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<mods:title id="ADB6974E6F40EE8E9A69EE91591E528C">Scorpions of Sri Lanka (Scorpiones Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat n</mods:title>
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<taxonomicName id="FFD84D4AA82EFFA9FF39FA769CE3FA04" ID-CoL="8LYZT" authority="(Karsch, 1879)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[249,641,1441,1465]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="57" pageNumber="56" phylum="Arthropoda" rank="species" species="basilicus" status="comb. nov.">
<emphasis id="0AACEADBA82EFFA9FF39FA769FBBFA04" bold="true" box="[249,473,1441,1465]" italics="true" pageId="57" pageNumber="56">Reddyanus basilicus</emphasis>
(
<bibRefCitation id="5C494B38A82EFFA9FE29FA759C1BFA04" author="Karsch" box="[489,633,1442,1465]" firstAuthor="Karsch" pageId="57" pageNumber="56" pagination="97 - 136" refId="ref41417" refString="KARSCH, F. 1879. Skorpionologische Beitrage I. and II. Mitteilungen des Munchener Entomologischen Vereins, 3: 6 - 22, 97 - 136." type="journal article" year="1879">Karsch, 1879</bibRefCitation>
)
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<taxonomicNameLabel id="119F57A0A82EFFA9FD47FA769C8AFA04" box="[647,744,1441,1465]" pageId="57" pageNumber="56" rank="species">comb. n.</taxonomicNameLabel>
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(
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,
<figureCitation id="A0E32A4CA82EFFA9FEFAFA179F00FA6A" box="[314,354,1472,1495]" captionStart="Figures 1619" captionStartId="8.[192,278,1853,1877]" captionTargetBox="[256,1384,176,1817]" captionTargetId="figure-11@8.[192,1439,170,1824]" captionTargetPageId="8" captionText="Figures 1619: Buthoscorpio sarasinorum. Figures 1617. Male from locality 15CF in dorsal (16) and ventral (17) views. Figures 1819. Female from locality 15CG in dorsal (18) and ventral (19) views." figureDoi="http://doi.org/10.5281/zenodo.7124478" httpUri="https://zenodo.org/record/7124478/files/figure.png" pageId="57" pageNumber="56">19 9</figureCitation>
,
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captionTargetPageId-19="23" captionTargetPageId-2="4" captionTargetPageId-20="24" captionTargetPageId-21="25" captionTargetPageId-22="26" captionTargetPageId-23="27" captionTargetPageId-24="30" captionTargetPageId-25="31" captionTargetPageId-26="33" captionTargetPageId-27="34" captionTargetPageId-28="35" captionTargetPageId-29="36" captionTargetPageId-3="6" captionTargetPageId-30="37" captionTargetPageId-31="38" captionTargetPageId-32="39" captionTargetPageId-33="40" captionTargetPageId-34="41" captionTargetPageId-35="42" captionTargetPageId-36="43" captionTargetPageId-37="44" captionTargetPageId-38="45" captionTargetPageId-39="46" captionTargetPageId-4="8" captionTargetPageId-40="47" captionTargetPageId-41="48" captionTargetPageId-42="49" captionTargetPageId-43="51" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="11" captionTargetPageId-8="12" captionTargetPageId-9="13" captionText-0="Figures 209224: Metasomal segments IIIII, lateral view of Sri Lankan Reddyanus and Isometrus species. Figures 209210. Reddyanus basilicus, male (209) and female (210) from locality 15CR. Figures 211212. R. besucheti, male holotype (211) and female from locality 15CG (212). Figures 213214. R. ceylonensis sp. n., male holotype (213) and female paratype from locality 15CI (214). Figures 215216. R. jayarathnei sp. n., male (215) and female (216) paratypes. Figures 217218. R. loebli, male (217) and female (218) from locality 15CG. Figures 219220. R. ranawanai sp. n., male holotype (219) and female paratype (220). Figures 221222. Isometrus maculatus, male from locality 15CP (221) and female from locality 15CI (222). Figures 223 224. I. thwaitesi, male from locality 15CH (223) and female from locality 15CO (224)." captionText-1="Figures 14: Figure 1. Sri Lankan scorpion expedition 2015 research team, the official photo. Figure 2. Collecting in the locality 15CA, within Peradeniya Univesity land. Figure 3. The main building of the Faculty of Science, Univesity of Peradeniya. Figure 4. F. Kovařík taking photo of Heterometrus swammerdami in the locality 15CB." captionText-10="Figures 4753: Figure 47. Left hemispermatophore of Buthoscorpio sarasinorum from locality 15CF. Figures 4853. Charmus laneus from locality 15CO. Figure 48. Left hemispermatophore. Figures 4953. Capsule region and flagellum. External (49), dorsal (50), internal (51) and ventral (52) views of left hemispermatophore, and externo-dorsal (53) view of right hemispermatophore presented as mirror image for comparison. Scale bars: 1 mm for 47, 500 µm for 48, 250 µm for 4952, and 200 µm for 53. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; pr, pars recta of flagellum; prf, pars reflecta of flagellum; t, trunk." captionText-11="Figures 5459: Charmus laneus from locality 15CO. Figures 5455. Male 14.7 mm long in dorsal (54) and ventral (55) views. Figures 5657. Juvenile male 7.5 mm long in dorsal (56) and ventral (57) views. Figures 5859. Female 20.1 mm long in dorsal (58) and ventral (59) views." captionText-12="Figures 6063: Charmus laneus from locality 15CO. Figures 60, 62. Male, chelicerae, carapace and tergites IIII (60) and sternopectinal region and sternites IIIVII (62). Figures 61, 63. Female, chelicerae, carapace and tergites IIII (61) and sternopectinal region and sternite III (63)." captionText-13="Figures 6465: Charmus laneus from locality 15CO, showing granulation and color pattern of male. Chelicerae, carapace and tergites (64). Coxosternal region and sternites (65). Scale bar: 500 µm." captionText-14="Figures 6667: Left chelicera of male Charmus laneus from locality 15CO. Figures 6667. Dorsal (66) and ventral (67a) view (white light + UV), and ventroexternal (67b) aspect (UV) showing 2 denticles on ventral surface of fixed finger (d1, d2). Scale bar: 200 µm." captionText-15="Figures 6870: Charmus laneus, male from locality 15CO. Metasoma V and telson under UV fluorescence, showing granulation, punctuation and setation in dorsal (68), lateral (69) and ventral (70) views. Scale bar: 500 µm." captionText-16="Figures 7173: Metasoma and telson of Charmus laneus male from locality 15CO, showing granulation, punctuation, setation and color pattern in dorsal (71), lateral (72) and ventral (73) views. Scale bar: 1 mm." captionText-17="Figures 7479: Charmus laneus from locality 15CO. Figures 7476. Male, metasoma and telson, lateral (74), ventral (75), and dorsal (76) views. Figures 7779. Female, metasoma and telson, lateral (77), ventral (78), and dorsal (79) views." captionText-18="Figures 8084: Comparing Sri Lankan Charmus female metasomas. Figures 8082. Female holotype of Charmus laneus, metasoma and telson, lateral (80), ventral (81), and dorsal (82) views. Figures 83. Female of Charmus laneus from locality 15CO, metasoma and telson dorsal. Figures 84. Female holotype of Charmus saradieli sp. n. (MHNG) metasoma and telson dorsal." captionText-19="Figures 8587: Charmus. Figures 8586. Male (85) and female (86) of Charmus laneus from locality 15CO. Figure 87. Male paratype of Charmus saradieli sp. n. (UPSL)." captionText-2="Figures 511: Figure 5. The reserarch team at a field station in the locality 15CB. Figure 6. Tea time in Girithale Wildlife Training Center close to the locality 15CD. From left P. Devasurendra, F. Kovařík and K. B. Ranawana. Figure 7. P. Rajkumar (Environmental Research, Jaffna), F. Kovařík and local people in the locality 15CK video recording Hottentotta tamulus. Figure 8. S. Jayarathne taking a picture of a dangerous snake Hypnale hypnale in the locality 15CD. Figure 9. Herpetologist S. Goonewardene and F. Kovařík in the locality 15CN. Figure 10. Meeting chief monk, in Monaragala &quot;Sri Sambodhi Viharaya&quot;, who has helped to research scorpions in the locality 15CQ. Figure 11. Profesor K. B. Ranawana in his office at University of Peradeniya." captionText-20="Figures 8891: Charmus saradieli sp. n. Figures 8889. Male paratype (MHNG) in dorsal (88) and ventral (89) views, and original labels. Figures 9091. Female holotype in dorsal (90) and ventral (91) views, and original labels. Scale bar: 5 mm." captionText-21="Figures 92100: Charmus. Figures 9295, 99100. Charmus saradieli sp. n., male paratype (MHNG), chelicerae, carapace and tergites IIV (92), sternopectinal region and sternites IIIIV (94), and telson lateral (99); female holotype, chelicerae, carapace and tergites IIII (93), sternopectinal region and sternite III (95), and telson lateral (100). Figures 9698. Charmus laneus, female holotype, chelicera and carapace (96), original label (97), and sternopectinal region and sternites IIIIV (98)." captionText-22="Figures 101106: Charmus saradieli sp. n. Figures 101103. Male paratype (MHNG), metasoma and telson, lateral (101), ventral (102), and dorsal (103) views. Figures 104106. Female holotype, metasoma and telson, lateral (104), ventral (105), and dorsal (106) views." captionText-23="Figures 107119: Charmus. Figures 107114. Charmus saradieli sp. n., female holotype. Pedipalp movable finger (107). Pedipalp chela, dorsal (108), external (109), and ventral (110) views. Pedipalp patella, dorsal (111) and external (112) views. Pedipalp femur and trochanter, internal (113) and dorsal (114) views. The trichobothrial pattern is indicated in Figures 108a 114a. Figures 115117. Charmus saradieli sp. n., male paratype (MHNG), chelicerae (115), pedipalp dorsal (116), and fourth leg dorsal (117). Figures 118119. Charmus laneus from locality 15CO. Pedipalp dorsal of male (118) and female (119)." captionText-24="Figures 120124: Hottentotta tamulus, male from locality 15CK, dorsal (120) and ventral (121) views, chelicerae, carapace and tergites IIII (122), sternopectinal region and sternites IIIIV (123), and telson lateral (124)." captionText-25="Figures 125126: Hottentotta tamulus from locality 15CK, female (125) and male (126)." captionText-26="Figures 127130: Isometrus maculatus. Figures 127128. Male from locality 15CP in dorsal (127) and ventral (128) views. Figures 129130. Female from locality 15CI in dorsal (129) and ventral (130) views." captionText-27="Figures 131134: Isometrus maculatus. Figures 131, 133. Male from locality 15CP, chelicerae, carapace and tergites IIII (131) and sternopectinal region and sternite III (133). Figures 132, 134. Female from locality 15CI, chelicerae, carapace and tergites IIII (132) and sternopectinal region and sternite III (134)." captionText-28="Figures 135137: Isometrus maculatus, female from locality 15CH (135) with newborns before first ecdysis (136), and with juveniles after first ecdysis (137)." captionText-29="Figures 138141: Isometrus thwaitesi. Figures 138139. Male from locality 15CH in dorsal (138) and ventral (139) views. Figures 140141. Female from locality 15CO in dorsal (140) and ventral (141) views." captionText-3="Figures 1215: Maps showing the known geographical distribution of the Sri Lankan species." captionText-30="Figures 142145: Isometrus thwaitesi. Figures 142, 144. Male from locality 15CH, chelicerae, carapace and tergites IIII (142) and sternopectinal region and sternite III (144). Figures 143, 145. Female from locality 15CO, chelicerae, carapace and tergites IIII (143) and sternopectinal region and sternite III (145)." captionText-31="Figures 146149: Left hemispermatophore of Isometrus thwaitesi from locality 15CH. External (146), dorsal (147), internal (148) and ventral (149) views. Note: distal part of flagellum and basal part of trunk (foot) were damaged during dissection. Scale bar: 500 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; pr, pars recta of flagellum; prf, pars reflecta of flagellum; t, trunk." captionText-32="Figures 150151: Isometrus thwaitesi from locality 15CF, male (150) and female (151)." captionText-33="Figures 152155: Lychas srilankensis from locality 15CJ. Figures 152153. Male in dorsal (152) and ventral (153) views. Figures 154155. Female in dorsal (154) and ventral (155) views." captionText-34="Figures 156157: Lychas srilankensis, male from locality 15CO in dorsal (156) and ventral (157) views." captionText-35="Figures 158165: Lychas srilankensis from locality 15CJ. Figures 158, 160, 162165. Male, chelicerae, carapace and tergites IIII (158), sternopectinal region and sternite III (160), and distal segments of legs IIV (162165), retroventral view. Figures 159, 161. Female, chelicerae, carapace and tergites IIII (159) and sternopectinal region and sternite III (161)." captionText-36="Figures 166171: Lychas srilankensis from locality 15CJ. Figures 166168. Male, metasoma and telson, lateral (166), ventral (167), and dorsal (168) views. Figures 169171. Female, metasoma and telson, lateral (169), ventral (170), and dorsal (171) views." captionText-37="Figures 172182: Lychas srilankensis from locality 15CJ. Figures 174180, 182. Male. Pedipalp chela, dorsal (174), external (175), and ventral (176) views. Pedipalp patella, dorsal (177) and external (178) views. Pedipalp femur and trochanter dorsal (179) view. The trichobothrial pattern is indicated in Figures 175179. Pedipalp movable finger (180). Complete pedipalp (182). Figures 172173, 181. Female. Pedipalp chela, dorsal (172) and external (173) views. Complete pedipalp (181)." captionText-38="Figures 183186: Right hemispermatophore of Lychas srilankensis from locality 15CN (presented as mirror image for comparison to other species). External (183), dorsal (184), internal (185) and ventral (186) views. Extracted after 3rd ecdysis. Hemispermatophores examined from a second male after 4th ecdysis were similar. Scale bar: 500 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; pr, pars recta of flagellum; prf, pars reflecta of flagellum; t, trunk." captionText-39="Figures 187190: Lychas srilankensis. Figures 187. Male after 5th ecdysis at locality 15CJ. Figures 188. Female after 5th ecdysis at locality 15CO. Figures 189. Male after 3rd ecdysis at locality 15CN. Figures 190. Female after 5th ecdysis at locality 15CJ." captionText-4="Figures 1619: Buthoscorpio sarasinorum. Figures 1617. Male from locality 15CF in dorsal (16) and ventral (17) views. Figures 1819. Female from locality 15CG in dorsal (18) and ventral (19) views." captionText-40="Figures 191192: Lychas srilankensis, female from locality 15CN with newborns before first ecdysis (191), and with juveniles after first ecdysis (192)." captionText-41="Figures 193200: Distal segments of leg IV, retroventral view of Sri Lankan Buthidae and Chaerilidae genera. Buthoscorpio sarasinorum, male from locality 15CF (193), Charmus laneus, male from locality 15CO (194), Hottentotta tamulus, male from locality 15CK (195), Isometrus maculatus, female from locality 15CI (196), Isometrus thwaitesi, female from locality 15CO (197), Lychas srilankensis, male from locality 15CJ (198), Reddyanus basilicus, male from locality 15CS (199), and Chaerilus ceylonensis, male from locality 15CD (200)." captionText-42="Figures 201208: Distal segments of legs, retroventral view of Sri Lankan Reddyanus. Figures 201204. R. ceylonensis sp. n., male holotype from locality 15CI, legs IIV. Figures 205208. Leg IV. R. besucheti, male from locality 15CD (205), R. jayarathnei sp. n., male paratype (206), R. loebli, male from locality 15CG (207), and R. ranawanai sp. n., male holotype (208)." captionText-43="Figures 225240: Pedipalp, dorsal view of Sri Lankan Reddyanus and Isometrus species. Figures 225226. Reddyanus basilicus, male (225) and female (226) from locality 15CR. Figures 227228. R. besucheti, male from locality 15CD (227) and female from locality 15CG (228). Figures 229230. R. ceylonensis sp. n., male holotype (229) and female paratype from locality 15CI (230). Figures 231232. R. jayarathnei sp. n., male (231) and female (232) paratypes. Figures 233234. R. loebli, male (233) and female (234) from locality 15CG. Figures 235236. R. ranawanai sp. n., male holotype (235) and female paratype (236). Figures 237238. Isometrus maculatus, male from locality 15CP (237) and female from locality 15CI (238). Figures 239240. I. thwaitesi, male from locality 15CH (239) and female from locality 15CO (240)." captionText-5="Figures 2023: Buthoscorpio sarasinorum. Figures 20, 22. Male from locality 15CF, chelicerae, carapace and tergites IIII (20) and sternopectinal region and sternite III (22). Figures 21, 23. Female from locality 15CG, chelicerae, carapace and tergites IIII (21) and sternopectinal region and sternite III (23)." captionText-6="Figures 2429: Buthoscorpio sarasinorum. Figures 2426. Male from locality 15CF, metasoma and telson, lateral (24), ventral (25), and dorsal (26) views. Figures 2729. Female from locality 15CG, metasoma and telson, lateral (27), ventral (28), and dorsal (29) views." captionText-7="Figures 3033: Capsule region and flagellum of left hemispermatophore of Buthoscorpio sarasinorum from locality 15CF. External (30), dorsal (31), internal (32) and ventral (33) views. Note: damaged section of pars reflecta of flagellum is interpolated as a dotted outline. Scale bar: 500 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; pr, pars recta of flagellum; prf, pars reflecta of flagellum; t, trunk." captionText-8="Figures 3438: Buthoscorpio sarasinorum. Figures 34. Male at locality 15CF. Figures 35. Female at locality 15CS under UV. Figures 3638. Female from locality 15CF (36) with newborns (37), and with juveniles after first ecdysis (38)." captionText-9="Figures 3946: Chela of pedipalps (3945) and movable finger of chela (46). Figures 3940. Buthoscorpio sarasinorum, male from locality 15CF (39) and female from locality 15CG (40). Figures 4143. Charmus laneus, male (41) and female (42) from locality 15CO, and female holotype (43). Figure 44. Charmus saradieli, sp. n., female holotype. Figures 4546. 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<figureCitation id="A0E32A4CA82EFFA9FD85FA179CC7FA6A" box="[581,677,1472,1495]" captionStart="Figures 241259" captionStartId="52.[192,278,1733,1757]" captionTargetBox="[238,1387,153,1699]" captionTargetId="figure-11@52.[234,1392,152,1704]" captionTargetPageId="52" captionText="Figures 241259: Pedipalp chela dorsal (241), movable finger (242250), and fixed finger (251259) of Sri Lankan Isometrus, Lychas, and Reddyanus species. Figures 241242, 252. Isometrus maculatus, female from locality 15CO (241) and male from locality 15CP (242, 252). Figures 243244, 253. I. thwaitesi, male from locality 15CH (243) and female from locality 15CO (244, 253). Figures 245, 254. Reddyanus basilicus, male from locality 15CS. Figures 246, 255. R. besucheti, male from locality 15CD. Figures 247, 256. R. ceylonensis sp. n., male holotype. Figures 248, 257. R. jayarathnei sp. n., male paratype. Figures 249, 258. R. loebli, male from locality 15CG. Figures 250, 259. R. ranawanai sp. n., male holotype. Figure 251. Lychas srilankensis, male from locality 15CO." figureDoi="http://doi.org/10.5281/zenodo.7124626" httpUri="https://zenodo.org/record/7124626/files/figure.png" pageId="57" pageNumber="56">245, 254</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FD6FFA179D6FFA6A" box="[687,781,1472,1495]" captionStart-0="Figures 260263" captionStart-1="Figures 264269" captionStart-2="Figures 270275" captionStart-3="Figures 276282" captionStart-4="Figures 283286" captionStart-5="Figures 287290" captionStartId-0="53.[192,278,1632,1656]" captionStartId-1="55.[192,278,1408,1432]" captionStartId-2="56.[192,278,1839,1863]" captionStartId-3="57.[192,278,892,916]" captionStartId-4="58.[192,278,1625,1649]" captionStartId-5="59.[192,278,1867,1891]" captionTargetBox-0="[196,1427,148,1596]" captionTargetBox-1="[191,1440,167,1380]" captionTargetBox-2="[215,1432,184,1819]" captionTargetBox-3="[198,1431,222,869]" captionTargetBox-4="[201,1430,178,1594]" captionTargetBox-5="[191,1440,167,1838]" captionTargetId-0="figure-62@53.[192,1439,144,1603]" captionTargetId-1="figure-377@55.[192,1439,168,1379]" captionTargetId-2="figure-11@56.[192,1439,173,1824]" captionTargetId-3="figure-6@57.[192,1439,199,888]" captionTargetId-4="figure-11@58.[192,1439,168,1596]" captionTargetId-5="figure-47@59.[192,1439,168,1837]" captionTargetPageId-0="53" captionTargetPageId-1="55" captionTargetPageId-2="56" captionTargetPageId-3="57" captionTargetPageId-4="58" captionTargetPageId-5="59" captionText-0="Figures 260263: Reddyanus basilicus from locality 15CS. Figures 260261. Male in dorsal (260) and ventral (261) views. Figures 262263. Female in dorsal (262) and ventral (263) views. Scale bar: 10 mm." captionText-1="Figures 264269: Reddyanus basilicus. Figures 264, 266. Male from locality 15CS, chelicerae, carapace and tergites IIII (264), and sternopectinal region and sternite III (266). Figures 265, 267. Female from locality 15CS, chelicerae, carapace and tergites IIII (265) and sternopectinal region and sternites IIIIV (267). Figures 268269. Male from locality 15CR, chelicera dorsal (268) and ventral (269) aspects." captionText-2="Figures 270275: Reddyanus basilicus from locality 15CS. Figures 270272. Male, metasoma and telson, lateral (270), ventral (271), and dorsal (272) views. Figures 273275. Female, metasoma and telson, lateral (273), ventral (274), and dorsal (275) views. Scale bar: 10 mm." captionText-3="Figures 276282: Reddyanus basilicus from locality 15CS. Figures 276277. Female. Pedipalp chela, dorsal (276) and external (277) views. Figures 278282. Male. Pedipalp chela, dorsal (278) and external (279) views. Pedipalp patella, dorsal (280) and external (281) views. Pedipalp femur and trochanter dorsal view (282). The trichobothrial pattern is indicated in Figures 279282." captionText-4="Figures 283286: Left hemispermatophore of Reddyanus basilicus from locality 15CR. External (283), dorsal (284), internal (285) and ventral (286) views. Scale bar: 500 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; t, trunk." captionText-5="Figures 287290: Reddyanus basilicus from locality 15CS, female (287), female with newborns before first ecdysis (288), female with juveniles after first ecdysis (289), and male (290)." figureDoi-0="http://doi.org/10.5281/zenodo.7124628" figureDoi-1="http://doi.org/10.5281/zenodo.7124630" figureDoi-2="http://doi.org/10.5281/zenodo.7124632" figureDoi-3="http://doi.org/10.5281/zenodo.7124634" figureDoi-4="http://doi.org/10.5281/zenodo.7124636" figureDoi-5="http://doi.org/10.5281/zenodo.7124638" httpUri-0="https://zenodo.org/record/7124628/files/figure.png" httpUri-1="https://zenodo.org/record/7124630/files/figure.png" httpUri-2="https://zenodo.org/record/7124632/files/figure.png" httpUri-3="https://zenodo.org/record/7124634/files/figure.png" httpUri-4="https://zenodo.org/record/7124636/files/figure.png" httpUri-5="https://zenodo.org/record/7124638/files/figure.png" pageId="57" pageNumber="56">260290</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FEECFA089FEBFA4B" box="[300,393,1503,1526]" captionStart="Figures 403429" captionStartId="85.[192,278,744,768]" captionTargetId="figure-11@84.[172,1464,185,1878]" captionTargetPageId="84" captionText="Figures 403429: Telson, lateral view of Sri Lankan Buthidae and Chaerilidae species. Figures 403404. Isometrus maculatus, male from locality 15CP (403) and female from locality 15CI (404). Figures 405406. I. thwaitesi, male from locality 15CH (405) and female from locality 15CO (406). Figures 407408. Lychas srilankensis, male (407) and female (408) from locality 15CJ. Figures 409410. Reddyanus basilicus, male (409) and female (410) from locality 15CR. Figures 411412. R. besucheti, male holotype (411) and female from locality 15CG (412). Figures 413414. R. ceylonensis sp. n., male holotype (413) and female paratype (414) from locality 15CI. Figures 415416. R. jayarathnei sp. n., male (415) and female (416) paratypes. Figures 417418. R. loebli, male (417) and female (418) from locality 15CG. Figures 419420. R. ranawanai sp. n., male holotype (419) and female paratype (420). Figures 421422: Buthoscorpio sarasinorum, male from locality 15CF (421) and female from locality 15CG (422). Figures 423424. Charmus laneus, male (423) and female (424) from locality 15CO. Figures 425426. Charmus saradieli sp. n. male paratype (425) (MHNG) and female holotype (426). Figurs 427. Hottentotta tamulus, male from locality 15CK (427). Figures 428429. Chaerilus ceylonensis, male (428) and female (429) from locality 15CD." figureDoi="http://doi.org/10.5281/zenodo.7124696" httpUri="https://zenodo.org/record/7124696/files/figure.png" pageId="57" pageNumber="56">409410</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FE57FA089F96FA4B" box="[407,500,1503,1526]" captionStart="Figures 547554" captionStartId="114.[192,278,810,834]" captionTargetBox="[191,1440,191,782]" captionTargetId="figure-538@114.[192,1440,192,781]" captionTargetPageId="114" captionText="Figures 547554: The mitotic metaphases of buthid males from Sri Lanka. Buthoscorpio sarasinorum from locality 15CF (2n=14) (547), Charmus laneus from locality 15CO (2n=9) (548), Isometrus thwaitesi from locality 15CF (2n=8) (549), Lychas srilankensis from locality 15CN (2n=16) (550), Reddyanus basilicus from locality 15CS (2n=16) (551), Reddyanus basilicus from locality 15CR (2n=15) (552), Reddyanus ceylonensis from locality 15CI (2n=16) (553), Reddyanus loebli from locality 15CH (2n=17) (554). Arrowheads indicate extra large odd chromosomes. Scale bar = 5 µm for 547554." figureDoi="http://doi.org/10.5281/zenodo.7124765" httpUri="https://zenodo.org/record/7124765/files/figure.png" pageId="57" pageNumber="56">551552</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FDC2FA089C48FA4B" box="[514,554,1503,1526]" captionStart="Figures 555570" captionStartId="115.[192,278,1733,1757]" captionTargetBox="[233,1393,170,1705]" captionTargetId="figure-151@115.[234,1392,171,1704]" captionTargetPageId="115" captionText="Figures 555570: Key characters for distinguishing males of Sri Lankan Isometrus and Reddyanus. Figures 555556, 565 566. Pedipalp chela dorsal view, I. maculatus from locality 15CP (555), I. thwaitesi from locality 15CH (556), R. basilicus, holotype (565), and R. besucheti, holotype (566). Figures 557558. Sternopectinal area, I. maculatus from locality 15CP (557), and I. thwaitesi from locality 15CH (558). Figures 559560, 567570. Sternite V, I. maculatus from locality 15CP (559), I. thwaitesi from locality 15CH (560), R. besucheti, from locality 15CF (567), R. ceylonensis sp. n., holotype (568), R. jayarathnei sp. n., paratype (569), and R. ranawanai sp. n., paratype (570). Figures 561564. Metasomal segment II lateral (561562) and pedipalp patella dorsal (563564), R. loebli from locality 15CG (561, 563) and R. besucheti, from locality 15CF (562, 564)." figureDoi="http://doi.org/10.5281/zenodo.7124767" httpUri="https://zenodo.org/record/7124767/files/figure.png" pageId="57" pageNumber="56">565</figureCitation>
,
<tableCitation id="755A0372A82EFFA9FDF7FA089CF1FA4B" box="[567,659,1503,1526]" captionStart="Table 3" captionStartId="86.[192,257,777,801]" captionTargetPageId="86" captionText="Table 3: Comparative measurements of adults of Reddyanus basilicus, R. besucheti and R. ceylonensis sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/6CA76641A841FFC6FF00FCDE9DD7FC84" pageId="57" pageNumber="56" tableUuid="6CA76641A841FFC6FF00FCDE9DD7FC84">Tables 3</tableCitation>
,
<tableCitation id="755A0372A82EFFA9FD5FFA089CCEFA4B" box="[671,684,1503,1526]" captionStart="Table 5" captionStartId="86.[192,257,1869,1893]" captionTargetPageId="86" captionText="Table 5: Comparison among Sri Lankan Reddyanus species (specimens), based upon selected morphometric ratios of adult males. Abbreviations: length (L), width (W), depth (D)." httpUri="http://table.plazi.org/id/6CA76641A841FFC6FF00F89A9CBAF8C0" pageId="57" pageNumber="56" tableUuid="6CA76641A841FFC6FF00F89A9CBAF8C0">5</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="70C26542A82EFFA9FF00F9CB9C21F970" pageId="57" pageNumber="56" type="reference_group">
<paragraph id="386736C9A82EFFA9FF00F9CB9C03F98E" blockId="57.[192,802,1564,1741]" box="[192,609,1564,1587]" pageId="57" pageNumber="56">
<treatmentCitationGroup id="18C811E7A82EFFA9FF00F9CB9C03F98E" box="[192,609,1564,1587]" pageId="57" pageNumber="56">
<treatmentCitation id="B97910D8A82EFFA9FF00F9CB9C38F98E" author="KARSCH" box="[192,602,1564,1587]" page="113" pageId="57" pageNumber="56" year="1879">
<taxonomicName id="FFD84D4AA82EFFA9FF00F9CB9C38F98E" authority="Karsch, 1879: 113" authorityName="Karsch" authorityPageNumber="113" authorityYear="1879" box="[192,602,1564,1587]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="57" pageNumber="56" phylum="Arthropoda" rank="species" species="basilicus">
<emphasis id="0AACEADBA82EFFA9FF00F9CB9FEDF98E" box="[192,399,1564,1587]" italics="true" pageId="57" pageNumber="56">Isometrus basilicus</emphasis>
<bibRefCitation id="5C494B38A82EFFA9FE55F9CB9C38F98E" author="Karsch" box="[405,602,1564,1587]" firstAuthor="Karsch" page="113" pageId="57" pageNumber="56" pagination="97 - 136" refId="ref41417" refString="KARSCH, F. 1879. Skorpionologische Beitrage I. and II. Mitteilungen des Munchener Entomologischen Vereins, 3: 6 - 22, 97 - 136." type="journal article" year="1879">Karsch, 1879: 113</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="386736C9A82EFFA9FF00F9EC9C21F970" blockId="57.[192,802,1564,1741]" pageId="57" pageNumber="56">
<treatmentCitationGroup id="18C811E7A82EFFA9FF00F9EC9C21F970" pageId="57" pageNumber="56">
<treatmentCitation id="B97910D8A82EFFA9FF00F9EC9D79F9EF" author="VACHON, M." box="[192,795,1595,1618]" page="92" pageId="57" pageNumber="56" year="1982">
<taxonomicName id="FFD84D4AA82EFFA9FF00F9EC9D79F9EF" authority=": Vachon, 1982: 91 - 92" authorityName="Vachon" authorityPageNumber="91 - 92" authorityYear="1982" box="[192,795,1595,1618]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="57" pageNumber="56" phylum="Arthropoda" rank="species" species="basilicus" subGenus="Reddyanus">
<emphasis id="0AACEADBA82EFFA9FF00F9EC9F4BF9EF" box="[192,297,1595,1618]" italics="true" pageId="57" pageNumber="56">Isometrus</emphasis>
(
<emphasis id="0AACEADBA82EFFA9FEFBF9EC9FD2F9EF" box="[315,432,1595,1618]" italics="true" pageId="57" pageNumber="56">Reddyanus</emphasis>
)
<emphasis id="0AACEADBA82EFFA9FE01F9EC9C42F9EF" box="[449,544,1595,1618]" italics="true" pageId="57" pageNumber="56">basilicus</emphasis>
:
<bibRefCitation id="5C494B38A82EFFA9FDF0F9EC9D79F9EF" author="Vachon" box="[560,795,1595,1618]" firstAuthor="Vachon" journalOrPublisher="Revue suisse de Zoologie" page="91 - 92" pageId="57" pageNumber="56" pagination="77 - 114" part="89" refId="ref45190" refString="VACHON, M. 1982. Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.). Revue suisse de Zoologie, 89: 77 - 114." title="Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.)" type="journal article" year="1982">Vachon, 1982: 9192</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 4046, 49?, 66, 8889;
<treatmentCitation id="B97910D8A82EFFA9FD83F98D9F79F932" author="FET, V. &amp; W. D. SISSOM &amp; G. LOWE &amp; M. E. BRAUNWALDER" page="151" pageId="57" pageNumber="56" year="2000">
<bibRefCitation id="5C494B38A82EFFA9FD83F98D9F79F932" author="Fet &amp; Lowe" bookContentInfo="689 pp." firstAuthor="Fet" journalOrPublisher="New York: The New York Entomological Society" page="151" pageId="57" pageNumber="56" refId="ref40873" refString="FET, V., W. D. SISSOM, G. LOWE &amp; M. E. BRAUNWALDER. 2000. Catalog of the Scorpions of the World (1758 - 1998). New York: The New York Entomological Society, 689 pp." title="Catalog of the Scorpions of the World (1758 - 1998)" type="book" year="2000">Fet &amp; Lowe, 2000: 151</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="B97910D8A82EFFA9FEEBF9AF9F8BF932" ID-DOI="http://doi.org/10.5281/zenodo.4672862" ID-Zenodo-Dep="4672862" author="Kovarik" box="[299,489,1656,1679]" httpUri="http://treatment.plazi.org/id/4703879F45192706FCA5ADBD262F817B" page="5" pageId="57" pageNumber="56" year="2003">
<bibRefCitation id="5C494B38A82EFFA9FEEBF9AF9F8BF932" author="Kovarik" box="[299,489,1656,1679]" firstAuthor="Kovarik" page="5" pageId="57" pageNumber="56" pagination="1 - 19" refId="ref41718" refString="KOVARIK, F. 2003. A review of the genus Isometrus Ehrenberg, 1828 (Scorpiones: Buthidae) with descriptions of four new species from Asia and Australia. Euscorpius, 10: 1 - 19." type="journal article" year="2003">Kovařík, 2003: 5</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="B97910D8A82EFFA9FDA2F9AF9FC8F913" author="KOVARIK, F &amp; A. A. OJANGUREN AFFILASTRO" page="185" pageId="57" pageNumber="56" year="2013">
<bibRefCitation id="5C494B38A82EFFA9FDA2F9AF9FC8F913" author="Kovarik &amp; Ojanguren" bookContentInfo="400 pp." firstAuthor="Kovarik" journalOrPublisher="Clairon Production, Prague" page="185" pageId="57" pageNumber="56" refId="ref42080" refString="KOVARIK, F &amp; A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Clairon Production, Prague, 400 pp." title="Illustrated catalog of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus" type="book" year="2013">Kovařík &amp; Ojanguren, 2013: 185</bibRefCitation>
</treatmentCitation>
, 350, 358, figs. 12961303, 1386 1392 (reference list until 2013).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="70C26542A82EFFA9FF00F9249DB3F98E" pageId="57" pageNumber="56" type="materials_examined">
<paragraph id="386736C9A82EFFA9FF00F9249FA0F894" blockId="57.[192,802,1779,1833]" pageId="57" pageNumber="56">
<typeStatus id="E763886BA82EFFA9FF00F9249E9AF8B7" box="[192,248,1779,1802]" pageId="57" pageNumber="56">TYPE</typeStatus>
LOCALITY AND
<typeStatus id="E763886BA82EFFA9FE65F9209FBBF8B7" box="[421,473,1783,1802]" pageId="57" pageNumber="56">TYPE</typeStatus>
REPOSITORY.
<collectingCountry id="40CF7659A82EFFA9FDB1F9249CA3F8B7" box="[625,705,1779,1802]" name="Sri Lanka" pageId="57" pageNumber="56">Ceylon</collectingCountry>
, now
<collectingCountry id="40CF7659A82EFFA9FCC3F9249F64F894" name="Sri Lanka" pageId="57" pageNumber="56">Sri Lanka</collectingCountry>
;
<collectionCode id="5EC9AE0CA82EFFA9FED1F8C59F3CF894" box="[273,350,1810,1833]" pageId="57" pageNumber="56">ZMHB</collectionCode>
No. 113.
</paragraph>
<paragraph id="386736C9A82EFFA9FCFEFBC49DFEFBDA" blockId="57.[830,1440,1043,1127]" pageId="57" pageNumber="56">
TYPE MATERIAL EXAMINED.
<materialsCitation id="88B03C94A82EFFA9FB43FBC49DF5FBDA" ID-GBIF-Occurrence="3913992353" collectionCode="ZMHB" collectorName="Nietner &amp; No." country="Sri Lanka" location="Ceylon" pageId="57" pageNumber="56" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="40CF7659A82EFFA9FB43FBC49AB2FB97" box="[1155,1232,1043,1066]" name="Sri Lanka" pageId="57" pageNumber="56">Ceylon</collectingCountry>
(now
<collectingCountry id="40CF7659A82EFFA9FAE1FBC49BF0FB97" box="[1313,1426,1043,1066]" name="Sri Lanka" pageId="57" pageNumber="56">Sri Lanka</collectingCountry>
),
<specimenCount id="2EDEFD40A82EFFA9FCFEFBE59D02FBF4" box="[830,864,1074,1097]" pageId="57" pageNumber="56" type="male">1♂</specimenCount>
(
<typeStatus id="E763886BA82EFFA9FCB3FBE59DB1FBF4" box="[883,979,1074,1097]" pageId="57" pageNumber="56" type="holotype">holotype</typeStatus>
,
<figureCitation id="A0E32A4CA82EFFA9FC22FBE59A20FBF4" box="[994,1090,1074,1097]" captionStart="Figures 555570" captionStartId="115.[192,278,1733,1757]" captionTargetBox="[233,1393,170,1705]" captionTargetId="figure-151@115.[234,1392,171,1704]" captionTargetPageId="115" captionText="Figures 555570: Key characters for distinguishing males of Sri Lankan Isometrus and Reddyanus. Figures 555556, 565 566. Pedipalp chela dorsal view, I. maculatus from locality 15CP (555), I. thwaitesi from locality 15CH (556), R. basilicus, holotype (565), and R. besucheti, holotype (566). Figures 557558. Sternopectinal area, I. maculatus from locality 15CP (557), and I. thwaitesi from locality 15CH (558). Figures 559560, 567570. Sternite V, I. maculatus from locality 15CP (559), I. thwaitesi from locality 15CH (560), R. besucheti, from locality 15CF (567), R. ceylonensis sp. n., holotype (568), R. jayarathnei sp. n., paratype (569), and R. ranawanai sp. n., paratype (570). Figures 561564. Metasomal segment II lateral (561562) and pedipalp patella dorsal (563564), R. loebli from locality 15CG (561, 563) and R. besucheti, from locality 15CF (562, 564)." figureDoi="http://doi.org/10.5281/zenodo.7124767" httpUri="https://zenodo.org/record/7124767/files/figure.png" pageId="57" pageNumber="56">Fig. 565</figureCitation>
,
<tableCitation id="755A0372A82EFFA9FB91FBE59AC4FBF4" box="[1105,1190,1074,1097]" captionStart="Table 3" captionStartId="86.[192,257,777,801]" captionTargetPageId="86" captionText="Table 3: Comparative measurements of adults of Reddyanus basilicus, R. besucheti and R. ceylonensis sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/6CA76641A841FFC6FF00FCDE9DD7FC84" pageId="57" pageNumber="56" tableUuid="6CA76641A841FFC6FF00FCDE9DD7FC84">Table 3</tableCitation>
), leg.
<collectorName id="952D531FA82EFFA9FB31FBE59B23FBF4" box="[1265,1345,1074,1097]" pageId="57" pageNumber="56">Nietner</collectorName>
,
<collectionCode id="5EC9AE0CA82EFFA9FA93FBE59BC2FBF4" box="[1363,1440,1074,1097]" pageId="57" pageNumber="56">ZMHB</collectionCode>
<collectorName id="952D531FA82EFFA9FCFEFB879D04FBDA" box="[830,870,1104,1127]" pageId="57" pageNumber="56">No.</collectorName>
113
</materialsCitation>
.
</paragraph>
<paragraph id="386736C9A82EFFA9FCFEFB599DB3F98E" blockId="57.[830,1441,1166,1587]" pageId="57" pageNumber="56">
OTHER MATERIAL EXAMINED.
<materialsCitation id="88B03C94A82EFFA9FB49FB599DE3FB5F" ID-GBIF-Occurrence="3913992323" collectionCode="FKCP" country="Sri Lanka" county="Hills" location="Namunukula" municipality="Badula District" pageId="57" pageNumber="56" specimenCount="1" specimenCount-male="1" stateProvince="Uva">
<collectingCountry id="40CF7659A82EFFA9FB49FB599A95FB18" box="[1161,1271,1166,1189]" name="Sri Lanka" pageId="57" pageNumber="56">Sri Lanka</collectingCountry>
,
<collectingRegion id="FA1CF82BA82EFFA9FAC5FB599BFEFB18" box="[1285,1436,1166,1189]" country="Sri Lanka" name="Uva" pageId="57" pageNumber="56">Uva Province</collectingRegion>
,
<collectingRegion id="FA1CF82BA82EFFA9FCFEFB7B9D09FB7E" box="[830,875,1196,1219]" country="Sri Lanka" name="Uva" pageId="57" pageNumber="56">Uva</collectingRegion>
<collectingCounty id="D1064E45A82EFFA9FCB8FB7B9DCCFB7E" box="[888,942,1196,1219]" pageId="57" pageNumber="56">Hills</collectingCounty>
,
<collectingMunicipality id="D803ACB3A82EFFA9FC00FB7B9A0EFB7E" box="[960,1132,1196,1219]" pageId="57" pageNumber="56">Badula District</collectingMunicipality>
,
<location id="3D076012A82EFFA9FBBEFB7B9B69FB7E" LSID="urn:lsid:plazi:treatment:B07187DFA82EFFACFF39FA769A36F896:3D076012A82EFFA9FBBEFB7B9B69FB7E" box="[1150,1291,1196,1219]" country="Sri Lanka" county="Hills" municipality="Badula District" name="Namunukula" pageId="57" pageNumber="56" stateProvince="Uva">Namunukula</location>
, 1970,
<specimenCount id="2EDEFD40A82EFFA9FAA5FB7B9BE4FB7E" box="[1381,1414,1196,1219]" pageId="57" pageNumber="56" type="male">1♂</specimenCount>
A,
<collectionCode id="5EC9AE0CA82EFFA9FCFEFB1C9DE3FB5F" box="[830,897,1227,1250]" pageId="57" pageNumber="56">FKCP</collectionCode>
</materialsCitation>
;
<materialsCitation id="88B03C94A82EFFA9FC50FB1C9DD6FABC" ID-GBIF-Occurrence="3913992333" collectionCode="FKCP" country="Sri Lanka" location="Kandy" pageId="57" pageNumber="56" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Uva">
<location id="3D076012A82EFFA9FC50FB1C9DB5FB5F" LSID="urn:lsid:plazi:treatment:B07187DFA82EFFACFF39FA769A36F896:3D076012A82EFFA9FC50FB1C9DB5FB5F" box="[912,983,1227,1250]" country="Sri Lanka" name="Kandy" pageId="57" pageNumber="56" stateProvince="Uva">Kandy</location>
,
<specimenCount id="2EDEFD40A82EFFA9FC25FB1C9A65FB5F" box="[997,1031,1227,1250]" pageId="57" pageNumber="56" type="male">1♂</specimenCount>
after 4th ecdysis
<specimenCount id="2EDEFD40A82EFFA9FB07FB1C9A8AFB5F" box="[1223,1256,1227,1250]" pageId="57" pageNumber="56" type="male">2♂</specimenCount>
after 5th ecdysis
<specimenCount id="2EDEFD40A82EFFA9FCFFFB3D9D02FABC" box="[831,864,1258,1281]" pageId="57" pageNumber="56" type="female">2♀</specimenCount>
,
<collectionCode id="5EC9AE0CA82EFFA9FCAEFB3D9DD6FABC" box="[878,948,1258,1281]" pageId="57" pageNumber="56">FKCP</collectionCode>
</materialsCitation>
;
<materialsCitation id="88B03C94A82EFFA9FC7FFB3D9A67FAC6" ID-GBIF-Occurrence="3913992311" collectingDate="2015-05-03" collectingDateMax="2015-05-04" collectingDateMin="2015-05-03" collectionCode="FKCP" country="Sri Lanka" county="Ampara District" elevation="40" latitude="6.8794446" location="Locality" longLatPrecision="15" longitude="81.722725" municipality="Lahugala Kitulana National Park" pageId="57" pageNumber="56" specimenCount="3" specimenCount-female="1" specimenCount-male="1" stateProvince="Eastern">
<collectingRegion id="FA1CF82BA82EFFA9FC7FFB3D9A1AFABC" box="[959,1144,1258,1281]" country="Sri Lanka" name="Eastern" pageId="57" pageNumber="56">Eastern Province</collectingRegion>
,
<collectingCounty id="D1064E45A82EFFA9FB42FB3D9B4DFABC" box="[1154,1327,1258,1281]" pageId="57" pageNumber="56">Ampara District</collectingCounty>
,
<collectingMunicipality id="D803ACB3A82EFFA9FAFDFB3D9A5FFAA2" pageId="57" pageNumber="56">Lahugala Kitulana National Park</collectingMunicipality>
,
<geoCoordinate id="5DEC500EA82EFFA9FB8FFADF9AAEFAA2" box="[1103,1228,1288,1311]" degrees="06" direction="north" minutes="52" orientation="latitude" pageId="57" pageNumber="56" precision="15" seconds="46" value="6.8794446">06°52'46&quot;N</geoCoordinate>
<geoCoordinate id="5DEC500EA82EFFA9FB17FADF9B14FAA2" box="[1239,1398,1288,1311]" degrees="081" direction="east" minutes="43" orientation="longitude" pageId="57" pageNumber="56" precision="1" seconds="21. 8" value="81.722725">081°43'21.8&quot;E</geoCoordinate>
,
<elevation id="B3F5D1FAA82EFFA9FA45FADF9DF0FA83" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="57" pageNumber="56" unit="m" value="40.0">
<quantity id="FF209B2CA82EFFA9FA45FADF9D31FA83" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="57" pageNumber="56" unit="m" value="40.0">40 m</quantity>
a.s.l.
</elevation>
(
<location id="3D076012A82EFFA9FC68FAF09A61FA83" LSID="urn:lsid:plazi:treatment:B07187DFA82EFFACFF39FA769A36F896:3D076012A82EFFA9FC68FAF09A61FA83" box="[936,1027,1319,1342]" country="Sri Lanka" county="Ampara District" latitude="6.8794446" longLatPrecision="15" longitude="81.722725" municipality="Lahugala Kitulana National Park" name="Locality" pageId="57" pageNumber="56" stateProvince="Eastern">Locality</location>
<emphasis id="0AACEADBA82EFFA9FBD1FAF19A31FA83" bold="true" box="[1041,1107,1318,1342]" pageId="57" pageNumber="56">15CR</emphasis>
,
<figureCitation id="A0E32A4CA82EFFA9FBA7FAF09AA8FA83" box="[1127,1226,1319,1342]" captionStart="Figures 595596" captionStartId="127.[192,278,1661,1685]" captionTargetBox="[191,1440,195,1633]" captionTargetId="figure-82@127.[192,1439,196,1632]" captionTargetPageId="127" captionText="Figures 595596: Figure 595. Locality 15CQ, Sri Lanka, Uva Province, Monaragala District, Monaragala, 06°52'30.7&quot;N 081°21'17&quot;E, 288 m a.s.l. During night collecting on 2.3.V.2015 (UV detection) we recorded Reddyanus loebli (under scales of bark and under bark) and Heterometrus serratus (females with juveniles inside oblique burrows). On the locality we recorded a temperature of 33.1 ºC at 5.00 p.m., which gradually dropped to 24.6 ºC (minimum temperature) before sunrise. Humidity during the night varied between 61% and 84%. Figure 596. Locality 15CR, Sri Lanka, Eastern Province, Ampara District, Lahugala Kitulana National Park, 06°52'46&quot;N 081°43'21.8&quot;E, 40 m a.s.l. During night collecting on 3.4.V.2015 (10.00 p.m. to 01.00 a.m.) we recorded Isometrus maculatus (running on branches, also sitting on leaves on trees), Reddyanus basilicus (on the ground among leaves) and Reddyanus loebli (hidden under scales of bark of standing tree trunks). On the locality we recorded between 10.00 p.m. on 3.V. and 01.00 a.m. on 4.V.2015 temperature 28 ºC, which gradually dropped to 26.9 ºC and humidity 73%." figureDoi="http://doi.org/10.5281/zenodo.7124794" httpUri="https://zenodo.org/record/7124794/files/figure.png" pageId="57" pageNumber="56">Fig. 596</figureCitation>
),
<date id="4C661009A82EFFA9FB25FAF09B08FA83" box="[1253,1386,1319,1342]" pageId="57" pageNumber="56" value="2015-05-03" valueMax="2015-05-04" valueMin="2015-05-03">
<collectingDate id="5C22E9E1A82EFFA9FB25FAF09B08FA83" box="[1253,1386,1319,1342]" pageId="57" pageNumber="56" value="2015-05-03" valueMax="2015-05-04" valueMin="2015-05-03">3.4.V.2015</collectingDate>
</date>
,
<specimenCount id="2EDEFD40A82EFFA9FABFFAF09BC2FA83" box="[1407,1440,1319,1342]" pageId="57" pageNumber="56" type="male">1♂</specimenCount>
(
<figureCitation id="A0E32A4CA82EFFA9FC87FA919DCDFAE0" box="[839,943,1350,1373]" captionStart="Figures 209224" captionStartId="50.[192,278,1736,1760]" captionTargetBox="[208,1434,174,1703]" captionTargetId="figure-11@50.[205,1452,168,1707]" captionTargetPageId="50" captionText="Figures 209224: Metasomal segments IIIII, lateral view of Sri Lankan Reddyanus and Isometrus species. Figures 209210. Reddyanus basilicus, male (209) and female (210) from locality 15CR. Figures 211212. R. besucheti, male holotype (211) and female from locality 15CG (212). Figures 213214. R. ceylonensis sp. n., male holotype (213) and female paratype from locality 15CI (214). Figures 215216. R. jayarathnei sp. n., male (215) and female (216) paratypes. Figures 217218. R. loebli, male (217) and female (218) from locality 15CG. Figures 219220. R. ranawanai sp. n., male holotype (219) and female paratype (220). Figures 221222. Isometrus maculatus, male from locality 15CP (221) and female from locality 15CI (222). Figures 223 224. I. thwaitesi, male from locality 15CH (223) and female from locality 15CO (224)." figureDoi="http://doi.org/10.5281/zenodo.7124622" httpUri="https://zenodo.org/record/7124622/files/figure.png" pageId="57" pageNumber="56">Figs. 209</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FC7AFA919D80FAE0" box="[954,994,1350,1373]" captionStart="Figures 225240" captionStartId="51.[192,278,1493,1517]" captionTargetBox="[193,1432,180,1451]" captionTargetId="figure-6@51.[192,1439,179,1464]" captionTargetPageId="51" captionText="Figures 225240: Pedipalp, dorsal view of Sri Lankan Reddyanus and Isometrus species. Figures 225226. Reddyanus basilicus, male (225) and female (226) from locality 15CR. Figures 227228. R. besucheti, male from locality 15CD (227) and female from locality 15CG (228). Figures 229230. R. ceylonensis sp. n., male holotype (229) and female paratype from locality 15CI (230). Figures 231232. R. jayarathnei sp. n., male (231) and female (232) paratypes. Figures 233234. R. loebli, male (233) and female (234) from locality 15CG. Figures 235236. R. ranawanai sp. n., male holotype (235) and female paratype (236). Figures 237238. Isometrus maculatus, male from locality 15CP (237) and female from locality 15CI (238). Figures 239240. I. thwaitesi, male from locality 15CH (239) and female from locality 15CO (240)." figureDoi="http://doi.org/10.5281/zenodo.7124624" httpUri="https://zenodo.org/record/7124624/files/figure.png" pageId="57" pageNumber="56">225</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FC31FA919A32FAE0" box="[1009,1104,1350,1373]" captionStart="Figures 264269" captionStartId="55.[192,278,1408,1432]" captionTargetBox="[191,1440,167,1380]" captionTargetId="figure-377@55.[192,1439,168,1379]" captionTargetPageId="55" captionText="Figures 264269: Reddyanus basilicus. Figures 264, 266. Male from locality 15CS, chelicerae, carapace and tergites IIII (264), and sternopectinal region and sternite III (266). Figures 265, 267. Female from locality 15CS, chelicerae, carapace and tergites IIII (265) and sternopectinal region and sternites IIIIV (267). Figures 268269. Male from locality 15CR, chelicera dorsal (268) and ventral (269) aspects." figureDoi="http://doi.org/10.5281/zenodo.7124630" httpUri="https://zenodo.org/record/7124630/files/figure.png" pageId="57" pageNumber="56">268269</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FB9DFA919AE7FAE0" box="[1117,1157,1350,1373]" captionStart="Figures 403429" captionStartId="85.[192,278,744,768]" captionTargetId="figure-11@84.[172,1464,185,1878]" captionTargetPageId="84" captionText="Figures 403429: Telson, lateral view of Sri Lankan Buthidae and Chaerilidae species. Figures 403404. Isometrus maculatus, male from locality 15CP (403) and female from locality 15CI (404). Figures 405406. I. thwaitesi, male from locality 15CH (405) and female from locality 15CO (406). Figures 407408. Lychas srilankensis, male (407) and female (408) from locality 15CJ. Figures 409410. Reddyanus basilicus, male (409) and female (410) from locality 15CR. Figures 411412. R. besucheti, male holotype (411) and female from locality 15CG (412). Figures 413414. R. ceylonensis sp. n., male holotype (413) and female paratype (414) from locality 15CI. Figures 415416. R. jayarathnei sp. n., male (415) and female (416) paratypes. Figures 417418. R. loebli, male (417) and female (418) from locality 15CG. Figures 419420. R. ranawanai sp. n., male holotype (419) and female paratype (420). Figures 421422: Buthoscorpio sarasinorum, male from locality 15CF (421) and female from locality 15CG (422). Figures 423424. Charmus laneus, male (423) and female (424) from locality 15CO. Figures 425426. Charmus saradieli sp. n. male paratype (425) (MHNG) and female holotype (426). Figurs 427. Hottentotta tamulus, male from locality 15CK (427). Figures 428429. Chaerilus ceylonensis, male (428) and female (429) from locality 15CD." figureDoi="http://doi.org/10.5281/zenodo.7124696" httpUri="https://zenodo.org/record/7124696/files/figure.png" pageId="57" pageNumber="56">409</figureCitation>
, 552)
<specimenCount id="2EDEFD40A82EFFA9FB0CFA919A8FFAE0" box="[1228,1261,1350,1373]" pageId="57" pageNumber="56" type="female">1♀</specimenCount>
(
<figureCitation id="A0E32A4CA82EFFA9FB3EFA919B01FAE0" box="[1278,1379,1350,1373]" captionStart="Figures 209224" captionStartId="50.[192,278,1736,1760]" captionTargetBox="[208,1434,174,1703]" captionTargetId="figure-11@50.[205,1452,168,1707]" captionTargetPageId="50" captionText="Figures 209224: Metasomal segments IIIII, lateral view of Sri Lankan Reddyanus and Isometrus species. Figures 209210. Reddyanus basilicus, male (209) and female (210) from locality 15CR. Figures 211212. R. besucheti, male holotype (211) and female from locality 15CG (212). Figures 213214. R. ceylonensis sp. n., male holotype (213) and female paratype from locality 15CI (214). Figures 215216. R. jayarathnei sp. n., male (215) and female (216) paratypes. Figures 217218. R. loebli, male (217) and female (218) from locality 15CG. Figures 219220. R. ranawanai sp. n., male holotype (219) and female paratype (220). Figures 221222. Isometrus maculatus, male from locality 15CP (221) and female from locality 15CI (222). Figures 223 224. I. thwaitesi, male from locality 15CH (223) and female from locality 15CO (224)." figureDoi="http://doi.org/10.5281/zenodo.7124622" httpUri="https://zenodo.org/record/7124622/files/figure.png" pageId="57" pageNumber="56">Figs. 210</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FAB1FA919BFBFAE0" box="[1393,1433,1350,1373]" captionStart="Figures 225240" captionStartId="51.[192,278,1493,1517]" captionTargetBox="[193,1432,180,1451]" captionTargetId="figure-6@51.[192,1439,179,1464]" captionTargetPageId="51" captionText="Figures 225240: Pedipalp, dorsal view of Sri Lankan Reddyanus and Isometrus species. Figures 225226. Reddyanus basilicus, male (225) and female (226) from locality 15CR. Figures 227228. R. besucheti, male from locality 15CD (227) and female from locality 15CG (228). Figures 229230. R. ceylonensis sp. n., male holotype (229) and female paratype from locality 15CI (230). Figures 231232. R. jayarathnei sp. n., male (231) and female (232) paratypes. Figures 233234. R. loebli, male (233) and female (234) from locality 15CG. Figures 235236. R. ranawanai sp. n., male holotype (235) and female paratype (236). Figures 237238. Isometrus maculatus, male from locality 15CP (237) and female from locality 15CI (238). Figures 239240. I. thwaitesi, male from locality 15CH (239) and female from locality 15CO (240)." figureDoi="http://doi.org/10.5281/zenodo.7124624" httpUri="https://zenodo.org/record/7124624/files/figure.png" pageId="57" pageNumber="56">226</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FC80FAB39D0BFAC6" box="[832,873,1380,1403]" captionStart="Figures 403429" captionStartId="85.[192,278,744,768]" captionTargetId="figure-11@84.[172,1464,185,1878]" captionTargetPageId="84" captionText="Figures 403429: Telson, lateral view of Sri Lankan Buthidae and Chaerilidae species. Figures 403404. Isometrus maculatus, male from locality 15CP (403) and female from locality 15CI (404). Figures 405406. I. thwaitesi, male from locality 15CH (405) and female from locality 15CO (406). Figures 407408. Lychas srilankensis, male (407) and female (408) from locality 15CJ. Figures 409410. Reddyanus basilicus, male (409) and female (410) from locality 15CR. Figures 411412. R. besucheti, male holotype (411) and female from locality 15CG (412). Figures 413414. R. ceylonensis sp. n., male holotype (413) and female paratype (414) from locality 15CI. Figures 415416. R. jayarathnei sp. n., male (415) and female (416) paratypes. Figures 417418. R. loebli, male (417) and female (418) from locality 15CG. Figures 419420. R. ranawanai sp. n., male holotype (419) and female paratype (420). Figures 421422: Buthoscorpio sarasinorum, male from locality 15CF (421) and female from locality 15CG (422). Figures 423424. Charmus laneus, male (423) and female (424) from locality 15CO. Figures 425426. Charmus saradieli sp. n. male paratype (425) (MHNG) and female holotype (426). Figurs 427. Hottentotta tamulus, male from locality 15CK (427). Figures 428429. Chaerilus ceylonensis, male (428) and female (429) from locality 15CD." figureDoi="http://doi.org/10.5281/zenodo.7124696" httpUri="https://zenodo.org/record/7124696/files/figure.png" pageId="57" pageNumber="56">410</figureCitation>
)
<specimenCount id="2EDEFD40A82EFFA9FCB9FAB39DCDFAC6" box="[889,943,1380,1403]" pageId="57" pageNumber="56" type="generic">1juv.</specimenCount>
,
<collectionCode id="5EC9AE0CA82EFFA9FC00FAB39A67FAC6" box="[960,1029,1380,1403]" pageId="57" pageNumber="56">FKCP</collectionCode>
</materialsCitation>
,
<materialsCitation id="88B03C94A82EFFA9FBD3FAB39B05FAC6" ID-GBIF-Occurrence="3913992331" box="[1043,1383,1380,1403]" collectionCode="UPSL" collectorName="Kovarik" country="Sri Lanka" pageId="57" pageNumber="56" specimenCount="1" specimenCount-male="1" stateProvince="Eastern">
<specimenCount id="2EDEFD40A82EFFA9FBD3FAB39A56FAC6" box="[1043,1076,1380,1403]" pageId="57" pageNumber="56" type="male">1♂</specimenCount>
,
<collectionCode id="5EC9AE0CA82EFFA9FB85FAB39AE4FAC6" box="[1093,1158,1380,1403]" pageId="57" pageNumber="56">UPSL</collectionCode>
, leg.
<collectorName id="952D531FA82EFFA9FB08FAB39B7DFAC6" box="[1224,1311,1380,1403]" pageId="57" pageNumber="56">Kovařík</collectorName>
et al.;
</materialsCitation>
<materialsCitation id="88B03C94A82EFFA9FAB1FAB39DB3F98E" ID-GBIF-Occurrence="3913992361" collectingDate="2015-05-04" collectionCode="FKCP" collectorName="Kovarik" country="Sri Lanka" county="Ampara District" elevation="56" latitude="7.3336945" location="Locality" longLatPrecision="1" longitude="81.699196" municipality="Ampara" pageId="57" pageNumber="56" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Eastern">
<collectingRegion id="FA1CF82BA82EFFA9FAB1FAB39DBBFA27" country="Sri Lanka" name="Eastern" pageId="57" pageNumber="56">Eastern Province</collectingRegion>
,
<collectingCounty id="D1064E45A82EFFA9FC2BFA549AC0FA27" box="[1003,1186,1411,1434]" pageId="57" pageNumber="56">Ampara District</collectingCounty>
,
<collectingMunicipality id="D803ACB3A82EFFA9FB74FA549B68FA27" box="[1204,1290,1411,1434]" pageId="57" pageNumber="56">Ampara</collectingMunicipality>
env.,
<geoCoordinate id="5DEC500EA82EFFA9FA9BFA549DE9FA04" degrees="07" direction="north" minutes="20" orientation="latitude" pageId="57" pageNumber="56" precision="1" seconds="01.3" value="7.3336945">07°20' 01.3&quot;N</geoCoordinate>
<geoCoordinate id="5DEC500EA82EFFA9FC55FA759A56FA04" box="[917,1076,1442,1465]" degrees="081" direction="east" minutes="41" orientation="longitude" pageId="57" pageNumber="56" precision="1" seconds="57. 1" value="81.699196">081°41'57.1&quot;E</geoCoordinate>
,
<elevation id="B3F5D1FAA82EFFA9FB81FA759AD6FA04" box="[1089,1204,1442,1465]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="57" pageNumber="56" unit="m" value="56.0">
<quantity id="FF209B2CA82EFFA9FB81FA759A18FA04" box="[1089,1146,1442,1465]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="57" pageNumber="56" unit="m" value="56.0">56 m</quantity>
a.s.l.
</elevation>
(
<location id="3D076012A82EFFA9FB05FA759B42FA04" LSID="urn:lsid:plazi:treatment:B07187DFA82EFFACFF39FA769A36F896:3D076012A82EFFA9FB05FA759B42FA04" box="[1221,1312,1442,1465]" country="Sri Lanka" county="Ampara District" latitude="7.3336945" longLatPrecision="1" longitude="81.699196" municipality="Ampara" name="Locality" pageId="57" pageNumber="56" stateProvince="Eastern">Locality</location>
<emphasis id="0AACEADBA82EFFA9FAE9FA769B04FA04" bold="true" box="[1321,1382,1441,1465]" pageId="57" pageNumber="56">15CS</emphasis>
,
<figureCitation id="A0E32A4CA82EFFA9FAB6FA759D05FA6A" captionStart="Figures 597598" captionStartId="128.[192,278,1657,1681]" captionTargetBox="[191,1440,191,1629]" captionTargetId="figure-140@128.[192,1439,192,1628]" captionTargetPageId="128" captionText="Figures 597598: Figure 597. Locality 15CS, Sri Lanka, Eastern Province, Ampara District, Ampara env., 07°20'01.3&quot;N 081°41'57.1&quot;E, 56 m a.s.l. During night collecting on 4.V.2015 (between 10.00 and 11.30 p.m.) we recorded Buthoscorpio sarasinorum (running on the ground among leaves and escaping to borrows), Reddyanus basilicus (on the ground among leaves), and Reddyanus loebli (hidden under scales of bark of standing tree trunks). Near to the locality we recorded maximum day temperature 39 ºC and 27 ºC night temperature at 10.00 p.m.; humidity 68%. Figure 598. Locality 15CT, Sri Lanka, Central Province, Kandy District, Tree centre Wildlif Trust Sri Lanka “Rantambe”, 07°12'22.1&quot;N 080°57'20.7&quot;E, 171 m a.s.l. During night collecting on 5.V.2015 (between 11.00 and 12.00 p.m.) during rain we recorded Buthoscorpio sarasinorum (running during rain on the ground among leaves) and Reddyanus loebli (hidden under scales of bark of standing tree trunks). On the locality we recorded night temperature 28.4 ºC 24.8 ºC (minimum temperature) and night humidity varied between 61% and 83%." figureDoi="http://doi.org/10.5281/zenodo.7124798" httpUri="https://zenodo.org/record/7124798/files/figure.png" pageId="57" pageNumber="56">Fig. 597</figureCitation>
),
<date id="4C661009A82EFFA9FCBEFA179D86FA6A" box="[894,996,1472,1495]" pageId="57" pageNumber="56" value="2015-05-04">
<collectingDate id="5C22E9E1A82EFFA9FCBEFA179D86FA6A" box="[894,996,1472,1495]" pageId="57" pageNumber="56" value="2015-05-04">4.V.2015</collectingDate>
</date>
,
<specimenCount id="2EDEFD40A82EFFA9FC31FA179A70FA6A" box="[1009,1042,1472,1495]" pageId="57" pageNumber="56" type="male">1♂</specimenCount>
(
<figureCitation id="A0E32A4CA82EFFA9FBE2FA179AEAFA6A" box="[1058,1160,1472,1495]" captionStart="Figures 193200" captionStartId="48.[192,278,1070,1094]" captionTargetBox="[213,1446,171,1037]" captionTargetId="figure-11@48.[205,1452,168,1041]" captionTargetPageId="48" captionText="Figures 193200: Distal segments of leg IV, retroventral view of Sri Lankan Buthidae and Chaerilidae genera. Buthoscorpio sarasinorum, male from locality 15CF (193), Charmus laneus, male from locality 15CO (194), Hottentotta tamulus, male from locality 15CK (195), Isometrus maculatus, female from locality 15CI (196), Isometrus thwaitesi, female from locality 15CO (197), Lychas srilankensis, male from locality 15CJ (198), Reddyanus basilicus, male from locality 15CS (199), and Chaerilus ceylonensis, male from locality 15CD (200)." figureDoi="http://doi.org/10.5281/zenodo.7124608" httpUri="https://zenodo.org/record/7124608/files/figure.png" pageId="57" pageNumber="56">Figs. 199</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FB59FA179A9BFA6A" box="[1177,1273,1472,1495]" captionStart="Figures 241259" captionStartId="52.[192,278,1733,1757]" captionTargetBox="[238,1387,153,1699]" captionTargetId="figure-11@52.[234,1392,152,1704]" captionTargetPageId="52" captionText="Figures 241259: Pedipalp chela dorsal (241), movable finger (242250), and fixed finger (251259) of Sri Lankan Isometrus, Lychas, and Reddyanus species. Figures 241242, 252. Isometrus maculatus, female from locality 15CO (241) and male from locality 15CP (242, 252). Figures 243244, 253. I. thwaitesi, male from locality 15CH (243) and female from locality 15CO (244, 253). Figures 245, 254. Reddyanus basilicus, male from locality 15CS. Figures 246, 255. R. besucheti, male from locality 15CD. Figures 247, 256. R. ceylonensis sp. n., male holotype. Figures 248, 257. R. jayarathnei sp. n., male paratype. Figures 249, 258. R. loebli, male from locality 15CG. Figures 250, 259. R. ranawanai sp. n., male holotype. Figure 251. Lychas srilankensis, male from locality 15CO." figureDoi="http://doi.org/10.5281/zenodo.7124626" httpUri="https://zenodo.org/record/7124626/files/figure.png" pageId="57" pageNumber="56">245, 254</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FAC5FA179B01FA6A" box="[1285,1379,1472,1495]" captionStart="Figures 260263" captionStartId="53.[192,278,1632,1656]" captionTargetBox="[196,1427,148,1596]" captionTargetId="figure-62@53.[192,1439,144,1603]" captionTargetPageId="53" captionText="Figures 260263: Reddyanus basilicus from locality 15CS. Figures 260261. Male in dorsal (260) and ventral (261) views. Figures 262263. Female in dorsal (262) and ventral (263) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7124628" httpUri="https://zenodo.org/record/7124628/files/figure.png" pageId="57" pageNumber="56">260261</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FAB3FA179D04FA4B" captionStart="Figures 264269" captionStartId="55.[192,278,1408,1432]" captionTargetBox="[191,1440,167,1380]" captionTargetId="figure-377@55.[192,1439,168,1379]" captionTargetPageId="55" captionText="Figures 264269: Reddyanus basilicus. Figures 264, 266. Male from locality 15CS, chelicerae, carapace and tergites IIII (264), and sternopectinal region and sternite III (266). Figures 265, 267. Female from locality 15CS, chelicerae, carapace and tergites IIII (265) and sternopectinal region and sternites IIIIV (267). Figures 268269. Male from locality 15CR, chelicera dorsal (268) and ventral (269) aspects." figureDoi="http://doi.org/10.5281/zenodo.7124630" httpUri="https://zenodo.org/record/7124630/files/figure.png" pageId="57" pageNumber="56">264, 266</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FCB8FA089DBAFA4B" box="[888,984,1503,1526]" captionStart="Figures 270275" captionStartId="56.[192,278,1839,1863]" captionTargetBox="[215,1432,184,1819]" captionTargetId="figure-11@56.[192,1439,173,1824]" captionTargetPageId="56" captionText="Figures 270275: Reddyanus basilicus from locality 15CS. Figures 270272. Male, metasoma and telson, lateral (270), ventral (271), and dorsal (272) views. Figures 273275. Female, metasoma and telson, lateral (273), ventral (274), and dorsal (275) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7124632" httpUri="https://zenodo.org/record/7124632/files/figure.png" pageId="57" pageNumber="56">270272</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FC26FA089A21FA4B" box="[998,1091,1503,1526]" captionStart-0="Figures 276282" captionStart-1="Figures 283286" captionStartId-0="57.[192,278,892,916]" captionStartId-1="58.[192,278,1625,1649]" captionTargetBox-0="[198,1431,222,869]" captionTargetBox-1="[201,1430,178,1594]" captionTargetId-0="figure-6@57.[192,1439,199,888]" captionTargetId-1="figure-11@58.[192,1439,168,1596]" captionTargetPageId-0="57" captionTargetPageId-1="58" captionText-0="Figures 276282: Reddyanus basilicus from locality 15CS. Figures 276277. Female. Pedipalp chela, dorsal (276) and external (277) views. Figures 278282. Male. Pedipalp chela, dorsal (278) and external (279) views. Pedipalp patella, dorsal (280) and external (281) views. Pedipalp femur and trochanter dorsal view (282). The trichobothrial pattern is indicated in Figures 279282." captionText-1="Figures 283286: Left hemispermatophore of Reddyanus basilicus from locality 15CR. External (283), dorsal (284), internal (285) and ventral (286) views. Scale bar: 500 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; t, trunk." figureDoi-0="http://doi.org/10.5281/zenodo.7124634" figureDoi-1="http://doi.org/10.5281/zenodo.7124636" httpUri-0="https://zenodo.org/record/7124634/files/figure.png" httpUri-1="https://zenodo.org/record/7124636/files/figure.png" pageId="57" pageNumber="56">278286</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FB94FA089A1DFA4B" box="[1108,1151,1503,1526]" captionStart="Figures 287290" captionStartId="59.[192,278,1867,1891]" captionTargetBox="[191,1440,167,1838]" captionTargetId="figure-47@59.[192,1439,168,1837]" captionTargetPageId="59" captionText="Figures 287290: Reddyanus basilicus from locality 15CS, female (287), female with newborns before first ecdysis (288), female with juveniles after first ecdysis (289), and male (290)." figureDoi="http://doi.org/10.5281/zenodo.7124638" httpUri="https://zenodo.org/record/7124638/files/figure.png" pageId="57" pageNumber="56">290</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FB4DFA089AD7FA4B" box="[1165,1205,1503,1526]" captionStart="Figures 547554" captionStartId="114.[192,278,810,834]" captionTargetBox="[191,1440,191,782]" captionTargetId="figure-538@114.[192,1440,192,781]" captionTargetPageId="114" captionText="Figures 547554: The mitotic metaphases of buthid males from Sri Lanka. Buthoscorpio sarasinorum from locality 15CF (2n=14) (547), Charmus laneus from locality 15CO (2n=9) (548), Isometrus thwaitesi from locality 15CF (2n=8) (549), Lychas srilankensis from locality 15CN (2n=16) (550), Reddyanus basilicus from locality 15CS (2n=16) (551), Reddyanus basilicus from locality 15CR (2n=15) (552), Reddyanus ceylonensis from locality 15CI (2n=16) (553), Reddyanus loebli from locality 15CH (2n=17) (554). Arrowheads indicate extra large odd chromosomes. Scale bar = 5 µm for 547554." figureDoi="http://doi.org/10.5281/zenodo.7124765" httpUri="https://zenodo.org/record/7124765/files/figure.png" pageId="57" pageNumber="56">551</figureCitation>
)
<specimenCount id="2EDEFD40A82EFFA9FB08FA089A88FA4B" box="[1224,1258,1503,1526]" pageId="57" pageNumber="56" type="female">2♀</specimenCount>
(
<figureCitation id="A0E32A4CA82EFFA9FB3DFA089BFBFA4B" box="[1277,1433,1503,1526]" captionStart="Figures 260263" captionStartId="53.[192,278,1632,1656]" captionTargetBox="[196,1427,148,1596]" captionTargetId="figure-62@53.[192,1439,144,1603]" captionTargetPageId="53" captionText="Figures 260263: Reddyanus basilicus from locality 15CS. Figures 260261. Male in dorsal (260) and ventral (261) views. Figures 262263. Female in dorsal (262) and ventral (263) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7124628" httpUri="https://zenodo.org/record/7124628/files/figure.png" pageId="57" pageNumber="56">Figs. 262263</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FCFEFA299DC7F9A8" box="[830,933,1534,1557]" captionStart="Figures 264269" captionStartId="55.[192,278,1408,1432]" captionTargetBox="[191,1440,167,1380]" captionTargetId="figure-377@55.[192,1439,168,1379]" captionTargetPageId="55" captionText="Figures 264269: Reddyanus basilicus. Figures 264, 266. Male from locality 15CS, chelicerae, carapace and tergites IIII (264), and sternopectinal region and sternite III (266). Figures 265, 267. Female from locality 15CS, chelicerae, carapace and tergites IIII (265) and sternopectinal region and sternites IIIIV (267). Figures 268269. Male from locality 15CR, chelicera dorsal (268) and ventral (269) aspects." figureDoi="http://doi.org/10.5281/zenodo.7124630" httpUri="https://zenodo.org/record/7124630/files/figure.png" pageId="57" pageNumber="56">265, 267</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FC7BFA299A79F9A8" box="[955,1051,1534,1557]" captionStart="Figures 270275" captionStartId="56.[192,278,1839,1863]" captionTargetBox="[215,1432,184,1819]" captionTargetId="figure-11@56.[192,1439,173,1824]" captionTargetPageId="56" captionText="Figures 270275: Reddyanus basilicus from locality 15CS. Figures 270272. Male, metasoma and telson, lateral (270), ventral (271), and dorsal (272) views. Figures 273275. Female, metasoma and telson, lateral (273), ventral (274), and dorsal (275) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7124632" httpUri="https://zenodo.org/record/7124632/files/figure.png" pageId="57" pageNumber="56">273275</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FBEFFA299AEDF9A8" box="[1071,1167,1534,1557]" captionStart="Figures 276282" captionStartId="57.[192,278,892,916]" captionTargetBox="[198,1431,222,869]" captionTargetId="figure-6@57.[192,1439,199,888]" captionTargetPageId="57" captionText="Figures 276282: Reddyanus basilicus from locality 15CS. Figures 276277. Female. Pedipalp chela, dorsal (276) and external (277) views. Figures 278282. Male. Pedipalp chela, dorsal (278) and external (279) views. Pedipalp patella, dorsal (280) and external (281) views. Pedipalp femur and trochanter dorsal view (282). The trichobothrial pattern is indicated in Figures 279282." figureDoi="http://doi.org/10.5281/zenodo.7124634" httpUri="https://zenodo.org/record/7124634/files/figure.png" pageId="57" pageNumber="56">276277</figureCitation>
,
<figureCitation id="A0E32A4CA82EFFA9FB63FA299B62F9A8" box="[1187,1280,1534,1557]" captionStart="Figures 287290" captionStartId="59.[192,278,1867,1891]" captionTargetBox="[191,1440,167,1838]" captionTargetId="figure-47@59.[192,1439,168,1837]" captionTargetPageId="59" captionText="Figures 287290: Reddyanus basilicus from locality 15CS, female (287), female with newborns before first ecdysis (288), female with juveniles after first ecdysis (289), and male (290)." figureDoi="http://doi.org/10.5281/zenodo.7124638" httpUri="https://zenodo.org/record/7124638/files/figure.png" pageId="57" pageNumber="56">287289</figureCitation>
),
<collectionCode id="5EC9AE0CA82EFFA9FAE0FA299B07F9A8" box="[1312,1381,1534,1557]" pageId="57" pageNumber="56">FKCP</collectionCode>
, leg.
<collectorName id="952D531FA82EFFA9FCFEF9CB9DF4F98E" box="[830,918,1564,1587]" pageId="57" pageNumber="56">Kovařík</collectorName>
et al.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="70C26542A82EFFACFCFEF98D9D16F93B" lastPageId="60" lastPageNumber="59" pageId="57" pageNumber="56" type="diagnosis">
<paragraph id="386736C9A82EFFAAFCFEF98D9DE2F8C4" blockId="57.[830,1441,1626,1833]" lastBlockId="58.[830,1440,1736,1913]" lastPageId="58" lastPageNumber="57" pageId="57" pageNumber="56">
DIAGNOSIS. Total length
<quantity id="FF209B2CA82EFFA9FB97F98D9AACF9CC" box="[1111,1230,1626,1649]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="4.6" metricValueMin="2.7" pageId="57" pageNumber="56" unit="mm" value="36.5" valueMax="46.0" valueMin="27.0">2746 mm</quantity>
. Male with longer metasomal segments and telson than female. Segments of pedipalps approximately the same length and width in both sexes. Ratio of pedipalp chela length/ width 3.41 3.79 in male. Pedipalp movable finger clearly longer than chela manus in both sexes, but in largest male specimens could be shorter than manus. Base coloration reddish. Pedipalps and legs with brown maculation, identical on femur and patella. First metasomal segment bears 10 carinae, second through fourth segments bear eight carinae, fifth segment bears five carinae in female and three to five in male. Terminal tubercle of each dorsal carina on metasomal segments of both sexes scarcely larger than preceding tubercles. Subaculear tooth wide and rounded, dorsally with granules in three or four rows; four or six symmetrical granules and one or two granules on tip. Glabrous zone along posterior margin of fifth sternite absent. Pectinal teeth number
<quantity id="FF209B2CA82DFFAAFACBF8949B08F8E7" box="[1291,1386,1859,1882]" metricMagnitude="-1" metricUnit="m" metricValue="3.937" metricValueMax="4.572" metricValueMin="3.302" pageId="58" pageNumber="57" unit="in" value="15.5" valueMax="18.0" valueMin="13.0">1318 in</quantity>
both sexes.
</paragraph>
<caption id="6CA76641A82DFFAAFF00F98E9F31F91E" ID-DOI="http://doi.org/10.5281/zenodo.7124636" ID-Zenodo-Dep="7124636" httpUri="https://zenodo.org/record/7124636/files/figure.png" pageId="58" pageNumber="57" startId="58.[192,278,1625,1649]" targetBox="[201,1430,178,1594]" targetPageId="58">
<paragraph id="386736C9A82DFFAAFF00F98E9F31F91E" blockId="58.[192,1441,1625,1699]" pageId="58" pageNumber="57">
<emphasis id="0AACEADBA82DFFAAFF00F98E9FE6F9CF" bold="true" box="[192,388,1625,1650]" pageId="58" pageNumber="57">Figures 283286:</emphasis>
Left hemispermatophore of
<taxonomicName id="FFD84D4AA82DFFAAFD60F98B9D04F9CC" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[672,870,1628,1649]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="58" pageNumber="57" phylum="Arthropoda" rank="species" species="basilicus">
<emphasis id="0AACEADBA82DFFAAFD60F98B9D04F9CC" box="[672,870,1628,1649]" italics="true" pageId="58" pageNumber="57">Reddyanus basilicus</emphasis>
</taxonomicName>
from locality 15CR. External (283), dorsal (284), internal (285) and ventral (286) views. Scale bar: 500 µm. Abbreviations:
<emphasis id="0AACEADBA82DFFAAFC85F9A29D35F937" box="[837,855,1653,1674]" italics="true" pageId="58" pageNumber="57">bl</emphasis>
, basal lobe;
<emphasis id="0AACEADBA82DFFAAFC13F9A29DBBF937" box="[979,985,1653,1674]" italics="true" pageId="58" pageNumber="57">f</emphasis>
, flagellum;
<emphasis id="0AACEADBA82DFFAAFB91F9A29A0AF937" box="[1105,1128,1653,1674]" italics="true" pageId="58" pageNumber="57">ml</emphasis>
, median lobe;
<emphasis id="0AACEADBA82DFFAAFB3AF9A29B7EF937" box="[1274,1308,1653,1674]" italics="true" pageId="58" pageNumber="57">mlc</emphasis>
, median lobe carina;
<emphasis id="0AACEADBA82DFFAAFEC7F9599F6CF91E" box="[263,270,1678,1699]" italics="true" pageId="58" pageNumber="57">t</emphasis>
, trunk.
</paragraph>
</caption>
<caption id="6CA76641A82CFFABFF00F89C9D7AF8C6" ID-DOI="http://doi.org/10.5281/zenodo.7124638" ID-Zenodo-Dep="7124638" httpUri="https://zenodo.org/record/7124638/files/figure.png" pageId="59" pageNumber="58" startId="59.[192,278,1867,1891]" targetBox="[191,1440,167,1838]" targetPageId="59">
<paragraph id="386736C9A82CFFABFF00F89C9D7AF8C6" blockId="59.[192,1440,1867,1915]" pageId="59" pageNumber="58">
<emphasis id="0AACEADBA82CFFABFF00F89C9FE4F8DE" bold="true" box="[192,390,1867,1891]" pageId="59" pageNumber="58">Figures 287290:</emphasis>
<taxonomicName id="FFD84D4AA82CFFABFE4FF8999C35F8DE" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[399,599,1870,1891]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="59" pageNumber="58" phylum="Arthropoda" rank="species" species="basilicus">
<emphasis id="0AACEADBA82CFFABFE4FF8999C35F8DE" box="[399,599,1870,1891]" italics="true" pageId="59" pageNumber="58">Reddyanus basilicus</emphasis>
</taxonomicName>
from locality 15CS, female (287), female with newborns before first ecdysis (288), female with juveniles after first ecdysis (289), and male (290).
</paragraph>
</caption>
<caption id="6CA76641A82BFFACFF00FA3A9A62F9A0" ID-DOI="http://doi.org/10.5281/zenodo.7124640" ID-Zenodo-Dep="7124640" httpUri="https://zenodo.org/record/7124640/files/figure.png" pageId="60" pageNumber="59" startId="60.[192,278,1517,1541]" targetBox="[207,1424,168,1482]" targetPageId="60">
<paragraph id="386736C9A82BFFACFF00FA3A9A62F9A0" blockId="60.[192,1443,1517,1566]" pageId="60" pageNumber="59">
<emphasis id="0AACEADBA82BFFACFF00FA3A9FE6F9B8" bold="true" box="[192,388,1517,1541]" pageId="60" pageNumber="59">Figures 2 91294:</emphasis>
<taxonomicName id="FFD84D4AA82BFFACFE4CFA389C3AF9B9" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Vachon" baseAuthorityYear="1982" box="[396,600,1519,1540]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="60" pageNumber="59" phylum="Arthropoda" rank="species" species="besucheti">
<emphasis id="0AACEADBA82BFFACFE4CFA389C3AF9B9" box="[396,600,1519,1540]" italics="true" pageId="60" pageNumber="59">Reddyanus besucheti</emphasis>
</taxonomicName>
.
<emphasis id="0AACEADBA82BFFACFDA5FA389D6CF9B9" bold="true" box="[613,782,1519,1541]" pageId="60" pageNumber="59">Figures 2 91292</emphasis>
. Male from locality 15CD in dorsal (291) and ventral (292) views.
<emphasis id="0AACEADBA82BFFACFF00F9DF9F0AF9A0" bold="true" box="[192,360,1544,1566]" pageId="60" pageNumber="59">Figures 293294</emphasis>
. Female from locality 15CG in dorsal (293) and ventral (294) views.
</paragraph>
</caption>
<paragraph id="386736C9A82BFFACFF00F9859D16F93B" blockId="60.[192,802,1618,1834]" lastBlockId="60.[830,1441,1618,1789]" pageId="60" pageNumber="59">
HEMISPERMATOPHORE (
<figureCitation id="A0E32A4CA82BFFACFE7EF9859C35F9D4" box="[446,599,1618,1641]" captionStart="Figures 283286" captionStartId="58.[192,278,1625,1649]" captionTargetBox="[201,1430,178,1594]" captionTargetId="figure-11@58.[192,1439,168,1596]" captionTargetPageId="58" captionText="Figures 283286: Left hemispermatophore of Reddyanus basilicus from locality 15CR. External (283), dorsal (284), internal (285) and ventral (286) views. Scale bar: 500 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; t, trunk." figureDoi="http://doi.org/10.5281/zenodo.7124636" httpUri="https://zenodo.org/record/7124636/files/figure.png" pageId="60" pageNumber="59">Figs. 283286</figureCitation>
). Trunk moderately narrow, elongate, more than twice as long as capsule region. Flagellum long, slender, laminiform, with broad hyaline fin along internal margin of cylindrical core, distally coiled. Median lobe narrow, distally truncate, with thin dorsal lamina near internal margin. Basal lobe greatly enlarged, a narrow, angular, pointed hook-like process arising dorsally, separated from base of median lobe.
</paragraph>
</subSubSection>
<subSubSection id="70C26542A82BFFACFCFEF95A9DF4F940" pageId="60" pageNumber="59" type="description">
<paragraph id="386736C9A82BFFACFCFEF95A9DF4F940" blockId="60.[830,1441,1618,1789]" pageId="60" pageNumber="59">
VARIABILITY. There is high variability in morphometric characters between small males
<quantity id="FF209B2CA82BFFACFB57F97C9B68F97F" box="[1175,1290,1707,1730]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.2" metricValueMin="2.7" pageId="60" pageNumber="59" unit="mm" value="29.5" valueMax="32.0" valueMin="27.0">2732 mm</quantity>
long after the fourth ecdysis, and males
<quantity id="FF209B2CA82BFFACFB9BF91E9AADF95D" box="[1115,1231,1737,1760]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" metricValueMax="4.6" metricValueMin="4.0" pageId="60" pageNumber="59" unit="mm" value="43.0" valueMax="46.0" valueMin="40.0">4046 mm</quantity>
long after the fifth ecdysis.
</paragraph>
</subSubSection>
<subSubSection id="70C26542A82BFFACFCFEF8C39A36F896" box="[830,1108,1812,1835]" pageId="60" pageNumber="59" type="distribution">
<paragraph id="386736C9A82BFFACFCFEF8C39A36F896" blockId="60.[830,1108,1812,1835]" box="[830,1108,1812,1835]" pageId="60" pageNumber="59">
DISTRIBUTION.
<collectingCountry id="40CF7659A82BFFACFC25F8C39A2CF896" box="[997,1102,1812,1835]" name="Sri Lanka" pageId="60" pageNumber="59">Sri Lanka</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>