500 lines
59 KiB
XML
500 lines
59 KiB
XML
<document id="675E6382AFD0B568B57272876A5A96B2" ID-DOI="10.5281/zenodo.200811" ID-GBIF-Dataset="76b2912f-8856-4c85-842d-c3312bc2650f" ID-ISSN="1175-5326" ID-Zenodo-Dep="200811" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460250427582" checkinUser="plazi" docAuthor="Mcdonald, Andrew T." docDate="2011" docId="E80287E9B114FFFDCDB95449FB43FDA9" docLanguage="en" docName="zt02783p068.pdf" docOrigin="Zootaxa 2783" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Uteodon aphanoecetes Carpenter & Wilson 2008, comb. nov." docType="treatment" docVersion="7" lastPageNumber="59" masterDocId="143BFF91B110FFF5CD2E537DFFE3FF95" masterDocTitle="The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda)" masterLastPageNumber="68" masterPageNumber="52" pageNumber="56" updateTime="1698242557538" updateUser="plazi">
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<mods:mods id="EBEF76F70C8A90E54250071438DC792B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BBC7C4AC5B310177778FDC52448757ED">
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<mods:title id="739E4A7467FBDBB92CA48A9F2F0C3DD4">The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda)</mods:title>
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<mods:name id="542DC3421F3F78B52ED0B9783CABF071" type="personal">
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<mods:namePart id="3406AAD0F72D34C7BF4089B26B3941B3">Mcdonald, Andrew T.</mods:namePart>
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<mods:typeOfResource id="07E89E6DF228181EC233F738B84633E5">text</mods:typeOfResource>
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<mods:title id="666A81FFFD5805399C1C955A364C384D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0067E7887DC14DE7426CAAFC3B544F6C">
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<mods:date id="0FCEC7FCD76074ADA04246ABA639B508">2011</mods:date>
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<mods:detail id="2B42E1BAC302D28A5C06820995E7C153" type="volume">
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<mods:number id="EF9120390C018DFFF33E6251F3366695">2783</mods:number>
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<mods:classification id="9C5EC6D9862ED59169805721A13D982A">journal article</mods:classification>
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<mods:identifier id="9CA6B658FC2FAFDEEA52F758A6B8084E" type="DOI">10.5281/zenodo.200811</mods:identifier>
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<mods:identifier id="262801D4A275495065E7BBE31F4E9D6F" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="6FACAA3012D09B0B2A069915CB0DE298" type="Zenodo-Dep">200811</mods:identifier>
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<treatment id="E80287E9B114FFFDCDB95449FB43FDA9" ID-DOI="http://doi.org/10.5281/zenodo.6185213" ID-GBIF-Taxon="119391381" ID-Zenodo-Dep="6185213" LSID="urn:lsid:plazi:treatment:E80287E9B114FFFDCDB95449FB43FDA9" httpUri="http://treatment.plazi.org/id/E80287E9B114FFFDCDB95449FB43FDA9" lastPageId="8" lastPageNumber="59" pageId="4" pageNumber="56">
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<subSubSection id="28B16574B114FFF1CDB95449FCD3F805" pageId="4" pageNumber="56" type="nomenclature">
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<paragraph id="601436FFB114FFF1CDB95449FC6EF8D8" blockId="4.[151,909,1843,1870]" box="[151,909,1843,1870]" pageId="4" pageNumber="56">
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<heading id="3B5C8193B114FFF1CDB95449FC6EF8D8" bold="true" box="[151,909,1843,1870]" fontSize="11" level="1" pageId="4" pageNumber="56" reason="1">
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<emphasis id="52DFEAEDB114FFF1CDB95449FC6EF8D8" bold="true" box="[151,909,1843,1870]" pageId="4" pageNumber="56">
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<taxonomicName id="A7AB4D7CB114FFF1CDB95449FD19F8D8" authority="Carpenter & Wilson, 2008" authorityName="Carpenter & Wilson" authorityYear="2008" box="[151,762,1843,1870]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="aphanoecetes" status="comb. nov.">
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<emphasis id="52DFEAEDB114FFF1CDB95449FE4BF8DB" bold="true" box="[151,424,1844,1870]" italics="true" pageId="4" pageNumber="56">Uteodon aphanoecetes</emphasis>
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<bibRefCitation id="043A4B0EB114FFF1CC81544EFD19F8D8" author="Carpenter" box="[431,762,1843,1869]" pageId="4" pageNumber="56" refString="Carpenter, K. & Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter & Wilson, 2008</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicNameLabel id="49EC5796B114FFF1CE29544EFC6EF8D8" box="[775,909,1843,1869]" pageId="4" pageNumber="56">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="601436FFB114FFF1CDB95406FCD3F805" blockId="4.[151,816,1914,1936]" box="[151,816,1914,1936]" pageId="4" pageNumber="56">
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2008
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<taxonomicName id="A7AB4D7CB114FFF1CDC95406FD03F805" authority="Carpenter & Wilson" authorityName="Carpenter & Wilson" box="[231,736,1914,1936]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="aphanoecetes">
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<emphasis id="52DFEAEDB114FFF1CDC95406FDE8F805" box="[231,523,1914,1936]" italics="true" pageId="4" pageNumber="56">Camptosaurus aphanoecetes</emphasis>
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Carpenter & Wilson
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</taxonomicName>
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, p. 232.
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</paragraph>
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</subSubSection>
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<subSubSection id="28B16574B114FFF3CDB954C0FABCF831" lastPageId="6" lastPageNumber="58" pageId="4" pageNumber="56" type="materials_examined">
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<paragraph id="601436FFB114FFF1CDB954C0FD07F840" blockId="4.[151,740,1981,2005]" box="[151,740,1981,2005]" pageId="4" pageNumber="56">
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<emphasis id="52DFEAEDB114FFF1CDB954C0FEE8F840" bold="true" box="[151,267,1981,2005]" pageId="4" pageNumber="56">
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<typeStatus id="BF10885DB114FFF1CDB954C0FEE5F840" box="[151,262,1981,2005]" pageId="4" pageNumber="56" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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CM 11337, nearly complete postcranium.
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</paragraph>
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<caption id="34D46677B115FFF0CDB954FEFB54F860" httpUri="https://zenodo.org/record/200814/files/figure.png" pageId="5" pageNumber="57" targetBox="[218,1348,194,1877]" targetPageId="5">
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<paragraph id="601436FFB115FFF0CDB954FEFB54F860" blockId="5.[151,1436,1923,2037]" pageId="5" pageNumber="57">
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<emphasis id="52DFEAEDB115FFF0CDB954FEFEF1F80D" bold="true" box="[151,274,1923,1946]" pageId="5" pageNumber="57">FIGURE 3.</emphasis>
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(
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<emphasis id="52DFEAEDB115FFF0CC0E54FEFED1F80D" bold="true" box="[288,306,1923,1944]" pageId="5" pageNumber="57">A</emphasis>
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) USNM 4282, left scapula of
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<taxonomicName id="A7AB4D7CB115FFF0CF4C54F9FCD8F80C" box="[610,827,1923,1945]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="5" pageNumber="57" phylum="Chordata" rank="species" species="dispar">
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<emphasis id="52DFEAEDB115FFF0CF4C54F9FCD8F80C" box="[610,827,1923,1945]" italics="true" pageId="5" pageNumber="57">Camptosaurus dispar</emphasis>
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</taxonomicName>
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in lateral view; (
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<emphasis id="52DFEAEDB115FFF0CEC854F9FC14F80C" bold="true" box="[998,1015,1924,1945]" pageId="5" pageNumber="57">B</emphasis>
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) CM 11337, cranial end of the right scapula of
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<taxonomicName id="A7AB4D7CB115FFF0CDCF54DFFE2EF82D" box="[225,461,1954,1976]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="5" pageNumber="57" phylum="Chordata" rank="species" species="aphanoecetes">
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<emphasis id="52DFEAEDB115FFF0CDCF54DFFE2EF82D" box="[225,461,1954,1976]" italics="true" pageId="5" pageNumber="57">Uteodon aphanoecetes</emphasis>
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</taxonomicName>
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in lateral view; (
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<emphasis id="52DFEAEDB115FFF0CFA254DCFD7DF82D" bold="true" box="[652,670,1953,1976]" pageId="5" pageNumber="57">C</emphasis>
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) CM 11337, left scapula of
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<taxonomicName id="A7AB4D7CB115FFF0CEF854DFFB21F82D" box="[982,1218,1954,1976]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="5" pageNumber="57" phylum="Chordata" rank="species" species="aphanoecetes">
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<emphasis id="52DFEAEDB115FFF0CEF854DFFB21F82D" box="[982,1218,1954,1976]" italics="true" pageId="5" pageNumber="57">Uteodon aphanoecetes</emphasis>
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</taxonomicName>
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in lateral view; (
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<emphasis id="52DFEAEDB115FFF0C8AF54DFFA70F822" bold="true" box="[1409,1427,1954,1975]" pageId="5" pageNumber="57">D</emphasis>
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) OXFUM J.3303, right scapula of
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<emphasis id="52DFEAEDB115FFF0CCC054BFFD2FF842" box="[494,716,1985,2007]" italics="true" pageId="5" pageNumber="57">
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Cumnoria
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<taxonomicName id="A7AB4D7CB115FFF0CF7254BCFD2FF842" box="[604,716,1985,2007]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="5" pageNumber="57" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
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</emphasis>
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in lateral view. Abbreviations: cap, convex cranial margin of the acromion process; tap, triangular acromion process. Scale bar in
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<emphasis id="52DFEAEDB115FFF0CFD5549DFCEEF860" bold="true" box="[763,781,2016,2037]" pageId="5" pageNumber="57">A</emphasis>
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equals 10 cm; scale bar in
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<emphasis id="52DFEAEDB115FFF0C90E549DFBD2F860" bold="true" box="[1056,1073,2016,2037]" pageId="5" pageNumber="57">B</emphasis>
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equals 5 cm.
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</paragraph>
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</caption>
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<caption id="34D46677B116FFF3CDB951ADFBD1FCD7" httpUri="https://zenodo.org/record/200815/files/figure.png" pageId="6" pageNumber="58" targetBox="[151,1436,193,698]" targetPageId="6">
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<paragraph id="601436FFB116FFF3CDB951ADFBD1FCD7" blockId="6.[151,1436,720,834]" pageId="6" pageNumber="58">
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<emphasis id="52DFEAEDB116FFF3CDB951ADFEF7FD70" bold="true" box="[151,276,720,743]" pageId="6" pageNumber="58">FIGURE 4.</emphasis>
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Ilia of
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<taxonomicName id="A7AB4D7CB116FFF3CC4D51ACFDA3FD73" box="[355,576,720,742]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="dispar">
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<emphasis id="52DFEAEDB116FFF3CC4D51ACFDA3FD73" box="[355,576,720,742]" italics="true" pageId="6" pageNumber="58">Camptosaurus dispar</emphasis>
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</taxonomicName>
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in lateral view. (
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<emphasis id="52DFEAEDB116FFF3CFDF51ADFCE0FD70" bold="true" box="[753,771,720,741]" pageId="6" pageNumber="58">A</emphasis>
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) right ilium of YPM 1877 (holotype) from Quarry 13; (
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<emphasis id="52DFEAEDB116FFF3C86551ACFABFFD73" bold="true" box="[1355,1372,721,742]" pageId="6" pageNumber="58">B</emphasis>
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) right ilium of YPM 8622 from Quarry 13; (
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<emphasis id="52DFEAEDB116FFF3CF065193FDD9FC90" bold="true" box="[552,570,750,773]" pageId="6" pageNumber="58">C</emphasis>
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) left ilium of USNM 5473 from Quarry 13; (
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<emphasis id="52DFEAEDB116FFF3C9345192FBCFFC91" bold="true" box="[1050,1068,751,772]" pageId="6" pageNumber="58">D</emphasis>
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) left ilium of YPM PU14553 from Cleveland-Lloyd Dinosaur Quarry. Abbreviations: mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process. Images in
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<emphasis id="52DFEAEDB116FFF3CCDF5051FDE0FCD4" bold="true" box="[497,515,812,833]" pageId="6" pageNumber="58">A</emphasis>
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,
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<emphasis id="52DFEAEDB116FFF3CF3E5051FDC2FCD4" bold="true" box="[528,545,812,833]" pageId="6" pageNumber="58">B</emphasis>
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, and
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<emphasis id="52DFEAEDB116FFF3CF775051FD88FCD4" bold="true" box="[601,619,812,833]" pageId="6" pageNumber="58">D</emphasis>
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are copyright YPM. Scale bar equals 10 cm.
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</paragraph>
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</caption>
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<caption id="34D46677B116FFF3CDB9564FFC2DFAFD" httpUri="https://zenodo.org/record/200816/files/figure.png" pageId="6" pageNumber="58" targetBox="[483,1104,894,1304]" targetPageId="6">
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<paragraph id="601436FFB116FFF3CDB9564FFC2DFAFD" blockId="6.[151,1436,1330,1384]" pageId="6" pageNumber="58">
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<emphasis id="52DFEAEDB116FFF3CDB9564FFEF6FADD" bold="true" box="[151,277,1330,1353]" pageId="6" pageNumber="58">FIGURE 5.</emphasis>
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YPM 7334, right pubis of
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<taxonomicName id="A7AB4D7CB116FFF3CF1D5649FCECFADC" box="[563,783,1331,1353]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="dispar">
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<emphasis id="52DFEAEDB116FFF3CF1D5649FCECFADC" box="[563,783,1331,1353]" italics="true" pageId="6" pageNumber="58">Camptosaurus dispar</emphasis>
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</taxonomicName>
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from Quarry 13 in lateral view. Abbreviations: pop, postpubic process; prp, prepubic process. Image is copyright YPM. Scale bar equals 10 cm.
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</paragraph>
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</caption>
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<paragraph id="601436FFB116FFF3CDEB56E8FC67FA39" blockId="6.[151,1436,1428,2029]" box="[197,900,1428,1453]" pageId="6" pageNumber="58">
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<emphasis id="52DFEAEDB116FFF3CDEB56E8FEA7FA38" bold="true" box="[197,324,1429,1453]" pageId="6" pageNumber="58">
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<typeStatus id="BF10885DB116FFF3CDEB56E8FEDCFA38" box="[197,319,1429,1453]" pageId="6" pageNumber="58" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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CM 15780, 41689, 79050;
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<collectionCode id="06BAAE3AB116FFF3CF5456E9FD23FA39" box="[634,704,1428,1452]" httpUri="http://grbio.org/cool/h4n0-drs5" name="Dinosaur National Monument" pageId="6" pageNumber="58">DINO</collectionCode>
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556, 1030, 1032.
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</paragraph>
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<paragraph id="601436FFB116FFF3CDEB56C5FA95FA45" blockId="6.[151,1436,1428,2029]" box="[197,1398,1464,1489]" pageId="6" pageNumber="58">
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<emphasis id="52DFEAEDB116FFF3CDEB56C5FE40FA44" bold="true" box="[197,419,1464,1489]" pageId="6" pageNumber="58">Referred material.</emphasis>
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Disarticulated elements held in CM and
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<collectionCode id="06BAAE3AB116FFF3CE4056C5FC57FA45" box="[878,948,1464,1488]" httpUri="http://grbio.org/cool/h4n0-drs5" name="Dinosaur National Monument" pageId="6" pageNumber="58">DINO</collectionCode>
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||
collections (
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<bibRefCitation id="043A4B0EB116FFF3C96D56C5FA88FA45" author="Carpenter" box="[1091,1387,1464,1488]" pageId="6" pageNumber="58" refString="Carpenter, K. & Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter & Wilson 2008</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="601436FFB116FFF3CDEB56A1FF00F9F5" blockId="6.[151,1436,1428,2029]" pageId="6" pageNumber="58">
|
||
<emphasis id="52DFEAEDB116FFF3CDEB56A1FE48FA60" bold="true" box="[197,427,1500,1525]" pageId="6" pageNumber="58">Generic etymology.</emphasis>
|
||
<taxonomicName id="A7AB4D7CB116FFF3CC9F56A3FE39FA60" box="[433,474,1502,1525]" class="Calcarea" family="Grantiidae" genus="Ute" kingdom="Animalia" order="Leucosolenida" pageId="6" pageNumber="58" phylum="Porifera" rank="genus">
|
||
<emphasis id="52DFEAEDB116FFF3CC9F56A3FE39FA60" box="[433,474,1502,1525]" italics="true" pageId="6" pageNumber="58">Ute</emphasis>
|
||
</taxonomicName>
|
||
(pronounced “yewt”) is the name of a Native American people who inhabit northeastern Utah, in the vicinity of Dinosaur National Monument.
|
||
<emphasis id="52DFEAEDB116FFF3CE68557DFC66F98D" box="[838,901,1536,1560]" italics="true" pageId="6" pageNumber="58">Odon</emphasis>
|
||
comes from the transliterated Greek meaning “tooth”, and is a common root in the generic names of basal iguanodonts. The gender of the generic name is masculine.
|
||
</paragraph>
|
||
<paragraph id="601436FFB116FFF3CDEB5511FBAAF93D" blockId="6.[151,1436,1428,2029]" pageId="6" pageNumber="58">
|
||
<emphasis id="52DFEAEDB116FFF3CDEB5511FE25F910" bold="true" box="[197,454,1644,1669]" pageId="6" pageNumber="58">Locality and horizon.</emphasis>
|
||
Carnegie Quarry at Dinosaur National Monument, Uintah County, Utah; Brushy Basin Member of the Morrison Formation, lower–middle Tithonian (
|
||
<bibRefCitation id="043A4B0EB116FFF3CE7D55EDFBDFF93D" author="Kowallis" box="[851,1084,1680,1704]" pageId="6" pageNumber="58" refString="Kowallis, B. J., Christiansen, E. H., Deino, A. L., Peterson, F., Turner, C. E., Kunk, M. J. & Obradovich, J. D. (1998) The age of the Morrison Formation. Modern Geology, 22, 235 - 260." type="journal article" year="1998">
|
||
Kowallis
|
||
<emphasis id="52DFEAEDB116FFF3CEEF55ECFC12F93D" box="[961,1009,1680,1704]" italics="true" pageId="6" pageNumber="58">et al</emphasis>
|
||
. 1998
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="601436FFB116FFF3CDEB55C9FCF3F958" blockId="6.[151,1436,1428,2029]" box="[197,784,1716,1741]" pageId="6" pageNumber="58">
|
||
<emphasis id="52DFEAEDB116FFF3CDEB55C9FE43F958" bold="true" box="[197,416,1716,1741]" pageId="6" pageNumber="58">Generic diagnosis.</emphasis>
|
||
<collectingCountry id="18BC766FB116FFF3CC8955C8FE25F958" box="[423,454,1717,1741]" name="American Samoa" pageId="6" pageNumber="58">As</collectingCountry>
|
||
for the species by monotypy.
|
||
</paragraph>
|
||
<paragraph id="601436FFB116FFF3CDEB55A5FABCF831" blockId="6.[151,1436,1428,2029]" pageId="6" pageNumber="58">
|
||
<emphasis id="52DFEAEDB116FFF3CDEB55A5FE42F964" bold="true" box="[197,417,1752,1777]" pageId="6" pageNumber="58">Specific diagnosis.</emphasis>
|
||
Basal styracosternan distinguished by a single autapomorphy: occipital condyle projects farther ventrally than the basal tubera (
|
||
<figureCitation id="F8902A7AB116FFF3CF605581FD70F881" box="[590,659,1788,1812]" captionStart="FIGURE 7" captionStartId="8.[151,250,1622,1645]" captionTargetBox="[472,1114,625,1600]" captionTargetId="figure@8.[472,1115,625,1601]" captionTargetPageId="8" captionText="FIGURE 7. Partial braincases of Camptosaurus dispar, Uteodon aphanoecetes, and Cumnoria prestwichii. (A) YPM 1880 in left lateral view (image copyright YPM); (B) CM 15780 in right lateral view (image courtesy of M. Lamanna); (C) OXFUM J. 3303 in right lateral view. Abbreviations: bt, basal tubera; oc, occipital condyle. Scale bar equals 5 cm." httpUri="https://zenodo.org/record/200818/files/figure.png" pageId="6" pageNumber="58">Fig. 7</figureCitation>
|
||
). Also distinguished from all other iguanodontians except
|
||
<emphasis id="52DFEAEDB116FFF3C8045580FEF0F8AD" italics="true" pageId="6" pageNumber="58">
|
||
Cumnoria
|
||
<taxonomicName id="A7AB4D7CB116FFF3CDB9545DFEF0F8AD" box="[151,275,1824,1848]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
|
||
</emphasis>
|
||
by the following unique combination of characters: scapula with acromion process that is convex along its cranial margin (
|
||
<figureCitation id="F8902A7AB116FFF3CC475439FE4FF8C9" box="[361,428,1860,1884]" captionStart="FIGURE 3" captionStartId="5.[151,250,1923,1946]" captionTargetBox="[218,1348,194,1877]" captionTargetId="figure@5.[213,1359,193,1878]" captionTargetPageId="5" captionText="FIGURE 3. (A) USNM 4282, left scapula of Camptosaurus dispar in lateral view; (B) CM 11337, cranial end of the right scapula of Uteodon aphanoecetes in lateral view; (C) CM 11337, left scapula of Uteodon aphanoecetes in lateral view; (D) OXFUM J. 3303, right scapula of Cumnoria prestwichii in lateral view. Abbreviations: cap, convex cranial margin of the acromion process; tap, triangular acromion process. Scale bar in A equals 10 cm; scale bar in B equals 5 cm." httpUri="https://zenodo.org/record/200814/files/figure.png" pageId="6" pageNumber="58">Fig. 3</figureCitation>
|
||
B), convex dorsal margin of scapula (
|
||
<figureCitation id="F8902A7AB116FFF3CE7E5439FC71F8C9" box="[848,914,1860,1884]" captionStart="FIGURE 3" captionStartId="5.[151,250,1923,1946]" captionTargetBox="[218,1348,194,1877]" captionTargetId="figure@5.[213,1359,193,1878]" captionTargetPageId="5" captionText="FIGURE 3. (A) USNM 4282, left scapula of Camptosaurus dispar in lateral view; (B) CM 11337, cranial end of the right scapula of Uteodon aphanoecetes in lateral view; (C) CM 11337, left scapula of Uteodon aphanoecetes in lateral view; (D) OXFUM J. 3303, right scapula of Cumnoria prestwichii in lateral view. Abbreviations: cap, convex cranial margin of the acromion process; tap, triangular acromion process. Scale bar in A equals 10 cm; scale bar in B equals 5 cm." httpUri="https://zenodo.org/record/200814/files/figure.png" pageId="6" pageNumber="58">Fig. 3</figureCitation>
|
||
C), convex dorsal margin of ilium, dorsal margin of ilium thickens mediolaterally towards
|
||
<emphasis id="52DFEAEDB116FFF3CFB65414FD4CF815" box="[664,687,1897,1920]" italics="true" pageId="6" pageNumber="58">M</emphasis>
|
||
.
|
||
<emphasis id="52DFEAEDB116FFF3CF915415FCA0F815" box="[703,835,1896,1920]" italics="true" pageId="6" pageNumber="58">iliocaudalis</emphasis>
|
||
platform, postacetabular process of ilium tapers to a point with break in slope along dorsal margin (
|
||
<figureCitation id="F8902A7AB116FFF3CF8D54F1FD06F831" box="[675,741,1932,1956]" captionStart="FIGURE 6" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1424,927,1709]" captionTargetId="figure@7.[151,1436,913,1723]" captionTargetPageId="7" captionText="FIGURE 6. Left ilium of CM 11337 (holotype of Uteodon aphanoecetes) in (A) ventrolateral view showing the convex dorsal margin, (B) lateral view, (C), caudal view. Sacrum and ilia of CM 11337 in (D) dorsal view. Right ilium of CM 11337 in (E) lateral view. Right ilium of OXFUM J. 3303 (holotype of Cumnoria prestwichii) in (F) lateral view. Abbreviations: dmi, dorsal margin of ilium; mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process." httpUri="https://zenodo.org/record/200817/files/figure.png" pageId="6" pageNumber="58">Fig. 6</figureCitation>
|
||
A), distal end of ischium forms cranially expanded boot.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="28B16574B116FFFDCDEB54CCFB43FDA9" lastPageId="8" lastPageNumber="60" pageId="6" pageNumber="58" type="discussion">
|
||
<paragraph id="601436FFB116FFF2CDEB54CCFD29FE5A" blockId="6.[151,1436,1428,2029]" lastBlockId="7.[151,1437,151,860]" lastPageId="7" lastPageNumber="59" pageId="6" pageNumber="58">
|
||
<emphasis id="52DFEAEDB116FFF3CDEB54CCFEDAF85C" bold="true" box="[197,313,1969,1993]" pageId="6" pageNumber="58">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="BF10885DB116FFF3CC5054CDFE03F85D" box="[382,480,1968,1992]" pageId="6" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="A7AB4D7CB116FFF3CF3C54CDFCF4F85D" box="[530,791,1968,1992]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="52DFEAEDB116FFF3CF3C54CDFCF4F85D" box="[530,791,1968,1992]" italics="true" pageId="6" pageNumber="58">Uteodon aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
, CM 11337, was originally referred to
|
||
<taxonomicName id="A7AB4D7CB116FFF3C9D754CCFA7FF85D" box="[1273,1436,1969,1992]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="58" phylum="Chordata" rank="genus">
|
||
<emphasis id="52DFEAEDB116FFF3C9D754CCFA7FF85D" box="[1273,1436,1969,1992]" italics="true" pageId="6" pageNumber="58">Camptosaurus</emphasis>
|
||
</taxonomicName>
|
||
“
|
||
<taxonomicName id="A7AB4D7CB116FFF3CD8D54A8FF11F878" box="[163,242,2005,2029]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="medius">
|
||
<emphasis id="52DFEAEDB116FFF3CD8D54A8FF11F878" box="[163,242,2005,2029]" italics="true" pageId="6" pageNumber="58">medius</emphasis>
|
||
</taxonomicName>
|
||
” (=
|
||
<taxonomicName id="A7AB4D7CB116FFF3CC0654A8FE72F878" box="[296,401,2005,2029]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="52DFEAEDB116FFF3CC0654A8FED8F879" box="[296,315,2005,2028]" italics="true" pageId="6" pageNumber="58">C</emphasis>
|
||
.
|
||
<emphasis id="52DFEAEDB116FFF3CC6454A8FE72F878" box="[330,401,2005,2029]" italics="true" pageId="6" pageNumber="58">dispar</emphasis>
|
||
</taxonomicName>
|
||
) by
|
||
<bibRefCitation id="043A4B0EB116FFF3CCE654A9FD9FF879" author="Gilmore" box="[456,636,2004,2028]" pageId="6" pageNumber="58" refString="Gilmore, C. W. (1925) Osteology of ornithopodous dinosaurs from the Dinosaur National Monument, Utah. Memoirs of the Carnegie Museum, 10, 385 - 411." type="journal article" year="1925">Gilmore (1925)</bibRefCitation>
|
||
.
|
||
<bibRefCitation id="043A4B0EB116FFF3CFA954A9FC5DF879" author="Carpenter" box="[647,958,2004,2028]" pageId="6" pageNumber="58" refString="Carpenter, K. & Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter & Wilson (2008)</bibRefCitation>
|
||
reassessed the specimen and found that it differed sufficiently from specimens of
|
||
<taxonomicName id="A7AB4D7CB117FFF2CF7953E5FD23FF25" box="[599,704,152,176]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="52DFEAEDB117FFF2CF7953E5FD89FF3A" box="[599,618,152,175]" italics="true" pageId="7" pageNumber="59">C</emphasis>
|
||
.
|
||
<emphasis id="52DFEAEDB117FFF2CF5753E5FD23FF25" box="[633,704,152,176]" italics="true" pageId="7" pageNumber="59">dispar</emphasis>
|
||
</taxonomicName>
|
||
to warrant the creation of the new species
|
||
<taxonomicName id="A7AB4D7CB117FFF2C98653E5FA82FF25" box="[1192,1377,152,176]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="52DFEAEDB117FFF2C98653E5FB58FF3A" box="[1192,1211,152,175]" italics="true" pageId="7" pageNumber="59">C</emphasis>
|
||
.
|
||
<emphasis id="52DFEAEDB117FFF2C9E753E5FA82FF25" box="[1225,1377,152,176]" italics="true" pageId="7" pageNumber="59">aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
. The new phylogenetic analysis supports the distinction of
|
||
<taxonomicName id="A7AB4D7CB117FFF2CFC053C1FC0EFF41" box="[750,1005,188,212]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="52DFEAEDB117FFF2CFC053C1FC0EFF41" box="[750,1005,188,212]" italics="true" pageId="7" pageNumber="59">Uteodon aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="A7AB4D7CB117FFF2C91F53C0FB7BFF41" box="[1073,1176,188,212]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="52DFEAEDB117FFF2C91F53C0FBA7FF41" box="[1073,1092,189,212]" italics="true" pageId="7" pageNumber="59">C</emphasis>
|
||
.
|
||
<emphasis id="52DFEAEDB117FFF2C97C53C1FB7BFF41" box="[1106,1176,188,212]" italics="true" pageId="7" pageNumber="59">dispar</emphasis>
|
||
</taxonomicName>
|
||
, but does not recover a monophyletic
|
||
<taxonomicName id="A7AB4D7CB117FFF2CC12539DFE3CFF62" box="[316,479,224,247]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="genus">
|
||
<emphasis id="52DFEAEDB117FFF2CC12539DFE3CFF62" box="[316,479,224,247]" italics="true" pageId="7" pageNumber="59">Camptosaurus</emphasis>
|
||
</taxonomicName>
|
||
as conceived by
|
||
<bibRefCitation id="043A4B0EB117FFF2CF9E53A2FC10FF62" author="Carpenter" box="[688,1011,223,247]" pageId="7" pageNumber="59" refString="Carpenter, K. & Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter & Wilson (2008)</bibRefCitation>
|
||
. Instead,
|
||
<taxonomicName id="A7AB4D7CB117FFF2C94A539DFA8BFF6D" box="[1124,1384,224,248]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="52DFEAEDB117FFF2C94A539DFA8BFF6D" box="[1124,1384,224,248]" italics="true" pageId="7" pageNumber="59">Uteodon aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="52DFEAEDB117FFF2CDB95278FE68FE89" box="[151,395,260,284]" italics="true" pageId="7" pageNumber="59">
|
||
Cumnoria
|
||
<taxonomicName id="A7AB4D7CB117FFF2CC3E5279FE68FE89" box="[272,395,260,284]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
|
||
</emphasis>
|
||
(see below) are more derived than
|
||
<taxonomicName id="A7AB4D7CB117FFF2CE355278FC60FE89" box="[795,899,260,284]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="52DFEAEDB117FFF2CE355278FCCDFE89" box="[795,814,261,284]" italics="true" pageId="7" pageNumber="59">C</emphasis>
|
||
.
|
||
<emphasis id="52DFEAEDB117FFF2CE125279FC60FE89" box="[828,899,260,284]" italics="true" pageId="7" pageNumber="59">dispar</emphasis>
|
||
</taxonomicName>
|
||
in the reduced consensus tree and are therefore the most basal members of Styracosterna, which is defined as those species more closely related to
|
||
<taxonomicName id="A7AB4D7CB117FFF2C8285255FEC4FEF1" class="Reptilia" family="Hadrosauridae" genus="Parasaurolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="walkeri">
|
||
<emphasis id="52DFEAEDB117FFF2C8285255FEC4FEF1" italics="true" pageId="7" pageNumber="59">Parasaurolophus walkeri</emphasis>
|
||
</taxonomicName>
|
||
than to
|
||
<taxonomicName id="A7AB4D7CB117FFF2CCAA5230FCFDFEF1" authority="Sereno 2005" authorityName="Sereno" authorityYear="2005" box="[388,798,332,356]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="52DFEAEDB117FFF2CCAA5230FD94FEF1" box="[388,631,332,356]" italics="true" pageId="7" pageNumber="59">Camptosaurus dispar</emphasis>
|
||
(
|
||
<bibRefCitation id="043A4B0EB117FFF2CFA85231FCF5FEF1" author="Sereno" box="[646,790,332,356]" pageId="7" pageNumber="59" refString="Sereno, P. C. (2005) Stem Archosauria version 1. 0, TaxonSearch. Available from: http: // www. taxonsearch. org / Archive / stemarchosauria- 1.0. php (21 April 2009)." type="journal article" year="2005">Sereno 2005</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="F8902A7AB117FFF2CE005231FC93FEF1" box="[814,880,332,356]" captionStart="FIGURE 1" captionStartId="2.[151,250,1854,1877]" captionTargetBox="[207,1262,450,1790]" captionTargetId="figure@2.[191,1274,445,1796]" captionTargetPageId="2" captionText="FIGURE 1. Reduced consensus tree of 11,850 MPTs of 358 steps each, following ordering of 22 multistate characters and safe taxonomic reduction. Camptosaurus dispar is highlighted in yellow and Uteodon aphanoecetes and Cumnoria prestwichii in blue. Numbers below and to the left of some nodes correspond to the following clade names: 1, Ornithopoda; 2, Iguanodontia; 3, Rhabdodontidae; 4, Dryomorpha; 5, Dryosauridae; 6, Ankylopollexia; 7, Styracosterna; 8, Hadrosauriformes; 9, Hadrosauroidea." httpUri="https://zenodo.org/record/200812/files/figure.png" pageId="7" pageNumber="59">Fig. 1</figureCitation>
|
||
). Furthermore,
|
||
<taxonomicName id="A7AB4D7CB117FFF2C90D5231FB60FEF1" box="[1059,1155,332,356]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="66" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="52DFEAEDB117FFF2C90D5231FB60FEF1" box="[1059,1155,332,356]" italics="true" pageId="7" pageNumber="59">Uteodon</emphasis>
|
||
</taxonomicName>
|
||
is also found to be more derived than
|
||
<taxonomicName id="A7AB4D7CB117FFF2CC04520DFE70FE1D" box="[298,403,368,392]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="52DFEAEDB117FFF2CC04520DFEDEFE12" box="[298,317,368,391]" italics="true" pageId="7" pageNumber="59">C</emphasis>
|
||
.
|
||
<emphasis id="52DFEAEDB117FFF2CC65520DFE70FE1D" box="[331,403,368,392]" italics="true" pageId="7" pageNumber="59">dispar</emphasis>
|
||
</taxonomicName>
|
||
in the agreement subtree (
|
||
<figureCitation id="F8902A7AB117FFF2CF935212FCE1FE12" box="[701,770,367,391]" captionStart="FIGURE 2" captionStartId="3.[151,250,1430,1453]" captionTargetBox="[296,1231,202,1382]" captionTargetId="figure@3.[275,1239,193,1396]" captionTargetPageId="3" captionText="FIGURE 2. Agreement subtree of 11,850 MPTs of 358 steps each, following ordering of 22 multistate characters and safe taxonomic reduction. Camptosaurus dispar is highlighted in yellow and Uteodon aphanoecetes in blue. Numbers below and to the left of some nodes correspond to the following clade names: 1, Iguanodontia; 2, Rhabdodontidae; 3, Dryomorpha; 4, Dryosauridae; 5, Ankylopollexia; 6, Styracosterna; 7, Hadrosauriformes; 8, Hadrosauroidea; 9, Hadrosauridae." httpUri="https://zenodo.org/record/200813/files/figure.png" pageId="7" pageNumber="59">Fig. 2</figureCitation>
|
||
). For the analysis,
|
||
<taxonomicName id="A7AB4D7CB117FFF2CEF9520DFB34FE1D" box="[983,1239,368,392]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="52DFEAEDB117FFF2CEF9520DFB34FE1D" box="[983,1239,368,392]" italics="true" pageId="7" pageNumber="59">Uteodon aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
was coded based upon the
|
||
<typeStatus id="BF10885DB117FFF2CC2C52E9FE84FE39" box="[258,359,404,428]" pageId="7" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
,
|
||
<typeStatus id="BF10885DB117FFF2CC5D52E8FE02FE38" box="[371,481,405,429]" pageId="7" pageNumber="59" type="paratype">paratypes</typeStatus>
|
||
, and referred specimens designated by
|
||
<bibRefCitation id="043A4B0EB117FFF2CE8E52E9FB38FE39" author="Carpenter" box="[928,1243,404,428]" pageId="7" pageNumber="59" refString="Carpenter, K. & Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter & Wilson (2008)</bibRefCitation>
|
||
. Only specimens in the CM collection could be examined firsthand.
|
||
</paragraph>
|
||
<paragraph id="601436FFB117FFF2CDE952A1FECAFCC9" blockId="7.[151,1437,151,860]" pageId="7" pageNumber="59">
|
||
<taxonomicName id="A7AB4D7CB117FFF2CDE952A1FE2BFE61" box="[199,456,476,500]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="52DFEAEDB117FFF2CDE952A1FE2BFE61" box="[199,456,476,500]" italics="true" pageId="7" pageNumber="59">Uteodon aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="52DFEAEDB117FFF2CF2D52A0FD1AFE61" box="[515,761,476,500]" italics="true" pageId="7" pageNumber="59">
|
||
Cumnoria
|
||
<taxonomicName id="A7AB4D7CB117FFF2CF5352A1FD1AFE61" box="[637,761,476,500]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
|
||
</emphasis>
|
||
are quite similar to each other and different from
|
||
<taxonomicName id="A7AB4D7CB117FFF2C81452A0FED2FD8D" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="52DFEAEDB117FFF2C81452A0FED2FD8D" italics="true" pageId="7" pageNumber="59">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<typeStatus id="BF10885DB117FFF2CC585282FE3BFD82" box="[374,472,511,535]" pageId="7" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
skeletons of
|
||
<taxonomicName id="A7AB4D7CB117FFF2CF5D517DFD37FD8D" box="[627,724,512,536]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="66" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="52DFEAEDB117FFF2CF5D517DFD37FD8D" box="[627,724,512,536]" italics="true" pageId="7" pageNumber="59">Uteodon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="52DFEAEDB117FFF2CE20517DFC9CFD82" box="[782,895,512,535]" italics="true" pageId="7" pageNumber="59">Cumnoria</emphasis>
|
||
both exhibit scapulae with convex dorsal margins, whereas scapulae of
|
||
<taxonomicName id="A7AB4D7CB117FFF2CC935158FD4CFDA9" box="[445,687,548,572]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="52DFEAEDB117FFF2CC935158FD4CFDA9" box="[445,687,548,572]" italics="true" pageId="7" pageNumber="59">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
are straight or somewhat concave along their dorsal margins (e.g., USNM 2210, 4282, 5473, 5954, 5955; YPM 8611) (
|
||
<figureCitation id="F8902A7AB117FFF2CFCE513AFCC7FDCA" box="[736,804,583,607]" captionStart="FIGURE 3" captionStartId="5.[151,250,1923,1946]" captionTargetBox="[218,1348,194,1877]" captionTargetId="figure@5.[213,1359,193,1878]" captionTargetPageId="5" captionText="FIGURE 3. (A) USNM 4282, left scapula of Camptosaurus dispar in lateral view; (B) CM 11337, cranial end of the right scapula of Uteodon aphanoecetes in lateral view; (C) CM 11337, left scapula of Uteodon aphanoecetes in lateral view; (D) OXFUM J. 3303, right scapula of Cumnoria prestwichii in lateral view. Abbreviations: cap, convex cranial margin of the acromion process; tap, triangular acromion process. Scale bar in A equals 10 cm; scale bar in B equals 5 cm." httpUri="https://zenodo.org/record/200814/files/figure.png" pageId="7" pageNumber="59">Fig. 3</figureCitation>
|
||
). Furthermore, as noted above, the ilia of
|
||
<emphasis id="52DFEAEDB117FFF2C9D55135FA8EFDCA" box="[1275,1389,584,607]" italics="true" pageId="7" pageNumber="59">Cumnoria</emphasis>
|
||
and
|
||
<taxonomicName id="A7AB4D7CB117FFF2CDB95111FF1BFD11" box="[151,248,620,644]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="66" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="52DFEAEDB117FFF2CDB95111FF1BFD11" box="[151,248,620,644]" italics="true" pageId="7" pageNumber="59">Uteodon</emphasis>
|
||
</taxonomicName>
|
||
exhibit convex dorsal margins (
|
||
<figureCitation id="F8902A7AB117FFF2CF5E5111FD54FD11" box="[624,695,620,644]" captionStart="FIGURE 6" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1424,927,1709]" captionTargetId="figure@7.[151,1436,913,1723]" captionTargetPageId="7" captionText="FIGURE 6. Left ilium of CM 11337 (holotype of Uteodon aphanoecetes) in (A) ventrolateral view showing the convex dorsal margin, (B) lateral view, (C), caudal view. Sacrum and ilia of CM 11337 in (D) dorsal view. Right ilium of CM 11337 in (E) lateral view. Right ilium of OXFUM J. 3303 (holotype of Cumnoria prestwichii) in (F) lateral view. Abbreviations: dmi, dorsal margin of ilium; mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process." httpUri="https://zenodo.org/record/200817/files/figure.png" pageId="7" pageNumber="59">Fig. 6</figureCitation>
|
||
), similar to some more derived iguanodonts such as
|
||
<taxonomicName id="A7AB4D7CB117FFF2C80D5111FEA2FD3D" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="52DFEAEDB117FFF2C80D5111FEA2FD3D" italics="true" pageId="7" pageNumber="59">Iguanodon bernissartensis</emphasis>
|
||
</taxonomicName>
|
||
(IRSNB 1534) and
|
||
<emphasis id="52DFEAEDB117FFF2CF0F51EDFD1BFD3D" box="[545,760,656,680]" italics="true" pageId="7" pageNumber="59">Equijubus normani</emphasis>
|
||
(
|
||
<bibRefCitation id="043A4B0EB117FFF2CE2851F2FC54FD32" author="You" box="[774,951,655,680]" pageId="7" pageNumber="59" refString="You, H., Luo, Z., Shubin, N. H., Witmer, L. M., Tang, Z. & Tang, F. (2003) The earliest-known duck-billed dinosaur from deposits of late Early Cretaceous age in northwest China and hadrosaur evolution. Cretaceous Research, 24, 347 - 355." type="journal article" year="2003">
|
||
You
|
||
<emphasis id="52DFEAEDB117FFF2CE1251ECFC8EFD3D" box="[828,877,656,680]" italics="true" pageId="7" pageNumber="59">et al</emphasis>
|
||
. 2003
|
||
</bibRefCitation>
|
||
), but different from the straight dorsal margin of the ilium of
|
||
<taxonomicName id="A7AB4D7CB117FFF2CC4351C8FD83FD59" box="[365,608,692,716]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="52DFEAEDB117FFF2CC4351C8FD83FD59" box="[365,608,692,716]" italics="true" pageId="7" pageNumber="59">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="F8902A7AB117FFF2CF5E51C9FD55FD59" box="[624,694,692,716]" captionStart="FIGURE 4" captionStartId="6.[151,250,720,743]" captionTargetBox="[151,1436,193,698]" captionTargetId="figure@6.[151,1436,193,699]" captionTargetPageId="6" captionText="FIGURE 4. Ilia of Camptosaurus dispar in lateral view. (A) right ilium of YPM 1877 (holotype) from Quarry 13; (B) right ilium of YPM 8622 from Quarry 13; (C) left ilium of USNM 5473 from Quarry 13; (D) left ilium of YPM PU 14553 from Cleveland-Lloyd Dinosaur Quarry. Abbreviations: mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process. Images in A, B, and D are copyright YPM. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200815/files/figure.png" pageId="7" pageNumber="59">Fig. 4</figureCitation>
|
||
). Furthermore,
|
||
<taxonomicName id="A7AB4D7CB117FFF2CE4851C9FC25FD59" box="[870,966,692,716]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="66" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="52DFEAEDB117FFF2CE4851C9FC25FD59" box="[870,966,692,716]" italics="true" pageId="7" pageNumber="59">Uteodon</emphasis>
|
||
</taxonomicName>
|
||
possesses an autapomorphy in the partial braincase of
|
||
<typeStatus id="BF10885DB117FFF2CC0951A5FE64FD65" box="[295,391,728,752]" pageId="7" pageNumber="59" type="paratype">paratype</typeStatus>
|
||
specimen CM 15780: the occipital condyle projects far ventrally relative to the basal tubera. This condition differs from the basioccipitals of
|
||
<taxonomicName id="A7AB4D7CB117FFF2CFE85180FC59FC81" box="[710,954,764,788]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="52DFEAEDB117FFF2CFE85180FC59FC81" box="[710,954,764,788]" italics="true" pageId="7" pageNumber="59">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
(USNM 5473, 5996, 5997; YPM 1856A, 1880) and
|
||
<emphasis id="52DFEAEDB117FFF2CC21505DFE62FCA2" box="[271,385,800,823]" italics="true" pageId="7" pageNumber="59">Cumnoria</emphasis>
|
||
(OXFUM J.3303), in which the basal tubera protrude farther ventrally than the occipital condyle (
|
||
<figureCitation id="F8902A7AB117FFF2CDF85039FEF8FCC9" box="[214,283,836,860]" captionStart="FIGURE 7" captionStartId="8.[151,250,1622,1645]" captionTargetBox="[472,1114,625,1600]" captionTargetId="figure@8.[472,1115,625,1601]" captionTargetPageId="8" captionText="FIGURE 7. Partial braincases of Camptosaurus dispar, Uteodon aphanoecetes, and Cumnoria prestwichii. (A) YPM 1880 in left lateral view (image copyright YPM); (B) CM 15780 in right lateral view (image courtesy of M. Lamanna); (C) OXFUM J. 3303 in right lateral view. Abbreviations: bt, basal tubera; oc, occipital condyle. Scale bar equals 5 cm." httpUri="https://zenodo.org/record/200818/files/figure.png" pageId="7" pageNumber="59">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="34D46677B117FFF2CDB955ADFAEAF8D7" httpUri="https://zenodo.org/record/200817/files/figure.png" pageId="7" pageNumber="59" targetBox="[151,1424,927,1709]" targetPageId="7">
|
||
<paragraph id="601436FFB117FFF2CDB955ADFAEAF8D7" blockId="7.[151,1436,1744,1858]" pageId="7" pageNumber="59">
|
||
<emphasis id="52DFEAEDB117FFF2CDB955ADFEF1F973" bold="true" box="[151,274,1744,1767]" pageId="7" pageNumber="59">FIGURE 6.</emphasis>
|
||
Left ilium of CM 11337 (holotype of
|
||
<taxonomicName id="A7AB4D7CB117FFF2CFB855ACFC9EF972" box="[662,893,1745,1767]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="52DFEAEDB117FFF2CFB855ACFC9EF972" box="[662,893,1745,1767]" italics="true" pageId="7" pageNumber="59">Uteodon aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
) in (
|
||
<emphasis id="52DFEAEDB117FFF2CE8055ACFC23F973" bold="true" box="[942,960,1745,1766]" pageId="7" pageNumber="59">A</emphasis>
|
||
) ventrolateral view showing the convex dorsal margin, (
|
||
<emphasis id="52DFEAEDB117FFF2CDDA558DFEE6F890" bold="true" box="[244,261,1776,1797]" pageId="7" pageNumber="59">B</emphasis>
|
||
) lateral view, (
|
||
<emphasis id="52DFEAEDB117FFF2CC8F5592FE50F893" bold="true" box="[417,435,1775,1798]" pageId="7" pageNumber="59">C</emphasis>
|
||
), caudal view. Sacrum and ilia of CM 11337 in (
|
||
<emphasis id="52DFEAEDB117FFF2CE83558DFC5CF890" bold="true" box="[941,959,1776,1797]" pageId="7" pageNumber="59">D</emphasis>
|
||
) dorsal view. Right ilium of CM 11337 in (
|
||
<emphasis id="52DFEAEDB117FFF2C8AD558DFA77F890" bold="true" box="[1411,1428,1776,1797]" pageId="7" pageNumber="59">E</emphasis>
|
||
) lateral view. Right ilium of OXFUM J.3303 (holotype of
|
||
<emphasis id="52DFEAEDB117FFF2CFF05472FC58F8B1" box="[734,955,1806,1828]" italics="true" pageId="7" pageNumber="59">
|
||
Cumnoria
|
||
<taxonomicName id="A7AB4D7CB117FFF2CE655473FC58F8B1" box="[843,955,1806,1828]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
|
||
</emphasis>
|
||
) in (
|
||
<emphasis id="52DFEAEDB117FFF2CEC25473FC18F8B6" bold="true" box="[1004,1019,1806,1827]" pageId="7" pageNumber="59">F</emphasis>
|
||
) lateral view. Abbreviations: dmi, dorsal margin of ilium; mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="601436FFB117FFFDCDE95412FB43FDA9" blockId="7.[151,1436,1903,2036]" lastBlockId="8.[151,1436,151,573]" lastPageId="8" lastPageNumber="60" pageId="7" pageNumber="59">
|
||
The results of the phylogenetic analysis and the morphological differences explicated above strongly suggest that
|
||
<taxonomicName id="A7AB4D7CB117FFF2CDE554E9FE2DF839" box="[203,462,1940,1964]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="52DFEAEDB117FFF2CDE554E9FE2DF839" box="[203,462,1940,1964]" italics="true" pageId="7" pageNumber="59">Uteodon aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="52DFEAEDB117FFF2CF2554E9FCE1F839" box="[523,770,1940,1964]" italics="true" pageId="7" pageNumber="59">
|
||
Cumnoria
|
||
<taxonomicName id="A7AB4D7CB117FFF2CFA954E9FCE1F839" box="[647,770,1940,1964]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
|
||
</emphasis>
|
||
should not be considered congeneric with
|
||
<taxonomicName id="A7AB4D7CB117FFF2C9D754E9FF3CF85A" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="52DFEAEDB117FFF2C9D754E9FF3CF85A" italics="true" pageId="7" pageNumber="59">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="A7AB4D7CB117FFF2CDC054CAFEADF85A" box="[238,334,1975,1999]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="66" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="52DFEAEDB117FFF2CDC054CAFEADF85A" box="[238,334,1975,1999]" italics="true" pageId="7" pageNumber="59">Uteodon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="52DFEAEDB117FFF2CCA854C5FE14F85A" box="[390,503,1976,1999]" italics="true" pageId="7" pageNumber="59">Cumnoria</emphasis>
|
||
are recovered as basal styracosternans more derived than
|
||
<taxonomicName id="A7AB4D7CB117FFF2C9A554C5FA9FF85A" box="[1163,1404,1975,1999]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="52DFEAEDB117FFF2C9A554C5FA9FF85A" box="[1163,1404,1975,1999]" italics="true" pageId="7" pageNumber="59">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
. It is thus a straightforward matter to reinstate the existing genus
|
||
<emphasis id="52DFEAEDB117FFF2CE7D54A1FC27F866" box="[851,964,2012,2035]" italics="true" pageId="7" pageNumber="59">Cumnoria</emphasis>
|
||
(
|
||
<bibRefCitation id="043A4B0EB117FFF2CEFD54A6FB83F866" author="Seeley" box="[979,1120,2011,2035]" pageId="7" pageNumber="59" refString="Seeley, H. G. (1888) On Cumnoria, an iguanodont genus founded upon the Iguanodon prestwichi, Hulke. Report of the British Association of the Advancement of Science, 57, 698." type="journal article" year="1888">Seeley 1888</bibRefCitation>
|
||
) for “
|
||
<taxonomicName id="A7AB4D7CB117FFF2C98A54A1FAA4F866" box="[1188,1351,2012,2035]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="genus">
|
||
<emphasis id="52DFEAEDB117FFF2C98A54A1FAA4F866" box="[1188,1351,2012,2035]" italics="true" pageId="7" pageNumber="59">Camptosaurus</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
<taxonomicName id="A7AB4D7CB117FFFDC87254A0FF3FFF25" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="8" lastPageNumber="60" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="prestwichii">
|
||
<emphasis id="52DFEAEDB117FFFDC87254A0FF3FFF25" italics="true" lastPageId="8" lastPageNumber="60" pageId="7" pageNumber="59">prestwichii</emphasis>
|
||
</taxonomicName>
|
||
(see below). However, a more difficult dilemma is the status of “
|
||
<taxonomicName id="A7AB4D7CB118FFFDCE9053E5FB82FF3A" box="[958,1121,152,175]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="genus">
|
||
<emphasis id="52DFEAEDB118FFFDCE9053E5FB82FF3A" box="[958,1121,152,175]" italics="true" pageId="8" pageNumber="60">Camptosaurus</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
<taxonomicName id="A7AB4D7CB118FFFDC95B53E5FAEEFF25" box="[1141,1293,152,176]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="52DFEAEDB118FFFDC95B53E5FAEEFF25" box="[1141,1293,152,176]" italics="true" pageId="8" pageNumber="60">aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
, which does not have a pre-existing genus into which it can be conveniently placed.
|
||
<collectingCountry id="18BC766FB118FFFDCEFD53C1FC11FF41" box="[979,1010,188,212]" name="American Samoa" pageId="8" pageNumber="60">As</collectingCountry>
|
||
described above, this species is very similar to
|
||
<emphasis id="52DFEAEDB118FFFDCC24539DFE1DFF6D" box="[266,510,224,248]" italics="true" pageId="8" pageNumber="60">
|
||
Cumnoria
|
||
<taxonomicName id="A7AB4D7CB118FFFDCCAD539DFE1DFF6D" box="[387,510,224,248]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
|
||
</emphasis>
|
||
, distinguished only by a single autapomorphy of the braincase. Thus, the possible taxonomic revision that immediately arises is to create the new combination
|
||
<emphasis id="52DFEAEDB118FFFDC9225278FAFCFE89" box="[1036,1311,260,284]" italics="true" pageId="8" pageNumber="60">
|
||
Cumnoria
|
||
<taxonomicName id="A7AB4D7CB118FFFDC9A95279FAFCFE89" box="[1159,1311,260,284]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="aphanoecetes">aphanoecetes</taxonomicName>
|
||
</emphasis>
|
||
. However, such a move presupposes that future phylogenetic analyses and discoveries will find a sister-taxa relationship of
|
||
<emphasis id="52DFEAEDB118FFFDCDB95230FE73FEF1" box="[151,400,332,356]" italics="true" pageId="8" pageNumber="60">
|
||
Cumnoria
|
||
<taxonomicName id="A7AB4D7CB118FFFDCC3A5231FE73FEF1" box="[276,400,332,356]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
|
||
</emphasis>
|
||
and “
|
||
<taxonomicName id="A7AB4D7CB118FFFDCCF25230FD9CFEF1" box="[476,639,333,356]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="genus">
|
||
<emphasis id="52DFEAEDB118FFFDCCF25230FD9CFEF1" box="[476,639,333,356]" italics="true" pageId="8" pageNumber="60">Camptosaurus</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
<taxonomicName id="A7AB4D7CB118FFFDCFB65231FCD2FEF1" box="[664,817,332,356]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="52DFEAEDB118FFFDCFB65231FCD2FEF1" box="[664,817,332,356]" italics="true" pageId="8" pageNumber="60">aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
. The topologies presented herein could change with additional new taxa and new material, most especially additional cranial elements of “
|
||
<taxonomicName id="A7AB4D7CB118FFFDC9B0520DFAA2FE12" box="[1182,1345,368,391]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="genus">
|
||
<emphasis id="52DFEAEDB118FFFDC9B0520DFAA2FE12" box="[1182,1345,368,391]" italics="true" pageId="8" pageNumber="60">Camptosaurus</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
<taxonomicName id="A7AB4D7CB118FFFDC875520DFF1BFE39" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="52DFEAEDB118FFFDC875520DFF1BFE39" italics="true" pageId="8" pageNumber="60">aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
(in addition to the known right dentary DINO 556 and partial braincase of CM 15780). It is this labile nature of the phylogeny that prompts the creation of the new genus and combination
|
||
<taxonomicName id="A7AB4D7CB118FFFDC97552C5FAB8FE45" box="[1115,1371,440,464]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="52DFEAEDB118FFFDC97552C5FAB8FE45" box="[1115,1371,440,464]" italics="true" pageId="8" pageNumber="60">Uteodon aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
. This species can thus occupy any position relative to
|
||
<emphasis id="52DFEAEDB118FFFDCF9D52A0FC45FE61" box="[691,934,476,500]" italics="true" pageId="8" pageNumber="60">
|
||
Cumnoria
|
||
<taxonomicName id="A7AB4D7CB118FFFDCE0552A1FC45FE61" box="[811,934,476,500]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="A7AB4D7CB118FFFDCEF352A0FB2CFE61" box="[989,1231,476,500]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="52DFEAEDB118FFFDCEF352A0FB2CFE61" box="[989,1231,476,500]" italics="true" pageId="8" pageNumber="60">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
in future analyses without necessitating still another taxonomic reassignment. From the standpoint of taxonomic stability, giving
|
||
<taxonomicName id="A7AB4D7CB118FFFDC844517CFE8DFDA9" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="52DFEAEDB118FFFDC844517CFE8DFDA9" italics="true" pageId="8" pageNumber="60">Uteodon aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
its own new genus is preferable to committing it to the genus
|
||
<emphasis id="52DFEAEDB118FFFDC9065158FB79FDA9" box="[1064,1178,549,572]" italics="true" pageId="8" pageNumber="60">Cumnoria</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |