureCitation id="D2AB2A4EFFDD2919FF4CFC27EF7CBEF2" box="[211,376,899,925]" captionStart="Figure 21" captionStartId="35.[115,190,1833,1857]" captionTargetBox="[131,1504,192,1811]" captionTargetId="figure-13@35.[131,1504,192,1811]" captionTargetPageId="35" captionText="Figure 21. Propeamussium (Parvamussium) mistensis n. sp., paired color images of molds in matrix and corresponding ammonium chloride-coated casts. A. Left valve exterior, holotype USNM 561849, length 8 mm. B. Left valve exterior, paratype USNM 561856, length 8.1 mm. C. Right valve exterior, paratype USNM 561850, length 5 mm. D. Right valve exterior on top of left valve, paratype USNM 561854, length 7 mm. E. Left valve interior, Paratype USNM 561852, length 6 mm." figureDoi="http://doi.org/10.5281/zenodo.10913529" httpUri="https://zenodo.org/record/10913529/files/figure.png" pageId="37" pageNumber="36">Figure 21AE</figureCitation>
illustrates the
<typeStatus id="952B8869FFDD2919FDB5FC27EC9CBEF2" box="[554,664,899,925]" pageId="37" pageNumber="36" type="holotype">holotype</typeStatus>
and four of the
<typeStatus id="952B8869FFDD2919FF03FC03EF1CBEAE" box="[156,280,935,961]" pageId="37" pageNumber="36" type="paratype">paratypes</typeStatus>
, each represented by a color photograph of the actual specimen paired with a black and white photograph of a latex cast coated with ammonium chloride.
<figureCitation id="D2AB2A4EFFDD2919FFEFFBB4EF13B945" box="[112,279,1040,1066]" captionStart="Figure 22" captionStartId="36.[114,189,1847,1871]" captionTargetBox="[144,764,195,1827]" captionTargetId="figure-438@36.[144,764,195,1827]" captionTargetPageId="36" captionText="Figure 22. Propeamussium (Parvamussium) mistensis n. sp., uncoated latex casts of three paratypes. A. Left valve exterior, USNM 561853. B. Left valve exterior, USNM 561856. Right valve exterior, USNM 561850. Scale bars=3 mm." figureDoi="http://doi.org/10.5281/zenodo.10913533" httpUri="https://zenodo.org/record/10913533/files/figure.png" pageId="37" pageNumber="36">Figure 22AC</figureCitation>
illustrates
<specimenCount id="5C96FD42FFDD2919FE02FBB4EC64B945" box="[413,608,1040,1066]" count="3" pageId="37" pageNumber="36" type="generic" typeStatus="paratype">three paratypes</specimenCount>
represented by photographs of uncoated latex casts. In some instances, photographs of actual specimens under different lighting reveal details useful for interpretation. In addition,
<figureCitation id="D2AB2A4EFFDD2919FFEFFB38EEFDB9D8" box="[112,249,1180,1207]" captionStart="Figure 23" captionStartId="37.[112,187,1860,1884]" captionTargetBox="[112,1521,1352,1834]" captionTargetId="figure-546@37.[112,1521,1352,1834]" captionTargetPageId="37" captionText="Figure 23. Propeamussium (Parvamussium) mistensis n. sp., reconstruction of major features of shell morphology based on all material examined. A. Left valve exterior. B. Left valve interior. C. Right valve exterior. D. Right valve interior. Dashed lines in C, D indicate extent of the larger left valve and uncalcified flexible apron of right valve." figureDoi="http://doi.org/10.5281/zenodo.10913535" httpUri="https://zenodo.org/record/10913535/files/figure.png" pageId="37" pageNumber="36">Fig. 23AD</figureCitation>
provides rendered drawings as a summary guide for interpreting future collections of fragments and imperfect specimens.
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFDD2919FF0FFAA2EBB7BDB2" box="[144,1459,195,1312]" pageId="37" pageNumber="36" type="etymology">
<paragraph id="4A2F36CBFFDD2919FF0FFAA2EBB7BDB2" blockId="37.[112,800,195,1312]" box="[144,1459,195,1312]" lastBlockId="37.[832,1521,195,1161]" pageId="37" pageNumber="36">
<emphasis id="78E4EAD9FFDD2919FF0FFAA2EF24B84F" bold="true" box="[144,288,1286,1312]" pageId="37" pageNumber="36">Etymology</emphasis>
—named for the classic crinoid locality, cold seep, and river bluff outcrops at Mist,
<collectingRegion id="8854F829FFDD2919FACEFF67EBABBDB2" box="[1361,1455,195,221]" country="United States of America" name="Oregon" pageId="37" pageNumber="36">Oregon</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFDD2919FCFFFF42EDC4B9E6" pageId="37" pageNumber="36" type="materials_examined">
<paragraph id="4A2F36CBFFDD2919FCFFFF42EB60BCBD" blockId="37.[832,1521,195,1161]" pageId="37" pageNumber="36">
<emphasis id="78E4EAD9FFDD2919FCFFFF42EA62BC6F" bold="true" box="[864,1126,230,256]" pageId="37" pageNumber="36">Material examined</emphasis>
—Although the primary
<typeStatus id="952B8869FFDD2919FA35FF43EBEBBC6E" box="[1450,1519,231,257]" pageId="37" pageNumber="36">types</typeStatus>
(collected by Harold Vokes) are all from the carbonate hardground of the main crinoid lagerstätte at Mist, specimens of the propeamussiid occur in float from a higher siliciclastic horizon with abundant spines and tests of two echinoid species. To avoid confusion, a new
<collectionCode id="2C81AE0EFFDD2919FA3BFE31EBF4BCC0" box="[1444,1520,405,431]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="37" pageNumber="36" type="University or college">UCMP</collectionCode>
locality number is not assigned (see below).
</paragraph>
<paragraph id="4A2F36CBFFDD2919FCFFFE7EEDE7BF77" blockId="37.[832,1521,195,1161]" pageId="37" pageNumber="36">
<materialsCitation id="FAF83C96FFDD2919FCFFFE7EEDE7BF77" collectionCode="USNM" pageId="37" pageNumber="36" specimenCode="USNM 561849" specimenCount="1" typeStatus="holotype">
<emphasis id="78E4EAD9FFDD2919FCFFFE7EEDDDBC9B" bold="true" box="[864,985,474,500]" pageId="37" pageNumber="36">
<typeStatus id="952B8869FFDD2919FCFFFE7EEDDDBC9B" box="[864,985,474,500]" pageId="37" pageNumber="36" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCode id="1A369EB0FFDD2919FC6BFE7EEAA3BC9A" box="[1012,1191,474,501]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="37" pageNumber="36" type="Museum">USNM 561849</specimenCode>
, (incomplete) length
<quantity id="8D689B2EFFDD2919FA3AFE7EEBE8BC9A" box="[1445,1516,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="37" pageNumber="36" unit="mm" value="8.0">8 mm</quantity>
, height
<quantity id="8D689B2EFFDD2919FC09FE5AEDDBBF77" box="[918,991,510,536]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="36" unit="mm" value="9.0">9 mm</quantity>
.
</materialsCitation>
</paragraph>
<paragraph id="4A2F36CBFFDD2919FCFFFD84EAEBBF87" blockId="37.[832,1521,195,1161]" pageId="37" pageNumber="36">
<emphasis id="78E4EAD9FFDD2919FCFFFD84EA58BF55" bold="true" box="[864,1116,544,570]" pageId="37" pageNumber="36">
Figured
<typeStatus id="952B8869FFDD2919FC4BFD84EA58BF55" box="[980,1116,544,570]" pageId="37" pageNumber="36" type="paratype">paratypes</typeStatus>
</emphasis>
<collectionCode id="2C81AE0EFFDD2919FBE7FD84EAC3BF55" box="[1144,1223,544,570]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="37" pageNumber="36" type="Museum">USNM</collectionCode>
561850, length
<quantity id="8D689B2EFFDD2919FA01FD84EBEFBF54" box="[1438,1515,544,571]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="36" unit="mm" value="5.0">5 mm</quantity>
, height
<quantity id="8D689B2EFFDD2919FC0BFDE7EDDABF32" box="[916,990,579,606]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="36" unit="mm" value="5.0">5 mm</quantity>
;
<collectionCode id="2C81AE0EFFDD2919FC77FDE7EA31BF32" box="[1000,1077,579,605]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="37" pageNumber="36" type="Museum">USNM</collectionCode>
561852, length
<quantity id="8D689B2EFFDD2919FB63FDE7EB47BF32" box="[1276,1347,579,605]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="37" pageNumber="36" unit="mm" value="6.0">6 mm</quantity>
, height
<quantity id="8D689B2EFFDD2919FA3DFDE7EBEFBF32" box="[1442,1515,579,605]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="37" pageNumber="36" unit="mm" value="6.0">6 mm</quantity>
;
<collectionCode id="2C81AE0EFFDD2919FCDFFDC2ED89BFEF" box="[832,909,614,640]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="37" pageNumber="36" type="Museum">USNM</collectionCode>
561853, length of hingline 3.3 mm;
<collectionCode id="2C81AE0EFFDD2919FAA6FDC2EB82BFEF" box="[1337,1414,614,640]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="37" pageNumber="36" type="Museum">USNM</collectionCode>
561854, length
<quantity id="8D689B2EFFDD2919FC0DFD2DEDDCBFCC" box="[914,984,649,675]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="37" pageNumber="36" unit="mm" value="7.0">7 mm</quantity>
, height
<quantity id="8D689B2EFFDD2919FBACFD2CEA7EBFCC" box="[1075,1146,648,675]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="37" pageNumber="36" unit="mm" value="6.0">6 mm</quantity>
;
<collectionCode id="2C81AE0EFFDD2919FB1CFD2CEAD4BFCD" box="[1155,1232,648,674]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="37" pageNumber="36" type="Museum">USNM</collectionCode>
561856, length 8.1 mm, height 8.3 mm. Measurements are approximate, greatest dimensions are all within
<quantity id="8D689B2EFFDD2919FB1CFD69EAEFBF87" box="[1155,1259,717,744]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="37" pageNumber="36" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">58 mm</quantity>
.
</paragraph>
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<emphasis id="78E4EAD9FFDD2919FCFFFD54EA9EBE65" bold="true" box="[864,1178,752,778]" pageId="37" pageNumber="36">
Unfigured
<materialsCitation id="FAF83C96FFDD2919FB9BFD54EB8BBE64" box="[1028,1423,752,779]" collectionCode="USNM" pageId="37" pageNumber="36" specimenCode="USNM 561851" specimenCount="1" typeStatus="paratype">
<typeStatus id="952B8869FFDD2919FB9BFD54EA9EBE65" box="[1028,1178,752,778]" pageId="37" pageNumber="36" type="paratype">paratypes</typeStatus>
<specimenCode id="1A369EB0FFDD2919FB25FD54EB8DBE64" box="[1210,1417,752,779]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="37" pageNumber="36" type="Museum">USNM 561851</specimenCode>
,
</materialsCitation>
</emphasis>
<materialsCitation id="FAF83C96FFDD2919FA02FD54EDACBE41" collectionCode="USNM" pageId="37" pageNumber="36" specimenCode="USNM 561855" specimenCount="1" typeStatus="paratype">
<specimenCode id="1A369EB0FFDD2919FA02FD54EDA7BE41" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="37" pageNumber="36" type="Museum">USNM 561855</specimenCode>
,
</materialsCitation>
<materialsCitation id="FAF83C96FFDD2919FC31FCB0EA62BE41" box="[942,1126,787,814]" collectionCode="USNM" pageId="37" pageNumber="36" specimenCode="USNM 561857" specimenCount="1" typeStatus="paratype">
<specimenCode id="1A369EB0FFDD2919FC31FCB0EA62BE41" box="[942,1126,787,814]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="37" pageNumber="36" type="Museum">USNM 561857</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="4A2F36CBFFDD2919FCFFFC92EAB8BED6" blockId="37.[832,1521,195,1161]" pageId="37" pageNumber="36">
<emphasis id="78E4EAD9FFDD2919FCFFFC92EB14BE3F" bold="true" box="[864,1296,822,848]" pageId="37" pageNumber="36">
Additional Unfigured
<materialsCitation id="FAF83C96FFDD2919FB0CFC92EABDBED6" collectionCode="UCMP" pageId="37" pageNumber="36" specimenCode="UCMP 110764" specimenCount="1" typeStatus="paratype">
<typeStatus id="952B8869FFDD2919FB0CFC92EB14BE3F" box="[1171,1296,822,848]" pageId="37" pageNumber="36" type="paratype">paratype</typeStatus>
<specimenCode id="1A369EB0FFDD2919FAB3FC92EBE8BE3F" box="[1324,1516,822,848]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="37" pageNumber="36" type="University or college">UCMP 110764</specimenCode>
. This specimen is from a float block with echinoid spines from above the main crinoid layer and is the only specimen preserving shell material
</materialsCitation>
</emphasis>
.
</paragraph>
<paragraph id="4A2F36CBFFDD2919FCFFFC65EDC4B9E6" blockId="37.[832,1521,195,1161]" pageId="37" pageNumber="36">
<emphasis id="78E4EAD9FFDD2919FCFFFC65EA0EBEB4" bold="true" box="[864,1034,961,987]" pageId="37" pageNumber="36">Type locality</emphasis>
<collectionCode id="2C81AE0EFFDD2919FBB9FC65EA6DBEB4" box="[1062,1129,961,987]" country="USA" httpUri="http://grbio.org/cool/3n47-ewde" name="U.S. Geological Survey" pageId="37" pageNumber="36">USGS</collectionCode>
15318=
<collectionCode id="2C81AE0EFFDD2919FB5AFC65EB1FBEB3" box="[1221,1307,961,988]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="37" pageNumber="36" type="University or college">UCMP</collectionCode>
A5018. Numbers refer to the generalized Mist locality, including a prominent bench adjacent to the river bluff that was removed by commercial quarrying of the crinoids. For an excellent description of the stratigraphy at Mist see
<bibRefCitation id="2E014B3AFFDD2919FA19FBE9EDBEB9E6" author="Burns, C. &amp; K. A. Campbell &amp; R. Mooi" pageId="37" pageNumber="36" pagination="210 - 231" refId="ref27894" refString="Burns, C., K. A. Campbell, and R. Mooi. 2005. Exceptional crinoid occurrences and associated carbonates of the Keasey Formation (early Oliogocene) at Mist, Oregon, USA. Palaeogeography, Palaeoclimatology, Palaeoecology 227: 210 - 231." type="journal article" year="2005">Burns et al. (2005)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>