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<document id="D094D5E8820B97966D0A4DBB2F856B85" ID-CLB-Dataset="24859" ID-DOI="10.11646/zootaxa.4700.3.1" ID-GBIF-Dataset="1b0654ea-eecb-4242-ad9a-850602f7d185" ID-ISSN="1175-5326" ID-Zenodo-Dep="3548509" ID-ZooBank="B0712990-7A3B-4042-9A4B-5CE96AFAECF8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1574256400520" checkinUser="plazi" docAuthor="Grogan, William L., Díaz, Florentina, Spinelli, Gustavo R. &amp; Ronderos, Maria M." docDate="2019" docId="DF72B52DDE01FF80FF541905FE2E0954" docLanguage="en" docName="zootaxa.4700.3.1.pdf" docOrigin="Zootaxa 4700 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dasyhelea aliciae Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos 2019, n. sp." docType="treatment" docVersion="10" lastPageNumber="310" masterDocId="234BCD55DE06FF89FFC31D26FF870D63" masterDocTitle="The Biting Midges of the Caribbean island Curaçao (Diptera: Ceratopogonidae) I. Species in the genus Dasyhelea Kieffer" masterLastPageNumber="325" masterPageNumber="301" pageNumber="308" updateTime="1698701345973" updateUser="plazi">
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<mods:title id="E063E1833540D5135240E46A59FD57A1">The Biting Midges of the Caribbean island Curaçao (Diptera: Ceratopogonidae) I. Species in the genus Dasyhelea Kieffer</mods:title>
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<mods:namePart id="858880EC26B3538B03C3315F2EAA37AC">Grogan, William L.</mods:namePart>
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<mods:namePart id="5F460B11ED282D582D02965CF75F59FF">Díaz, Florentina</mods:namePart>
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<mods:namePart id="7B278A19DFB64EEDFA320A93354F7735">Spinelli, Gustavo R.</mods:namePart>
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<mods:namePart id="7400652FD89274D9EF036BD5B2A9B867">Ronderos, Maria M.</mods:namePart>
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<treatment id="DF72B52DDE01FF80FF541905FE2E0954" ID-DOI="http://doi.org/10.5281/zenodo.5457136" ID-GBIF-Taxon="160458586" ID-Zenodo-Dep="5457136" LSID="urn:lsid:plazi:treatment:DF72B52DDE01FF80FF541905FE2E0954" httpUri="http://treatment.plazi.org/id/DF72B52DDE01FF80FF541905FE2E0954" lastPageId="9" lastPageNumber="310" pageId="7" pageNumber="308">
<subSubSection id="1FC157B0DE01FF8EFF541905FE31095E" box="[151,438,1059,1086]" pageId="7" pageNumber="308" type="nomenclature">
<paragraph id="5764043BDE01FF8EFF541905FE31095E" blockId="7.[151,438,1059,1121]" box="[151,438,1059,1086]" pageId="7" pageNumber="308">
<heading id="0C2CB357DE01FF8EFF541905FE31095E" bold="true" box="[151,438,1059,1086]" fontSize="11" level="1" pageId="7" pageNumber="308" reason="1">
<emphasis id="65AFD829DE01FF8EFF541905FE31095E" bold="true" box="[151,438,1059,1086]" pageId="7" pageNumber="308">
<taxonomicName id="90DB7FB8DE01FF8EFF541905FEEB095D" ID-CoL="8BQRW" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[151,364,1059,1086]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="aliciae" status="sp. nov.">
<emphasis id="65AFD829DE01FF8EFF541905FEEB095D" bold="true" box="[151,364,1059,1086]" italics="true" pageId="7" pageNumber="308">Dasyhelea aliciae</emphasis>
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<taxonomicNameLabel id="7E9C6552DE01FF8EFEB01905FE31095E" box="[371,438,1059,1085]" pageId="7" pageNumber="308" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="1FC157B0DE01FF8EFF541961FEE10902" box="[151,358,1094,1121]" pageId="7" pageNumber="308" type="description">
<paragraph id="5764043BDE01FF8EFF541961FEE10902" blockId="7.[151,438,1059,1121]" box="[151,358,1094,1121]" pageId="7" pageNumber="308">
(
<figureCitation id="CFE018BEDE01FF8EFF5C1961FE8C0902" box="[159,267,1094,1121]" captionStart="FIGURES 15" captionStartId="8.[151,264,1942,1967]" captionTargetBox="[168,1425,197,1894]" captionTargetId="figure@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 15. Dasyhelea aliciae n. sp. Holotype male. 6-10. Dasyhelea recurva n. sp. Holotype male. 1, 6 head. 2, 7. palpus. 3, 8 thorax and abdomen. 4, 9 wing. 5, 10 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548511" httpUri="https://zenodo.org/record/3548511/files/figure.png" pageId="7" pageNumber="308">Figs. 15</figureCitation>
,
<figureCitation id="CFE018BEDE01FF8EFED41960FED90902" box="[279,350,1094,1121]" captionStart="FIGURES 5269" captionStartId="20.[151,264,1944,1969]" captionTargetBox="[187,1405,192,1908]" captionTargetId="figure@20.[170,1417,181,1919]" captionTargetPageId="20" captionText="FIGURES 5269. Male Dasyhelea genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 5254 D. aliciae n. sp., 5557 D. recurva n. sp., 5860 D. latiala n. sp., 6163 D. cyrtostyla n. sp., 6466 D. curacaoensis n. sp., 6769 D. rhopaloparamera n. sp. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="7" pageNumber="308">5254</figureCitation>
)
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<subSubSection id="1FC157B0DE01FF80FF5419AEFE250F78" lastPageId="9" lastPageNumber="310" pageId="7" pageNumber="308" type="diagnosis">
<paragraph id="5764043BDE01FF8EFF5419AEFE800819" blockId="7.[151,1437,1160,1943]" pageId="7" pageNumber="308">
<emphasis id="65AFD829DE01FF8EFF5419AEFE8D09C0" bold="true" box="[151,266,1160,1187]" pageId="7" pageNumber="308">Diagnosis</emphasis>
. The only Neotropical species of the leptobranchia group with the following combination of characters. Males wing with a short costa (costal ratio 0.42); aedeagus stout, heavily sclerotized, H-shaped; gonocoxal apodemes and paramere asymmetrical, left gonocoxal apodeme barely contacting base of paramere; paramere sinuate with slender elongate recurved distal portion, the basal section of which is adpressed to the apex of the distal portion; sternite 9 moderately elongate, extending just below basal arch of aedeagus, distal margin slightly curved; and apex of tergite 9 broad, curved with very widely spaced elongate, divergent apicolateral processes. Female unknown.
</paragraph>
<paragraph id="5764043BDE01FF80FF0418A3FB0C0C94" blockId="7.[151,1437,1160,1943]" lastBlockId="9.[151,1437,153,1079]" lastPageId="9" lastPageNumber="310" pageId="7" pageNumber="308">
<emphasis id="65AFD829DE01FF8EFF0418A3FE8308FC" bold="true" box="[199,260,1413,1439]" pageId="7" pageNumber="308">Male</emphasis>
. Head (
<figureCitation id="CFE018BEDE01FF8EFE9918A2FE1D08FC" box="[346,410,1412,1439]" captionStart="FIGURES 15" captionStartId="8.[151,264,1942,1967]" captionTargetBox="[168,1425,197,1894]" captionTargetId="figure@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 15. Dasyhelea aliciae n. sp. Holotype male. 6-10. Dasyhelea recurva n. sp. Holotype male. 1, 6 head. 2, 7. palpus. 3, 8 thorax and abdomen. 4, 9 wing. 5, 10 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548511" httpUri="https://zenodo.org/record/3548511/files/figure.png" pageId="7" pageNumber="308">Fig. 1</figureCitation>
). Dark brown. Eyes contiguous for a distance equal to the width of 45 ommatidia. Antennal flagellum dark brown; flagellomeres 29 sculptured, rhomboidal, 1013 elongate; flagellomeres 210 with whorl of long sensillae chaetica comprising the moderately dense plume; antennal ratio 0.88; flagellum length
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. Frontal sclerite elliptical, with short pointed ventral projection, with folded lower marginal ear-like lobes, difficult to see. Clypeus with 5 pairs of setae. Palpus (
<figureCitation id="CFE018BEDE01FF8EFD521B32FD530B4C" box="[657,724,1556,1583]" captionStart="FIGURES 15" captionStartId="8.[151,264,1942,1967]" captionTargetBox="[168,1425,197,1894]" captionTargetId="figure@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 15. Dasyhelea aliciae n. sp. Holotype male. 6-10. Dasyhelea recurva n. sp. Holotype male. 1, 6 head. 2, 7. palpus. 3, 8 thorax and abdomen. 4, 9 wing. 5, 10 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548511" httpUri="https://zenodo.org/record/3548511/files/figure.png" pageId="7" pageNumber="308">Fig. 2</figureCitation>
) brown; segment 3 as long as 4+5 combined with 12 scattered hyaline capitate sensillae on distal 2/3 of mesal surface; palpal ratio 3.28. Thorax (
<figureCitation id="CFE018BEDE01FF8EFBFE1B1EFB050B30" box="[1085,1154,1592,1619]" captionStart="FIGURES 15" captionStartId="8.[151,264,1942,1967]" captionTargetBox="[168,1425,197,1894]" captionTargetId="figure@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 15. Dasyhelea aliciae n. sp. Holotype male. 6-10. Dasyhelea recurva n. sp. Holotype male. 1, 6 head. 2, 7. palpus. 3, 8 thorax and abdomen. 4, 9 wing. 5, 10 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548511" httpUri="https://zenodo.org/record/3548511/files/figure.png" pageId="7" pageNumber="308">Fig. 3</figureCitation>
). Dark brown; scutellum with 6 large, 3 smaller setae. Femora, tibiae dark brown, extreme bases slightly paler; tarsi with tarsomeres 14 light brown, tarsomeres 5 infuscated; hind tibial comb with 8 spines; prothoracic tarsal ratio 2.14, mesothoracic tarsal ratio 2.28, metathoracic tarsal ratio 2.11. Wing (
<figureCitation id="CFE018BEDE01FF8EFD7E1B82FC850BDC" box="[701,770,1700,1727]" captionStart="FIGURES 15" captionStartId="8.[151,264,1942,1967]" captionTargetBox="[168,1425,197,1894]" captionTargetId="figure@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 15. Dasyhelea aliciae n. sp. Holotype male. 6-10. Dasyhelea recurva n. sp. Holotype male. 1, 6 head. 2, 7. palpus. 3, 8 thorax and abdomen. 4, 9 wing. 5, 10 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548511" httpUri="https://zenodo.org/record/3548511/files/figure.png" pageId="7" pageNumber="308">Fig. 4</figureCitation>
) membrane hyaline with moderately dense macrotrichiae; radial cells without lumen, 2
<superScript id="A0AEA973DE01FF8EFE191BE1FE6D0BB6" attach="left" box="[474,490,1735,1749]" fontSize="6" pageId="7" pageNumber="308">nd</superScript>
radial cell very short; apices of costa, radius at perpendicular angle with wing margin; fork of CuA
<subScript id="CB5F067EDE01FF8EFE921BD8FEDE0A6F" attach="left" box="[337,345,1790,1804]" fontSize="6" pageId="7" pageNumber="308">1</subScript>
slightly distal to level of apex of 2
<superScript id="A0AEA973DE01FF8EFD181BCDFD6C0B9A" attach="left" box="[731,747,1771,1785]" fontSize="6" pageId="7" pageNumber="308">nd</superScript>
radial cell; anal lobe moderately developed; wing length
<quantity id="9023A9DEDE01FF8EFAA81BCAFF410A48" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="7" pageNumber="308" unit="mm" value="0.65">0.65 mm</quantity>
, width
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; costal ratio 0.42. Halter dark brown. Abdomen brown. Genitalia (
<figureCitation id="CFE018BEDE01FF8EFBB91A36FB4C0A48" box="[1146,1227,1808,1835]" captionStart="FIGURES 15" captionStartId="8.[151,264,1942,1967]" captionTargetBox="[168,1425,197,1894]" captionTargetId="figure@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 15. Dasyhelea aliciae n. sp. Holotype male. 6-10. Dasyhelea recurva n. sp. Holotype male. 1, 6 head. 2, 7. palpus. 3, 8 thorax and abdomen. 4, 9 wing. 5, 10 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548511" httpUri="https://zenodo.org/record/3548511/files/figure.png" pageId="7" pageNumber="308">Figs. 5</figureCitation>
,
<figureCitation id="CFE018BEDE01FF8EFB1A1A36FAA30A48" box="[1241,1316,1808,1835]" captionStart="FIGURES 5269" captionStartId="20.[151,264,1944,1969]" captionTargetBox="[187,1405,192,1908]" captionTargetId="figure@20.[170,1417,181,1919]" captionTargetPageId="20" captionText="FIGURES 5269. Male Dasyhelea genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 5254 D. aliciae n. sp., 5557 D. recurva n. sp., 5860 D. latiala n. sp., 6163 D. cyrtostyla n. sp., 6466 D. curacaoensis n. sp., 6769 D. rhopaloparamera n. sp. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="7" pageNumber="308">5254</figureCitation>
). Tergite 9 tapering progressively distally to rounded apex that extends to apices of gonocoxites; apicolateral processes very widely spaced, slender, elongate, cylindrical with medium-sized subapical seta; cercus very small, located at base of apicolateral process, with 2 small setae. Sternite 9 (
<figureCitation id="CFE018BEDE01FF8EFD341A5AFCC00AF4" box="[759,839,1916,1943]" captionStart="FIGURES 15" captionStartId="8.[151,264,1942,1967]" captionTargetBox="[168,1425,197,1894]" captionTargetId="figure@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 15. Dasyhelea aliciae n. sp. Holotype male. 6-10. Dasyhelea recurva n. sp. Holotype male. 1, 6 head. 2, 7. palpus. 3, 8 thorax and abdomen. 4, 9 wing. 5, 10 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548511" httpUri="https://zenodo.org/record/3548511/files/figure.png" pageId="7" pageNumber="308">Figs. 5</figureCitation>
,
<figureCitation id="CFE018BEDE01FF8EFC901A5AFCF70AF4" box="[851,880,1916,1943]" captionStart="FIGURES 5269" captionStartId="20.[151,264,1944,1969]" captionTargetBox="[187,1405,192,1908]" captionTargetId="figure@20.[170,1417,181,1919]" captionTargetPageId="20" captionText="FIGURES 5269. Male Dasyhelea genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 5254 D. aliciae n. sp., 5557 D. recurva n. sp., 5860 D. latiala n. sp., 6163 D. cyrtostyla n. sp., 6466 D. curacaoensis n. sp., 6769 D. rhopaloparamera n. sp. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="7" pageNumber="308">52</figureCitation>
) 0.30 length of greatest width; anterior margin curved; distal portion moderately long, sub-triangular, tapered to broad apex that extends just below basal arch of aedeagus. Gonocoxite straight, moderately slender, 2.2 x longer than broad, with a few large setae on apicomesal side; gonostylus 0.80 length of gonocoxite, distal 1/2 greatly curved, tapering to narrow pointed tip. Gonocoxal apodemes and paramere (
<figureCitation id="CFE018BEDE0FFF80FE1F1C23FDA90C7C" box="[476,558,260,287]" captionStart="FIGURES 15" captionStartId="8.[151,264,1942,1967]" captionTargetBox="[168,1425,197,1894]" captionTargetId="figure@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 15. Dasyhelea aliciae n. sp. Holotype male. 6-10. Dasyhelea recurva n. sp. Holotype male. 1, 6 head. 2, 7. palpus. 3, 8 thorax and abdomen. 4, 9 wing. 5, 10 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548511" httpUri="https://zenodo.org/record/3548511/files/figure.png" pageId="9" pageNumber="310">Figs. 5</figureCitation>
,
<figureCitation id="CFE018BEDE0FFF80FDFE1C22FDDD0C7C" box="[573,602,260,287]" captionStart="FIGURES 5269" captionStartId="20.[151,264,1944,1969]" captionTargetBox="[187,1405,192,1908]" captionTargetId="figure@20.[170,1417,181,1919]" captionTargetPageId="20" captionText="FIGURES 5269. Male Dasyhelea genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 5254 D. aliciae n. sp., 5557 D. recurva n. sp., 5860 D. latiala n. sp., 6163 D. cyrtostyla n. sp., 6466 D. curacaoensis n. sp., 6769 D. rhopaloparamera n. sp. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="9" pageNumber="310">53</figureCitation>
) asymmetrical; proximal portions of apodemes stout; basal arm of right apodeme recurved 120°, proximal portion broadly fused to paramere; basal arm of left apodeme broad with bluntly rounded apex,” distal portion slender, slightly recurved, barely contacting base of paramere; paramere stout, broad basally, tapering and curved distally, distal portion slender, abruptly recurved anteroventrally 180°, and adpressed to apex of proximal portion. Aedeagus (
<figureCitation id="CFE018BEDE0FFF80FD951CB3FD210CCC" box="[598,678,404,431]" captionStart="FIGURES 15" captionStartId="8.[151,264,1942,1967]" captionTargetBox="[168,1425,197,1894]" captionTargetId="figure@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 15. Dasyhelea aliciae n. sp. Holotype male. 6-10. Dasyhelea recurva n. sp. Holotype male. 1, 6 head. 2, 7. palpus. 3, 8 thorax and abdomen. 4, 9 wing. 5, 10 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548511" httpUri="https://zenodo.org/record/3548511/files/figure.png" pageId="9" pageNumber="310">Figs. 5</figureCitation>
,
<figureCitation id="CFE018BEDE0FFF80FD711CB2FD480CCC" box="[690,719,404,431]" captionStart="FIGURES 5269" captionStartId="20.[151,264,1944,1969]" captionTargetBox="[187,1405,192,1908]" captionTargetId="figure@20.[170,1417,181,1919]" captionTargetPageId="20" captionText="FIGURES 5269. Male Dasyhelea genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 5254 D. aliciae n. sp., 5557 D. recurva n. sp., 5860 D. latiala n. sp., 6163 D. cyrtostyla n. sp., 6466 D. curacaoensis n. sp., 6769 D. rhopaloparamera n. sp. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="9" pageNumber="310">54</figureCitation>
) stout, H-shaped, very heavily sclerotized, 0.71 length of greatest width; basal arch broad, extending 0.30 of total length of aedeagus, posterior margin nearly straight; basal arms short, moderately divergent; posterolateral arms slightly convergent, tips broadly rounded.
</paragraph>
<caption id="03A454B3DE0EFF81FF541AB0FC0C0AB1" ID-DOI="http://doi.org/10.5281/zenodo.3548511" ID-Zenodo-Dep="3548511" httpUri="https://zenodo.org/record/3548511/files/figure.png" pageId="8" pageNumber="309" startId="8.[151,264,1942,1967]" targetBox="[168,1425,197,1894]" targetPageId="8">
<paragraph id="5764043BDE0EFF81FF541AB0FC0C0AB1" blockId="8.[151,1437,1942,2003]" pageId="8" pageNumber="309">
<emphasis id="65AFD829DE0EFF81FF541AB0FEB30ACC" bold="true" box="[151,308,1942,1967]" pageId="8" pageNumber="309">FIGURES 15</emphasis>
.
<taxonomicName id="90DB7FB8DE0EFF81FE831AB0FE710ACD" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[320,502,1942,1966]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="aliciae" status="sp. nov.">
<emphasis id="65AFD829DE0EFF81FE831AB0FE710ACD" box="[320,502,1942,1966]" italics="true" pageId="8" pageNumber="309">Dasyhelea aliciae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE0EFF81FE381AB1FDB30ACD" box="[507,564,1943,1967]" pageId="8" pageNumber="309" rank="species">
<emphasis id="65AFD829DE0EFF81FE381AB1FD8E0ACC" bold="true" box="[507,521,1943,1967]" pageId="8" pageNumber="309">n</emphasis>
.
<emphasis id="65AFD829DE0EFF81FDD51AB1FDA90ACC" bold="true" box="[534,558,1943,1967]" pageId="8" pageNumber="309">sp</emphasis>
.
</taxonomicNameLabel>
Holotype male.
<emphasis id="65AFD829DE0EFF81FD181AB0FC8E0ACC" bold="true" box="[731,777,1942,1967]" pageId="8" pageNumber="309">6-10</emphasis>
.
<taxonomicName id="90DB7FB8DE0EFF81FCD51AB0FC520ACD" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[790,981,1942,1966]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="recurva" status="sp. nov.">
<emphasis id="65AFD829DE0EFF81FCD51AB0FC520ACD" box="[790,981,1942,1966]" italics="true" pageId="8" pageNumber="309">Dasyhelea recurva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE0EFF81FC181AB1FB940ACD" box="[987,1043,1943,1967]" pageId="8" pageNumber="309" rank="species">
<emphasis id="65AFD829DE0EFF81FC181AB1FC6E0ACC" bold="true" box="[987,1001,1943,1967]" pageId="8" pageNumber="309">n</emphasis>
.
<emphasis id="65AFD829DE0EFF81FC361AB1FB8A0ACC" bold="true" box="[1013,1037,1943,1967]" pageId="8" pageNumber="309">sp</emphasis>
.
</taxonomicNameLabel>
Holotype male. 1, 6 head. 2, 7. palpus. 3, 8 thorax and abdomen. 4, 9 wing. 5, 10 genitalia. Scale bars = 0.05 mm.
</paragraph>
</caption>
<paragraph id="5764043BDE0FFF80FF041F27FE250F78" blockId="9.[151,1437,153,1079]" box="[199,418,513,539]" pageId="9" pageNumber="310">
<emphasis id="65AFD829DE0FFF80FF041F27FE990F78" bold="true" box="[199,286,513,539]" pageId="9" pageNumber="310">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="1FC157B0DE0FFF80FF041F02FE4B0F5C" box="[199,460,548,575]" pageId="9" pageNumber="310" type="distribution">
<paragraph id="5764043BDE0FFF80FF041F02FE4B0F5C" blockId="9.[151,1437,153,1079]" box="[199,460,548,575]" pageId="9" pageNumber="310">
<emphasis id="65AFD829DE0FFF80FF041F02FEDE0F5C" bold="true" box="[199,345,548,575]" pageId="9" pageNumber="310">Distribution</emphasis>
.
<collectingCountry id="2FCC44ABDE0FFF80FEA41F03FE4F0F5C" box="[359,456,549,575]" name="Curaçao" pageId="9" pageNumber="310">Curaçao</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1FC157B0DE0FFF80FF041F6FFF530FAC" pageId="9" pageNumber="310" type="materials_examined">
<paragraph id="5764043BDE0FFF80FF041F6FFB270FE4" blockId="9.[151,1437,153,1079]" pageId="9" pageNumber="310">
<materialsCitation id="E7B30E66DE0FFF80FF041F6FFB270FE4" ID-GBIF-Occurrence="2556250904" collectingDate="2014-11-09" collectingMethod="BL trap" collectionCode="FSCA" collectorName="M. C. Thomas" country="Curaçao" latitude="12.344619" location="Christoffelberg" pageId="9" pageNumber="310" specimenCount="1" typeStatus="holotype">
<emphasis id="65AFD829DE0FFF80FF041F6FFEF40F00" bold="true" box="[199,371,584,611]" pageId="9" pageNumber="310">Type material</emphasis>
.
<typeStatus id="8860BA99DE0FFF80FE441F6FFE680F00" box="[391,495,585,611]" pageId="9" pageNumber="310" type="holotype">Holotype</typeStatus>
male, labeled “
<collectingCountry id="2FCC44ABDE0FFF80FD6E1F6FFC970F00" box="[685,784,585,611]" name="Curaçao" pageId="9" pageNumber="310">Curaçao</collectingCountry>
, Christoffel Nat. Pk.,
<location id="520452E0DE0FFF80FBD91F6FFB430F00" LSID="urn:lsid:plazi:treatment:DF72B52DDE01FF80FF541905FE2E0954:520452E0DE0FFF80FBD91F6FFB430F00" box="[1050,1220,585,611]" country="Curaçao" latitude="12.344619" name="Christoffelberg" pageId="9" pageNumber="310">Christoffelberg</location>
trailhead parking,
<geoCoordinate id="32EF62FCDE0FFF80FF541F4AFD8D0FE4" box="[151,522,620,647]" pageId="9" pageNumber="310">12°20ʹ40.63ʹʹ N, 69°67ʹ11.38ʹʹ W</geoCoordinate>
,
<date id="236522FBDE0FFF80FDD61F4BFD090FE4" box="[533,654,621,647]" pageId="9" pageNumber="310" value="2014-11-09">
<collectingDate id="3321DB13DE0FFF80FDD61F4BFD090FE4" box="[533,654,621,647]" pageId="9" pageNumber="310" value="2014-11-09">9-XI-2014</collectingDate>
</date>
,
<collectorName id="FA2E61EDDE0FFF80FD5A1F4BFCBA0FE4" box="[665,829,621,647]" pageId="9" pageNumber="310">M. C. Thomas</collectorName>
,
<collectingMethod id="8E9A7C2CDE0FFF80FC8B1F4BFC270FE4" box="[840,928,621,647]" pageId="9" pageNumber="310">BL trap</collectingMethod>
” (deposited in
<collectionCode id="31CA9CFEDE0FFF80FB8A1F4BFB130FE4" box="[1097,1172,621,647]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="9" pageNumber="310" type="Museum">FSCA</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="5764043BDE0FFF80FF041FB6FF530FAC" blockId="9.[151,1437,153,1079]" pageId="9" pageNumber="310">
<emphasis id="65AFD829DE0FFF80FF041FB6FDAE0FC8" bold="true" box="[199,553,656,683]" pageId="9" pageNumber="310">Derivation of specific epithet</emphasis>
. This species is named for Alicia Díaz, sister of the 2nd author, Florentina Díaz.
</paragraph>
</subSubSection>
<subSubSection id="1FC157B0DE0FFF80FF041FFEFE2E0954" pageId="9" pageNumber="310" type="discussion">
<paragraph id="5764043BDE0FFF80FF041FFEFE2E0954" blockId="9.[151,1437,153,1079]" pageId="9" pageNumber="310">
<emphasis id="65AFD829DE0FFF80FF041FFEFEC20F90" bold="true" box="[199,325,728,755]" pageId="9" pageNumber="310">Discussion</emphasis>
. This new species is similar to the
<typeStatus id="8860BA99DE0FFF80FD1D1FFFFCC70F90" box="[734,832,729,755]" pageId="9" pageNumber="310" type="holotype">holotype</typeStatus>
male of the recently described
<taxonomicName id="90DB7FB8DE0FFF80FB681FFCFE8E0E7B" authority="Grogan et al. (2016)" authorityName="Grogan" authorityYear="2016" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="waughi">
<emphasis id="65AFD829DE0FFF80FB681FFCFB380F90" box="[1195,1215,730,755]" italics="true" pageId="9" pageNumber="310">D</emphasis>
.
<emphasis id="65AFD829DE0FFF80FB131FFFFAA50F90" box="[1232,1314,729,755]" italics="true" pageId="9" pageNumber="310">waughi</emphasis>
<bibRefCitation id="334A79CADE0FFF80FAE81FFFFE8E0E7B" author="Grogan, W. L. Jr. &amp; Diaz, F. &amp; Spinelli, G. R. &amp; Ronderos, M. M." pageId="9" pageNumber="310" pagination="201 - 254" refId="ref19484" refString="Grogan, W. L. Jr., Diaz, F., Spinelli, G. R. &amp; Ronderos, M. M. (2016) The biting and predaceous midges of Guadeloupe (Diptera: Ceratopogonidae). II. Species of the subfamily Dasyheleinae. Zootaxa, 4184 (2), 201 - 254." type="journal article" year="2016">
Grogan
<emphasis id="65AFD829DE0FFF80FA4B1FFCFF2A0E74" italics="true" pageId="9" pageNumber="310">et al</emphasis>
. (2016)
</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="2FCC44ABDE0FFF80FE911FDBFE5E0E74" box="[338,473,765,791]" name="Guadeloupe" pageId="9" pageNumber="310">Guadeloupe</collectingCountry>
which also has a stout H-shaped aedeagus; asymmetrical gonocoxal apodemes and paramere; dark brown scutum, femora and tibiae; wing with a moderately developed anal lobe; and a clypeus with 5 pairs of setae. However, the male of
<taxonomicName id="90DB7FB8DE0FFF80FD871E60FD3F0E3C" box="[580,696,837,863]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="waughi">
<emphasis id="65AFD829DE0FFF80FD871E60FDDF0E3C" box="[580,600,838,863]" italics="true" pageId="9" pageNumber="310">D</emphasis>
.
<emphasis id="65AFD829DE0FFF80FDA51E63FD3F0E3C" box="[614,696,837,863]" italics="true" pageId="9" pageNumber="310">waughi</emphasis>
</taxonomicName>
differs from this new species in having a wing with a longer costa (costal ratio 0.50 vs.
<quantity id="9023A9DEDE0FFF80FE421E4FFE480EE0" box="[385,463,873,899]" metricMagnitude="-2" metricUnit="m" metricValue="1.0668" pageId="9" pageNumber="310" unit="in" value="0.42">0.42 in</quantity>
<taxonomicName id="90DB7FB8DE0FFF80FE151E4CFDC30EE0" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[470,580,873,899]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="aliciae" status="sp. nov.">
<emphasis id="65AFD829DE0FFF80FE151E4CFE6D0EE0" box="[470,490,874,899]" italics="true" pageId="9" pageNumber="310">D</emphasis>
.
<emphasis id="65AFD829DE0FFF80FE3B1E4FFDC30EE0" box="[504,580,873,899]" italics="true" pageId="9" pageNumber="310">aliciae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE0FFF80FD881E4FFD0E0EE0" box="[587,649,873,899]" pageId="9" pageNumber="310" rank="species">n. sp.</taxonomicNameLabel>
), a longer sternite 9 that is narrowly tapered distally, an aedeagus with a broadly concave basal arch and the medio-distal margin has 3 small teeth, and, the apical margin of tergite 9 is more narrowly tapered and the apicolateral processes are parallel, not divergent. In addition, in the
<typeStatus id="8860BA99DE0FFF80FB3B1E97FADD0EA8" box="[1272,1370,945,971]" pageId="9" pageNumber="310" type="holotype">holotype</typeStatus>
of
<taxonomicName id="90DB7FB8DE0FFF80FA421E94FF6E0E8C" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="waughi">
<emphasis id="65AFD829DE0FFF80FA421E94FA120EA8" box="[1409,1429,946,971]" italics="true" pageId="9" pageNumber="310">D</emphasis>
.
<emphasis id="65AFD829DE0FFF80FF541EF3FF6E0E8C" box="[151,233,981,1007]" italics="true" pageId="9" pageNumber="310">waughi</emphasis>
</taxonomicName>
, the right gonocoxal apodeme is only narrowly fused with the base of the paramere, but the basal portion of the paramere is much narrower, and the recurved apical section is shaped like a willow leaf and is much broader than in this new species.
</paragraph>
</subSubSection>
</treatment>
</document>