560 lines
71 KiB
XML
560 lines
71 KiB
XML
<document ID-DOI="10.18590/euscorpius.2016.vol2016.iss220.1" ID-GBIF-Dataset="03196918-10d5-44d2-ab42-d38f88314c7b" ID-ISSN="1536-9307" ID-Zenodo-Dep="7124466" checkinTime="1638249082081" checkinUser="felipe" docAuthor="Kovařík, František, Lowe, Graeme, Ranawana, Kithsiri B., Hoferek, David & Š, V. A." docDate="2016" docId="B07187DFA87BFFE0FEE1FAB89FB4F90F" docLanguage="en" docName="Euscorpius.220.1-133.pdf" docOrigin="Euscorpius 220" docSource="https://doi.org/10.18590/euscorpius.2016.vol2016.iss220.1" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Heterometrus serratus : Takashima 1945" docType="treatment" docVersion="6" lastPageNumber="111" masterDocId="4C48FFA7A817FF90FFC0FFD79E62FFBD" masterDocTitle="Scorpions of Sri Lanka (Scorpiones Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat n" masterLastPageNumber="133" masterPageNumber="1" pageNumber="107" updateTime="1664453448004" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Scorpions of Sri Lanka (Scorpiones Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat n</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kovařík, František</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lowe, Graeme</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ranawana, Kithsiri B.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hoferek, David</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Š, V. A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Euscorpius</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2016-12-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>220</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>133</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>https://doi.org/10.18590/euscorpius.2016.vol2016.iss220.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.18590/euscorpius.2016.vol2016.iss220.1</mods:identifier>
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<mods:identifier type="GBIF-Dataset">03196918-10d5-44d2-ab42-d38f88314c7b</mods:identifier>
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<mods:identifier type="ISSN">1536-9307</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7124466</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7124598" ID-Zenodo-Dep="7124598" LSID="urn:lsid:plazi:treatment:B07187DFA87BFFE0FEE1FAB89FB4F90F" httpUri="http://treatment.plazi.org/id/B07187DFA87BFFE0FEE1FAB89FB4F90F" lastPageId="112" lastPageNumber="111" pageId="108" pageNumber="107">
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<subSubSection box="[289,705,1391,1414]" pageId="108" pageNumber="107" type="nomenclature">
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||
<paragraph blockId="108.[289,705,1391,1445]" box="[289,705,1391,1414]" pageId="108" pageNumber="107">
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<heading box="[289,705,1391,1414]" centered="true" fontSize="10" level="2" pageId="108" pageNumber="107" reason="4">
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<taxonomicName authority="(Pocock, 1900)" authorityName=": Takashima" authorityYear="1945" baseAuthorityName="Pocock" baseAuthorityYear="1900" box="[289,705,1391,1414]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="108" pageNumber="107" phylum="Arthropoda" rank="species" species="serratus">
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<emphasis bold="true" box="[289,535,1391,1414]" italics="true" pageId="108" pageNumber="107">Heterometrus serratus</emphasis>
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(
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<bibRefCitation author="Pocock" bookContentInfo="279 pp" box="[551,698,1391,1414]" firstAuthor="Pocock" journalOrPublisher="London: W. T. Blandford" pageId="108" pageNumber="107" part="xii" refId="ref43790" refString="POCOCK, R. I. 1900. Arachnida. The fauna of British India, including Ceylon and Burma. Published un- der the authority of the Secretary of State for India in Council. London: W. T. Blandford, xii, 279 pp" title="Arachnida. The fauna of British India, including Ceylon and Burma. Published un- der the authority of the Secretary of State for India in Council" type="book" year="1900">Pocock, 1900</bibRefCitation>
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)
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||
</taxonomicName>
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||
</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection box="[338,655,1422,1445]" pageId="108" pageNumber="107" type="description">
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||
<paragraph blockId="108.[289,705,1391,1445]" box="[338,655,1422,1445]" pageId="108" pageNumber="107">
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||
(
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<figureCitation box="[347,433,1422,1445]" captionStart="Figures 12–15" captionStartId="6.[192,278,1885,1909]" captionTargetBox="[191,1440,167,1856]" captionTargetId="figure-11@6.[192,1439,168,1855]" captionTargetPageId="6" captionText="Figures 12–15: Maps showing the known geographical distribution of the Sri Lankan species." figureDoi="http://doi.org/10.5281/zenodo.7124476" httpUri="https://zenodo.org/record/7124476/files/figure.png" pageId="108" pageNumber="107">Figs. 15</figureCitation>
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,
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<figureCitation box="[446,540,1422,1445]" captionStart-0="Figures 504–511" captionStart-1="Figures 512–515" captionStart-2="Figures 516–519" captionStart-3="Figures 520–525" captionStart-4="Figures 526–529" captionStart-5="Figures 530–534" captionStartId-0="104.[192,278,1369,1393]" captionStartId-1="105.[192,278,1618,1642]" captionStartId-2="106.[192,278,1829,1853]" captionStartId-3="107.[192,278,1460,1484]" captionStartId-4="108.[192,278,1026,1050]" captionStartId-5="109.[192,278,1865,1889]" captionTargetBox-0="[195,1433,177,1322]" captionTargetBox-1="[204,1432,173,1576]" captionTargetBox-2="[213,1417,180,1801]" captionTargetBox-3="[197,1430,202,1414]" captionTargetBox-4="[199,1425,181,994]" captionTargetBox-5="[191,1440,191,1836]" captionTargetId-0="figure-11@104.[192,1439,168,1340]" captionTargetId-1="figure-6@105.[192,1439,168,1589]" captionTargetId-2="figure-47@106.[214,1416,181,1800]" captionTargetId-3="figure-6@107.[192,1439,192,1431]" captionTargetId-4="figure-11@108.[192,1439,168,997]" captionTargetId-5="figure-12@109.[192,1439,192,1836]" captionTargetPageId-0="104" captionTargetPageId-1="105" captionTargetPageId-2="106" captionTargetPageId-3="107" captionTargetPageId-4="108" captionTargetPageId-5="109" captionText-0="Figures 504–511: Pedipalp chela. Figures 504–507: Heterometrus indus from locality 15CA, pedipalp chela dorsal (504) view of female, and pedipalp chela dorsal (505), external (506) and ventral (507) views of male. Figures 508–511: Heterometrus serratus from locality 15CP, pedipalp chela dorsal (508) view of female, and pedipalp chela dorsal (509), external (510) and ventral (511) views of male." captionText-1="Figures 512–515: Heterometrus serratus from locality 15CP. Figures 512–513. Male in dorsal (512) and ventral (513) views. Figures 514–515. Female, dorsal (514) and ventral (515) views. Scale bar: 10 mm." captionText-2="Figures 516–519: Heterometrus serratus from locality 15CP. Figures 516, 518. Male, chelicerae, carapace and tergites I–II (516), and sternopectinal region (518). Figures 517, 519. Female, chelicerae, carapace and tergites I–II (517), and sternopectinal region (519)." captionText-3="Figures 520–525: Heterometrus serratus from locality 15CP. Figures 520–522. Male, metasoma and telson, lateral (520), ventral (521), and dorsal (522) views. Figures 523–525. Female, metasoma and telson, lateral (523), ventral (524), and dorsal (525) views. Scale bar: 10 mm." captionText-4="Figures 526–529: Heterometrus serratus, female from locality 15CP, tarsomeres of legs I–IV, ventral view." captionText-5="Figures 530–534: Heterometrus serratus. Figures 530–533. Juvenile from locality 15CQ in phases of ecdysis. Figures 534. Female at locality 15CP." figureDoi-0="http://doi.org/10.5281/zenodo.7124738" figureDoi-1="http://doi.org/10.5281/zenodo.7124740" figureDoi-2="http://doi.org/10.5281/zenodo.7124742" figureDoi-3="http://doi.org/10.5281/zenodo.7124746" figureDoi-4="http://doi.org/10.5281/zenodo.7124750" figureDoi-5="http://doi.org/10.5281/zenodo.7124752" httpUri-0="https://zenodo.org/record/7124738/files/figure.png" httpUri-1="https://zenodo.org/record/7124740/files/figure.png" httpUri-2="https://zenodo.org/record/7124742/files/figure.png" httpUri-3="https://zenodo.org/record/7124746/files/figure.png" httpUri-4="https://zenodo.org/record/7124750/files/figure.png" httpUri-5="https://zenodo.org/record/7124752/files/figure.png" pageId="108" pageNumber="107">508–534</figureCitation>
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,
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<figureCitation box="[553,648,1422,1445]" captionStart="Figures 571–574" captionStartId="116.[192,278,786,810]" captionTargetBox="[195,1423,184,735]" captionTargetId="figure-11@116.[192,1439,168,757]" captionTargetPageId="116" captionText="Figures 571–574: Key characters for distinguishing Sri Lankan Heterometrus. Figures 571–572. Pedipalp chela dorsal view of male, Heterometrus serratus from locality 15CP (571) and Heterometrus gravimanus from locality 15CE (572). Figures 573– 574. Metasomal segments IV-II lateral view of female, Heterometrus indus from locality 15CA (573) and Heterometrus serratus from locality 15CP (574)." figureDoi="http://doi.org/10.5281/zenodo.7124769" httpUri="https://zenodo.org/record/7124769/files/figure.png" pageId="108" pageNumber="107">571, 574</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="108" pageNumber="107" type="reference_group">
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<paragraph blockId="108.[192,802,1489,1757]" box="[192,658,1489,1512]" pageId="108" pageNumber="107">
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<treatmentCitationGroup box="[192,658,1489,1512]" pageId="108" pageNumber="107">
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<treatmentCitation author="POCOCK, R. I." box="[192,611,1489,1512]" page="86" pageId="108" pageNumber="107" year="1900">
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<taxonomicName authority="Pocock, 1900: 86" authorityName="Pocock" authorityPageNumber="86" authorityYear="1900" box="[192,611,1489,1512]" class="Arachnida" family="Scorpionidae" genus="Palamnaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="108" pageNumber="107" phylum="Arthropoda" rank="species" species="serratus">
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<emphasis box="[192,418,1489,1512]" italics="true" pageId="108" pageNumber="107">Palamnaeus serratus</emphasis>
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<bibRefCitation author="Pocock" bookContentInfo="279 pp" box="[425,611,1489,1512]" firstAuthor="Pocock" journalOrPublisher="London: W. T. Blandford" page="86" pageId="108" pageNumber="107" part="xii" refId="ref43790" refString="POCOCK, R. I. 1900. Arachnida. The fauna of British India, including Ceylon and Burma. Published un- der the authority of the Secretary of State for India in Council. London: W. T. Blandford, xii, 279 pp" title="Arachnida. The fauna of British India, including Ceylon and Burma. Published un- der the authority of the Secretary of State for India in Council" type="book" year="1900">Pocock, 1900: 86</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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, 97.
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</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph blockId="108.[192,802,1489,1757]" box="[192,678,1520,1543]" pageId="108" pageNumber="107">
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||
<treatmentCitationGroup box="[192,678,1520,1543]" pageId="108" pageNumber="107">
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<treatmentCitation author="TAKASHIMA, H." box="[192,672,1520,1543]" page="94" pageId="108" pageNumber="107" year="1945">
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<taxonomicName authority=": Takashima, 1945: 94" authorityName=": Takashima" authorityPageNumber="94" authorityYear="1945" baseAuthorityName="Pocock" baseAuthorityYear="1900" box="[192,672,1520,1543]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="108" pageNumber="107" phylum="Arthropoda" rank="species" species="serratus">
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<emphasis box="[192,433,1520,1543]" italics="true" pageId="108" pageNumber="107">Heterometrus serratus</emphasis>
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:
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<bibRefCitation author="Takashima" box="[447,672,1520,1543]" firstAuthor="Takashima" journalOrPublisher="Acta Arachnologica, Tokyo" page="94" pageId="108" pageNumber="107" pagination="68 - 106" part="9" refId="ref44598" refString="TAKASHIMA, H. 1945. Scorpions of Eastern Asia. Acta Arachnologica, Tokyo, 9: 68 - 106." title="Scorpions of Eastern Asia" type="journal article" year="1945">Takashima, 1945: 94</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="108.[192,802,1489,1757]" pageId="108" pageNumber="107">
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<treatmentCitationGroup pageId="108" pageNumber="107">
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<treatmentCitation author="TIKADER B. K. & D. B. BASTAWADE" page="561" pageId="108" pageNumber="107" year="1983">
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<taxonomicName authority=": Tikader & Bastawade, 1983: 555 - 561" authorityName=": Tikader & Bastawade" authorityPageNumber="555 - 561" authorityYear="1983" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="108" pageNumber="107" phylum="Arthropoda" rank="species" species="serratus" subGenus="Srilankametrus">
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<emphasis box="[192,339,1550,1573]" italics="true" pageId="108" pageNumber="107">Heterometrus</emphasis>
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(
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<emphasis box="[368,531,1550,1573]" italics="true" pageId="108" pageNumber="107">Srilankametrus</emphasis>
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)
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<emphasis box="[561,649,1550,1573]" italics="true" pageId="108" pageNumber="107">serratus</emphasis>
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:
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<bibRefCitation author="Tikader & Bastawade" bookContentInfo="668 pp." firstAuthor="Tikader" journalOrPublisher="Zoological Survey of India, Calcutta" page="555 - 561" pageId="108" pageNumber="107" refId="ref44789" refString="TIKADER B. K. & D. B. BASTAWADE. 1983. The Fauna of India: Scorpions. Scorpionida, Arachnida. Vol III. Zoological Survey of India, Calcutta, 668 pp." title="The Fauna of India: Scorpions. Scorpionida, Arachnida. Vol III" type="book" year="1983">Tikader & Bastawade, 1983: 555–561</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, figs. 1483–1496;
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<treatmentCitation author="FET" page="448" pageId="108" pageNumber="107" year="2000">
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<bibRefCitation author="Fet" firstAuthor="Fet" page="448" pageId="108" pageNumber="107" refId="ref40873" refString="FET, V., W. D. SISSOM, G. LOWE & M. E. BRAUNWALDER. 2000. Catalog of the Scorpions of the World (1758 - 1998). New York: The New York Entomological Society, 689 pp." type="book" year="2000">Fet, 2000: 448</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="108.[192,802,1489,1757]" pageId="108" pageNumber="107">
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<taxonomicName authority="indus" authorityName="indus: Couzijn" authorityYear="1981" box="[192,470,1642,1665]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="108" pageNumber="107" phylum="Arthropoda" rank="subSpecies" species="indus" subSpecies="indus">
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<emphasis box="[192,470,1642,1665]" italics="true" pageId="108" pageNumber="107">Heterometrus indus indus</emphasis>
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</taxonomicName>
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(in part):
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<bibRefCitation author="COUZIJN, H. W. C." journalOrPublisher="Zoologische Verhandelingen" pageId="108" pageNumber="107" pagination="1 - 196" part="184" refId="ref40696" refString="COUZIJN, H. W. C. 1981. Revision of the genus Heterometrus. Zoologische Verhandelingen, 184: 1 - 196." title="Revision of the genus Heterometrus" type="journal article" year="1981">Couzijn, 1981: 121– 123</bibRefCitation>
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, fig. 35 (in part).
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</paragraph>
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<paragraph blockId="108.[192,802,1489,1757]" pageId="108" pageNumber="107">
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<treatmentCitationGroup pageId="108" pageNumber="107">
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<taxonomicName authorityName=": Fet" authorityYear="2000" baseAuthorityName="De Geer" baseAuthorityYear="1778" box="[192,404,1704,1727]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="108" pageNumber="107" phylum="Arthropoda" rank="species" species="indus">
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<emphasis box="[192,404,1704,1727]" italics="true" pageId="108" pageNumber="107">Heterometrus indus</emphasis>
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</taxonomicName>
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(in part):
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<treatmentCitation author="KOVARIK" box="[515,751,1704,1727]" page="17" pageId="108" pageNumber="107" year="2004">
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<bibRefCitation author="Kovarik" box="[515,751,1704,1727]" firstAuthor="Kovarik" page="17 - 20" pageId="108" pageNumber="107" pagination="1 - 60" refId="ref41757" refString="KOVARIK, F. 2004. A review of the genus Heterometrus Ehrenberg, 1828, with descriptions of seven new species (Scorpiones, Scorpionidae). Euscorpius, 15: 1 - 60." type="journal article" year="2004">Kovařík, 2004: 17–20</bibRefCitation>
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</treatmentCitation>
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, fig. 16;
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<treatmentCitation author="KOVARIK, F." box="[281,476,1734,1757]" page="38" pageId="108" pageNumber="107" year="2009">
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||
<bibRefCitation author="Kovarik" bookContentInfo="170 pp." box="[281,476,1734,1757]" firstAuthor="Kovarik" journalOrPublisher="Clairon Production, Prague" page="38" pageId="108" pageNumber="107" refId="ref41793" refString="KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Clairon Production, Prague, 170 pp." title="Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species" type="book" year="2009">Kovařík, 2009: 38</bibRefCitation>
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</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="108" pageNumber="107" type="materials_examined">
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||
<paragraph blockId="108.[192,802,1802,1856]" pageId="108" pageNumber="107">
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||
T YPE LOCALITY AND
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<typeStatus box="[421,473,1806,1825]" pageId="108" pageNumber="107">TYPE</typeStatus>
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REPOSITORY.
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||
<collectingCountry box="[625,705,1802,1825]" name="Sri Lanka" pageId="108" pageNumber="107">Ceylon</collectingCountry>
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, now
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<collectingCountry name="Sri Lanka" pageId="108" pageNumber="107">Sri Lanka</collectingCountry>
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; BMNH.
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||
</paragraph>
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||
<paragraph blockId="108.[830,1440,1103,1187]" pageId="108" pageNumber="107">
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||
TYPE MATERIAL EXAMINED.
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||
<materialsCitation collectionCode="BMNH" collectorName="Dr. Ondaatje" country="Sri Lanka" location="Ceylon" municipality="Ceylon" pageId="108" pageNumber="107" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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||
<collectingCountry box="[1140,1247,1103,1126]" name="Sri Lanka" pageId="108" pageNumber="107">Sri Lanka</collectingCountry>
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||
,
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||
<collectingMunicipality box="[1262,1342,1103,1126]" pageId="108" pageNumber="107">Ceylon</collectingMunicipality>
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||
,
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||
<specimenCount box="[1356,1387,1103,1126]" pageId="108" pageNumber="107" type="male">1♂</specimenCount>
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||
(
|
||
<typeStatus pageId="108" pageNumber="107">holotype</typeStatus>
|
||
, fig.
|
||
<quantity box="[959,1016,1134,1157]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="108" pageNumber="107" unit="in" value="16.0">16 in</quantity>
|
||
<bibRefCitation author="Kovarik" box="[1024,1222,1134,1157]" firstAuthor="Kovarik" page="19" pageId="108" pageNumber="107" pagination="1 - 60" refId="ref41757" refString="KOVARIK, F. 2004. A review of the genus Heterometrus Ehrenberg, 1828, with descriptions of seven new species (Scorpiones, Scorpionidae). Euscorpius, 15: 1 - 60." type="journal article" year="2004">Kovařík, 2004: 19</bibRefCitation>
|
||
), leg.
|
||
<collectorName box="[1296,1433,1134,1157]" pageId="108" pageNumber="107">Dr. Ondaatje</collectorName>
|
||
,
|
||
<collectionCode box="[832,910,1164,1187]" pageId="108" pageNumber="107">BMNH</collectionCode>
|
||
No. 1888.55.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="108.[830,1442,1222,1491]" pageId="108" pageNumber="107">
|
||
OTHER MATERIAL EXAMINED.
|
||
<materialsCitation collectingDate="2015-04-30" collectionCode="FKCP, UPSL" collectorName="Kovarik" country="Sri Lanka" elevation="40" latitude="5.9745" location="Kekanadura village" longLatPrecision="1" longitude="80.60569" municipality="Matara District" pageId="108" pageNumber="107" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Southern">
|
||
<collectingCountry box="[1197,1319,1222,1245]" name="Sri Lanka" pageId="108" pageNumber="107">Sri Lanka</collectingCountry>
|
||
,
|
||
<collectingRegion country="Sri Lanka" name="Southern" pageId="108" pageNumber="107">Southern Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality box="[952,1130,1253,1276]" pageId="108" pageNumber="107">Matara District</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:B07187DFA87BFFE0FEE1FAB89FB4F90F:3D076012A87BFFFCFB42FB329B02FB41" box="[1154,1376,1253,1276]" country="Sri Lanka" latitude="5.9745" longLatPrecision="1" longitude="80.60569" municipality="Matara District" name="Kekanadura village" pageId="108" pageNumber="107" stateProvince="Southern">Kekanadura village</location>
|
||
,
|
||
<geoCoordinate degrees="05" direction="north" minutes="58" orientation="latitude" pageId="108" pageNumber="107" precision="1" seconds="28.2" value="5.9745">05° 58'28.2"N</geoCoordinate>
|
||
<geoCoordinate box="[951,1110,1284,1307]" degrees="080" direction="east" minutes="36" orientation="longitude" pageId="108" pageNumber="107" precision="1" seconds="20. 5" value="80.60569">080°36'20.5"E</geoCoordinate>
|
||
,
|
||
<elevation box="[1126,1248,1284,1307]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="108" pageNumber="107" unit="m" value="40.0">
|
||
<quantity box="[1126,1187,1284,1307]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="108" pageNumber="107" unit="m" value="40.0">40 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
(Locality
|
||
<emphasis bold="true" box="[1371,1434,1283,1307]" pageId="108" pageNumber="107">15CP</emphasis>
|
||
,
|
||
<figureCitation box="[830,926,1314,1337]" captionStart="Figures 592–594" captionStartId="126.[192,278,1617,1641]" captionTargetBox="[213,1417,167,1588]" captionTargetId="figure-251@126.[214,1416,168,1587]" captionTargetPageId="126" captionText="Figures 592–594: Figure 592. Locality 15CO, Sri Lanka, North Central Province, Puttalam District, Eluwankulam, 08°17'15"N 079°50'38.7"E, 38 m a.s.l. During night collecting from 10.30 p.m. on 28.IV.2015 to 4.00 a.m. on 29.IV.2015 (UV detection) we recorded Charmus laneus (running on the ground among leaves), Isometrus thwaitesii (running on branches of bushes, also sitting on leaves), Lychas srilankensis (on the ground under branches, leaves or in their vicinity), Reddyanus ceylonensis sp. n. (on the ground), Chaerilus ceylonensis (inside oblique usually not curved burrows 15 to 30 cm long) and Heterometrus swammerdami (inside long burrows). On the locality we recorded between 10.00 p.m. and 4.00 a.m. temperature 25.9 ºC and humidity between 58% and 70%. On the locality we recorded 6 scorpion species and genera of all three Sri Lankan families. Figures 593–594. Locality 15CP, Sri Lanka, Southern Province, Matara District, Kekanadura village, 05°58'28.2"N 080°36'20.5"E, 40 m a.s.l. During night collecting on 30.IV.2015 (UV detection) we recorded Isometrus maculatus (inside a house, the species occupy the roof of the house, Fig. 593) and Heterometrus serratus (inside almost straight burrows in a ground wall). On the locality we recorded night temperature 28.4 ºC– 25.2 ºC (minimum temperature) and night humidity varied between 63% and 89%." figureDoi="http://doi.org/10.5281/zenodo.7124791" httpUri="https://zenodo.org/record/7124791/files/figure.png" pageId="108" pageNumber="107">Fig. 594</figureCitation>
|
||
),
|
||
<date box="[955,1077,1314,1337]" pageId="108" pageNumber="107" value="2015-04-30">
|
||
<collectingDate box="[955,1077,1314,1337]" pageId="108" pageNumber="107" value="2015-04-30">30.IV.2015</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount box="[1097,1131,1314,1337]" pageId="108" pageNumber="107" type="male">1♂</specimenCount>
|
||
(
|
||
<figureCitation box="[1153,1313,1314,1337]" captionStart-0="Figures 504–511" captionStart-1="Figures 512–515" captionStartId-0="104.[192,278,1369,1393]" captionStartId-1="105.[192,278,1618,1642]" captionTargetBox-0="[195,1433,177,1322]" captionTargetBox-1="[204,1432,173,1576]" captionTargetId-0="figure-11@104.[192,1439,168,1340]" captionTargetId-1="figure-6@105.[192,1439,168,1589]" captionTargetPageId-0="104" captionTargetPageId-1="105" captionText-0="Figures 504–511: Pedipalp chela. Figures 504–507: Heterometrus indus from locality 15CA, pedipalp chela dorsal (504) view of female, and pedipalp chela dorsal (505), external (506) and ventral (507) views of male. Figures 508–511: Heterometrus serratus from locality 15CP, pedipalp chela dorsal (508) view of female, and pedipalp chela dorsal (509), external (510) and ventral (511) views of male." captionText-1="Figures 512–515: Heterometrus serratus from locality 15CP. Figures 512–513. Male in dorsal (512) and ventral (513) views. Figures 514–515. Female, dorsal (514) and ventral (515) views. Scale bar: 10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7124738" figureDoi-1="http://doi.org/10.5281/zenodo.7124740" httpUri-0="https://zenodo.org/record/7124738/files/figure.png" httpUri-1="https://zenodo.org/record/7124740/files/figure.png" pageId="108" pageNumber="107">Figs. 509–513</figureCitation>
|
||
,
|
||
<figureCitation box="[1333,1433,1314,1337]" captionStart="Figures 516–519" captionStartId="106.[192,278,1829,1853]" captionTargetBox="[213,1417,180,1801]" captionTargetId="figure-47@106.[214,1416,181,1800]" captionTargetPageId="106" captionText="Figures 516–519: Heterometrus serratus from locality 15CP. Figures 516, 518. Male, chelicerae, carapace and tergites I–II (516), and sternopectinal region (518). Figures 517, 519. Female, chelicerae, carapace and tergites I–II (517), and sternopectinal region (519)." figureDoi="http://doi.org/10.5281/zenodo.7124742" httpUri="https://zenodo.org/record/7124742/files/figure.png" pageId="108" pageNumber="107">516, 518</figureCitation>
|
||
,
|
||
<figureCitation box="[830,924,1345,1368]" captionStart="Figures 520–525" captionStartId="107.[192,278,1460,1484]" captionTargetBox="[197,1430,202,1414]" captionTargetId="figure-6@107.[192,1439,192,1431]" captionTargetPageId="107" captionText="Figures 520–525: Heterometrus serratus from locality 15CP. Figures 520–522. Male, metasoma and telson, lateral (520), ventral (521), and dorsal (522) views. Figures 523–525. Female, metasoma and telson, lateral (523), ventral (524), and dorsal (525) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7124746" httpUri="https://zenodo.org/record/7124746/files/figure.png" pageId="108" pageNumber="107">520–522</figureCitation>
|
||
,
|
||
<figureCitation box="[939,980,1345,1368]" captionStart="Figures 571–574" captionStartId="116.[192,278,786,810]" captionTargetBox="[195,1423,184,735]" captionTargetId="figure-11@116.[192,1439,168,757]" captionTargetPageId="116" captionText="Figures 571–574: Key characters for distinguishing Sri Lankan Heterometrus. Figures 571–572. Pedipalp chela dorsal view of male, Heterometrus serratus from locality 15CP (571) and Heterometrus gravimanus from locality 15CE (572). Figures 573– 574. Metasomal segments IV-II lateral view of female, Heterometrus indus from locality 15CA (573) and Heterometrus serratus from locality 15CP (574)." figureDoi="http://doi.org/10.5281/zenodo.7124769" httpUri="https://zenodo.org/record/7124769/files/figure.png" pageId="108" pageNumber="107">571</figureCitation>
|
||
)
|
||
<specimenCount box="[997,1031,1345,1368]" pageId="108" pageNumber="107" type="female">1♀</specimenCount>
|
||
(
|
||
<figureCitation box="[1049,1153,1345,1368]" captionStart="Figures 504–511" captionStartId="104.[192,278,1369,1393]" captionTargetBox="[195,1433,177,1322]" captionTargetId="figure-11@104.[192,1439,168,1340]" captionTargetPageId="104" captionText="Figures 504–511: Pedipalp chela. Figures 504–507: Heterometrus indus from locality 15CA, pedipalp chela dorsal (504) view of female, and pedipalp chela dorsal (505), external (506) and ventral (507) views of male. Figures 508–511: Heterometrus serratus from locality 15CP, pedipalp chela dorsal (508) view of female, and pedipalp chela dorsal (509), external (510) and ventral (511) views of male." figureDoi="http://doi.org/10.5281/zenodo.7124738" httpUri="https://zenodo.org/record/7124738/files/figure.png" pageId="108" pageNumber="107">Figs. 508</figureCitation>
|
||
,
|
||
<figureCitation box="[1166,1262,1345,1368]" captionStart="Figures 512–515" captionStartId="105.[192,278,1618,1642]" captionTargetBox="[204,1432,173,1576]" captionTargetId="figure-6@105.[192,1439,168,1589]" captionTargetPageId="105" captionText="Figures 512–515: Heterometrus serratus from locality 15CP. Figures 512–513. Male in dorsal (512) and ventral (513) views. Figures 514–515. Female, dorsal (514) and ventral (515) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7124740" httpUri="https://zenodo.org/record/7124740/files/figure.png" pageId="108" pageNumber="107">514–515</figureCitation>
|
||
,
|
||
<figureCitation box="[1275,1374,1345,1368]" captionStart="Figures 516–519" captionStartId="106.[192,278,1829,1853]" captionTargetBox="[213,1417,180,1801]" captionTargetId="figure-47@106.[214,1416,181,1800]" captionTargetPageId="106" captionText="Figures 516–519: Heterometrus serratus from locality 15CP. Figures 516, 518. Male, chelicerae, carapace and tergites I–II (516), and sternopectinal region (518). Figures 517, 519. Female, chelicerae, carapace and tergites I–II (517), and sternopectinal region (519)." figureDoi="http://doi.org/10.5281/zenodo.7124742" httpUri="https://zenodo.org/record/7124742/files/figure.png" pageId="108" pageNumber="107">517, 519</figureCitation>
|
||
,
|
||
<figureCitation captionStart-0="Figures 520–525" captionStart-1="Figures 526–529" captionStartId-0="107.[192,278,1460,1484]" captionStartId-1="108.[192,278,1026,1050]" captionTargetBox-0="[197,1430,202,1414]" captionTargetBox-1="[199,1425,181,994]" captionTargetId-0="figure-6@107.[192,1439,192,1431]" captionTargetId-1="figure-11@108.[192,1439,168,997]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionText-0="Figures 520–525: Heterometrus serratus from locality 15CP. Figures 520–522. Male, metasoma and telson, lateral (520), ventral (521), and dorsal (522) views. Figures 523–525. Female, metasoma and telson, lateral (523), ventral (524), and dorsal (525) views. Scale bar: 10 mm." captionText-1="Figures 526–529: Heterometrus serratus, female from locality 15CP, tarsomeres of legs I–IV, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.7124746" figureDoi-1="http://doi.org/10.5281/zenodo.7124750" httpUri-0="https://zenodo.org/record/7124746/files/figure.png" httpUri-1="https://zenodo.org/record/7124750/files/figure.png" pageId="108" pageNumber="107">523– 529</figureCitation>
|
||
,
|
||
<figureCitation box="[885,925,1376,1399]" captionStart="Figures 571–574" captionStartId="116.[192,278,786,810]" captionTargetBox="[195,1423,184,735]" captionTargetId="figure-11@116.[192,1439,168,757]" captionTargetPageId="116" captionText="Figures 571–574: Key characters for distinguishing Sri Lankan Heterometrus. Figures 571–572. Pedipalp chela dorsal view of male, Heterometrus serratus from locality 15CP (571) and Heterometrus gravimanus from locality 15CE (572). Figures 573– 574. Metasomal segments IV-II lateral view of female, Heterometrus indus from locality 15CA (573) and Heterometrus serratus from locality 15CP (574)." figureDoi="http://doi.org/10.5281/zenodo.7124769" httpUri="https://zenodo.org/record/7124769/files/figure.png" pageId="108" pageNumber="107">574</figureCitation>
|
||
),
|
||
<collectionCode box="[947,1014,1376,1399]" pageId="108" pageNumber="107">FKCP</collectionCode>
|
||
,
|
||
<specimenCount box="[1027,1061,1376,1399]" pageId="108" pageNumber="107" type="female">1♀</specimenCount>
|
||
im.,
|
||
<collectionCode box="[1109,1177,1376,1399]" pageId="108" pageNumber="107">UPSL</collectionCode>
|
||
, leg.
|
||
<collectorName box="[1235,1322,1376,1399]" pageId="108" pageNumber="107">Kovařík</collectorName>
|
||
et al
|
||
</materialsCitation>
|
||
.;
|
||
<materialsCitation collectingDate="2015-05-02" collectingDateMax="2015-05-03" collectingDateMin="2015-05-02" collectionCode="FKCP" collectorName="Kovarik" country="Sri Lanka" elevation="288" latitude="6.8751945" location="Monaragala" longLatPrecision="15" longitude="81.35472" municipality="Monaragala District" pageId="108" pageNumber="107" specimenCount="2" specimenCount-female="2" stateProvince="Uva">
|
||
<collectingRegion country="Sri Lanka" name="Uva" pageId="108" pageNumber="107">Uva Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality box="[955,1185,1406,1429]" pageId="108" pageNumber="107">Monaragala District</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:B07187DFA87BFFE0FEE1FAB89FB4F90F:3D076012A87BFFFCFB7FFAA99B5CFA28" box="[1215,1342,1406,1429]" country="Sri Lanka" latitude="6.8751945" longLatPrecision="15" longitude="81.35472" municipality="Monaragala District" name="Monaragala" pageId="108" pageNumber="107" stateProvince="Uva">Monaragala</location>
|
||
,
|
||
<geoCoordinate degrees="06" direction="north" minutes="52" orientation="latitude" pageId="108" pageNumber="107" precision="1" seconds="30.7" value="6.8751945">06°52' 30.7"N</geoCoordinate>
|
||
<geoCoordinate box="[917,1056,1437,1460]" degrees="081" direction="east" minutes="21" orientation="longitude" pageId="108" pageNumber="107" precision="15" seconds="17" value="81.35472">081°21'17"E</geoCoordinate>
|
||
,
|
||
<elevation box="[1069,1198,1437,1460]" metricMagnitude="2" metricUnit="m" metricValue="2.88" pageId="108" pageNumber="107" unit="m" value="288.0">
|
||
<quantity box="[1069,1140,1437,1460]" metricMagnitude="2" metricUnit="m" metricValue="2.88" pageId="108" pageNumber="107" unit="m" value="288.0">288 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
(Locality
|
||
<emphasis bold="true" box="[1315,1382,1436,1460]" pageId="108" pageNumber="107">15CQ</emphasis>
|
||
,
|
||
<figureCitation captionStart="Figures 595–596" captionStartId="127.[192,278,1661,1685]" captionTargetBox="[191,1440,195,1633]" captionTargetId="figure-82@127.[192,1439,196,1632]" captionTargetPageId="127" captionText="Figures 595–596: Figure 595. Locality 15CQ, Sri Lanka, Uva Province, Monaragala District, Monaragala, 06°52'30.7"N 081°21'17"E, 288 m a.s.l. During night collecting on 2.–3.V.2015 (UV detection) we recorded Reddyanus loebli (under scales of bark and under bark) and Heterometrus serratus (females with juveniles inside oblique burrows). On the locality we recorded a temperature of 33.1 ºC at 5.00 p.m., which gradually dropped to 24.6 ºC (minimum temperature) before sunrise. Humidity during the night varied between 61% and 84%. Figure 596. Locality 15CR, Sri Lanka, Eastern Province, Ampara District, Lahugala Kitulana National Park, 06°52'46"N 081°43'21.8"E, 40 m a.s.l. During night collecting on 3.–4.V.2015 (10.00 p.m. to 01.00 a.m.) we recorded Isometrus maculatus (running on branches, also sitting on leaves on trees), Reddyanus basilicus (on the ground among leaves) and Reddyanus loebli (hidden under scales of bark of standing tree trunks). On the locality we recorded between 10.00 p.m. on 3.V. and 01.00 a.m. on 4.V.2015 temperature 28 ºC, which gradually dropped to 26.9 ºC and humidity 73%." figureDoi="http://doi.org/10.5281/zenodo.7124794" httpUri="https://zenodo.org/record/7124794/files/figure.png" pageId="108" pageNumber="107">Fig. 595</figureCitation>
|
||
),
|
||
<date box="[893,1028,1468,1491]" pageId="108" pageNumber="107" value="2015-05-02" valueMax="2015-05-03" valueMin="2015-05-02">
|
||
<collectingDate box="[893,1028,1468,1491]" pageId="108" pageNumber="107" value="2015-05-02" valueMax="2015-05-03" valueMin="2015-05-02">2.–3.V.2015</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount box="[1038,1072,1468,1491]" pageId="108" pageNumber="107" type="female">2♀</specimenCount>
|
||
,
|
||
<collectionCode box="[1085,1152,1468,1491]" pageId="108" pageNumber="107">FKCP</collectionCode>
|
||
, leg.
|
||
<collectorName box="[1211,1299,1468,1491]" pageId="108" pageNumber="107">Kovařík</collectorName>
|
||
et al
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="111" lastPageNumber="110" pageId="108" pageNumber="107" type="diagnosis">
|
||
<paragraph blockId="108.[830,1441,1526,1856]" lastBlockId="111.[192,802,1452,1659]" lastPageId="111" lastPageNumber="110" pageId="108" pageNumber="107">
|
||
DIAGNOSIS. Total length
|
||
<quantity box="[1117,1264,1526,1549]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="108" pageNumber="107" unit="mm" value="115.0" valueMax="130.0" valueMin="100.0">100–130 mm</quantity>
|
||
long. Color of adults uniformly reddish black to greenish black. Pectinal teeth number
|
||
<quantity box="[1059,1163,1587,1610]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" metricValueMax="3.81" metricValueMin="3.048" pageId="108" pageNumber="107" unit="in" value="13.5" valueMax="15.0" valueMin="12.0">12–15 in</quantity>
|
||
both sexes. Male with slightly narrower chela than female; chela length/ width ratio 1.79–1.94 in males, 1.69 in female. Chela hirsute, lobiform, without carinae on dorsoexternal surface, but may bear rows of granules. Chela length/ width ratio 1.8–2.0 in adults. Entire manus covered by rounded granules that may merge and appear as rows. Pedipalp patella without pronounced internal tubercle. Carapace smooth, glossy medially, with granules at margins. Second metasomal segment approximately as long as wide. Fifth segment of metasoma about as long pedipalp femur, fourth segment of metasoma shorter than pedipalp femur. Dorsal and dorsolateral carinae of metasomal segments granulated. Vesicle of telson usually longer than aculeus. Spination formula of tarsomeres II of legs: 3-4/4-6: 4/4-5: 4/5-6: 4/5-7.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7124752" ID-Zenodo-Dep="7124752" httpUri="https://zenodo.org/record/7124752/files/figure.png" pageId="109" pageNumber="108" startId="109.[192,278,1865,1889]" targetBox="[191,1440,191,1836]" targetPageId="109">
|
||
<paragraph blockId="109.[192,1440,1865,1913]" pageId="109" pageNumber="108">
|
||
<emphasis bold="true" box="[192,389,1865,1889]" pageId="109" pageNumber="108">Figures 530–534:</emphasis>
|
||
<taxonomicName authorityName=": Tikader & Bastawade" authorityYear="1983" box="[399,616,1867,1888]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="109" pageNumber="108" phylum="Arthropoda" rank="species" species="serratus">
|
||
<emphasis box="[399,616,1867,1888]" italics="true" pageId="109" pageNumber="108">Heterometrus serratus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" box="[630,800,1867,1889]" pageId="109" pageNumber="108">Figures 530–533</emphasis>
|
||
. Juvenile from locality 15CQ in phases of ecdysis.
|
||
<emphasis bold="true" box="[1313,1434,1867,1889]" pageId="109" pageNumber="108">Figures 534</emphasis>
|
||
. Female at locality 15CP.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7124754" ID-Zenodo-Dep="7124754" httpUri="https://zenodo.org/record/7124754/files/figure.png" pageId="110" pageNumber="109" startId="110.[192,278,1349,1373]" targetBox="[213,1421,158,1315]" targetPageId="110">
|
||
<paragraph blockId="110.[192,1441,1349,1397]" pageId="110" pageNumber="109">
|
||
<emphasis bold="true" box="[192,387,1349,1373]" pageId="110" pageNumber="109">Figures 535–536:</emphasis>
|
||
<taxonomicName authorityName="Simon" authorityYear="1872" box="[394,673,1351,1372]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="110" pageNumber="109" phylum="Arthropoda" rank="species" species="swammerdami">
|
||
<emphasis box="[394,673,1351,1372]" italics="true" pageId="110" pageNumber="109">Heterometrus swammerdami</emphasis>
|
||
</taxonomicName>
|
||
from locality 15CB, female in dorsal (535) and ventral (536) views. Scale bar: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="110" pageNumber="109">
|
||
<table box="[383,1229,1455,1832]" gridcols="4" gridrows="9" pageId="110" pageNumber="109">
|
||
<tr box="[383,1229,1455,1509]" gridrow="0" pageId="110" pageNumber="109">
|
||
<th box="[383,696,1455,1509]" gridcol="0" gridrow="0" pageId="110" pageNumber="109">
|
||
<emphasis bold="true" box="[529,612,1455,1479]" pageId="110" pageNumber="109">Species</emphasis>
|
||
</th>
|
||
<th box="[796,824,1455,1509]" gridcol="1" gridrow="0" pageId="110" pageNumber="109">
|
||
<emphasis bold="true" box="[796,824,1455,1479]" pageId="110" pageNumber="109">2n</emphasis>
|
||
</th>
|
||
<th box="[861,1076,1455,1509]" gridcol="2" gridrow="0" pageId="110" pageNumber="109">
|
||
<emphasis bold="true" pageId="110" pageNumber="109">Relative Size of Chromosomes</emphasis>
|
||
</th>
|
||
<th box="[1135,1229,1455,1509]" gridcol="3" gridrow="0" pageId="110" pageNumber="109">
|
||
<emphasis bold="true" box="[1135,1229,1455,1479]" pageId="110" pageNumber="109">Locality</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[383,1229,1529,1552]" gridrow="1" pageId="110" pageNumber="109">
|
||
<th box="[383,696,1529,1552]" gridcol="0" gridrow="1" pageId="110" pageNumber="109">
|
||
<taxonomicName authorityName=": Fet" authorityYear="1997" box="[383,668,1529,1552]" class="Arachnida" family="Buthidae" genus="Buthoscorpio" kingdom="Animalia" order="Scorpiones" pageId="110" pageNumber="109" phylum="Arthropoda" rank="species" species="sarasinorum">
|
||
<emphasis box="[383,668,1529,1552]" italics="true" pageId="110" pageNumber="109">Buthoscorpio sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[796,824,1529,1552]" gridcol="1" gridrow="1" pageId="110" pageNumber="109">14</td>
|
||
<td box="[861,1076,1529,1552]" gridcol="2" gridrow="1" pageId="110" pageNumber="109">2x 11.2 + 7.4 – 5.7</td>
|
||
<td box="[1135,1229,1529,1552]" gridcol="3" gridrow="1" pageId="110" pageNumber="109">15CF</td>
|
||
</tr>
|
||
<tr box="[383,1229,1569,1592]" gridrow="2" pageId="110" pageNumber="109">
|
||
<th box="[383,696,1569,1592]" gridcol="0" gridrow="2" pageId="110" pageNumber="109">
|
||
<taxonomicName authorityName="Karsch" authorityYear="1879" box="[383,557,1569,1592]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="110" pageNumber="109" phylum="Arthropoda" rank="species" species="laneus">
|
||
<emphasis box="[383,557,1569,1592]" italics="true" pageId="110" pageNumber="109">Charmus laneus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[796,824,1569,1592]" gridcol="1" gridrow="2" pageId="110" pageNumber="109">9</td>
|
||
<td box="[861,1076,1569,1592]" gridcol="2" gridrow="2" pageId="110" pageNumber="109">19.2 + 12.9 – 6.7</td>
|
||
<td box="[1135,1229,1569,1592]" gridcol="3" gridrow="2" pageId="110" pageNumber="109">15CO</td>
|
||
</tr>
|
||
<tr box="[383,1229,1609,1632]" gridrow="3" pageId="110" pageNumber="109">
|
||
<th box="[383,696,1609,1632]" gridcol="0" gridrow="3" pageId="110" pageNumber="109">
|
||
<taxonomicName authorityName=": Vachon" authorityYear="1982" box="[383,591,1609,1632]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="110" pageNumber="109" phylum="Arthropoda" rank="species" species="thwaitesi">
|
||
<emphasis box="[383,591,1609,1632]" italics="true" pageId="110" pageNumber="109">Isometrus thwaitesi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[796,824,1609,1632]" gridcol="1" gridrow="3" pageId="110" pageNumber="109">8</td>
|
||
<td box="[861,1076,1609,1632]" gridcol="2" gridrow="3" pageId="110" pageNumber="109">1 6.4 – 13.0 + 2x 6.3</td>
|
||
<td box="[1135,1229,1609,1632]" gridcol="3" gridrow="3" pageId="110" pageNumber="109">15CF</td>
|
||
</tr>
|
||
<tr box="[383,1229,1649,1672]" gridrow="4" pageId="110" pageNumber="109">
|
||
<th box="[383,696,1649,1672]" gridcol="0" gridrow="4" pageId="110" pageNumber="109">
|
||
<taxonomicName authorityName="Lourenco" authorityYear="1997" box="[383,592,1649,1672]" class="Arachnida" family="Buthidae" genus="Lychas" kingdom="Animalia" order="Scorpiones" pageId="110" pageNumber="109" phylum="Arthropoda" rank="species" species="srilankensis">
|
||
<emphasis box="[383,592,1649,1672]" italics="true" pageId="110" pageNumber="109">Lychas srilankensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[796,824,1649,1672]" gridcol="1" gridrow="4" pageId="110" pageNumber="109">16</td>
|
||
<td box="[861,1076,1649,1672]" gridcol="2" gridrow="4" pageId="110" pageNumber="109">2x 9.2 + 7.0 – 4.2</td>
|
||
<td box="[1135,1229,1649,1672]" gridcol="3" gridrow="4" pageId="110" pageNumber="109">15CN</td>
|
||
</tr>
|
||
<tr box="[383,1229,1689,1712]" gridrow="5" pageId="110" pageNumber="109">
|
||
<th box="[383,696,1689,1712]" gridcol="0" gridrow="5" pageId="110" pageNumber="109">
|
||
<taxonomicName authorityName="Kovařík & Lowe & Ranawana & Hoferek & Š" authorityYear="2016" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[383,602,1689,1712]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="110" pageNumber="109" phylum="Arthropoda" rank="species" species="basilicus">
|
||
<emphasis box="[383,602,1689,1712]" italics="true" pageId="110" pageNumber="109">Reddyanus basilicus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[796,824,1689,1712]" gridcol="1" gridrow="5" pageId="110" pageNumber="109">16</td>
|
||
<td box="[861,1076,1689,1712]" gridcol="2" gridrow="5" pageId="110" pageNumber="109">8.0 – 4.6</td>
|
||
<td box="[1135,1229,1689,1712]" gridcol="3" gridrow="5" pageId="110" pageNumber="109">15CS</td>
|
||
</tr>
|
||
<tr box="[383,1229,1729,1752]" gridrow="6" pageId="110" pageNumber="109">
|
||
<th box="[383,696,1729,1752]" gridcol="0" gridrow="6" pageId="110" pageNumber="109">
|
||
<taxonomicName authorityName="Kovařík & Lowe & Ranawana & Hoferek & Š" authorityYear="2016" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[383,602,1729,1752]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="110" pageNumber="109" phylum="Arthropoda" rank="species" species="basilicus">
|
||
<emphasis box="[383,602,1729,1752]" italics="true" pageId="110" pageNumber="109">Reddyanus basilicus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[796,824,1729,1752]" gridcol="1" gridrow="6" pageId="110" pageNumber="109">15</td>
|
||
<td box="[861,1076,1729,1752]" gridcol="2" gridrow="6" pageId="110" pageNumber="109">11.5 + 8.0 – 4.6</td>
|
||
<td box="[1135,1229,1729,1752]" gridcol="3" gridrow="6" pageId="110" pageNumber="109">15CR</td>
|
||
</tr>
|
||
<tr box="[383,1229,1769,1793]" gridrow="7" pageId="110" pageNumber="109">
|
||
<th box="[383,696,1769,1793]" gridcol="0" gridrow="7" pageId="110" pageNumber="109">
|
||
<taxonomicName authority="Kovařík & Lowe & Ranawana & Hoferek & Š, 2016" authorityName="Kovařík & Lowe & Ranawana & Hoferek & Š" authorityYear="2016" box="[383,630,1769,1792]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="110" pageNumber="109" phylum="Arthropoda" rank="species" species="ceylonensis" status="sp. n.">
|
||
<emphasis box="[383,630,1769,1792]" italics="true" pageId="110" pageNumber="109">Reddyanus ceylonensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[636,696,1769,1793]" pageId="110" pageNumber="109">
|
||
<taxonomicNameLabel box="[636,696,1769,1793]" pageId="110" pageNumber="109" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<td box="[796,824,1769,1793]" gridcol="1" gridrow="7" pageId="110" pageNumber="109">16</td>
|
||
<td box="[861,1076,1769,1793]" gridcol="2" gridrow="7" pageId="110" pageNumber="109">8.9 – 4.2</td>
|
||
<td box="[1135,1229,1769,1793]" gridcol="3" gridrow="7" pageId="110" pageNumber="109">15CI</td>
|
||
</tr>
|
||
<tr box="[383,1229,1809,1832]" gridrow="8" pageId="110" pageNumber="109">
|
||
<th box="[383,696,1809,1832]" gridcol="0" gridrow="8" pageId="110" pageNumber="109">
|
||
<taxonomicName authorityName="Kovařík & Lowe & Ranawana & Hoferek & Š" authorityYear="2016" baseAuthorityName="Vachon" baseAuthorityYear="1982" box="[383,568,1809,1832]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="110" pageNumber="109" phylum="Arthropoda" rank="species" species="loebli">
|
||
<emphasis box="[383,568,1809,1832]" italics="true" pageId="110" pageNumber="109">Reddyanus loebli</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[796,824,1809,1832]" gridcol="1" gridrow="8" pageId="110" pageNumber="109">1 7</td>
|
||
<td box="[861,1076,1809,1832]" gridcol="2" gridrow="8" pageId="110" pageNumber="109">7.7 – 5.0 + 3.5</td>
|
||
<td box="[1135,1229,1809,1832]" gridcol="3" gridrow="8" pageId="110" pageNumber="109">15CH</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption ID-Table-UUID="6CA76641A879FFFEFF00F8909B1AF8E2" box="[192,1400,1863,1887]" httpUri="http://table.plazi.org/id/6CA76641A879FFFEFF00F8909B1AF8E2" pageId="110" pageNumber="109" startId="110.[192,257,1863,1887]" targetBox="[383,1229,1455,1832]" targetIsTable="true" targetPageId="110">
|
||
<paragraph blockId="110.[192,1400,1863,1887]" box="[192,1400,1863,1887]" pageId="110" pageNumber="109">
|
||
<emphasis bold="true" box="[192,286,1863,1887]" pageId="110" pageNumber="109">Table 6:</emphasis>
|
||
The diploid numbers and the relative size of the diploid set (in %) of chromosomes of Sri Lankan buthid scorpions.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7124756" ID-Zenodo-Dep="7124756" httpUri="https://zenodo.org/record/7124756/files/figure.png" pageId="111" pageNumber="110" startId="111.[192,278,1350,1374]" targetBox="[191,1440,191,1322]" targetPageId="111">
|
||
<paragraph blockId="111.[192,1440,1350,1399]" pageId="111" pageNumber="110">
|
||
<emphasis bold="true" box="[192,388,1350,1374]" pageId="111" pageNumber="110">Figures 537–538:</emphasis>
|
||
<taxonomicName authorityName="Simon" authorityYear="1872" box="[396,675,1353,1374]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="111" pageNumber="110" phylum="Arthropoda" rank="species" species="swammerdami">
|
||
<emphasis box="[396,675,1353,1374]" italics="true" pageId="111" pageNumber="110">Heterometrus swammerdami</emphasis>
|
||
</taxonomicName>
|
||
from locality 15CB, female, chelicerae, carapace and tergites I–III (537), and sternopectinal region with sternites III–IV (538).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection lastPageId="112" lastPageNumber="111" pageId="111" pageNumber="110" type="discussion">
|
||
<paragraph blockId="111.[192,802,1697,1873]" lastBlockId="112.[192,802,1390,1659]" lastPageId="112" lastPageNumber="111" pageId="111" pageNumber="110">
|
||
COMMENTS.
|
||
<taxonomicName authority="Pocock, 1900" authorityName="Pocock" authorityYear="1900" box="[331,712,1697,1720]" class="Arachnida" family="Scorpionidae" genus="Palamnaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="111" pageNumber="110" phylum="Arthropoda" rank="species" species="serratus">
|
||
<emphasis box="[331,558,1697,1720]" italics="true" pageId="111" pageNumber="110">Palamnaeus serratus</emphasis>
|
||
<bibRefCitation author="Pocock" bookContentInfo="279 pp" box="[567,712,1697,1720]" firstAuthor="Pocock" journalOrPublisher="London: W. T. Blandford" pageId="111" pageNumber="110" part="xii" refId="ref43790" refString="POCOCK, R. I. 1900. Arachnida. The fauna of British India, including Ceylon and Burma. Published un- der the authority of the Secretary of State for India in Council. London: W. T. Blandford, xii, 279 pp" title="Arachnida. The fauna of British India, including Ceylon and Burma. Published un- der the authority of the Secretary of State for India in Council" type="book" year="1900">Pocock, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
was synonymized with
|
||
<taxonomicName authorityName=": Fet" authorityYear="2000" baseAuthorityName="De Geer" baseAuthorityYear="1778" box="[379,473,1728,1751]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="111" pageNumber="110" phylum="Arthropoda" rank="species" species="indus">
|
||
<emphasis box="[379,473,1728,1751]" italics="true" pageId="111" pageNumber="110">H. indus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="COUZIJN, H. W. C." box="[520,743,1728,1751]" journalOrPublisher="Zoologische Verhandelingen" pageId="111" pageNumber="110" pagination="1 - 196" part="184" refId="ref40696" refString="COUZIJN, H. W. C. 1981. Revision of the genus Heterometrus. Zoologische Verhandelingen, 184: 1 - 196." title="Revision of the genus Heterometrus" type="journal article" year="1981">Couzijn (1981: 121)</bibRefCitation>
|
||
. The first author (
|
||
<bibRefCitation author="Kovarik" box="[332,533,1758,1781]" firstAuthor="Kovarik" page="17" pageId="111" pageNumber="110" pagination="1 - 60" refId="ref41757" refString="KOVARIK, F. 2004. A review of the genus Heterometrus Ehrenberg, 1828, with descriptions of seven new species (Scorpiones, Scorpionidae). Euscorpius, 15: 1 - 60." type="journal article" year="2004">Kovařík, 2004: 17</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Kovarik" bookContentInfo="170 pp." box="[592,793,1758,1781]" firstAuthor="Kovarik" journalOrPublisher="Clairon Production, Prague" page="38" pageId="111" pageNumber="110" refId="ref41793" refString="KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Clairon Production, Prague, 170 pp." title="Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species" type="book" year="2009">Kovařík, 2009: 38</bibRefCitation>
|
||
) accepted the synonymy, whereas
|
||
<bibRefCitation author="Tikader & Bastawade" bookContentInfo="668 pp." firstAuthor="Tikader" journalOrPublisher="Zoological Survey of India, Calcutta" page="555" pageId="111" pageNumber="110" refId="ref44789" refString="TIKADER B. K. & D. B. BASTAWADE. 1983. The Fauna of India: Scorpions. Scorpionida, Arachnida. Vol III. Zoological Survey of India, Calcutta, 668 pp." title="The Fauna of India: Scorpions. Scorpionida, Arachnida. Vol III" type="book" year="1983">Tikader & Bastawade (1983: 555)</bibRefCitation>
|
||
considered
|
||
<taxonomicName authorityName="Pocock" authorityYear="1900" box="[475,605,1820,1843]" class="Arachnida" family="Scorpionidae" genus="Palamnaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="111" pageNumber="110" phylum="Arthropoda" rank="species" species="serratus">
|
||
<emphasis box="[475,605,1820,1843]" italics="true" pageId="111" pageNumber="110">H. serratus</emphasis>
|
||
</taxonomicName>
|
||
a valid species. Examination of additional specimens collected during the
|
||
<collectingRegion box="[872,903,1452,1475]" country="United Kingdom" name="Surrey" pageId="111" pageNumber="110">Sri</collectingRegion>
|
||
Lankan expedition of 2015 convinced us that
|
||
<taxonomicName authorityName="Pocock" authorityYear="1900" class="Arachnida" family="Scorpionidae" genus="Palamnaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="111" pageNumber="110" phylum="Arthropoda" rank="species" species="serratus">
|
||
<emphasis italics="true" pageId="111" pageNumber="110">H. serratus</emphasis>
|
||
</taxonomicName>
|
||
is in fact a valid species.
|
||
<bibRefCitation author="Pocock" bookContentInfo="279 pp" box="[1232,1440,1482,1505]" firstAuthor="Pocock" journalOrPublisher="London: W. T. Blandford" page="86" pageId="111" pageNumber="110" part="xii" refId="ref43790" refString="POCOCK, R. I. 1900. Arachnida. The fauna of British India, including Ceylon and Burma. Published un- der the authority of the Secretary of State for India in Council. London: W. T. Blandford, xii, 279 pp" title="Arachnida. The fauna of British India, including Ceylon and Burma. Published un- der the authority of the Secretary of State for India in Council" type="book" year="1900">Pocock (1900: 86)</bibRefCitation>
|
||
differentiated
|
||
<taxonomicName authorityName="Pocock" authorityYear="1900" box="[986,1112,1513,1536]" class="Arachnida" family="Scorpionidae" genus="Palamnaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="111" pageNumber="110" phylum="Arthropoda" rank="species" species="serratus">
|
||
<emphasis box="[986,1112,1513,1536]" italics="true" pageId="111" pageNumber="110">H. serratus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName=": Fet" authorityYear="2000" baseAuthorityName="De Geer" baseAuthorityYear="1778" box="[1175,1272,1513,1536]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="111" pageNumber="110" phylum="Arthropoda" rank="species" species="indus">
|
||
<emphasis box="[1175,1272,1513,1536]" italics="true" pageId="111" pageNumber="110">H. indus</emphasis>
|
||
</taxonomicName>
|
||
as follows:
|
||
<emphasis bold="true" box="[1418,1440,1512,1536]" pageId="111" pageNumber="110">1)</emphasis>
|
||
dorsal and dorsolateral carinae of metasomal segments granulated in
|
||
<taxonomicName authorityName="Pocock" authorityYear="1900" box="[981,1102,1574,1597]" class="Arachnida" family="Scorpionidae" genus="Palamnaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="111" pageNumber="110" phylum="Arthropoda" rank="species" species="serratus">
|
||
<emphasis box="[981,1102,1574,1597]" italics="true" pageId="111" pageNumber="110">H. serratus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[1119,1208,1574,1597]" captionStart="Figures 571–574" captionStartId="116.[192,278,786,810]" captionTargetBox="[195,1423,184,735]" captionTargetId="figure-11@116.[192,1439,168,757]" captionTargetPageId="116" captionText="Figures 571–574: Key characters for distinguishing Sri Lankan Heterometrus. Figures 571–572. Pedipalp chela dorsal view of male, Heterometrus serratus from locality 15CP (571) and Heterometrus gravimanus from locality 15CE (572). Figures 573– 574. Metasomal segments IV-II lateral view of female, Heterometrus indus from locality 15CA (573) and Heterometrus serratus from locality 15CP (574)." figureDoi="http://doi.org/10.5281/zenodo.7124769" httpUri="https://zenodo.org/record/7124769/files/figure.png" pageId="111" pageNumber="110">Fig. 574</figureCitation>
|
||
), smooth in
|
||
<taxonomicName authorityName=": Fet" authorityYear="2000" baseAuthorityName="De Geer" baseAuthorityYear="1778" box="[1350,1440,1574,1597]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="111" pageNumber="110" phylum="Arthropoda" rank="species" species="indus">
|
||
<emphasis box="[1350,1440,1574,1597]" italics="true" pageId="111" pageNumber="110">H. indus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[838,930,1605,1628]" captionStart="Figures 571–574" captionStartId="116.[192,278,786,810]" captionTargetBox="[195,1423,184,735]" captionTargetId="figure-11@116.[192,1439,168,757]" captionTargetPageId="116" captionText="Figures 571–574: Key characters for distinguishing Sri Lankan Heterometrus. Figures 571–572. Pedipalp chela dorsal view of male, Heterometrus serratus from locality 15CP (571) and Heterometrus gravimanus from locality 15CE (572). Figures 573– 574. Metasomal segments IV-II lateral view of female, Heterometrus indus from locality 15CA (573) and Heterometrus serratus from locality 15CP (574)." figureDoi="http://doi.org/10.5281/zenodo.7124769" httpUri="https://zenodo.org/record/7124769/files/figure.png" pageId="111" pageNumber="110">Fig. 573</figureCitation>
|
||
). We found other minor differences:
|
||
<emphasis bold="true" box="[1343,1365,1604,1628]" pageId="111" pageNumber="110">2)</emphasis>
|
||
sexual dimorphism in proportions of pedipalps not noticeable in
|
||
<taxonomicName authorityName=": Fet" authorityYear="2000" baseAuthorityName="De Geer" baseAuthorityYear="1778" box="[830,923,1666,1689]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="111" pageNumber="110" phylum="Arthropoda" rank="species" species="indus">
|
||
<emphasis box="[830,923,1666,1689]" italics="true" pageId="111" pageNumber="110">H. indus</emphasis>
|
||
</taxonomicName>
|
||
, male with slightly narrower chela than female in
|
||
<taxonomicName authorityName="Pocock" authorityYear="1900" box="[861,983,1697,1720]" class="Arachnida" family="Scorpionidae" genus="Palamnaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="111" pageNumber="110" phylum="Arthropoda" rank="species" species="serratus">
|
||
<emphasis box="[861,983,1697,1720]" italics="true" pageId="111" pageNumber="110">H. serratus</emphasis>
|
||
</taxonomicName>
|
||
(chela length to width ratio 1.79–1.94 in males, 1.69 in the females);
|
||
<emphasis bold="true" box="[1167,1189,1727,1751]" pageId="111" pageNumber="110">3)</emphasis>
|
||
carapace smooth and glossy, only occasionally with granules at margins in
|
||
<taxonomicName authorityName=": Fet" authorityYear="2000" baseAuthorityName="De Geer" baseAuthorityYear="1778" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="111" pageNumber="110" phylum="Arthropoda" rank="species" species="indus">
|
||
<emphasis italics="true" pageId="111" pageNumber="110">H. indus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[913,1082,1789,1812]" captionStart="Figures 486–489" captionStartId="99.[192,278,1844,1868]" captionTargetBox="[213,1417,167,1815]" captionTargetId="figure-33@99.[214,1416,168,1814]" captionTargetPageId="99" captionText="Figures 486–489: Heterometrus indus from locality 15CA. Figures 486, 488. Male, chelicerae, carapace and tergites I–III (486), and sternopectinal region (488). Figures 487, 489. Female, chelicerae, carapace and tergites I–II (487), and sternopectinal region (489)." figureDoi="http://doi.org/10.5281/zenodo.7124726" httpUri="https://zenodo.org/record/7124726/files/figure.png" pageId="111" pageNumber="110">Figs. 486–487</figureCitation>
|
||
); carapace smooth and glossy medially, always with more granules at margins in
|
||
<taxonomicName authorityName="Pocock" authorityYear="1900" class="Arachnida" family="Scorpionidae" genus="Palamnaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="111" pageNumber="110" phylum="Arthropoda" rank="species" species="serratus">
|
||
<emphasis italics="true" pageId="111" pageNumber="110">H. serratus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[940,1094,1850,1873]" captionStart="Figures 516–519" captionStartId="106.[192,278,1829,1853]" captionTargetBox="[213,1417,180,1801]" captionTargetId="figure-47@106.[214,1416,181,1800]" captionTargetPageId="106" captionText="Figures 516–519: Heterometrus serratus from locality 15CP. Figures 516, 518. Male, chelicerae, carapace and tergites I–II (516), and sternopectinal region (518). Figures 517, 519. Female, chelicerae, carapace and tergites I–II (517), and sternopectinal region (519)." figureDoi="http://doi.org/10.5281/zenodo.7124742" httpUri="https://zenodo.org/record/7124742/files/figure.png" pageId="111" pageNumber="110">Figs. 516–517</figureCitation>
|
||
);
|
||
<emphasis bold="true" box="[1122,1144,1849,1873]" pageId="111" pageNumber="110">4)</emphasis>
|
||
spination formula of tarsomeres II of legs: 3/4: 2-4/3-4: 4/4-5: 4/
|
||
<quantity box="[655,696,1390,1413]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="112" pageNumber="111" unit="in" value="5.0">5 in</quantity>
|
||
<taxonomicName authorityName=": Fet" authorityYear="2000" baseAuthorityName="De Geer" baseAuthorityYear="1778" box="[704,794,1390,1413]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="112" pageNumber="111" phylum="Arthropoda" rank="species" species="indus">
|
||
<emphasis box="[704,794,1390,1413]" italics="true" pageId="112" pageNumber="111">H. indus</emphasis>
|
||
</taxonomicName>
|
||
; 3-4/4-6: 4/4-5: 4/5-6: 4/
|
||
<quantity box="[466,530,1421,1444]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" metricValueMax="1.778" metricValueMin="1.27" pageId="112" pageNumber="111" unit="in" value="6.0" valueMax="7.0" valueMin="5.0">5-7 in</quantity>
|
||
<taxonomicName authorityName="Pocock" authorityYear="1900" box="[536,655,1421,1444]" class="Arachnida" family="Scorpionidae" genus="Palamnaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="112" pageNumber="111" phylum="Arthropoda" rank="species" species="serratus">
|
||
<emphasis box="[536,655,1421,1444]" italics="true" pageId="112" pageNumber="111">H. serratus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7124761" ID-Zenodo-Dep="7124761" httpUri="https://zenodo.org/record/7124761/files/figure.png" pageId="112" pageNumber="111" startId="112.[192,278,1298,1322]" targetBox="[192,1439,153,1265]" targetPageId="112">
|
||
<paragraph blockId="112.[192,1441,1298,1347]" pageId="112" pageNumber="111">
|
||
<emphasis bold="true" box="[192,389,1298,1322]" pageId="112" pageNumber="111">Figures 539–543:</emphasis>
|
||
<taxonomicName authorityName="Simon" authorityYear="1872" box="[399,679,1301,1322]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="112" pageNumber="111" phylum="Arthropoda" rank="species" species="swammerdami">
|
||
<emphasis box="[399,679,1301,1322]" italics="true" pageId="112" pageNumber="111">Heterometrus swammerdami</emphasis>
|
||
</taxonomicName>
|
||
from locality 15CB, female, pedipalp chela dorsal (539), external (540) and ventral (541) views, tarsomeres of third (542) and fourth (543) legs retroventral views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="112.[192,802,1390,1659]" pageId="112" pageNumber="111">
|
||
The distribution of
|
||
<taxonomicName authorityName="Pocock" authorityYear="1900" box="[400,520,1452,1475]" class="Arachnida" family="Scorpionidae" genus="Palamnaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="112" pageNumber="111" phylum="Arthropoda" rank="species" species="serratus">
|
||
<emphasis box="[400,520,1452,1475]" italics="true" pageId="112" pageNumber="111">H. serratus</emphasis>
|
||
</taxonomicName>
|
||
was previously unknown, as the type locality was imprecise and the
|
||
<typeStatus box="[650,743,1482,1505]" pageId="112" pageNumber="111">holotype</typeStatus>
|
||
male was the only known specimen. We collected additional specimens at localities cited here as 15CP (
|
||
<figureCitation box="[656,747,1544,1567]" captionStart="Figures 592–594" captionStartId="126.[192,278,1617,1641]" captionTargetBox="[213,1417,167,1588]" captionTargetId="figure-251@126.[214,1416,168,1587]" captionTargetPageId="126" captionText="Figures 592–594: Figure 592. Locality 15CO, Sri Lanka, North Central Province, Puttalam District, Eluwankulam, 08°17'15"N 079°50'38.7"E, 38 m a.s.l. During night collecting from 10.30 p.m. on 28.IV.2015 to 4.00 a.m. on 29.IV.2015 (UV detection) we recorded Charmus laneus (running on the ground among leaves), Isometrus thwaitesii (running on branches of bushes, also sitting on leaves), Lychas srilankensis (on the ground under branches, leaves or in their vicinity), Reddyanus ceylonensis sp. n. (on the ground), Chaerilus ceylonensis (inside oblique usually not curved burrows 15 to 30 cm long) and Heterometrus swammerdami (inside long burrows). On the locality we recorded between 10.00 p.m. and 4.00 a.m. temperature 25.9 ºC and humidity between 58% and 70%. On the locality we recorded 6 scorpion species and genera of all three Sri Lankan families. Figures 593–594. Locality 15CP, Sri Lanka, Southern Province, Matara District, Kekanadura village, 05°58'28.2"N 080°36'20.5"E, 40 m a.s.l. During night collecting on 30.IV.2015 (UV detection) we recorded Isometrus maculatus (inside a house, the species occupy the roof of the house, Fig. 593) and Heterometrus serratus (inside almost straight burrows in a ground wall). On the locality we recorded night temperature 28.4 ºC– 25.2 ºC (minimum temperature) and night humidity varied between 63% and 89%." figureDoi="http://doi.org/10.5281/zenodo.7124791" httpUri="https://zenodo.org/record/7124791/files/figure.png" pageId="112" pageNumber="111">Fig. 594</figureCitation>
|
||
) and 15CQ (
|
||
<figureCitation box="[275,374,1574,1597]" captionStart="Figures 595–596" captionStartId="127.[192,278,1661,1685]" captionTargetBox="[191,1440,195,1633]" captionTargetId="figure-82@127.[192,1439,196,1632]" captionTargetPageId="127" captionText="Figures 595–596: Figure 595. Locality 15CQ, Sri Lanka, Uva Province, Monaragala District, Monaragala, 06°52'30.7"N 081°21'17"E, 288 m a.s.l. During night collecting on 2.–3.V.2015 (UV detection) we recorded Reddyanus loebli (under scales of bark and under bark) and Heterometrus serratus (females with juveniles inside oblique burrows). On the locality we recorded a temperature of 33.1 ºC at 5.00 p.m., which gradually dropped to 24.6 ºC (minimum temperature) before sunrise. Humidity during the night varied between 61% and 84%. Figure 596. Locality 15CR, Sri Lanka, Eastern Province, Ampara District, Lahugala Kitulana National Park, 06°52'46"N 081°43'21.8"E, 40 m a.s.l. During night collecting on 3.–4.V.2015 (10.00 p.m. to 01.00 a.m.) we recorded Isometrus maculatus (running on branches, also sitting on leaves on trees), Reddyanus basilicus (on the ground among leaves) and Reddyanus loebli (hidden under scales of bark of standing tree trunks). On the locality we recorded between 10.00 p.m. on 3.V. and 01.00 a.m. on 4.V.2015 temperature 28 ºC, which gradually dropped to 26.9 ºC and humidity 73%." figureDoi="http://doi.org/10.5281/zenodo.7124794" httpUri="https://zenodo.org/record/7124794/files/figure.png" pageId="112" pageNumber="111">Fig. 595</figureCitation>
|
||
). Our data suggest that
|
||
<taxonomicName authorityName="Pocock" authorityYear="1900" box="[648,772,1574,1597]" class="Arachnida" family="Scorpionidae" genus="Palamnaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="112" pageNumber="111" phylum="Arthropoda" rank="species" species="serratus">
|
||
<emphasis box="[648,772,1574,1597]" italics="true" pageId="112" pageNumber="111">H. serratus</emphasis>
|
||
</taxonomicName>
|
||
is distributed in the southern part, and
|
||
<taxonomicName authorityName=": Fet" authorityYear="2000" baseAuthorityName="De Geer" baseAuthorityYear="1778" box="[622,720,1605,1628]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="112" pageNumber="111" phylum="Arthropoda" rank="species" species="indus">
|
||
<emphasis box="[622,720,1605,1628]" italics="true" pageId="112" pageNumber="111">H. indus</emphasis>
|
||
</taxonomicName>
|
||
in the central part of
|
||
<collectingCountry box="[348,453,1636,1659]" name="Sri Lanka" pageId="112" pageNumber="111">Sri Lanka</collectingCountry>
|
||
(
|
||
<figureCitation box="[468,544,1636,1659]" captionStart="Figures 12–15" captionStartId="6.[192,278,1885,1909]" captionTargetBox="[191,1440,167,1856]" captionTargetId="figure-11@6.[192,1439,168,1855]" captionTargetPageId="6" captionText="Figures 12–15: Maps showing the known geographical distribution of the Sri Lankan species." figureDoi="http://doi.org/10.5281/zenodo.7124476" httpUri="https://zenodo.org/record/7124476/files/figure.png" pageId="112" pageNumber="111">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[192,470,1691,1714]" pageId="112" pageNumber="111" type="distribution">
|
||
<paragraph blockId="112.[192,470,1691,1714]" box="[192,470,1691,1714]" pageId="112" pageNumber="111">
|
||
DISTRIBUTION.
|
||
<collectingCountry box="[359,463,1691,1714]" name="Sri Lanka" pageId="112" pageNumber="111">Sri Lanka</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |