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<document id="6B83CC2904BDB4F1FA53AE545411953A" ID-CLB-Dataset="153629" ID-DOI="10.18590/euscorpius.2016.vol2016.iss220.1" ID-GBIF-Dataset="03196918-10d5-44d2-ab42-d38f88314c7b" ID-ISSN="1536-9307" ID-Zenodo-Dep="7124466" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638249082081" checkinUser="felipe" docAuthor="Kovařík, František, Lowe, Graeme, Ranawana, Kithsiri B., Hoferek, David &amp; Š, V. A." docDate="2016" docId="B07187DFA854FFDDFC5CFA7D9B74F8D1" docLanguage="en" docName="Euscorpius.220.1-133.pdf" docOrigin="Euscorpius 220" docSource="https://doi.org/10.18590/euscorpius.2016.vol2016.iss220.1" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Reddyanus jayarathnei Kovarik 2016, sp. n." docType="treatment" docUuid="FE1515A9-8302-46D4-8298-11BFB4555882" docUuidSource="ZooBank" docVersion="9" lastPageNumber="76" masterDocId="4C48FFA7A817FF90FFC0FFD79E62FFBD" masterDocTitle="Scorpions of Sri Lanka (Scorpiones Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat n" masterLastPageNumber="133" masterPageNumber="1" pageNumber="66" updateTime="1699471154095" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="257F7F57F60EC6D2BAAF5D69A1FA6FAF">Scorpions of Sri Lanka (Scorpiones Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat n</mods:title>
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<mods:namePart id="781B604A095C58733BBEAA8265C5CD81">Kovařík, František</mods:namePart>
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<mods:namePart id="517FF5362509DAD1CC3C56867C95712A">Lowe, Graeme</mods:namePart>
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<mods:namePart id="BCB52F2AB718A427F2975C2407D657AF">Ranawana, Kithsiri B.</mods:namePart>
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<mods:namePart id="FA0A824E1DB46F3091C575D9C9BDDD6E">Hoferek, David</mods:namePart>
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<mods:namePart id="E413B3C228FA4D935C5FAF72B2442D28">Š, V. A.</mods:namePart>
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<treatment id="B07187DFA854FFDDFC5CFA7D9B74F8D1" ID-DOI="http://doi.org/10.5281/zenodo.7124553" ID-GBIF-Taxon="201134869" ID-Zenodo-Dep="7124553" LSID="urn:lsid:zoobank.org:act:FE1515A9-8302-46D4-8298-11BFB4555882" httpUri="http://treatment.plazi.org/id/B07187DFA854FFDDFC5CFA7D9B74F8D1" lastPageId="77" lastPageNumber="76" pageId="67" pageNumber="66">
<subSubSection id="70C26542A854FFD3FC5CFA7D9B20FA7F" box="[924,1346,1450,1475]" pageId="67" pageNumber="66" type="nomenclature">
<paragraph id="386736C9A854FFD3FC5CFA7D9B20FA7F" blockId="67.[838,1432,1450,1598]" box="[924,1346,1450,1475]" pageId="67" pageNumber="66">
<heading id="632F81A5A854FFD3FC5CFA7D9B20FA7F" box="[924,1346,1450,1475]" centered="true" fontSize="11" level="2" pageId="67" pageNumber="66" reason="2">
<taxonomicName id="FFD84D4AA854FFD3FC5CFA7D9A9BFA7F" ID-CoL="4RP7B" authority="Kovarik" authorityName="Kovarik" authorityYear="2016" box="[924,1273,1450,1474]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="67" pageNumber="66" phylum="Arthropoda" rank="species" species="jayarathnei" status="sp. nov.">
<emphasis id="0AACEADBA854FFD3FC5CFA7D9AF9FA7F" bold="true" box="[924,1179,1450,1474]" italics="true" pageId="67" pageNumber="66">Reddyanus jayarathnei</emphasis>
Kovařík
</taxonomicName>
,
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.
<emphasis id="0AACEADBA854FFD3FAECFA7C9B59FA7E" bold="true" box="[1324,1339,1451,1475]" pageId="67" pageNumber="66">n</emphasis>
.
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<paragraph id="386736C9A854FFD3FC8CFA1D9A9DF9BD" blockId="67.[838,1432,1450,1598]" pageId="67" pageNumber="66">
(
<figureCitation id="A0E32A4CA854FFD3FC95FA1D9DC9FA5C" box="[853,939,1482,1505]" captionStart="Figures 1215" captionStartId="6.[192,278,1885,1909]" captionTargetBox="[191,1440,167,1856]" captionTargetId="figure-11@6.[192,1439,168,1855]" captionTargetPageId="6" captionText="Figures 1215: Maps showing the known geographical distribution of the Sri Lankan species." figureDoi="http://doi.org/10.5281/zenodo.7124476" httpUri="https://zenodo.org/record/7124476/files/figure.png" pageId="67" pageNumber="66">Figs. 14</figureCitation>
,
<figureCitation id="A0E32A4CA854FFD3FC78FA1D9D82FA5C" box="[952,992,1482,1505]" captionStart="Figures 201208" captionStartId="49.[192,278,1057,1081]" captionTargetBox="[204,1438,173,1022]" captionTargetId="figure-6@49.[192,1439,168,1028]" captionTargetPageId="49" captionText="Figures 201208: Distal segments of legs, retroventral view of Sri Lankan Reddyanus. Figures 201204. R. ceylonensis sp. n., male holotype from locality 15CI, legs IIV. Figures 205208. Leg IV. R. besucheti, male from locality 15CD (205), R. jayarathnei sp. n., male paratype (206), R. loebli, male from locality 15CG (207), and R. ranawanai sp. n., male holotype (208)." figureDoi="http://doi.org/10.5281/zenodo.7124612" httpUri="https://zenodo.org/record/7124612/files/figure.png" pageId="67" pageNumber="66">206</figureCitation>
,
<figureCitation id="A0E32A4CA854FFD3FC2EFA1D9A29FA5C" box="[1006,1099,1482,1505]" captionStart="Figures 209224" captionStartId="50.[192,278,1736,1760]" captionTargetBox="[208,1434,174,1703]" captionTargetId="figure-11@50.[205,1452,168,1707]" captionTargetPageId="50" captionText="Figures 209224: Metasomal segments IIIII, lateral view of Sri Lankan Reddyanus and Isometrus species. Figures 209210. Reddyanus basilicus, male (209) and female (210) from locality 15CR. Figures 211212. R. besucheti, male holotype (211) and female from locality 15CG (212). Figures 213214. R. ceylonensis sp. n., male holotype (213) and female paratype from locality 15CI (214). Figures 215216. R. jayarathnei sp. n., male (215) and female (216) paratypes. Figures 217218. R. loebli, male (217) and female (218) from locality 15CG. Figures 219220. R. ranawanai sp. n., male holotype (219) and female paratype (220). Figures 221222. Isometrus maculatus, male from locality 15CP (221) and female from locality 15CI (222). Figures 223 224. I. thwaitesi, male from locality 15CH (223) and female from locality 15CO (224)." figureDoi="http://doi.org/10.5281/zenodo.7124622" httpUri="https://zenodo.org/record/7124622/files/figure.png" pageId="67" pageNumber="66">215216</figureCitation>
,
<figureCitation id="A0E32A4CA854FFD3FB98FA1D9AD4FA5C" box="[1112,1206,1482,1505]" captionStart="Figures 225240" captionStartId="51.[192,278,1493,1517]" captionTargetBox="[193,1432,180,1451]" captionTargetId="figure-6@51.[192,1439,179,1464]" captionTargetPageId="51" captionText="Figures 225240: Pedipalp, dorsal view of Sri Lankan Reddyanus and Isometrus species. Figures 225226. Reddyanus basilicus, male (225) and female (226) from locality 15CR. Figures 227228. R. besucheti, male from locality 15CD (227) and female from locality 15CG (228). Figures 229230. R. ceylonensis sp. n., male holotype (229) and female paratype from locality 15CI (230). Figures 231232. R. jayarathnei sp. n., male (231) and female (232) paratypes. Figures 233234. R. loebli, male (233) and female (234) from locality 15CG. Figures 235236. R. ranawanai sp. n., male holotype (235) and female paratype (236). Figures 237238. Isometrus maculatus, male from locality 15CP (237) and female from locality 15CI (238). Figures 239240. I. thwaitesi, male from locality 15CH (239) and female from locality 15CO (240)." figureDoi="http://doi.org/10.5281/zenodo.7124624" httpUri="https://zenodo.org/record/7124624/files/figure.png" pageId="67" pageNumber="66">231232</figureCitation>
,
<figureCitation id="A0E32A4CA854FFD3FB03FA1D9B41FA5C" box="[1219,1315,1482,1505]" captionStart="Figures 241259" captionStartId="52.[192,278,1733,1757]" captionTargetBox="[238,1387,153,1699]" captionTargetId="figure-11@52.[234,1392,152,1704]" captionTargetPageId="52" captionText="Figures 241259: Pedipalp chela dorsal (241), movable finger (242250), and fixed finger (251259) of Sri Lankan Isometrus, Lychas, and Reddyanus species. Figures 241242, 252. Isometrus maculatus, female from locality 15CO (241) and male from locality 15CP (242, 252). Figures 243244, 253. I. thwaitesi, male from locality 15CH (243) and female from locality 15CO (244, 253). Figures 245, 254. Reddyanus basilicus, male from locality 15CS. Figures 246, 255. R. besucheti, male from locality 15CD. Figures 247, 256. R. ceylonensis sp. n., male holotype. Figures 248, 257. R. jayarathnei sp. n., male paratype. Figures 249, 258. R. loebli, male from locality 15CG. Figures 250, 259. R. ranawanai sp. n., male holotype. Figure 251. Lychas srilankensis, male from locality 15CO." figureDoi="http://doi.org/10.5281/zenodo.7124626" httpUri="https://zenodo.org/record/7124626/files/figure.png" pageId="67" pageNumber="66">248, 257</figureCitation>
,
<figureCitation id="A0E32A4CA854FFD3FAEEFA1D9BE9FA5C" box="[1326,1419,1482,1505]" captionStart-0="Figures 334337" captionStart-1="Figures 338348" captionStart-2="Figures 349354" captionStart-3="Figures 355357" captionStartId-0="70.[192,278,1786,1810]" captionStartId-1="71.[192,278,1729,1753]" captionStartId-2="72.[192,278,1600,1624]" captionStartId-3="73.[192,278,1872,1896]" captionTargetBox-0="[201,1437,224,1752]" captionTargetBox-1="[191,1440,191,1701]" captionTargetBox-2="[206,1421,175,1560]" captionTargetBox-3="[191,1440,167,1843]" captionTargetId-0="figure-11@70.[192,1439,216,1757]" captionTargetId-1="figure-68@71.[192,1439,192,1700]" captionTargetId-2="figure-11@72.[192,1439,168,1571]" captionTargetId-3="figure-59@73.[192,1439,168,1842]" captionTargetPageId-0="70" captionTargetPageId-1="71" captionTargetPageId-2="72" captionTargetPageId-3="73" captionText-0="Figures 334337: Reddyanus jayarathnei sp. n., paratypes. Figures 334335. Male in dorsal (334) and ventral (335) views. Figures 336337. Female in dorsal (336) and ventral (337) views. Scale bar: 10 mm." captionText-1="Figures 338348: Reddyanus jayarathnei sp. n., paratypes. Figures 338, 340, 343348. Male, chelicerae, carapace and tergites IIII (338), sternopectinal region (340). Pedipalp chela, dorsal (343), external (344) and ventral (345) views. Pedipalp patella, dorsal (346) and external (347) views. Pedipalp femur and trochanter dorsal (348) view. The trichobothrial pattern is indicated in Figures 344348. Figures 339, 341342. Female, chelicerae, carapace and tergites IIII (339), sternopectinal region and sternite III (341), and pedipalp chela dorsal view (342)." captionText-2="Figures 349354: Reddyanus jayarathnei sp. n., paratypes. Figures 349351. Male, metasoma and telson, lateral (349), ventral (350), and dorsal (351) views. Figures 352354. Female, metasoma and telson, lateral (352), ventral (353), and dorsal (354) views. Scale bar: 10 mm." captionText-3="Figures 355357: Reddyanus jayarathnei sp. n., female paratype (355) and male holotype (356) at the type locality (357), Galle district, Kanneliya Rain Forest, 06°15'04&quot; N 80°20'18&quot; E." figureDoi-0="http://doi.org/10.5281/zenodo.7124661" figureDoi-1="http://doi.org/10.5281/zenodo.7124663" figureDoi-2="http://doi.org/10.5281/zenodo.7124665" figureDoi-3="http://doi.org/10.5281/zenodo.7124668" httpUri-0="https://zenodo.org/record/7124661/files/figure.png" httpUri-1="https://zenodo.org/record/7124663/files/figure.png" httpUri-2="https://zenodo.org/record/7124665/files/figure.png" httpUri-3="https://zenodo.org/record/7124668/files/figure.png" pageId="67" pageNumber="66">334357</figureCitation>
,
<figureCitation id="A0E32A4CA854FFD3FC20FA3E9A5FF9BD" box="[992,1085,1513,1536]" captionStart="Figures 403429" captionStartId="85.[192,278,744,768]" captionTargetId="figure-11@84.[172,1464,185,1878]" captionTargetPageId="84" captionText="Figures 403429: Telson, lateral view of Sri Lankan Buthidae and Chaerilidae species. Figures 403404. Isometrus maculatus, male from locality 15CP (403) and female from locality 15CI (404). Figures 405406. I. thwaitesi, male from locality 15CH (405) and female from locality 15CO (406). Figures 407408. Lychas srilankensis, male (407) and female (408) from locality 15CJ. Figures 409410. Reddyanus basilicus, male (409) and female (410) from locality 15CR. Figures 411412. R. besucheti, male holotype (411) and female from locality 15CG (412). Figures 413414. R. ceylonensis sp. n., male holotype (413) and female paratype (414) from locality 15CI. Figures 415416. R. jayarathnei sp. n., male (415) and female (416) paratypes. Figures 417418. R. loebli, male (417) and female (418) from locality 15CG. Figures 419420. R. ranawanai sp. n., male holotype (419) and female paratype (420). Figures 421422: Buthoscorpio sarasinorum, male from locality 15CF (421) and female from locality 15CG (422). Figures 423424. Charmus laneus, male (423) and female (424) from locality 15CO. Figures 425426. Charmus saradieli sp. n. male paratype (425) (MHNG) and female holotype (426). Figurs 427. Hottentotta tamulus, male from locality 15CK (427). Figures 428429. Chaerilus ceylonensis, male (428) and female (429) from locality 15CD." figureDoi="http://doi.org/10.5281/zenodo.7124696" httpUri="https://zenodo.org/record/7124696/files/figure.png" pageId="67" pageNumber="66">415416</figureCitation>
, 569,
<tableCitation id="755A0372A854FFD3FB40FA3E9A94F9BD" box="[1152,1270,1513,1536]" captionStart-0="Table 4" captionStart-1="Table 5" captionStartId-0="86.[192,257,1428,1452]" captionStartId-1="86.[192,257,1869,1893]" captionTargetPageId-0="86" captionTargetPageId-1="86" captionText-0="Table 4: Comparative measurements of adults of Reddyanus ranawanai sp. n. and R. jayarathnei sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-1="Table 5: Comparison among Sri Lankan Reddyanus species (specimens), based upon selected morphometric ratios of adult males. Abbreviations: length (L), width (W), depth (D)." httpUri-0="http://table.plazi.org/id/6CA76641A841FFC6FF00FA439D16FA79" httpUri-1="http://table.plazi.org/id/6CA76641A841FFC6FF00F89A9CBAF8C0" pageId="67" pageNumber="66" tableUuid-0="6CA76641A841FFC6FF00FA439D16FA79" tableUuid-1="6CA76641A841FFC6FF00F89A9CBAF8C0">Tables 45</tableCitation>
)
</paragraph>
<paragraph id="386736C9A854FFD3FC86F9D09B3FF983" blockId="67.[838,1432,1450,1598]" pageId="67" pageNumber="66">
<emphasis id="0AACEADBA854FFD3FC86F9D09B3FF983" bold="true" pageId="67" pageNumber="66">
http://www.zoobank.org/urn:lsid:zoobank.org:act:F
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</emphasis>
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<typeStatus id="E763886BA854FFD3FCFEF9B49D15F9C7" box="[830,887,1635,1658]" pageId="67" pageNumber="66">TYPE</typeStatus>
LOCALITY AND
<typeStatus id="E763886BA854FFD3FBE2F9B09A35F9C7" box="[1058,1111,1639,1658]" pageId="67" pageNumber="66">TYPE</typeStatus>
REPOSITORY.
<collectingCountry id="40CF7659A854FFD3FB2FF9B49B39F9C7" box="[1263,1371,1635,1658]" name="Sri Lanka" pageId="67" pageNumber="66">Sri Lanka</collectingCountry>
,
<location id="3D076012A854FFD3FAA6F9B49DEAF924" LSID="urn:lsid:plazi:treatment:B07187DFA854FFDDFC5CFA7D9B74F8D1:3D076012A854FFD3FAA6F9B49DEAF924" country="Sri Lanka" latitude="6.251111" longLatPrecision="21" longitude="80.33833" municipality="Kanneliya Rain Forest" name="Galle district" pageId="67" pageNumber="66">Galle district</location>
,
<collectingMunicipality id="D803ACB3A854FFD3FC5BF9559AF6F924" box="[923,1172,1666,1689]" pageId="67" pageNumber="66">Kanneliya Rain Forest</collectingMunicipality>
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TYPE MATERIAL.
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</paragraph>
</subSubSection>
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<paragraph id="386736C9A853FFD4FF00F90A9FB2F882" blockId="68.[192,1441,1757,1855]" pageId="68" pageNumber="67">
<emphasis id="0AACEADBA853FFD4FF00F90A9FE6F948" bold="true" box="[192,388,1757,1781]" pageId="68" pageNumber="67">Figures 327330:</emphasis>
Left hemispermatophore of
<taxonomicName id="FFD84D4AA853FFD4FD5BF9089D1BF949" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[667,889,1759,1780]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="68" pageNumber="67" phylum="Arthropoda" rank="species" species="ceylonensis" status="sp. nov.">
<emphasis id="0AACEADBA853FFD4FD5BF9089D1BF949" box="[667,889,1759,1780]" italics="true" pageId="68" pageNumber="67">Reddyanus ceylonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A853FFD4FC41F9089DD4F949" box="[897,950,1759,1780]" pageId="68" pageNumber="67" rank="species">
<emphasis id="0AACEADBA853FFD4FC41F9089DFAF949" bold="true" box="[897,920,1759,1780]" pageId="68" pageNumber="67">sp</emphasis>
.
<emphasis id="0AACEADBA853FFD4FC63F9089DD2F949" bold="true" box="[931,944,1759,1780]" pageId="68" pageNumber="67">n</emphasis>
.
</taxonomicNameLabel>
male paratype from locality 15CI. External (327), dorsal (328), internal (329) and ventral (330) views. Extracted after 4th ecdysis. A second hemispermatophore examined from a second male after 5th ecdysis was similar. Scale bar: 250 µm. Abbreviations:
<emphasis id="0AACEADBA853FFD4FC7FF8C69DB2F89B" box="[959,976,1809,1830]" italics="true" pageId="68" pageNumber="67">bl</emphasis>
, basal lobe;
<emphasis id="0AACEADBA853FFD4FB8FF8C69A36F89B" box="[1103,1108,1809,1830]" italics="true" pageId="68" pageNumber="67">f</emphasis>
, flagellum;
<emphasis id="0AACEADBA853FFD4FB0EF8C69A86F89B" box="[1230,1252,1809,1830]" italics="true" pageId="68" pageNumber="67">ml</emphasis>
, median lobe;
<emphasis id="0AACEADBA853FFD4FAB9F8C69BF8F89B" box="[1401,1434,1809,1830]" italics="true" pageId="68" pageNumber="67">mlc</emphasis>
, median lobe carina;
<emphasis id="0AACEADBA853FFD4FE43F8FD9FE8F882" box="[387,394,1834,1855]" italics="true" pageId="68" pageNumber="67">t</emphasis>
, trunk.
</paragraph>
</caption>
<caption id="6CA76641A852FFD5FF00FA029D6FF9BB" ID-DOI="http://doi.org/10.5281/zenodo.7124659" ID-Zenodo-Dep="7124659" httpUri="https://zenodo.org/record/7124659/files/figure.png" pageId="69" pageNumber="68" startId="69.[192,278,1493,1517]" targetBox="[191,1440,175,1465]" targetPageId="69">
<paragraph id="386736C9A852FFD5FF00FA029D6FF9BB" blockId="69.[192,1442,1493,1542]" pageId="69" pageNumber="68">
<emphasis id="0AACEADBA852FFD5FF00FA029FE6FA53" bold="true" box="[192,388,1493,1518]" pageId="69" pageNumber="68">Figures 331333:</emphasis>
<taxonomicName id="FFD84D4AA852FFD5FE4CFA0F9C0EFA50" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[396,620,1496,1517]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="69" pageNumber="68" phylum="Arthropoda" rank="species" species="ceylonensis" status="sp. nov.">
<emphasis id="0AACEADBA852FFD5FE4CFA0F9C0EFA50" box="[396,620,1496,1517]" italics="true" pageId="69" pageNumber="68">Reddyanus ceylonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A852FFD5FDB5FA0F9CC8FA50" box="[629,682,1496,1517]" pageId="69" pageNumber="68" rank="species">
<emphasis id="0AACEADBA852FFD5FDB5FA0F9CEEFA50" bold="true" box="[629,652,1496,1517]" pageId="69" pageNumber="68">sp</emphasis>
.
<emphasis id="0AACEADBA852FFD5FD58FA0F9CC7FA50" bold="true" box="[664,677,1496,1517]" pageId="69" pageNumber="68">n</emphasis>
.
</taxonomicNameLabel>
, female from locality 15CH (331), female from locality 15CI with newborns before first ecdysis (332), and male from locality 15CI (333).
</paragraph>
</caption>
<subSubSection id="70C26542A852FFD5FF00F9619FA0F956" pageId="69" pageNumber="68" type="etymology">
<paragraph id="386736C9A852FFD5FF00F9619FA0F956" blockId="69.[192,802,1718,1771]" pageId="69" pageNumber="68">
ETYMOLOGY. Named after V. A. Sanjeewa Jayarathne who collected the
<typeStatus id="E763886BA852FFD5FE43F9039FD9F956" box="[387,443,1748,1771]" pageId="69" pageNumber="68">types</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="70C26542A852FFD8FF00F8C59C3CF8ED" lastPageId="72" lastPageNumber="71" pageId="69" pageNumber="68" type="diagnosis">
<paragraph id="386736C9A852FFD8FF00F8C59C3CF8ED" blockId="69.[192,802,1810,1894]" lastBlockId="72.[192,803,1726,1872]" lastPageId="72" lastPageNumber="71" pageId="69" pageNumber="68">
DIAGNOSIS. Total length 37.1 mm (female) 45.5 mm (male). Male with slightly longer metasomal segments and telson than female. Pedipalp segments approximately the same length and width in both sexes. Pedipalp movable finger longer than manus of chela in both sexes. Pedipalps and legs with brown maculation, identical on femur and patella. First metasomal segment bears 10 carinae, second through fourth segments bear eight carinae, fifth segment bears five carinae in female and three to five in male. Posterior terminal tubercle of each dorsal carina on metasomal segments of both sexes scarcely larger than preceding tubercles. Subaculear tooth wide and rounded, dorsally with granules in four rows; six symmetrical granules in three rows and one or two granules on tip. Ratio of metasomal segment II length/ width 1.81 in male. Glabrous zone on posterior part of fifth sternite present medially in male. Pectinal teeth number
<quantity id="FF209B2CA85FFFD8FE93F8EE9FEBF8ED" box="[339,393,1849,1872]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="72" pageNumber="71" unit="in" value="12.0">12 in</quantity>
female,
<quantity id="FF209B2CA85FFFD8FE26F8EE9C7EF8ED" box="[486,540,1849,1872]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="72" pageNumber="71" unit="in" value="14.0">14 in</quantity>
male.
</paragraph>
</subSubSection>
<caption id="6CA76641A851FFD6FF00F92D9D90F896" ID-DOI="http://doi.org/10.5281/zenodo.7124661" ID-Zenodo-Dep="7124661" httpUri="https://zenodo.org/record/7124661/files/figure.png" pageId="70" pageNumber="69" startId="70.[192,278,1786,1810]" targetBox="[201,1437,224,1752]" targetPageId="70">
<paragraph id="386736C9A851FFD6FF00F92D9D90F896" blockId="70.[192,1441,1786,1835]" pageId="70" pageNumber="69">
<emphasis id="0AACEADBA851FFD6FF00F92D9FE7F8AF" bold="true" box="[192,389,1786,1810]" pageId="70" pageNumber="69">Figures 334337:</emphasis>
<taxonomicName id="FFD84D4AA851FFD6FE4DF92A9C0CF8AF" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[397,622,1789,1810]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="69" phylum="Arthropoda" rank="species" species="jayarathnei" status="sp. nov.">
<emphasis id="0AACEADBA851FFD6FE4DF92A9C0CF8AF" box="[397,622,1789,1810]" italics="true" pageId="70" pageNumber="69">Reddyanus jayarathnei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A851FFD6FDB7F92B9CCEF8AF" box="[631,684,1788,1810]" pageId="70" pageNumber="69" rank="species">
<emphasis id="0AACEADBA851FFD6FDB7F92B9CECF8AC" bold="true" box="[631,654,1788,1809]" pageId="70" pageNumber="69">sp</emphasis>
.
<emphasis id="0AACEADBA851FFD6FD5AF92B9CC5F8AC" bold="true" box="[666,679,1788,1809]" pageId="70" pageNumber="69">n</emphasis>
.
</taxonomicNameLabel>
, paratypes.
<emphasis id="0AACEADBA851FFD6FCE6F92B9DADF8AC" bold="true" box="[806,975,1788,1810]" pageId="70" pageNumber="69">Figures 334335</emphasis>
. Male in dorsal (334) and ventral (335) views.
<emphasis id="0AACEADBA851FFD6FF00F8C29F0AF897" bold="true" box="[192,360,1813,1835]" pageId="70" pageNumber="69">Figures 336337</emphasis>
. Female in dorsal (336) and ventral (337) views. Scale bar: 10 mm.
</paragraph>
</caption>
<caption id="6CA76641A850FFD7FF00F9169CEAF880" ID-DOI="http://doi.org/10.5281/zenodo.7124663" ID-Zenodo-Dep="7124663" httpUri="https://zenodo.org/record/7124663/files/figure.png" pageId="71" pageNumber="70" startId="71.[192,278,1729,1753]" targetBox="[191,1440,191,1701]" targetPageId="71">
<paragraph id="386736C9A850FFD7FF00F9169CEAF880" blockId="71.[192,1440,1729,1853]" pageId="71" pageNumber="70">
<emphasis id="0AACEADBA850FFD7FF00F9169FE1F967" bold="true" box="[192,387,1729,1754]" pageId="71" pageNumber="70">Figures 338348:</emphasis>
<taxonomicName id="FFD84D4AA850FFD7FE49F9139C0AF964" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[393,616,1732,1753]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="71" pageNumber="70" phylum="Arthropoda" rank="species" species="jayarathnei" status="sp. nov.">
<emphasis id="0AACEADBA850FFD7FE49F9139C0AF964" box="[393,616,1732,1753]" italics="true" pageId="71" pageNumber="70">Reddyanus jayarathnei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A850FFD7FDAEF9139CC0F964" box="[622,674,1732,1753]" pageId="71" pageNumber="70" rank="species">
<emphasis id="0AACEADBA850FFD7FDAEF9139CE7F964" bold="true" box="[622,645,1732,1753]" pageId="71" pageNumber="70">sp</emphasis>
.
<emphasis id="0AACEADBA850FFD7FD50F9139CFFF964" bold="true" box="[656,669,1732,1753]" pageId="71" pageNumber="70">n</emphasis>
.
</taxonomicNameLabel>
, paratypes.
<emphasis id="0AACEADBA850FFD7FCD8F9149A42F965" bold="true" box="[792,1056,1731,1753]" pageId="71" pageNumber="70">Figures 338, 340, 343348</emphasis>
. Male, chelicerae, carapace and tergites IIII (338), sternopectinal region (340). Pedipalp chela, dorsal (343), external (344) and ventral (345) views. Pedipalp patella, dorsal (346) and external (347) views. Pedipalp femur and trochanter dorsal (348) view. The trichobothrial pattern is indicated in Figures 344348.
<emphasis id="0AACEADBA850FFD7FEB1F8D99C2BF89E" bold="true" box="[369,585,1806,1828]" pageId="71" pageNumber="70">Figures 339, 341342</emphasis>
. Female, chelicerae, carapace and tergites IIII (339), sternopectinal region and sternite III (341), and pedipalp chela dorsal view (342).
</paragraph>
</caption>
<caption id="6CA76641A85FFFD8FF00F9979F8CF934" ID-DOI="http://doi.org/10.5281/zenodo.7124665" ID-Zenodo-Dep="7124665" httpUri="https://zenodo.org/record/7124665/files/figure.png" pageId="72" pageNumber="71" startId="72.[192,278,1600,1624]" targetBox="[206,1421,175,1560]" targetPageId="72">
<paragraph id="386736C9A85FFFD8FF00F9979F8CF934" blockId="72.[192,1440,1600,1673]" pageId="72" pageNumber="71">
<emphasis id="0AACEADBA85FFFD8FF00F9979FEAF9E5" bold="true" box="[192,392,1600,1624]" pageId="72" pageNumber="71">Figures 349354:</emphasis>
<taxonomicName id="FFD84D4AA85FFFD8FE53F9959C14F9EA" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[403,630,1602,1623]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="72" pageNumber="71" phylum="Arthropoda" rank="species" species="jayarathnei" status="sp. nov.">
<emphasis id="0AACEADBA85FFFD8FE53F9959C14F9EA" box="[403,630,1602,1623]" italics="true" pageId="72" pageNumber="71">Reddyanus jayarathnei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A85FFFD8FD41F9959CD9F9EA" box="[641,699,1602,1623]" pageId="72" pageNumber="71" rank="species">
<emphasis id="0AACEADBA85FFFD8FD41F9959CFAF9EA" bold="true" box="[641,664,1602,1623]" pageId="72" pageNumber="71">sp</emphasis>
.
<emphasis id="0AACEADBA85FFFD8FD69F9959CD4F9EA" bold="true" box="[681,694,1602,1623]" pageId="72" pageNumber="71">n</emphasis>
.
</taxonomicNameLabel>
, paratypes.
<emphasis id="0AACEADBA85FFFD8FCFBF9959D85F9EA" bold="true" box="[827,999,1602,1624]" pageId="72" pageNumber="71">Figures 349351</emphasis>
. Male, metasoma and telson, lateral (349), ventral (350), and dorsal (351) views.
<emphasis id="0AACEADBA85FFFD8FDFEF98D9C85F9CD" bold="true" box="[574,743,1626,1648]" pageId="72" pageNumber="71">Figures 352354</emphasis>
. Female, metasoma and telson, lateral (352), ventral (353), and dorsal (354) views. Scale bar: 10 mm.
</paragraph>
</caption>
<subSubSection id="70C26542A85FFFDDFCFEF9699B74F8D1" lastPageId="77" lastPageNumber="76" pageId="72" pageNumber="71" type="description">
<paragraph id="386736C9A85FFFDAFCFEF9699E9AF963" blockId="72.[830,1447,1726,1872]" lastBlockId="74.[192,809,1704,1909]" lastPageId="74" lastPageNumber="73" pageId="72" pageNumber="71">
DESCRIPTION. Total length 37.1 mm (female
<typeStatus id="E763886BA85FFFD8FAECF9699BEAF968" box="[1324,1416,1726,1749]" pageId="72" pageNumber="71" type="paratype">paratype</typeStatus>
) 45.5 mm (male
<typeStatus id="E763886BA85FFFD8FC2EF90A9A32F949" box="[1006,1104,1757,1780]" pageId="72" pageNumber="71" type="holotype">holotype</typeStatus>
). The habitus is shown in
<figureCitation id="A0E32A4CA85FFFD8FAB2F90A9DC0F8AF" captionStart="Figures 334337" captionStartId="70.[192,278,1786,1810]" captionTargetBox="[201,1437,224,1752]" captionTargetId="figure-11@70.[192,1439,216,1757]" captionTargetPageId="70" captionText="Figures 334337: Reddyanus jayarathnei sp. n., paratypes. Figures 334335. Male in dorsal (334) and ventral (335) views. Figures 336337. Female in dorsal (336) and ventral (337) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7124661" httpUri="https://zenodo.org/record/7124661/files/figure.png" pageId="72" pageNumber="71">Figs. 334337</figureCitation>
. For measurements and ratios see
<tableCitation id="755A0372A85FFFD8FAE6F92C9BC2F8AF" box="[1318,1440,1787,1810]" captionStart-0="Table 4" captionStart-1="Table 5" captionStartId-0="86.[192,257,1428,1452]" captionStartId-1="86.[192,257,1869,1893]" captionTargetPageId-0="86" captionTargetPageId-1="86" captionText-0="Table 4: Comparative measurements of adults of Reddyanus ranawanai sp. n. and R. jayarathnei sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-1="Table 5: Comparison among Sri Lankan Reddyanus species (specimens), based upon selected morphometric ratios of adult males. Abbreviations: length (L), width (W), depth (D)." httpUri-0="http://table.plazi.org/id/6CA76641A841FFC6FF00FA439D16FA79" httpUri-1="http://table.plazi.org/id/6CA76641A841FFC6FF00F89A9CBAF8C0" pageId="72" pageNumber="71" tableUuid-0="6CA76641A841FFC6FF00FA439D16FA79" tableUuid-1="6CA76641A841FFC6FF00F89A9CBAF8C0">Tables 45</tableCitation>
. For position and distribution of trichobothria of pedipalps see
<figureCitation id="A0E32A4CA85FFFD8FCB0F8EE9A6FF8ED" box="[880,1037,1849,1872]" captionStart="Figures 338348" captionStartId="71.[192,278,1729,1753]" captionTargetBox="[191,1440,191,1701]" captionTargetId="figure-68@71.[192,1439,192,1700]" captionTargetPageId="71" captionText="Figures 338348: Reddyanus jayarathnei sp. n., paratypes. Figures 338, 340, 343348. Male, chelicerae, carapace and tergites IIII (338), sternopectinal region (340). Pedipalp chela, dorsal (343), external (344) and ventral (345) views. Pedipalp patella, dorsal (346) and external (347) views. Pedipalp femur and trochanter dorsal (348) view. The trichobothrial pattern is indicated in Figures 344348. Figures 339, 341342. Female, chelicerae, carapace and tergites IIII (339), sternopectinal region and sternite III (341), and pedipalp chela dorsal view (342)." figureDoi="http://doi.org/10.5281/zenodo.7124663" httpUri="https://zenodo.org/record/7124663/files/figure.png" pageId="72" pageNumber="71">Figs. 344348</figureCitation>
. The male has slightly longer metasomal segments and telson than the female (
<figureCitation id="A0E32A4CA85DFFDAFD73F97F9E88F963" captionStart="Figures 349354" captionStartId="72.[192,278,1600,1624]" captionTargetBox="[206,1421,175,1560]" captionTargetId="figure-11@72.[192,1439,168,1571]" captionTargetPageId="72" captionText="Figures 349354: Reddyanus jayarathnei sp. n., paratypes. Figures 349351. Male, metasoma and telson, lateral (349), ventral (350), and dorsal (351) views. Figures 352354. Female, metasoma and telson, lateral (352), ventral (353), and dorsal (354) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7124665" httpUri="https://zenodo.org/record/7124665/files/figure.png" pageId="74" pageNumber="73">Figs. 349 354</figureCitation>
).
</paragraph>
<caption id="6CA76641A85EFFD9FF00F8879D47F83D" ID-DOI="http://doi.org/10.5281/zenodo.7124668" ID-Zenodo-Dep="7124668" httpUri="https://zenodo.org/record/7124668/files/figure.png" pageId="73" pageNumber="72" startId="73.[192,278,1872,1896]" targetBox="[191,1440,167,1843]" targetPageId="73">
<paragraph id="386736C9A85EFFD9FF00F8879D47F83D" blockId="73.[192,1440,1872,1920]" pageId="73" pageNumber="72">
<emphasis id="0AACEADBA85EFFD9FF00F8879FE4F8D5" bold="true" box="[192,390,1872,1896]" pageId="73" pageNumber="72">Figures 355357:</emphasis>
<taxonomicName id="FFD84D4AA85EFFD9FE4FF8859C12F8DA" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[399,624,1874,1895]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="73" pageNumber="72" phylum="Arthropoda" rank="species" species="jayarathnei" status="sp. nov.">
<emphasis id="0AACEADBA85EFFD9FE4FF8859C12F8DA" box="[399,624,1874,1895]" italics="true" pageId="73" pageNumber="72">Reddyanus jayarathnei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A85EFFD9FDB9F8859CD3F8DA" box="[633,689,1874,1895]" pageId="73" pageNumber="72" rank="species">
<emphasis id="0AACEADBA85EFFD9FDB9F8859CF2F8DA" bold="true" box="[633,656,1874,1895]" pageId="73" pageNumber="72">sp</emphasis>
.
<emphasis id="0AACEADBA85EFFD9FD5FF8859CCEF8DA" bold="true" box="[671,684,1874,1895]" pageId="73" pageNumber="72">n</emphasis>
.
</taxonomicNameLabel>
, female paratype (355) and male holotype (356) at the type locality (357), Galle district, Kanneliya Rain Forest, 06°15'04&quot; N 80°20'18&quot; E.
</paragraph>
</caption>
<caption id="6CA76641A85DFFDAFF00F99B9D5CF9C1" ID-DOI="http://doi.org/10.5281/zenodo.7124670" ID-Zenodo-Dep="7124670" httpUri="https://zenodo.org/record/7124670/files/figure.png" pageId="74" pageNumber="73" startId="74.[192,278,1612,1636]" targetBox="[210,1425,149,1578]" targetPageId="74">
<paragraph id="386736C9A85DFFDAFF00F99B9D5CF9C1" blockId="74.[192,1441,1612,1660]" pageId="74" pageNumber="73">
<emphasis id="0AACEADBA85DFFDAFF00F99B9FE4F9D9" bold="true" box="[192,390,1612,1636]" pageId="74" pageNumber="73">Figures 358361:</emphasis>
<taxonomicName id="FFD84D4AA85DFFDAFE50F9999C59F9DE" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Vachon" baseAuthorityYear="1982" box="[400,571,1614,1635]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="74" pageNumber="73" phylum="Arthropoda" rank="species" species="loebli">
<emphasis id="0AACEADBA85DFFDAFE50F9999C59F9DE" box="[400,571,1614,1635]" italics="true" pageId="74" pageNumber="73">Reddyanus loebli</emphasis>
</taxonomicName>
from locality 15CG.
<emphasis id="0AACEADBA85DFFDAFCD7F99A9DA0F9DF" bold="true" box="[791,962,1613,1635]" pageId="74" pageNumber="73">Figures 358359</emphasis>
. Male in dorsal (358) and ventral (359) views.
<emphasis id="0AACEADBA85DFFDAFF00F9B19F05F9C6" bold="true" box="[192,359,1638,1660]" pageId="74" pageNumber="73">Figures 360361</emphasis>
. Female in dorsal (360) and ventral (361) views.
</paragraph>
</caption>
<paragraph id="386736C9A85DFFDAFF00F9329D9CF8A5" blockId="74.[192,809,1704,1909]" lastBlockId="74.[830,1441,1703,1908]" pageId="74" pageNumber="73">
<emphasis id="0AACEADBA85DFFDAFF00F9329F59F940" bold="true" box="[192,315,1765,1789]" pageId="74" pageNumber="73">Coloration</emphasis>
(
<figureCitation id="A0E32A4CA85DFFDAFE8BF9319F8AF940" box="[331,488,1766,1789]" captionStart="Figures 334337" captionStartId="70.[192,278,1786,1810]" captionTargetBox="[201,1437,224,1752]" captionTargetId="figure-11@70.[192,1439,216,1757]" captionTargetPageId="70" captionText="Figures 334337: Reddyanus jayarathnei sp. n., paratypes. Figures 334335. Male in dorsal (334) and ventral (335) views. Figures 336337. Female in dorsal (336) and ventral (337) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7124661" httpUri="https://zenodo.org/record/7124661/files/figure.png" pageId="74" pageNumber="73">Figs. 334337</figureCitation>
). Entire mesosoma and carapace dark, almost black with yellow to reddish spots. Chelicera strongly reticulated, with spotted fingers. Pedipalps dorsally and laterally reddish, with brown to black spots, identical on femur and patella. Pedipalp chela fingers reddish black. Legs with same color and pattern as pedipalp femur and patella. Metasomal segments reddish brown with the spots. Telson reddish brown with spots.
</paragraph>
<paragraph id="386736C9A85DFFDDFCFEF8C89FC2FA4F" blockId="74.[830,1441,1703,1908]" lastBlockId="77.[192,803,1342,1898]" lastPageId="77" lastPageNumber="76" pageId="74" pageNumber="73">
<emphasis id="0AACEADBA85DFFDAFCFEF8C89A04F88A" bold="true" box="[830,1126,1823,1847]" pageId="74" pageNumber="73">Mesosoma and carapace</emphasis>
(
<figureCitation id="A0E32A4CA85DFFDAFBBCF8F79B41F88A" box="[1148,1315,1824,1847]" captionStart="Figures 338348" captionStartId="71.[192,278,1729,1753]" captionTargetBox="[191,1440,191,1701]" captionTargetId="figure-68@71.[192,1439,192,1700]" captionTargetPageId="71" captionText="Figures 338348: Reddyanus jayarathnei sp. n., paratypes. Figures 338, 340, 343348. Male, chelicerae, carapace and tergites IIII (338), sternopectinal region (340). Pedipalp chela, dorsal (343), external (344) and ventral (345) views. Pedipalp patella, dorsal (346) and external (347) views. Pedipalp femur and trochanter dorsal (348) view. The trichobothrial pattern is indicated in Figures 344348. Figures 339, 341342. Female, chelicerae, carapace and tergites IIII (339), sternopectinal region and sternite III (341), and pedipalp chela dorsal view (342)." figureDoi="http://doi.org/10.5281/zenodo.7124663" httpUri="https://zenodo.org/record/7124663/files/figure.png" pageId="74" pageNumber="73">Figs. 338341</figureCitation>
). Carapace without carinae but with large granules. Mesosoma with one granulated median carina. Tergite VII pentacarinate. Fifth sternite with glabrous zone on posterior medial part in the male. Seventh sternite with four incomplete carinae, sparsely granulate. Pectinal tooth count is
<quantity id="FF209B2CA85AFFDDFD02FAAA9C98FA29" box="[706,762,1405,1428]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="77" pageNumber="76" unit="in" value="12.0">12 in</quantity>
the female,
<quantity id="FF209B2CA85AFFDDFEDCFA4B9F38FA0E" box="[284,346,1436,1459]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="77" pageNumber="76" unit="in" value="14.0">14 in</quantity>
the male. Pectines with three marginal lamellae and seven middle lamellae. The lamellae bear numerous pale setae.
</paragraph>
<caption id="6CA76641A85CFFDBFF00F8F89FA9F8C5" ID-DOI="http://doi.org/10.5281/zenodo.7124672" ID-Zenodo-Dep="7124672" httpUri="https://zenodo.org/record/7124672/files/figure.png" pageId="75" pageNumber="74" startId="75.[192,278,1839,1863]" targetBox="[213,1417,167,1810]" targetPageId="75">
<paragraph id="386736C9A85CFFDBFF00F8F89FA9F8C5" blockId="75.[192,1441,1839,1912]" pageId="75" pageNumber="74">
<emphasis id="0AACEADBA85CFFDBFF00F8F89FE6F8FA" bold="true" box="[192,388,1839,1863]" pageId="75" pageNumber="74">Figures 362365:</emphasis>
<taxonomicName id="FFD84D4AA85CFFDBFE4AF8E59C53F8FA" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Vachon" baseAuthorityYear="1982" box="[394,561,1842,1863]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="75" pageNumber="74" phylum="Arthropoda" rank="species" species="loebli">
<emphasis id="0AACEADBA85CFFDBFE4AF8E59C53F8FA" box="[394,561,1842,1863]" italics="true" pageId="75" pageNumber="74">Reddyanus loebli</emphasis>
</taxonomicName>
from locality 15CG.
<emphasis id="0AACEADBA85CFFDBFCC2F8E69DC9F8FB" bold="true" box="[770,939,1841,1863]" pageId="75" pageNumber="74">Figures 362, 364</emphasis>
. Male, chelicerae, carapace and tergites IIII (362), sternopectinal region and sternite III (364).
<emphasis id="0AACEADBA85CFFDBFD4AF89E9D5CF8E2" bold="true" box="[650,830,1865,1887]" pageId="75" pageNumber="74">Figures 363, 365</emphasis>
. Female, chelicerae, carapace and tergites IIII (363), and sternopectinal region (365).
</paragraph>
</caption>
<caption id="6CA76641A85BFFDCFF00F8E79E9CF8C7" ID-DOI="http://doi.org/10.5281/zenodo.7124674" ID-Zenodo-Dep="7124674" httpUri="https://zenodo.org/record/7124674/files/figure.png" pageId="76" pageNumber="75" startId="76.[192,278,1840,1864]" targetBox="[205,1406,172,1808]" targetPageId="76">
<paragraph id="386736C9A85BFFDCFF00F8E79E9CF8C7" blockId="76.[192,1440,1840,1914]" pageId="76" pageNumber="75">
<emphasis id="0AACEADBA85BFFDCFF00F8E79FE6F8F5" bold="true" box="[192,388,1840,1864]" pageId="76" pageNumber="75">Figures 366371:</emphasis>
<taxonomicName id="FFD84D4AA85BFFDCFE4CF8E49C56F8F5" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Vachon" baseAuthorityYear="1982" box="[396,564,1843,1864]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="76" pageNumber="75" phylum="Arthropoda" rank="species" species="loebli">
<emphasis id="0AACEADBA85BFFDCFE4CF8E49C56F8F5" box="[396,564,1843,1864]" italics="true" pageId="76" pageNumber="75">Reddyanus loebli</emphasis>
</taxonomicName>
from locality 15CG.
<emphasis id="0AACEADBA85BFFDCFCCAF8E59DD3F8FA" bold="true" box="[778,945,1842,1864]" pageId="76" pageNumber="75">Figures 366368</emphasis>
. Male, metasoma and telson, lateral (366), ventral (367), and dorsal (368) views.
<emphasis id="0AACEADBA85BFFDCFE37F89C9CC3F8DD" bold="true" box="[503,673,1867,1889]" pageId="76" pageNumber="75">Figures 369371</emphasis>
. Female, metasoma and telson, lateral (369), ventral (370), and dorsal (371) v iews.
</paragraph>
</caption>
<caption id="6CA76641A85AFFDDFF00FBA29FB2FAB4" ID-DOI="http://doi.org/10.5281/zenodo.7124676" ID-Zenodo-Dep="7124676" httpUri="https://zenodo.org/record/7124676/files/figure.png" pageId="77" pageNumber="76" startId="77.[192,278,1141,1165]" targetBox="[202,1423,152,1109]" targetPageId="77">
<paragraph id="386736C9A85AFFDDFF00FBA29FB2FAB4" blockId="77.[192,1440,1141,1289]" pageId="77" pageNumber="76">
<emphasis id="0AACEADBA85AFFDDFF00FBA29FE4FB30" bold="true" box="[192,390,1141,1165]" pageId="77" pageNumber="76">Figures 372375:</emphasis>
Distal trunk, capsule region and flagellum of left hemispermatophore of
<taxonomicName id="FFD84D4AA85AFFDDFBADFBA09B75FB31" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Vachon" baseAuthorityYear="1982" box="[1133,1303,1143,1164]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="loebli">
<emphasis id="0AACEADBA85AFFDDFBADFBA09B75FB31" box="[1133,1303,1143,1164]" italics="true" pageId="77" pageNumber="76">Reddyanus loebli</emphasis>
</taxonomicName>
from locality 15CH. External (372), dorsal (373), internal (374) and ventral (375) views. Extracted after 5th ecdysis. Note: this hemispermatopore was folded along a dorsal line and flattened; dorsal and ventral views show it as unfolded and flat, hence the broad appearance of the trunk. Compare with Figs. 283286 and 327330 which show typical
<taxonomicName id="FFD84D4AA85AFFDDFBF3FB159AFEFB6A" authorityName="Vachon" authorityYear="1972" box="[1075,1180,1218,1239]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="77" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA85AFFDDFBF3FB159AFEFB6A" box="[1075,1180,1218,1239]" italics="true" pageId="77" pageNumber="76">Reddyanus</emphasis>
</taxonomicName>
hemispermatophores with trunk in natural curved conformation. Scale bar: 500 µm. Abbreviations:
<emphasis id="0AACEADBA85AFFDDFC68FB0C9DD9FB4D" box="[936,955,1243,1264]" italics="true" pageId="77" pageNumber="76">bl</emphasis>
, basal lobe;
<emphasis id="0AACEADBA85AFFDDFB81FB0C9A2AFB4D" box="[1089,1096,1243,1264]" italics="true" pageId="77" pageNumber="76">f</emphasis>
, flagellum;
<emphasis id="0AACEADBA85AFFDDFB04FB0C9ABEFB4D" box="[1220,1244,1243,1264]" italics="true" pageId="77" pageNumber="76">ml</emphasis>
, median lobe;
<emphasis id="0AACEADBA85AFFDDFAB7FB0C9BFBFB4D" box="[1399,1433,1243,1264]" italics="true" pageId="77" pageNumber="76">mlc</emphasis>
, median lobe carina;
<emphasis id="0AACEADBA85AFFDDFE43FB239FE8FAB4" box="[387,394,1268,1289]" italics="true" pageId="77" pageNumber="76">t</emphasis>
, trunk.
</paragraph>
</caption>
<paragraph id="386736C9A85AFFDDFF00FA2D9A58FAC9" blockId="77.[192,803,1342,1898]" lastBlockId="77.[830,1441,1342,1667]" pageId="77" pageNumber="76">
<emphasis id="0AACEADBA85AFFDDFF00FA2D9FD9F9AF" bold="true" box="[192,443,1530,1554]" pageId="77" pageNumber="76">Metasoma and telson</emphasis>
(
<figureCitation id="A0E32A4CA85AFFDDFE0EFA2C9C0CF9AF" box="[462,622,1531,1554]" captionStart="Figures 209224" captionStartId="50.[192,278,1736,1760]" captionTargetBox="[208,1434,174,1703]" captionTargetId="figure-11@50.[205,1452,168,1707]" captionTargetPageId="50" captionText="Figures 209224: Metasomal segments IIIII, lateral view of Sri Lankan Reddyanus and Isometrus species. Figures 209210. Reddyanus basilicus, male (209) and female (210) from locality 15CR. Figures 211212. R. besucheti, male holotype (211) and female from locality 15CG (212). Figures 213214. R. ceylonensis sp. n., male holotype (213) and female paratype from locality 15CI (214). Figures 215216. R. jayarathnei sp. n., male (215) and female (216) paratypes. Figures 217218. R. loebli, male (217) and female (218) from locality 15CG. Figures 219220. R. ranawanai sp. n., male holotype (219) and female paratype (220). Figures 221222. Isometrus maculatus, male from locality 15CP (221) and female from locality 15CI (222). Figures 223 224. I. thwaitesi, male from locality 15CH (223) and female from locality 15CO (224)." figureDoi="http://doi.org/10.5281/zenodo.7124622" httpUri="https://zenodo.org/record/7124622/files/figure.png" pageId="77" pageNumber="76">Figs. 215216</figureCitation>
,
<figureCitation id="A0E32A4CA85AFFDDFDBDFA2C9CBEF9AF" box="[637,732,1531,1554]" captionStart="Figures 349354" captionStartId="72.[192,278,1600,1624]" captionTargetBox="[206,1421,175,1560]" captionTargetId="figure-11@72.[192,1439,168,1571]" captionTargetPageId="72" captionText="Figures 349354: Reddyanus jayarathnei sp. n., paratypes. Figures 349351. Male, metasoma and telson, lateral (349), ventral (350), and dorsal (351) views. Figures 352354. Female, metasoma and telson, lateral (352), ventral (353), and dorsal (354) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7124665" httpUri="https://zenodo.org/record/7124665/files/figure.png" pageId="77" pageNumber="76">349354</figureCitation>
,
<figureCitation id="A0E32A4CA85AFFDDFD2CFA2C9E8AF98C" captionStart="Figures 403429" captionStartId="85.[192,278,744,768]" captionTargetId="figure-11@84.[172,1464,185,1878]" captionTargetPageId="84" captionText="Figures 403429: Telson, lateral view of Sri Lankan Buthidae and Chaerilidae species. Figures 403404. Isometrus maculatus, male from locality 15CP (403) and female from locality 15CI (404). Figures 405406. I. thwaitesi, male from locality 15CH (405) and female from locality 15CO (406). Figures 407408. Lychas srilankensis, male (407) and female (408) from locality 15CJ. Figures 409410. Reddyanus basilicus, male (409) and female (410) from locality 15CR. Figures 411412. R. besucheti, male holotype (411) and female from locality 15CG (412). Figures 413414. R. ceylonensis sp. n., male holotype (413) and female paratype (414) from locality 15CI. Figures 415416. R. jayarathnei sp. n., male (415) and female (416) paratypes. Figures 417418. R. loebli, male (417) and female (418) from locality 15CG. Figures 419420. R. ranawanai sp. n., male holotype (419) and female paratype (420). Figures 421422: Buthoscorpio sarasinorum, male from locality 15CF (421) and female from locality 15CG (422). Figures 423424. Charmus laneus, male (423) and female (424) from locality 15CO. Figures 425426. Charmus saradieli sp. n. male paratype (425) (MHNG) and female holotype (426). Figurs 427. Hottentotta tamulus, male from locality 15CK (427). Figures 428429. Chaerilus ceylonensis, male (428) and female (429) from locality 15CD." figureDoi="http://doi.org/10.5281/zenodo.7124696" httpUri="https://zenodo.org/record/7124696/files/figure.png" pageId="77" pageNumber="76">415 416</figureCitation>
). The first metasomal segment bears 10 carinae and the second to the fourth segments bear eight carinae, the fifth segment bears five carinae well developed in the female and only indicated or absent in the male. Lateral inframedian carinae may be indicated on second metasomal segment. Ventral carina present on telson. Intercarinal surfaces of metasoma granulated including dorsal surface mainly in the female. In both sexes, posterior terminal tubercle of each dorsal carina on metasomal segments not enlarged. Telson elongate, with subaculear tooth wide and rounded, dorsally with granules in four rows; six symmetrical granules in three rows and one or two granules on the tip.
</paragraph>
<paragraph id="386736C9A85AFFDDFCFEFAAC9A8CF994" blockId="77.[830,1441,1342,1667]" pageId="77" pageNumber="76">
<emphasis id="0AACEADBA85AFFDDFCFEFAAC9DCFFA2E" bold="true" box="[830,941,1403,1427]" pageId="77" pageNumber="76">Pedipalps</emphasis>
(
<figureCitation id="A0E32A4CA85AFFDDFC7EFAAB9A3AFA2E" box="[958,1112,1404,1427]" captionStart="Figures 225240" captionStartId="51.[192,278,1493,1517]" captionTargetBox="[193,1432,180,1451]" captionTargetId="figure-6@51.[192,1439,179,1464]" captionTargetPageId="51" captionText="Figures 225240: Pedipalp, dorsal view of Sri Lankan Reddyanus and Isometrus species. Figures 225226. Reddyanus basilicus, male (225) and female (226) from locality 15CR. Figures 227228. R. besucheti, male from locality 15CD (227) and female from locality 15CG (228). Figures 229230. R. ceylonensis sp. n., male holotype (229) and female paratype from locality 15CI (230). Figures 231232. R. jayarathnei sp. n., male (231) and female (232) paratypes. Figures 233234. R. loebli, male (233) and female (234) from locality 15CG. Figures 235236. R. ranawanai sp. n., male holotype (235) and female paratype (236). Figures 237238. Isometrus maculatus, male from locality 15CP (237) and female from locality 15CI (238). Figures 239240. I. thwaitesi, male from locality 15CH (239) and female from locality 15CO (240)." figureDoi="http://doi.org/10.5281/zenodo.7124624" httpUri="https://zenodo.org/record/7124624/files/figure.png" pageId="77" pageNumber="76">Figs. 231232</figureCitation>
,
<figureCitation id="A0E32A4CA85AFFDDFBA8FAAB9AA7FA2E" box="[1128,1221,1404,1427]" captionStart="Figures 338348" captionStartId="71.[192,278,1729,1753]" captionTargetBox="[191,1440,191,1701]" captionTargetId="figure-68@71.[192,1439,192,1700]" captionTargetPageId="71" captionText="Figures 338348: Reddyanus jayarathnei sp. n., paratypes. Figures 338, 340, 343348. Male, chelicerae, carapace and tergites IIII (338), sternopectinal region (340). Pedipalp chela, dorsal (343), external (344) and ventral (345) views. Pedipalp patella, dorsal (346) and external (347) views. Pedipalp femur and trochanter dorsal (348) view. The trichobothrial pattern is indicated in Figures 344348. Figures 339, 341342. Female, chelicerae, carapace and tergites IIII (339), sternopectinal region and sternite III (341), and pedipalp chela dorsal view (342)." figureDoi="http://doi.org/10.5281/zenodo.7124663" httpUri="https://zenodo.org/record/7124663/files/figure.png" pageId="77" pageNumber="76">342348</figureCitation>
). Femur and patella only very sparsely hirsute, with complete carinae, granulated. Dorsal carinae indicated on chela manus in both sexes. Sixth row of granules on movable finger with one external granule. The seventh row of granules on the fixed finger without additional granules.
</paragraph>
<paragraph id="386736C9A85AFFDDFCFEF9F89DE9F93E" blockId="77.[830,1441,1342,1667]" pageId="77" pageNumber="76">
<emphasis id="0AACEADBA85AFFDDFCFEF9F89D11F9FA" bold="true" box="[830,883,1583,1607]" pageId="77" pageNumber="76">Legs</emphasis>
(
<figureCitation id="A0E32A4CA85AFFDDFC45F9E79D80F9FA" box="[901,994,1584,1607]" captionStart="Figures 201208" captionStartId="49.[192,278,1057,1081]" captionTargetBox="[204,1438,173,1022]" captionTargetId="figure-6@49.[192,1439,168,1028]" captionTargetPageId="49" captionText="Figures 201208: Distal segments of legs, retroventral view of Sri Lankan Reddyanus. Figures 201204. R. ceylonensis sp. n., male holotype from locality 15CI, legs IIV. Figures 205208. Leg IV. R. besucheti, male from locality 15CD (205), R. jayarathnei sp. n., male paratype (206), R. loebli, male from locality 15CG (207), and R. ranawanai sp. n., male holotype (208)." figureDoi="http://doi.org/10.5281/zenodo.7124612" httpUri="https://zenodo.org/record/7124612/files/figure.png" pageId="77" pageNumber="76">Fig. 206</figureCitation>
). Femur and patella bear complete carinae and are granulated. Legs hirsute without bristle combs.
</paragraph>
<paragraph id="386736C9A85AFFDDFCFEF9769B74F8D1" blockId="77.[830,1440,1697,1900]" pageId="77" pageNumber="76">
AFFINITIES. The described features distinguish
<taxonomicName id="FFD84D4AA85AFFDDFA81F9769DE3F96B" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="jayarathnei" status="sp. nov.">
<emphasis id="0AACEADBA85AFFDDFA81F9769DE3F96B" italics="true" pageId="77" pageNumber="76">R. jayarathnei</emphasis>
</taxonomicName>
<emphasis id="0AACEADBA85AFFDDFC4EF9699DB3F96B" bold="true" box="[910,977,1726,1750]" pageId="77" pageNumber="76">
<taxonomicNameLabel id="119F57A0A85AFFDDFC4EF9699DB3F96B" box="[910,977,1726,1750]" pageId="77" pageNumber="76" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
from all other species of the genus.
<taxonomicName id="FFD84D4AA85AFFDDFA49F9689DDBF949" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="jayarathnei" status="sp. nov.">
<emphasis id="0AACEADBA85AFFDDFA49F9689DDBF949" italics="true" pageId="77" pageNumber="76">R. jayarathnei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A85AFFDDFC03F90A9A62F949" box="[963,1024,1757,1780]" pageId="77" pageNumber="76" rank="species">sp. n.</taxonomicNameLabel>
is well characterized by the glabrous zone on the posterior medial part of the fifth sternite in the male, which is not as extensive in
<taxonomicName id="FFD84D4AA85AFFDDFB26F8CE9B15F88D" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[1254,1399,1817,1840]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="ranawanai" status="sp. nov.">
<emphasis id="0AACEADBA85AFFDDFB26F8CE9B15F88D" box="[1254,1399,1817,1840]" italics="true" pageId="77" pageNumber="76">R. ranawanai</emphasis>
</taxonomicName>
<emphasis id="0AACEADBA85AFFDDFA42F8CE9D31F8F2" bold="true" pageId="77" pageNumber="76">
<taxonomicNameLabel id="119F57A0A85AFFDDFA42F8CE9D31F8F2" pageId="77" pageNumber="76" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A0E32A4CA85AFFDDFCA5F8E09DA0F8F3" box="[869,962,1847,1870]" captionStart="Figures 555570" captionStartId="115.[192,278,1733,1757]" captionTargetBox="[233,1393,170,1705]" captionTargetId="figure-151@115.[234,1392,171,1704]" captionTargetPageId="115" captionText="Figures 555570: Key characters for distinguishing males of Sri Lankan Isometrus and Reddyanus. Figures 555556, 565 566. Pedipalp chela dorsal view, I. maculatus from locality 15CP (555), I. thwaitesi from locality 15CH (556), R. basilicus, holotype (565), and R. besucheti, holotype (566). Figures 557558. Sternopectinal area, I. maculatus from locality 15CP (557), and I. thwaitesi from locality 15CH (558). Figures 559560, 567570. Sternite V, I. maculatus from locality 15CP (559), I. thwaitesi from locality 15CH (560), R. besucheti, from locality 15CF (567), R. ceylonensis sp. n., holotype (568), R. jayarathnei sp. n., paratype (569), and R. ranawanai sp. n., paratype (570). Figures 561564. Metasomal segment II lateral (561562) and pedipalp patella dorsal (563564), R. loebli from locality 15CG (561, 563) and R. besucheti, from locality 15CF (562, 564)." figureDoi="http://doi.org/10.5281/zenodo.7124767" httpUri="https://zenodo.org/record/7124767/files/figure.png" pageId="77" pageNumber="76">Fig. 570</figureCitation>
versus
<figureCitation id="A0E32A4CA85AFFDDFBDAF8E09A14F8F3" box="[1050,1142,1847,1870]" captionStart="Figures 555570" captionStartId="115.[192,278,1733,1757]" captionTargetBox="[233,1393,170,1705]" captionTargetId="figure-151@115.[234,1392,171,1704]" captionTargetPageId="115" captionText="Figures 555570: Key characters for distinguishing males of Sri Lankan Isometrus and Reddyanus. Figures 555556, 565 566. Pedipalp chela dorsal view, I. maculatus from locality 15CP (555), I. thwaitesi from locality 15CH (556), R. basilicus, holotype (565), and R. besucheti, holotype (566). Figures 557558. Sternopectinal area, I. maculatus from locality 15CP (557), and I. thwaitesi from locality 15CH (558). Figures 559560, 567570. Sternite V, I. maculatus from locality 15CP (559), I. thwaitesi from locality 15CH (560), R. besucheti, from locality 15CF (567), R. ceylonensis sp. n., holotype (568), R. jayarathnei sp. n., paratype (569), and R. ranawanai sp. n., paratype (570). Figures 561564. Metasomal segment II lateral (561562) and pedipalp patella dorsal (563564), R. loebli from locality 15CG (561, 563) and R. besucheti, from locality 15CF (562, 564)." figureDoi="http://doi.org/10.5281/zenodo.7124767" httpUri="https://zenodo.org/record/7124767/files/figure.png" pageId="77" pageNumber="76">Fig. 569</figureCitation>
) and is absent in other
<collectingRegion id="FA1CF82BA85AFFDDFA41F8E09BC2F8F3" box="[1409,1440,1847,1870]" country="United Kingdom" name="Surrey" pageId="77" pageNumber="76">Sri</collectingRegion>
Lankan
<taxonomicName id="FFD84D4AA85AFFDDFC55F8829A3CF8D1" box="[917,1118,1877,1900]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="77" pageNumber="76" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="0AACEADBA85AFFDDFC55F8829A68F8D1" box="[917,1034,1877,1900]" italics="true" pageId="77" pageNumber="76">Reddyanus</emphasis>
species
</taxonomicName>
(
<figureCitation id="A0E32A4CA85AFFDDFBACF8829B69F8D1" box="[1132,1291,1877,1900]" captionStart="Figures 555570" captionStartId="115.[192,278,1733,1757]" captionTargetBox="[233,1393,170,1705]" captionTargetId="figure-151@115.[234,1392,171,1704]" captionTargetPageId="115" captionText="Figures 555570: Key characters for distinguishing males of Sri Lankan Isometrus and Reddyanus. Figures 555556, 565 566. Pedipalp chela dorsal view, I. maculatus from locality 15CP (555), I. thwaitesi from locality 15CH (556), R. basilicus, holotype (565), and R. besucheti, holotype (566). Figures 557558. Sternopectinal area, I. maculatus from locality 15CP (557), and I. thwaitesi from locality 15CH (558). Figures 559560, 567570. Sternite V, I. maculatus from locality 15CP (559), I. thwaitesi from locality 15CH (560), R. besucheti, from locality 15CF (567), R. ceylonensis sp. n., holotype (568), R. jayarathnei sp. n., paratype (569), and R. ranawanai sp. n., paratype (570). Figures 561564. Metasomal segment II lateral (561562) and pedipalp patella dorsal (563564), R. loebli from locality 15CG (561, 563) and R. besucheti, from locality 15CF (562, 564)." figureDoi="http://doi.org/10.5281/zenodo.7124767" httpUri="https://zenodo.org/record/7124767/files/figure.png" pageId="77" pageNumber="76">Figs. 567568</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>