303 lines
32 KiB
XML
303 lines
32 KiB
XML
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<mods:title id="0A6302877560A980DABA31C0882ED855">Johnwellsia, a new intertidal genus of Parastenheliidae (Copepoda, Harpacticoida) from the Taiwan Strait, China, including a review of the family and key to genera</mods:title>
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<mods:namePart id="EF5CBF786A71ABC42CB06F112E296EAE">Huys, Rony</mods:namePart>
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<mods:affiliation id="091DB95A716D23754E165DBB4F8FBD02">Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K.</mods:affiliation>
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<mods:namePart id="12BD830A064A546FA6F8809E8D93616B">Mu, Fanghong</mods:namePart>
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<mods:affiliation id="FDD52A38F38D3E414003BFB37E8E9739">College of Marine Life Science, Ocean University of China, 5 Yushan Road, Qingdao 266003, China.</mods:affiliation>
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<mods:title id="CD11E0E9469BD64D554624CCC3CE56C5">Zootaxa</mods:title>
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<mods:date id="2E5009C1B685BF9DE1B62CB629B4A5A0">2021</mods:date>
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<treatment id="951887EAFFF8FFA2FF51D277E690FD7B" ID-DOI="http://doi.org/10.5281/zenodo.5579343" ID-GBIF-Taxon="189091332" ID-Zenodo-Dep="5579343" LSID="urn:lsid:zoobank.org:act:3541F2B6-4847-4ECE-B77A-C4D55732718F" httpUri="http://treatment.plazi.org/id/951887EAFFF8FFA2FF51D277E690FD7B" lastPageId="60" lastPageNumber="296" pageId="59" pageNumber="295">
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<paragraph id="1D0E36FCFFF8FFA5FF51D277E5D4FE52" blockId="59.[151,886,367,423]" box="[151,404,367,393]" pageId="59" pageNumber="295">
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<heading id="46468190FFF8FFA5FF51D277E5D4FE52" bold="true" box="[151,404,367,393]" fontSize="11" level="1" pageId="59" pageNumber="295" reason="1">
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<emphasis id="2FC5EAEEFFF8FFA5FF51D277E5D4FE52" bold="true" box="[151,404,367,393]" pageId="59" pageNumber="295">
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<taxonomicName id="DAB14D7FFFF8FFA5FF51D277E55EFE52" authority="Huys & Mu, 2021" box="[151,286,367,393]" class="Hexanauplia" family="Parastenheliidae" genus="Karaytugia" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="295" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="2FC5EAEEFFF8FFA5FF51D277E55EFE52" bold="true" box="[151,286,367,393]" italics="true" pageId="59" pageNumber="295">Karaytugia</emphasis>
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<taxonomicNameLabel id="34F65795FFF8FFA5FEE0D277E5D4FE52" box="[294,404,367,393]" pageId="59" pageNumber="295" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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<paragraph id="1D0E36FCFFF8FFA5FF51D297E736FE7C" blockId="59.[151,886,367,423]" box="[151,886,399,423]" pageId="59" pageNumber="295">
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urn:lsid:zoobank.org:act:
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</uri>
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<subSubSection id="55AB6577FFF8FFA5FF51D2CEE5C7FBAC" pageId="59" pageNumber="295" type="description">
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<paragraph id="1D0E36FCFFF8FFA5FF51D2CEE615FD30" blockId="59.[151,1437,469,2008]" pageId="59" pageNumber="295">
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<materialsCitation id="ADD93CA1FFF8FFA5FF51D2CEE611FD30" ID-GBIF-Occurrence="3387337359" collectionCode="V" county="They" location="Turkish" municipality="The" pageId="59" pageNumber="295" specimenCount="6" specimenCount-female="3" specimenCount-male="3">
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<bibRefCitation id="79204B0DFFF8FFA5FF51D2CEE5FBFE2B" author="Kuru, S. & Karaytug, S." box="[151,443,469,496]" pageId="59" pageNumber="295" pagination="121 - 127" refId="ref75081" refString="Kuru, S. & Karaytug, S. (2015) A new species of Parastenhelia Thompson & A. Scott, 1903 (Copepoda, Harpacticoida, Parastenheliidae) from Turkey. Biharean Biologist, 9, 121 - 127." type="journal article" year="2015">Kuru & Karaytuğ (2015)</bibRefCitation>
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described a new species,
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<taxonomicName id="DAB14D7FFFF8FFA5FD34D2CDE7A0FE34" authorityName="Kuru & Karaytug" authorityYear="2015" baseAuthorityName="Kuru & Karaytug" baseAuthorityYear="2015" box="[754,992,469,495]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="295" phylum="Arthropoda" rank="species" species="aydini">
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<emphasis id="2FC5EAEEFFF8FFA5FD34D2CDE7A0FE34" box="[754,992,469,495]" italics="true" pageId="59" pageNumber="295">Parastenhelia aydini</emphasis>
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</taxonomicName>
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, from a
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beach in the
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<location id="186E6027FFF8FFA5FA8ED2CEE57AFDC8" LSID="urn:lsid:plazi:treatment:951887EAFFF8FFA2FF51D277E690FD7B:186E6027FFF8FFA5FA8ED2CEE57AFDC8" county="They" municipality="The" name="Eastern Mediterranean" pageId="59" pageNumber="295">Eastern Mediterranean</location>
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and considered it morphologically most similar to
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<taxonomicName id="DAB14D7FFFF8FFA5FCABD2E2E7AFFDC8" baseAuthorityName="Lang" baseAuthorityYear="1948" box="[877,1007,506,531]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="295" phylum="Arthropoda" rank="species" species="spinosa">
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<emphasis id="2FC5EAEEFFF8FFA5FCABD2E2E7AFFDC8" box="[877,1007,506,531]" italics="true" pageId="59" pageNumber="295">Pa. spinosa</emphasis>
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</taxonomicName>
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.
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<collectingMunicipality id="FD6AAC86FFF8FFA5FC3DD2E1E067FDC8" box="[1019,1063,505,531]" pageId="59" pageNumber="295">The</collectingMunicipality>
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authors refer to their unpublished observations but it is not clear which “population” they identified as
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<taxonomicName id="DAB14D7FFFF8FFA5FC77D106E076FDEC" baseAuthorityName="Lang" baseAuthorityYear="1948" box="[945,1078,542,567]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="295" phylum="Arthropoda" rank="species" species="spinosa">
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<emphasis id="2FC5EAEEFFF8FFA5FC77D106E076FDEC" box="[945,1078,542,567]" italics="true" pageId="59" pageNumber="295">Pa. spinosa</emphasis>
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</taxonomicName>
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.
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<collectingCounty id="F46F4E70FFF8FFA5FB83D105E03FFDEC" box="[1093,1151,541,567]" pageId="59" pageNumber="295">They</collectingCounty>
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differentiated
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<taxonomicName id="DAB14D7FFFF8FFA5FAE1D106E1DCFDEC" authorityName="Kuru & Karaytug" authorityYear="2015" baseAuthorityName="Kuru & Karaytug" baseAuthorityYear="2015" box="[1319,1436,541,567]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="295" phylum="Arthropoda" rank="species" species="aydini">
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<emphasis id="2FC5EAEEFFF8FFA5FAE1D106E1DCFDEC" box="[1319,1436,541,567]" italics="true" pageId="59" pageNumber="295">Pa. aydini</emphasis>
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</taxonomicName>
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from the latter by the combination of the following characters: (a)
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<specimenCount id="0BB7FD75FFF8FFA5FCBDD15AE7D0FD87" box="[891,912,578,604]" pageId="59" pageNumber="295" type="female">♀</specimenCount>
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antennule 9-segmented; (b) spinular rows near outer margin of P1–P4 bases significantly reduced; (c) P4 enp-3 with four setae in both sexes; (d) ornamentation of endopodal and exopodal setae of
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<specimenCount id="0BB7FD75FFF8FFA5FDC1D192E65CFD7F" box="[519,540,650,676]" pageId="59" pageNumber="295" type="female">♀</specimenCount>
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P5 different; (e) caudal ramus seta
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<collectionCode id="7BA0AE39FFF8FFA5FC67D192E7F5FD7F" box="[929,949,650,676]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="59" pageNumber="295" type="Museum">V</collectionCode>
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of
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<specimenCount id="0BB7FD75FFF8FFA5FC11D192E7ACFD7F" box="[983,1004,650,676]" pageId="59" pageNumber="295" type="female">♀</specimenCount>
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not swollen near the base; (f) modified element on the seventh segment of
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<specimenCount id="0BB7FD75FFF8FFA5FDEDD1B6E600FD13" box="[555,576,686,712]" pageId="59" pageNumber="295" type="male">♂</specimenCount>
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antennule not present; (g) apophysis on
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<specimenCount id="0BB7FD75FFF8FFA5FBD4D1B6E067FD13" box="[1042,1063,686,712]" pageId="59" pageNumber="295" type="male">♂</specimenCount>
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P3 enp-3 notched; and (h)
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<specimenCount id="0BB7FD75FFF8FFA5FAA7D1B6E136FD13" box="[1377,1398,686,712]" pageId="59" pageNumber="295" type="male">♂</specimenCount>
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P5 exopod 2-segmented, five setae in total
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="1D0E36FCFFF8FFA5FF01D1EDE5C7FBAC" blockId="59.[151,1437,469,2008]" pageId="59" pageNumber="295">
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Unique characters displayed by
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<taxonomicName id="DAB14D7FFFF8FFA5FDEFD1EEE6DBFCD4" authorityName="Kuru & Karaytug" authorityYear="2015" baseAuthorityName="Kuru & Karaytug" baseAuthorityYear="2015" box="[553,667,757,783]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="295" phylum="Arthropoda" rank="species" species="aydini">
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<emphasis id="2FC5EAEEFFF8FFA5FDEFD1EEE6DBFCD4" box="[553,667,757,783]" italics="true" pageId="59" pageNumber="295">Pa. aydini</emphasis>
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</taxonomicName>
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include (a) the modification of the endopodal claw on the maxilliped; the distal half does not taper to a sharp tip as in other members of the family, forming instead a spatulate structure fringed with a series of densely arranged serrations around the inner margin and rounded apex; (b) P1 exp-3 with one very long naked seta and three claws of different lengths, the longest one being 0.8 times the length of exp-2; (c) P1 enp-2 claws radically divergent in size, with large one nearly three times longer than short one; (d) inner seta of P2–P3 enp-1 reduced; (e) spinous apophysis on
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<specimenCount id="0BB7FD75FFF8FFA5FCD6D0B2E765FC1F" box="[784,805,938,964]" pageId="59" pageNumber="295" type="male">♂</specimenCount>
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P3 endopod with foot-shaped tip; and (f)
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||
<specimenCount id="0BB7FD75FFF8FFA5FAC0D0B2E15BFC1F" box="[1286,1307,938,964]" pageId="59" pageNumber="295" type="male">♂</specimenCount>
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||
P5 exopod 2-segmented, with one very long outer seta on exp-1 and four elements on exp-2; the only other species in the
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<taxonomicName id="DAB14D7FFFF8FFA5FF51D0E9E50FFBD0" baseAuthorityName="Willen" baseAuthorityYear="2000" box="[151,335,1009,1035]" class="Hexanauplia" family="Parastenheliidae" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="295" phylum="Arthropoda" rank="family">Parastenheliidae</taxonomicName>
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||
that display a 2-segmented exopod are
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||
<taxonomicName id="DAB14D7FFFF8FFA5FCDBD0E9E035FBD0" box="[797,1141,1009,1035]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="295" phylum="Arthropoda" rank="species" species="willemvervoorti" status="sp. nov.">
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||
<emphasis id="2FC5EAEEFFF8FFA5FCDBD0E9E035FBD0" box="[797,1141,1009,1035]" italics="true" pageId="59" pageNumber="295">Parastenhelia willemvervoorti</emphasis>
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||
</taxonomicName>
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||
<emphasis id="2FC5EAEEFFF8FFA5FBB9D0E9E09BFBD0" bold="true" box="[1151,1243,1009,1035]" pageId="59" pageNumber="295">
|
||
<taxonomicNameLabel id="34F65795FFF8FFA5FBB9D0E9E09BFBD0" box="[1151,1243,1009,1035]" pageId="59" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
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||
and
|
||
<taxonomicName id="DAB14D7FFFF8FFA5FAD0D0E9E4B9FBF4" class="Hexanauplia" family="Parastenheliidae" genus="Johnwellsia" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="295" phylum="Arthropoda" rank="species" species="bipartita" status="gen. et sp. nov.">
|
||
<emphasis id="2FC5EAEEFFF8FFA5FAD0D0E9E4B9FBF4" italics="true" pageId="59" pageNumber="295">Johnwellsia bipartita</emphasis>
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||
</taxonomicName>
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||
<emphasis id="2FC5EAEEFFF8FFA5FEC6D70DE5EDFBF4" bold="true" box="[256,429,1045,1071]" pageId="59" pageNumber="295">
|
||
<taxonomicNameLabel id="34F65795FFF8FFA5FEC6D70DE5EDFBF4" box="[256,429,1045,1071]" pageId="59" pageNumber="295" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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</emphasis>
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which have six short elements, and one spine and two setae, respectively, on exp-2. Based on this combination of apomorphic character states,
|
||
<taxonomicName id="DAB14D7FFFF8FFA5FD33D722E72BFB88" authorityName="Kuru & Karaytug" authorityYear="2015" baseAuthorityName="Kuru & Karaytug" baseAuthorityYear="2015" box="[757,875,1081,1107]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="295" phylum="Arthropoda" rank="species" species="aydini">
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<emphasis id="2FC5EAEEFFF8FFA5FD33D722E72BFB88" box="[757,875,1081,1107]" italics="true" pageId="59" pageNumber="295">Pa. aydini</emphasis>
|
||
</taxonomicName>
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is her fixed as the
|
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<typeStatus id="C20A885EFFF8FFA5FB89D722E03FFB8F" box="[1103,1151,1082,1108]" pageId="59" pageNumber="295">type</typeStatus>
|
||
species of a new genus,
|
||
<taxonomicName id="DAB14D7FFFF8FFA5FF51D746E557FBAC" box="[151,279,1118,1143]" class="Hexanauplia" family="Parastenheliidae" genus="Karaytugia" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="295" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="2FC5EAEEFFF8FFA5FF51D746E557FBAC" box="[151,279,1118,1143]" italics="true" pageId="59" pageNumber="295">Karaytugia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2FC5EAEEFFF8FFA5FED8D745E5C7FBAC" bold="true" box="[286,391,1117,1143]" pageId="59" pageNumber="295">
|
||
<taxonomicNameLabel id="34F65795FFF8FFA5FED8D745E5C7FBAC" box="[286,391,1117,1143]" pageId="59" pageNumber="295" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55AB6577FFF8FFA2FF01D799E754FE30" lastPageId="60" lastPageNumber="296" pageId="59" pageNumber="295" type="diagnosis">
|
||
<paragraph id="1D0E36FCFFF8FFA5FF01D799E6B9FA8B" blockId="59.[151,1437,469,2008]" pageId="59" pageNumber="295">
|
||
<emphasis id="2FC5EAEEFFF8FFA5FF01D799E502FB40" bold="true" box="[199,322,1153,1179]" pageId="59" pageNumber="295">Diagnosis.</emphasis>
|
||
<materialsCitation id="ADD93CA1FFF8FFA5FE94D799E6B2FA8B" ID-GBIF-Occurrence="3387337346" collectionCode="V" pageId="59" pageNumber="295" specimenCount="1" specimenCount-female="1">
|
||
<taxonomicName id="DAB14D7FFFF8FFA5FE94D799E64DFB40" baseAuthorityName="Willen" baseAuthorityYear="2000" box="[338,525,1153,1179]" class="Hexanauplia" family="Parastenheliidae" kingdom="Animalia" order="Harpacticoida" pageId="59" pageNumber="295" phylum="Arthropoda" rank="family">Parastenheliidae</taxonomicName>
|
||
. Sexual dimorphism in antennule, P2–P3 endopods, P5–P6 and urosomal segmentation. Body subcylindrical, short; posterior margin of cephalothorax and somites bearing P2–P5 with plain hyaline frills; remaining urosomites (except anal somite) with serrate hyaline frills. Rostrum defined at base, of moderate size, reaching to about halfway second segment of antennule; elongate triangular. Anal operculum semicircular, bordered with fine spinules. Caudal ramus wider than long, with conspicuous spinular ornamentation; with seven setae, setae IV–
|
||
<collectionCode id="7BA0AE39FFF8FFA5FE01D62EE59BFA8B" box="[455,475,1334,1360]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="59" pageNumber="295" type="Museum">V</collectionCode>
|
||
not basally inflated in
|
||
<specimenCount id="0BB7FD75FFF8FFA5FD1AD62EE6B2FA8B" box="[732,754,1334,1360]" pageId="59" pageNumber="295" type="female">♀</specimenCount>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1D0E36FCFFF8FFA5FF01D641E0BFF963" blockId="59.[151,1437,469,2008]" pageId="59" pageNumber="295">
|
||
<materialsCitation id="ADD93CA1FFF8FFA5FF01D641E0BCF963" ID-GBIF-Occurrence="3387337336" collectionCode="VIII, IX, XII, XIII, XIV, XVII, XVIII, XIX, XX, XXI, XXII, XXIII, XXIV, XXV, XXVI, XXVIII" pageId="59" pageNumber="295" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
|
||
Antennule elongate and 9-segmented in
|
||
<specimenCount id="0BB7FD75FFF8FFA5FD51D642E6ECFAAF" box="[663,684,1370,1396]" pageId="59" pageNumber="295" type="female">♀</specimenCount>
|
||
, segment 1 not elongate, segments 7–8 shortest, with aesthetascs on segments 4 and 9; haplocer and 11-segmented in
|
||
<specimenCount id="0BB7FD75FFF8FFA5FD33D666E74CFA43" box="[757,780,1406,1432]" pageId="59" pageNumber="295" type="male">♂</specimenCount>
|
||
, with geniculation between segments 7 and 8, segment 5 swollen and with aesthetasc; segmental homologies in
|
||
<specimenCount id="0BB7FD75FFF8FFA5FCCDD6BAE763FA67" box="[779,803,1442,1468]" pageId="59" pageNumber="295" type="male">♂</specimenCount>
|
||
: I, II–
|
||
<collectionCode id="7BA0AE39FFF8FFA5FCAAD6BAE7DEFA67" box="[876,926,1442,1468]" pageId="59" pageNumber="295">VIII</collectionCode>
|
||
,
|
||
<collectionCode id="7BA0AE39FFF8FFA5FC6DD6BAE789FA67" box="[939,969,1442,1468]" pageId="59" pageNumber="295">IX</collectionCode>
|
||
–
|
||
<collectionCode id="7BA0AE39FFF8FFA5FC1ED6BAE040FA67" box="[984,1024,1442,1468]" pageId="59" pageNumber="295">XII</collectionCode>
|
||
,
|
||
<collectionCode id="7BA0AE39FFF8FFA5FBCBD6BAE000FA67" box="[1037,1088,1442,1468]" pageId="59" pageNumber="295">XIII</collectionCode>
|
||
,
|
||
<collectionCode id="7BA0AE39FFF8FFA5FB88D6BAE0C1FA67" box="[1102,1153,1442,1468]" pageId="59" pageNumber="295">XIV</collectionCode>
|
||
–
|
||
<collectionCode id="7BA0AE39FFF8FFA5FB49D6BAE08BFA67" box="[1167,1227,1442,1468]" pageId="59" pageNumber="295">XVII</collectionCode>
|
||
,
|
||
<collectionCode id="7BA0AE39FFF8FFA5FB1FD6BAE160FA67" box="[1241,1312,1442,1468]" pageId="59" pageNumber="295">XVIII</collectionCode>
|
||
,
|
||
<collectionCode id="7BA0AE39FFF8FFA5FAEBD6BAE120FA67" box="[1325,1376,1442,1468]" pageId="59" pageNumber="295">XIX</collectionCode>
|
||
–
|
||
<collectionCode id="7BA0AE39FFF8FFA5FAA9D6BAE1D8FA67" box="[1391,1432,1442,1468]" pageId="59" pageNumber="295">XX</collectionCode>
|
||
,
|
||
<collectionCode id="7BA0AE39FFF8FFA5FF51D6DEE48AFA3B" box="[151,202,1478,1504]" pageId="59" pageNumber="295">XXI</collectionCode>
|
||
–
|
||
<collectionCode id="7BA0AE39FFF8FFA5FF1FD6DEE555FA3B" box="[217,277,1478,1504]" pageId="59" pageNumber="295">XXII</collectionCode>
|
||
,
|
||
<collectionCode id="7BA0AE39FFF8FFA5FEE5D6DEE52AFA3B" box="[291,362,1478,1504]" pageId="59" pageNumber="295">XXIII</collectionCode>
|
||
,
|
||
<collectionCode id="7BA0AE39FFF8FFA5FEBED6DEE5FEFA3B" box="[376,446,1478,1504]" pageId="59" pageNumber="295">XXIV</collectionCode>
|
||
–
|
||
<collectionCode id="7BA0AE39FFF8FFA5FE0DD6DEE647FA3B" box="[459,519,1478,1504]" pageId="59" pageNumber="295">XXV</collectionCode>
|
||
,
|
||
<collectionCode id="7BA0AE39FFF8FFA5FDD3D6DEE61CFA3B" box="[533,604,1478,1504]" pageId="59" pageNumber="295">XXVI</collectionCode>
|
||
–
|
||
<collectionCode id="7BA0AE39FFF8FFA5FDADD6DEE685FA3B" box="[619,709,1478,1504]" pageId="59" pageNumber="295">XXVIII</collectionCode>
|
||
. Antenna not sexually dimorphic; proximal endopodal segment largely separated from basis, with pinnate seta on abexopodal margin; exopod 2-segmented, proximal segment with two setae, distal segment with two lateral and three apical elements; distal endopodal segment without penicillate elements. Mandible with three elements on basis; endopod with eight setae; exopod 1-segmented, with two setae. Maxillulary coxal epipodite represented by one seta. Maxilla with three endites on syncoxa; endopod discrete, with three setae. Maxilliped with two setae on syncoxa; basis with two setae and longitudinal spinular row on palmar margin; endopod represented by conspicuously serrated claw, accompanied by three accessory setae
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1D0E36FCFFF8FFA5FF01D5D9E612F80C" blockId="59.[151,1437,469,2008]" pageId="59" pageNumber="295">
|
||
P1 inner basal spine not sexually dimorphic. P1 exopod 3-segmented; exp-2 much longer than others, nearly three times as long as exp-1 and about half as long as enp-1, with short inner seta; exp-3 small, with two pinnate spines, one geniculate seta and one naked seta. P1 endopod 2-segmented; enp-1 elongate, about 1.5 times length of exopod, with short, proximally inserted, pinnate inner seta, segment margins without area of reduced chitinization; enp-2 very small, with one naked minute seta and two dentate claws. P2–P4 rami 3-segmented; inner seta of P2–P3 exp-1 reduced in size, absent in P4. P2–P3 endopods
|
||
<specimenCount id="0BB7FD75FFF8FFA5FD39D46EE754F84B" box="[767,788,1910,1936]" pageId="59" pageNumber="295" type="male">♂</specimenCount>
|
||
3-segmented. P2 slightly more robust in
|
||
<specimenCount id="0BB7FD75FFF8FFA5FB36D46EE148F84B" box="[1264,1288,1910,1936]" pageId="59" pageNumber="295" type="male">♂</specimenCount>
|
||
. P3 endopod
|
||
<specimenCount id="0BB7FD75FFF8FFA5FF51D482E4ECF86F" box="[151,172,1946,1972]" pageId="59" pageNumber="295" type="male">♂</specimenCount>
|
||
without inner setae (
|
||
<emphasis id="2FC5EAEEFFF8FFA5FE5BD482E5F4F868" box="[413,436,1946,1971]" italics="true" pageId="59" pageNumber="295">vs</emphasis>
|
||
two in
|
||
<specimenCount id="0BB7FD75FFF8FFA5FDCBD482E660F86F" box="[525,544,1946,1972]" pageId="59" pageNumber="295" type="female">♀</specimenCount>
|
||
) on enp-3 and outer spine modified into apically truncated spinous apophysis. Armature formula of P2–P4 as follows:
|
||
</paragraph>
|
||
<paragraph id="1D0E36FCFFFFFFA2FD8BD3BAE00CFEED" pageId="60" pageNumber="296">
|
||
<table id="6FB1C45CFFFF0061FEA6D3BAE157FEED" box="[352,1303,162,310]" gridcols="3" gridrows="4" pageId="60" pageNumber="296" rowsContinueFrom="58.[352,1347,588,736]" rowsContinueIn="61.[352,1403,1843,1992]">
|
||
<tr id="A38134BEFFFF0061FEA6D3BAE157FF67" box="[352,1303,162,188]" gridrow="0" pageId="60" pageNumber="296" rowspan-0="1">
|
||
<th id="E0505DC2FFFF0061FD8BD3BAE6E2FF67" box="[589,674,162,188]" gridcol="1" gridrow="0" pageId="60" pageNumber="296">exopod</th>
|
||
<th id="E0505DC2FFFF0061FC3FD3BAE157FF67" box="[1017,1303,162,188]" gridcol="2" gridrow="0" pageId="60" pageNumber="296">endopod</th>
|
||
</tr>
|
||
<tr id="A38134BEFFFF0061FEA6D3D3E157FF3E" box="[352,1303,203,229]" gridrow="1" pageId="60" pageNumber="296">
|
||
<th id="E0505DC2FFFF0061FEA6D3D3E53AFF3E" box="[352,378,203,229]" gridcol="0" gridrow="1" pageId="60" pageNumber="296">P2</th>
|
||
<td id="E0505DC2FFFF0061FD8BD3D3E6E2FF3E" box="[589,674,203,229]" gridcol="1" gridrow="1" pageId="60" pageNumber="296">1.1.123</td>
|
||
<td id="E0505DC2FFFF0061FC3FD3D3E157FF3E" box="[1017,1303,203,229]" gridcol="2" gridrow="1" pageId="60" pageNumber="296">1.1.121</td>
|
||
</tr>
|
||
<tr id="A38134BEFFFF0061FEA6D3EBE157FED5" box="[352,1303,243,270]" gridrow="2" pageId="60" pageNumber="296">
|
||
<th id="E0505DC2FFFF0061FEA6D3EBE53AFED5" box="[352,378,243,270]" gridcol="0" gridrow="2" pageId="60" pageNumber="296">P3</th>
|
||
<td id="E0505DC2FFFF0061FD8BD3EBE6E2FED5" box="[589,674,243,270]" gridcol="1" gridrow="2" pageId="60" pageNumber="296">1.1.323</td>
|
||
<td id="E0505DC2FFFF0061FC3FD3EBE157FED5" box="[1017,1303,243,270]" gridcol="2" gridrow="2" pageId="60" pageNumber="296">1.1.221 [♂: 1.1.02 + apo]</td>
|
||
</tr>
|
||
<tr id="A38134BEFFFF0061FEA6D204E157FEED" box="[352,1303,284,310]" gridrow="3" pageId="60" pageNumber="296">
|
||
<th id="E0505DC2FFFF0061FEA6D204E53AFEED" box="[352,378,284,310]" gridcol="0" gridrow="3" pageId="60" pageNumber="296">P4</th>
|
||
<td id="E0505DC2FFFF0061FD8BD204E6E2FEED" box="[589,674,284,310]" gridcol="1" gridrow="3" pageId="60" pageNumber="296">0.1.323</td>
|
||
<td id="E0505DC2FFFF0061FC3FD204E157FEED" box="[1017,1303,284,310]" gridcol="2" gridrow="3" pageId="60" pageNumber="296">1.1.121</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="1D0E36FCFFFFFFA2FF01D27DE754FE30" blockId="60.[151,1437,357,672]" pageId="60" pageNumber="296">
|
||
P
|
||
<specimenCount id="0BB7FD75FFFFFFA2FF1ED27DE545FEA4" box="[216,261,357,384]" pageId="60" pageNumber="296" type="female">5 ♀</specimenCount>
|
||
endopodal lobe with five setae, outermost well developed, innermost shortest; inner margin without transverse striae. P
|
||
<specimenCount id="0BB7FD75FFFFFFA2FEB7D291E5DDFE78" box="[369,413,393,420]" pageId="60" pageNumber="296" type="female">5 ♀</specimenCount>
|
||
exopod elongate, with six elements. P
|
||
<specimenCount id="0BB7FD75FFFFFFA2FC98D291E7CAFE78" box="[862,906,393,420]" pageId="60" pageNumber="296" type="male">5 ♂</specimenCount>
|
||
endopodal lobe with two elements, outer one shortest; exopod 2-segmented, with one long outer seta on exp-1 and four elements on exp-2. Vestigial P
|
||
<specimenCount id="0BB7FD75FFFFFFA2FAA9D2B5E1DCFE1C" box="[1391,1436,429,455]" pageId="60" pageNumber="296" type="female">6 ♀</specimenCount>
|
||
represented by three minute setae. P
|
||
<specimenCount id="0BB7FD75FFFFFFA2FDEAD2C9E616FE30" box="[556,598,465,491]" pageId="60" pageNumber="296" type="male">6 ♂</specimenCount>
|
||
with three setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55AB6577FFFFFFA2FF01D2EDE7E1FD8C" pageId="60" pageNumber="296" type="etymology">
|
||
<paragraph id="1D0E36FCFFFFFFA2FF01D2EDE7E1FD8C" blockId="60.[151,1437,357,672]" pageId="60" pageNumber="296">
|
||
<emphasis id="2FC5EAEEFFFFFFA2FF01D2EDE50EFDD4" bold="true" box="[199,334,501,527]" pageId="60" pageNumber="296">Etymology.</emphasis>
|
||
The genus name is dedicated to Prof. Süphan Karaytuğ (
|
||
<collectingRegion id="DF75F81EFFFFFFA2FC04D2EDE053FDD4" box="[962,1043,501,527]" country="Turkey" name="Mersin" pageId="60" pageNumber="296">Mersin</collectingRegion>
|
||
University,
|
||
<collectingCountry id="65A6766CFFFFFFA2FB5DD2EDE0AFFDCB" box="[1179,1263,501,528]" name="Turkey" pageId="60" pageNumber="296">Turkey</collectingCountry>
|
||
), in recognition of his contributions to the harpacticoid fauna of
|
||
<collectingCountry id="65A6766CFFFFFFA2FD1CD101E76BFDE8" box="[730,811,537,563]" name="Turkey" pageId="60" pageNumber="296">Turkey</collectingCountry>
|
||
, and under whose mentorship the next generation of harpacticoid experts in his home country emerged. Gender: feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55AB6577FFFFFFA2FF01D179E690FD7B" pageId="60" pageNumber="296" type="type_taxon">
|
||
<paragraph id="1D0E36FCFFFFFFA2FF01D179E690FD7B" blockId="60.[151,1437,357,672]" pageId="60" pageNumber="296">
|
||
<emphasis id="2FC5EAEEFFFFFFA2FF01D179E591FDA0" bold="true" box="[199,465,609,635]" pageId="60" pageNumber="296">
|
||
<typeStatus id="C20A885EFFFFFFA2FF01D179E542FDA0" box="[199,258,609,635]" pageId="60" pageNumber="296">Type</typeStatus>
|
||
and only species.
|
||
</emphasis>
|
||
<taxonomicName id="DAB14D7FFFFFFFA2FE1ED179E795FDA7" authority="Kuru & Karaytug, 2015" authorityName="Kuru & Karaytug" authorityYear="2015" baseAuthorityName="Kuru & Karaytug" baseAuthorityYear="2015" box="[472,981,609,636]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="296" phylum="Arthropoda" rank="species" species="aydini">
|
||
<emphasis id="2FC5EAEEFFFFFFA2FE1ED179E682FDA0" box="[472,706,609,635]" italics="true" pageId="60" pageNumber="296">Parastenhelia aydini</emphasis>
|
||
<bibRefCitation id="79204B0DFFFFFFA2FD0FD179E795FDA7" author="Kuru, S. & Karaytug, S." box="[713,981,609,636]" pageId="60" pageNumber="296" pagination="121 - 127" refId="ref75081" refString="Kuru, S. & Karaytug, S. (2015) A new species of Parastenhelia Thompson & A. Scott, 1903 (Copepoda, Harpacticoida, Parastenheliidae) from Turkey. Biharean Biologist, 9, 121 - 127." type="journal article" year="2015">Kuru & Karaytuğ, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
=
|
||
<taxonomicName id="DAB14D7FFFFFFFA2FC35D17AE499FD7B" authority="(Kuru & Karaytug, 2015)" baseAuthorityName="Kuru & Karaytug" baseAuthorityYear="2015" class="Hexanauplia" family="Parastenheliidae" genus="Karaytugia" kingdom="Animalia" order="Harpacticoida" pageId="60" pageNumber="296" phylum="Arthropoda" rank="species" species="aydini" status="comb. nov.">
|
||
<emphasis id="2FC5EAEEFFFFFFA2FC35D17AE0FFFDA0" box="[1011,1215,609,635]" italics="true" pageId="60" pageNumber="296">Karaytugia aydini</emphasis>
|
||
(
|
||
<bibRefCitation id="79204B0DFFFFFFA2FB08D179E491FD7B" author="Kuru, S. & Karaytug, S." pageId="60" pageNumber="296" pagination="121 - 127" refId="ref75081" refString="Kuru, S. & Karaytug, S. (2015) A new species of Parastenhelia Thompson & A. Scott, 1903 (Copepoda, Harpacticoida, Parastenheliidae) from Turkey. Biharean Biologist, 9, 121 - 127." type="journal article" year="2015">Kuru & Karaytuğ, 2015</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="2FC5EAEEFFFFFFA2FF26D19DE520FD44" bold="true" box="[224,352,645,671]" pageId="60" pageNumber="296">
|
||
<taxonomicNameLabel id="34F65795FFFFFFA2FF26D19DE520FD44" box="[224,352,645,671]" pageId="60" pageNumber="296" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(by original designation herein).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |