219 lines
33 KiB
XML
219 lines
33 KiB
XML
<document id="59736501515C221DCFDEE8F869E6448C" ID-CLB-Dataset="8250" ID-DOI="10.11646/zootaxa.4860.4.2" ID-GBIF-Dataset="7fc75baa-2e48-490b-9a6e-256378b4531e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4414254" ID-ZooBank="C5095CD0-1DF5-4BE9-9B31-0CDB9443CF7F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609706152222" checkinUser="plazi" docAuthor="Lambkin, Kevin J." docDate="2020" docId="231AE47EFF85FFFF63F2FF2A2308F968" docLanguage="en" docName="zootaxa.4860.4.2.pdf" docOrigin="Zootaxa 4860 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ipsviciidae Tillyard 1919" docType="treatment" docVersion="10" lastPageNumber="506" masterDocId="DF239C06FF86FFFC6365FFBD2142FFF5" masterDocTitle="Revision of the Ipsviciidae of the Late Triassic of Queensland (Hemiptera Cicadomorpha: Scytinopteroidea)" masterLastPageNumber="520" masterPageNumber="503" pageNumber="506" updateTime="1698852020800" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="97AD301BE2905885173DAB6A91801250">Revision of the Ipsviciidae of the Late Triassic of Queensland (Hemiptera Cicadomorpha: Scytinopteroidea)</mods:title>
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<mods:namePart id="E4F0DD01953AE5C7709D8E7480F7171C">Lambkin, Kevin J.</mods:namePart>
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<mods:title id="789D293724809F26991EC9CFC93B6A47">Zootaxa</mods:title>
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<mods:part id="860FFE6E015390691B8FA266E1690A86">
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<mods:date id="E0274122208CA6858C408AC56BF9629B">2020</mods:date>
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<mods:number id="4AF8BB7C13B8E0D55F447C071799C8D6">2020-10-14</mods:number>
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<mods:number id="BE24853A27EB8E7A128F16DA12B82637">4860</mods:number>
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<mods:identifier id="CE8D149037DEC5DAC007BEF26BBF540D" type="DOI">10.11646/zootaxa.4860.4.2</mods:identifier>
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<treatment id="231AE47EFF85FFFF63F2FF2A2308F968" ID-DOI="http://doi.org/10.5281/zenodo.4537869" ID-GBIF-Taxon="177323946" ID-Zenodo-Dep="4537869" LSID="urn:lsid:plazi:treatment:231AE47EFF85FFFF63F2FF2A2308F968" httpUri="http://treatment.plazi.org/id/231AE47EFF85FFFF63F2FF2A2308F968" lastPageNumber="506" pageId="3" pageNumber="506">
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<subSubSection id="E3A906E3FF85FFFF63F2FF2A2376FF44" box="[151,564,151,177]" pageId="3" pageNumber="506" type="nomenclature">
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<paragraph id="AB0C5568FF85FFFF63F2FF2A2376FF44" blockId="3.[151,564,151,177]" box="[151,564,151,177]" pageId="3" pageNumber="506">
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<heading id="F044E204FF85FFFF63F2FF2A2376FF44" bold="true" box="[151,564,151,177]" fontSize="11" level="1" pageId="3" pageNumber="506" reason="1">
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<emphasis id="99C7897AFF85FFFF63F2FF2A2376FF44" bold="true" box="[151,564,151,177]" pageId="3" pageNumber="506">
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Family
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<taxonomicName id="6CB32EEBFF85FFFF6392FF2A2376FF44" authority="Tillyard, 1919" authorityName="Tillyard" authorityYear="1919" box="[247,564,151,177]" class="Insecta" family="Ipsviciidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="family">Ipsviciidae Tillyard, 1919</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E3A906E3FF85FFFF63F2FF612270FEE7" pageId="3" pageNumber="506" type="reference_group">
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<paragraph id="AB0C5568FF85FFFF63F2FF612242FF01" blockId="3.[151,818,219,397]" box="[151,768,219,244]" pageId="3" pageNumber="506">
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<taxonomicName id="6CB32EEBFF85FFFF63F2FF612090FF06" authority="Tillyard, 1919: 866" authorityName="Tillyard" authorityPageNumber="866" authorityYear="1919" box="[151,466,219,244]" class="Insecta" family="Ipsviciidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="family">Ipsviciidae Tillyard, 1919: 866</taxonomicName>
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, 878, 860, 862, 865, 891, 892
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</paragraph>
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<paragraph id="AB0C5568FF85FFFF63F2FF472270FEE7" blockId="3.[151,818,219,397]" box="[151,818,250,274]" pageId="3" pageNumber="506">
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Willcoxiidae: Tillyard, 1919: 865 (
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<emphasis id="99C7897AFF85FFFF629DFF4623C5FEE7" box="[504,647,250,274]" italics="true" pageId="3" pageNumber="506">nomen nudum</emphasis>
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) (
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<emphasis id="99C7897AFF85FFFF61FBFF472268FEE7" box="[670,810,250,274]" italics="true" pageId="3" pageNumber="506">lapsus calami</emphasis>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="E3A906E3FF85FFFF63F2FEA42353FE9B" pageId="3" pageNumber="506" type="description">
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<paragraph id="AB0C5568FF85FFFF63F2FEA423F5FEC4" blockId="3.[151,818,219,397]" box="[151,695,281,305]" pageId="3" pageNumber="506">
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Ipswicidae (
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<emphasis id="99C7897AFF85FFFF6271FEA72072FEC4" box="[276,304,282,305]" italics="true" pageId="3" pageNumber="506">sic</emphasis>
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):
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<bibRefCitation id="CF222899FF85FFFF6223FEA420B8FEC4" author="Jeannel, R." box="[326,506,281,305]" pageId="3" pageNumber="506" pagination="1 - 101" refId="ref7992" refString="Jeannel, R. (1947) Paleontologie et peuplement de la Terre. Introduction a l'entomologie, 1 (3), 1 - 101, 14 pls." type="journal article" year="1947">Jeannel, 1947: 72</bibRefCitation>
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;
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<bibRefCitation id="CF222899FF85FFFF6160FEA423F5FEC4" author="Jeannel, R." box="[517,695,281,305]" pageId="3" pageNumber="506" refId="ref8105" refString="Jeannel, R. (1979) Paleontologie et peuplement de la Terre. Deuxieme edition. Societe nouvelle des editions Boubee & Cie, Paris, 101 pp., 14 pls." type="book" year="1979">Jeannel, 1979: 72</bibRefCitation>
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</paragraph>
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<paragraph id="AB0C5568FF85FFFF63F2FE8520BCFEA5" blockId="3.[151,818,219,397]" box="[151,510,312,336]" pageId="3" pageNumber="506">
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Ipswiciidae (
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<emphasis id="99C7897AFF85FFFF627EFE852075FEBA" box="[283,311,312,335]" italics="true" pageId="3" pageNumber="506">sic</emphasis>
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):
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<bibRefCitation id="CF222899FF85FFFF6228FE8520BCFEA5" author="Jeannel, R." box="[333,510,312,336]" pageId="3" pageNumber="506" pagination="1 - 110" refId="ref8022" refString="Jeannel, R. (1949) Les insectes. Classification et phylogenie. Les insectes fossiles. In: Grasse, P. - P. (Ed.), Traite de zoologie. Insectes (Paleontologie, geonemie, insectes inferieurs et Coleopteres). 9. Masson et Cie, Paris, pp. 1 - 110." type="book chapter" year="1949">Jeannel, 1949: 82</bibRefCitation>
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</paragraph>
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<paragraph id="AB0C5568FF85FFFF63F2FEEB2353FE9B" blockId="3.[151,818,219,397]" box="[151,529,342,366]" pageId="3" pageNumber="506">
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Ipswichidae (
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<emphasis id="99C7897AFF85FFFF6245FEEA207EFE9B" box="[288,316,343,366]" italics="true" pageId="3" pageNumber="506">sic</emphasis>
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):
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<bibRefCitation id="CF222899FF85FFFF6237FEEB2353FE9B" author="Jeannel, R." box="[338,529,342,366]" pageId="3" pageNumber="506" refId="ref8084" refString="Jeannel, R. (1960) Introduction to entomology. Hutchison & Co., London, 344 pp." type="book" year="1960">Jeannel, 1960: 309</bibRefCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="E3A906E3FF85FFFF63F2FEC823EDFE78" box="[151,687,373,397]" pageId="3" pageNumber="506" type="reference_group">
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<paragraph id="AB0C5568FF85FFFF63F2FEC823EDFE78" blockId="3.[151,818,219,397]" box="[151,687,373,397]" pageId="3" pageNumber="506">
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<treatmentCitationGroup id="8BA37246FF85FFFF63F2FEC823EDFE78" box="[151,687,373,397]" pageId="3" pageNumber="506">
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<taxonomicName id="6CB32EEBFF85FFFF63F2FEC820DFFE78" authority="Tillyard, 1920" authorityName="Tillyard" authorityYear="1920" box="[151,413,373,397]" class="Insecta" family="Ipsviciidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="family">
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Ipsviciidae
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<bibRefCitation id="CF222899FF85FFFF6269FEC820DFFE78" author="Tillyard, R. J." box="[268,413,373,397]" pageId="3" pageNumber="506" refId="ref9717" refString="Tillyard, R. J. (1920) [Letter to Benjamin Dunstan, 19 March 1920], as above." type="book" year="1920">Tillyard, 1920</bibRefCitation>
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</taxonomicName>
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(
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<emphasis id="99C7897AFF85FFFF62CEFECB2085FE78" box="[427,455,374,397]" italics="true" pageId="3" pageNumber="506">sic</emphasis>
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||
):
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<treatmentCitation id="2A127379FF85FFFF62B8FEC823EDFE78" author="Fujiyama, I." box="[477,687,373,397]" page="372" pageId="3" pageNumber="506" year="1973">
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<bibRefCitation id="CF222899FF85FFFF62B8FEC823EDFE78" author="Fujiyama, I." box="[477,687,373,397]" pageId="3" pageNumber="506" pagination="331 - 386" refId="ref7660" refString="Fujiyama, I. (1973) Mesozoic insect fauna of east Asia Part I. Introduction and Upper Triassic faunas. Bulletin of the National Science Museum of Tokyo 16, 331 - 386." type="journal article" year="1973">Fujiyama, 1973: 372</bibRefCitation>
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="E3A906E3FF85FFFF63F2FE062556FCEF" pageId="3" pageNumber="506" type="diagnosis">
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<paragraph id="AB0C5568FF85FFFF63F2FE062556FCEF" blockId="3.[151,1437,443,1693]" pageId="3" pageNumber="506">
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<emphasis id="99C7897AFF85FFFF63F2FE0620CAFE20" bold="true" box="[151,392,443,469]" pageId="3" pageNumber="506">Emended diagnosis.</emphasis>
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Tegmen tuberculate/punctate, with sculpture quite coarse basally merging to slightly finer apically; costal margin convex, apex narrowly rounded, posterior margin straight; well-developed marginal membrane, somewhat widened postero-apically; without a hypocostal pit, lower carina of the basal costal space, or nodal groove; bSc variably weakly to strongly convex; costal fracture forwardly curved, arising from apex of basal cell or from extreme base of R; dSc a long, shallow groove running along a broad costal space from near base of R just beyond costal fracture and petering out at around level of
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<emphasis id="99C7897AFF85FFFF6079FDD22210FD7C" box="[796,850,623,649]" italics="true" pageId="3" pageNumber="506">1r-m</emphasis>
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, crossed by 4–6 costal veinlets; basal cell triangular, closed by vein fusion at a single point; R with numerous variable terminal branches extending to tegmen apex; M and CuA fused for a variable length beyond basal cell; M not strongly bent towards CuA at nodal level, with 3 or 4 branches terminating on postero-apical margin; apical venation very variable; at least two
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<emphasis id="99C7897AFF85FFFF67EFFD6125F0FD00" box="[1162,1202,732,757]" italics="true" pageId="3" pageNumber="506">r-m</emphasis>
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; CuA simple; clavus occupying no more than 0.7 tegmen length; space behind PCu+1A very narrow.
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</paragraph>
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</subSubSection>
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<subSubSection id="E3A906E3FF85FFFF63A2FC9E2308F968" pageId="3" pageNumber="506" type="description">
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<paragraph id="AB0C5568FF85FFFF63A2FC9E2384FBA8" blockId="3.[151,1437,443,1693]" pageId="3" pageNumber="506">
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<emphasis id="99C7897AFF85FFFF63A2FC9E201BFCC8" bold="true" box="[199,345,803,829]" pageId="3" pageNumber="506">Description.</emphasis>
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Head apparently small, ovate, dorsal surface tuberculate to irregularly rugose (
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<figureCitation id="338849EDFF85FFFF67DCFC9E244BFCC8" box="[1209,1289,803,829]" captionStart="FIGURES 36–39" captionStartId="12.[151,264,1944,1968]" captionTargetBox="[157,1427,181,1920]" captionTargetId="figure-25@12.[157,1431,181,1920]" captionTargetPageId="12" captionText="FIGURES 36–39. 36–38, Ipsviciopsis elegans Tillyard, body specimens: 36, QMF60203; 37, QMF60202; 38, QMF60199a; 39, possible ipsviciid nymph, QMF60198." figureDoi="http://doi.org/10.5281/zenodo.4414268" httpUri="https://zenodo.org/record/4414268/files/figure.png" pageId="3" pageNumber="506">Fig. 36</figureCitation>
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, and possible ipsviciid nymph—Fig. 39), rostrum (in possible ipsviciid nymph—Fig. 39) very long, extending beyond abdominal sternite 5. Pronotum two times wider than long, expanded antero-laterally as a pair of paranotal-like lobes, anterior and posterior margins more or less straight, the whole densely tuberculate to irregularly rugose, with a pair of small oval areas antero-medially somewhat smoother (
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<figureCitation id="338849EDFF85FFFF61D9FC0E2273FC38" box="[700,817,947,973]" captionStart="FIGURES 7–10" captionStartId="7.[151,264,2009,2033]" captionTargetBox="[220,1366,181,1985]" captionTargetId="figure-15@7.[219,1368,181,1985]" captionTargetPageId="7" captionText="FIGURES 7–10. Ipsvicia jonesi Tillyard, body specimens: 7, QMF60157a; 8, QMF60158; 9, QMF60159; 10, ACCI.69." figureDoi="http://doi.org/10.5281/zenodo.4414260" httpUri="https://zenodo.org/record/4414260/files/figure.png" pageId="3" pageNumber="506">Figs 8, 10</figureCitation>
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,
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<figureCitation id="338849EDFF85FFFF605BFC0E221FFC38" box="[830,861,947,973]" captionStart="FIGURES 36–39" captionStartId="12.[151,264,1944,1968]" captionTargetBox="[157,1427,181,1920]" captionTargetId="figure-25@12.[157,1431,181,1920]" captionTargetPageId="12" captionText="FIGURES 36–39. 36–38, Ipsviciopsis elegans Tillyard, body specimens: 36, QMF60203; 37, QMF60202; 38, QMF60199a; 39, possible ipsviciid nymph, QMF60198." figureDoi="http://doi.org/10.5281/zenodo.4414268" httpUri="https://zenodo.org/record/4414268/files/figure.png" pageId="3" pageNumber="506">37</figureCitation>
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); pronotum completely covering mesoscutum with only mesoscutellum exposed (
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<figureCitation id="338849EDFF85FFFF628BFC6A2306FC07" box="[494,580,983,1010]" captionStart="FIGURES 36–39" captionStartId="12.[151,264,1944,1968]" captionTargetBox="[157,1427,181,1920]" captionTargetId="figure-25@12.[157,1431,181,1920]" captionTargetPageId="12" captionText="FIGURES 36–39. 36–38, Ipsviciopsis elegans Tillyard, body specimens: 36, QMF60203; 37, QMF60202; 38, QMF60199a; 39, possible ipsviciid nymph, QMF60198." figureDoi="http://doi.org/10.5281/zenodo.4414268" httpUri="https://zenodo.org/record/4414268/files/figure.png" pageId="3" pageNumber="506">Fig. 36</figureCitation>
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); mesoscutum smooth with a pair of oblique ridges converging posteriorly; mesoscutellum very small, coarsely tuberculate with a small rugose area subapically and a pair of small, smoother depressed areas anteriorly (
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<figureCitation id="338849EDFF85FFFF62AFFBA22351FBCF" box="[458,531,1055,1082]" captionStart="FIGURES 7–10" captionStartId="7.[151,264,2009,2033]" captionTargetBox="[220,1366,181,1985]" captionTargetId="figure-15@7.[219,1368,181,1985]" captionTargetPageId="7" captionText="FIGURES 7–10. Ipsvicia jonesi Tillyard, body specimens: 7, QMF60157a; 8, QMF60158; 9, QMF60159; 10, ACCI.69." figureDoi="http://doi.org/10.5281/zenodo.4414260" httpUri="https://zenodo.org/record/4414260/files/figure.png" pageId="3" pageNumber="506">Figs 7</figureCitation>
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,
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<figureCitation id="338849EDFF85FFFF617AFBA2237DFBCC" box="[543,575,1055,1081]" captionStart="FIGURES 36–39" captionStartId="12.[151,264,1944,1968]" captionTargetBox="[157,1427,181,1920]" captionTargetId="figure-25@12.[157,1431,181,1920]" captionTargetPageId="12" captionText="FIGURES 36–39. 36–38, Ipsviciopsis elegans Tillyard, body specimens: 36, QMF60203; 37, QMF60202; 38, QMF60199a; 39, possible ipsviciid nymph, QMF60198." figureDoi="http://doi.org/10.5281/zenodo.4414268" httpUri="https://zenodo.org/record/4414268/files/figure.png" pageId="3" pageNumber="506">37</figureCitation>
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). Abdominal tergites smooth, spiracle-bearing pleurites finely punctate (
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<figureCitation id="338849EDFF85FFFF660FFBA221E5FBAB" captionStart="FIGURES 7–10" captionStartId="7.[151,264,2009,2033]" captionTargetBox="[220,1366,181,1985]" captionTargetId="figure-15@7.[219,1368,181,1985]" captionTargetPageId="7" captionText="FIGURES 7–10. Ipsvicia jonesi Tillyard, body specimens: 7, QMF60157a; 8, QMF60158; 9, QMF60159; 10, ACCI.69." figureDoi="http://doi.org/10.5281/zenodo.4414260" httpUri="https://zenodo.org/record/4414260/files/figure.png" pageId="3" pageNumber="506">Figs 7</figureCitation>
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,
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<figureCitation id="338849EDFF85FFFF63D6FBFE2190FBA8" box="[179,210,1091,1117]" captionStart="FIGURES 36–39" captionStartId="12.[151,264,1944,1968]" captionTargetBox="[157,1427,181,1920]" captionTargetId="figure-25@12.[157,1431,181,1920]" captionTargetPageId="12" captionText="FIGURES 36–39. 36–38, Ipsviciopsis elegans Tillyard, body specimens: 36, QMF60203; 37, QMF60202; 38, QMF60199a; 39, possible ipsviciid nymph, QMF60198." figureDoi="http://doi.org/10.5281/zenodo.4414268" httpUri="https://zenodo.org/record/4414268/files/figure.png" pageId="3" pageNumber="506">37</figureCitation>
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), female with siphonate ovipositor (
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<figureCitation id="338849EDFF85FFFF6103FBFE23F8FBA8" box="[614,698,1091,1117]" captionStart="FIGURES 36–39" captionStartId="12.[151,264,1944,1968]" captionTargetBox="[157,1427,181,1920]" captionTargetId="figure-25@12.[157,1431,181,1920]" captionTargetPageId="12" captionText="FIGURES 36–39. 36–38, Ipsviciopsis elegans Tillyard, body specimens: 36, QMF60203; 37, QMF60202; 38, QMF60199a; 39, possible ipsviciid nymph, QMF60198." figureDoi="http://doi.org/10.5281/zenodo.4414268" httpUri="https://zenodo.org/record/4414268/files/figure.png" pageId="3" pageNumber="506">Fig. 38</figureCitation>
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).
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</paragraph>
|
||
<paragraph id="AB0C5568FF85FFFF63A2FBDA2308F968" blockId="3.[151,1437,443,1693]" pageId="3" pageNumber="506">
|
||
<emphasis id="99C7897AFF85FFFF63A2FBDA2053FB74" bold="true" box="[199,273,1127,1153]" pageId="3" pageNumber="506">Notes.</emphasis>
|
||
The above diagnosis is a listing of the tegmen character states of
|
||
<taxonomicName id="6CB32EEBFF85FFFF6085FBD5257BFB74" authorityName="Tillyard" authorityYear="1919" box="[992,1081,1128,1153]" class="Insecta" family="Ipsviciidae" genus="Ipsvicia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="99C7897AFF85FFFF6085FBD5257BFB74" box="[992,1081,1128,1153]" italics="true" pageId="3" pageNumber="506">Ipsvicia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6CB32EEBFF85FFFF6709FBD525B2FB74" authorityName="Tillyard" authorityYear="1922" box="[1132,1264,1128,1153]" class="Insecta" family="Ipsviciidae" genus="Ipsviciopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="99C7897AFF85FFFF6709FBD525B2FB74" box="[1132,1264,1128,1153]" italics="true" pageId="3" pageNumber="506">Ipsviciopsis</emphasis>
|
||
</taxonomicName>
|
||
compared with the diagnostic characters recorded by
|
||
<bibRefCitation id="CF222899FF85FFFF615AFB36225EFB50" author="Shcherbakov, D. E." box="[575,796,1163,1189]" pageId="3" pageNumber="506" pagination="89 - 101" refId="ref9055" refString="Shcherbakov, D. E. (1984) Sistema i filogeniya permskikh Cicadomorpha (Cimicida, Cicadina). Palaeontologicheskii Zhurnal, 18 (2), 87 - 97. [in Russian, publishjed in English as: Shcherbakov, D. E. (1984) A system and the phylogeny of Permian Cicadomorpha (Cimicida, Cicadina). Paleontological Zhurnal, 2, 89 - 101.]" type="journal article" year="1984">Shcherbakov (1984</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="CF222899FF85FFFF6042FB362222FB50" author="Shcherbakov, D. E." box="[807,864,1163,1189]" pageId="3" pageNumber="506" pagination="1 - 26" refId="ref9241" refString="Shcherbakov, D. E. (2011) New and little-known families of Hemiptera Cicadomorpha from the Triassic of Central Asia - early analogs of treehoppers and planthoppers. Zootaxa, 2836 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 2836.1.1" type="journal article" year="2011">2011</bibRefCitation>
|
||
) for the other families of Scytinopteroidea—Scyti-nopteridae,
|
||
<taxonomicName id="6CB32EEBFF85FFFF627AFB122094FB3C" baseAuthorityName="Shcherbakov" baseAuthorityYear="1984" box="[287,470,1199,1225]" class="Insecta" family="Serpentivenidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="family">Serpentivenidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="6CB32EEBFF85FFFF6283FB1223C3FB3C" authorityName="Evans" authorityYear="1956" box="[486,641,1199,1225]" class="Insecta" family="Stenoviciidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="family">Stenoviciidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="6CB32EEBFF85FFFF61F5FB122202FB3C" authorityName="Evans" authorityYear="1950" box="[656,832,1199,1225]" class="Insecta" family="Paraknightiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="family">Paraknightiidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="6CB32EEBFF85FFFF601BFB122573FB3C" authorityName="Brauckmann, Martins-Neto and Gallego" authorityYear="2006" box="[894,1073,1199,1225]" class="Insecta" family="Saaloscytinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="family">Saaloscytinidae</taxonomicName>
|
||
, as illustrated by
|
||
<bibRefCitation id="CF222899FF85FFFF6661FB0D24D5FB3C" author="Evans, J. W." box="[1284,1431,1199,1226]" pageId="3" pageNumber="506" pagination="180 - 198" refId="ref7341" refString="Evans, J. W. (1943) Upper Permian Homoptera from New South Wales. Records of the Australian Museum, 21, 180 - 198. https: // doi. org / 10.3853 / j. 0067 - 1975.21.1943.534" type="journal article" year="1943">Evans (1943</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="CF222899FF85FFFF63F2FB6E2190FB1B" author="Evans, J. W." box="[151,210,1235,1262]" pageId="3" pageNumber="506" pagination="246 - 250" refId="ref7435" refString="Evans, J. W. (1950) A re-examination of an Upper Permian insect, Paraknightia magnifica Ev. Records of the Australian Museum, 22, 246 - 250. https: // doi. org / 10.3853 / j. 0067 - 1975.22.1950.605" type="journal article" year="1950">1950</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="CF222899FF85FFFF6384FB6E205EFB1B" author="Evans, J. W." box="[225,284,1235,1262]" pageId="3" pageNumber="506" pagination="17 - 23" refId="ref7521" refString="Evans, J. W. (1963) The systematic position of the Ipsviciidae (Upper Triassic Hemiptera) and some new Upper Permian and Middle Triassic Homoptera from Australia (Insecta). Journal of the Entomological Society of Queensland, 2, 17 - 23. https: // doi. org / 10.1111 / j. 1440 - 6055.1963. tb 00384. x" type="journal article" year="1963">1963</bibRefCitation>
|
||
),
|
||
<bibRefCitation id="CF222899FF85FFFF6251FB6E232BFB1B" author="Becker-Migdisova, E. E." box="[308,617,1235,1262]" pageId="3" pageNumber="506" pagination="1991" refId="ref6989" refString="Becker-Migdisova, E. E. (1962 a) Superorder Rhynchota. Insects with proboscis. In: Rohdendorf, B. B. (Ed.), Fundamentals of Paleontology. Vol. 9. Arthropoda, Tracheata, Chelicerata. Akademiya Nauk SSSR, Moscow, pp. 216 - 317. [in Russian; English translation 1991, Smithsonian Institution Libraries and The National Science Foundation, Washington, D. C.]" type="book chapter" year="1962">Becker-Migdisova (1962a)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="CF222899FF85FFFF611DFB6E22ECFB1B" author="Martins-Neto, R. G. & Brauckmann, C. & Gallego, O. & Carmona, M. J." box="[632,942,1235,1262]" pageId="3" pageNumber="506" pagination="1 - 9" refId="ref8682" refString="Martins-Neto, R. G., Brauckmann, C., Gallego, O. F & Carmona, M. J. (2006) The Triassic insect fauna from Argentina. - Blattoptera, Glosselytrodea, Miomoptera, Auchenorrhyncha, and Coleoptera from the Los Rastros Formation (Bermejo Basin), Los Chanares locality (La Rioja Province). Clausthaler Geowissenschaften, 5, 1 - 9." type="journal article" year="2006">
|
||
Martins-Neto
|
||
<emphasis id="99C7897AFF85FFFF607FFB69220DFB18" box="[794,847,1235,1261]" italics="true" pageId="3" pageNumber="506">et al</emphasis>
|
||
. (2006)
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="CF222899FF85FFFF60DBFB6E2539FB1B" author="Lambkin, K. J." box="[958,1147,1235,1262]" pageId="3" pageNumber="506" pagination="580 - 590" refId="ref8312" refString="Lambkin, K. J. (2016) Revision of the Scytinopteridae (Hemiptera: Cicadomorpha: Scytinopteroidea) of the Queensland Triassic. Zootaxa, 4117 (4), 580 - 590. https: // doi. org / 10.11646 / zootaxa. 4117.4.9" type="journal article" year="2016">Lambkin (2016)</bibRefCitation>
|
||
, as well as
|
||
<bibRefCitation id="CF222899FF85FFFF666FFB6E2199FAE4" author="Shcherbakov, D. E." pageId="3" pageNumber="506" pagination="89 - 101" refId="ref9055" refString="Shcherbakov, D. E. (1984) Sistema i filogeniya permskikh Cicadomorpha (Cimicida, Cicadina). Palaeontologicheskii Zhurnal, 18 (2), 87 - 97. [in Russian, publishjed in English as: Shcherbakov, D. E. (1984) A system and the phylogeny of Permian Cicadomorpha (Cimicida, Cicadina). Paleontological Zhurnal, 2, 89 - 101.]" type="journal article" year="1984">Shcherbakov (1984</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="CF222899FF85FFFF6382FB4A2061FAE4" author="Shcherbakov, D. E." box="[231,291,1271,1297]" pageId="3" pageNumber="506" pagination="1 - 26" refId="ref9241" refString="Shcherbakov, D. E. (2011) New and little-known families of Hemiptera Cicadomorpha from the Triassic of Central Asia - early analogs of treehoppers and planthoppers. Zootaxa, 2836 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 2836.1.1" type="journal article" year="2011">2011</bibRefCitation>
|
||
). Of this list, the stand-out diagnostic characters of the family are the unique form of dSc, the R field occupying the apex of the tegmen, the complex and variable apical venation, and the apparently simple CuA. Of particular note is the absence of the hypocostal pit at the base of the costal margin. The pit grasped the mesipemeral knob to lock the tegmen in place (
|
||
<bibRefCitation id="CF222899FF85FFFF6177FADE23ABFA8B" author="Shcherbakov, D. E." box="[530,745,1379,1406]" pageId="3" pageNumber="506" pagination="31 - 45" refId="ref9129" refString="Shcherbakov, D. E. (1996) Origin and evolution of the Auchenorrhyncha as shown by the fossil record. In: Schaefer, C. W. (Ed.), Studies on Hemipteran Phylogeny. Entomological Society of America, Lanham, Maryland, pp. 31 - 45." type="book chapter" year="1996">Shcherbakov 1996</bibRefCitation>
|
||
), and is characteristic of the
|
||
<taxonomicName id="6CB32EEBFF85FFFF674DFADE24DAFA8B" authority="(Lambkin 2016)" baseAuthorityName="Lambkin" baseAuthorityYear="2016" box="[1064,1432,1379,1406]" class="Insecta" family="Scytinopteridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="family">
|
||
Scytinopteridae (
|
||
<bibRefCitation id="CF222899FF85FFFF6783FADE24D2FA8B" author="Lambkin, K. J." box="[1254,1424,1379,1406]" pageId="3" pageNumber="506" pagination="580 - 590" refId="ref8312" refString="Lambkin, K. J. (2016) Revision of the Scytinopteridae (Hemiptera: Cicadomorpha: Scytinopteroidea) of the Queensland Triassic. Zootaxa, 4117 (4), 580 - 590. https: // doi. org / 10.11646 / zootaxa. 4117.4.9" type="journal article" year="2016">Lambkin 2016</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, but is absent also in the
|
||
<taxonomicName id="6CB32EEBFF85FFFF62CEFA3A2208FA54" authority="(Shcherbakov 2011)" baseAuthorityName="Shcherbakov" baseAuthorityYear="2011" box="[427,842,1415,1441]" class="Insecta" family="Saaloscytinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="family">
|
||
Saaloscytinidae (
|
||
<bibRefCitation id="CF222899FF85FFFF610FFA3A2201FA54" author="Shcherbakov, D. E." box="[618,835,1415,1441]" pageId="3" pageNumber="506" pagination="1 - 26" refId="ref9241" refString="Shcherbakov, D. E. (2011) New and little-known families of Hemiptera Cicadomorpha from the Triassic of Central Asia - early analogs of treehoppers and planthoppers. Zootaxa, 2836 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 2836.1.1" type="journal article" year="2011">Shcherbakov 2011</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. With the exception of
|
||
<taxonomicName id="6CB32EEBFF85FFFF6731FA352428FA54" authorityName="Barth, Ansorge et Brauckmann" authorityYear="2011" box="[1108,1386,1415,1441]" class="Insecta" family="Ipsviciidae" genus="Ipsvicia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="langenbergensis">
|
||
<emphasis id="99C7897AFF85FFFF6731FA352428FA54" box="[1108,1386,1415,1441]" italics="true" pageId="3" pageNumber="506">Ipsvicia langenbergensis</emphasis>
|
||
</taxonomicName>
|
||
, the other confirmed members of the family (see discussion above) have not been considered in the diagnosis due to their inadequate preservation. Little is known of the head, thorax and abdomen in the other five scytinoperoid families.
|
||
<taxonomicName id="6CB32EEBFF85FFFF63F2FA4E2247F9FB" authority="Evans, 1943" authorityName="Evans" authorityYear="1943" box="[151,773,1523,1550]" class="Insecta" family="Paraknightiidae" genus="Paraknightia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="magnifica">
|
||
<emphasis id="99C7897AFF85FFFF63F2FA4E20E6F9F8" box="[151,420,1523,1549]" italics="true" pageId="3" pageNumber="506">Paraknightia magnifica</emphasis>
|
||
<bibRefCitation id="CF222899FF85FFFF62C8FA492378F9F8" author="Evans, J. W." box="[429,570,1523,1550]" pageId="3" pageNumber="506" pagination="180 - 198" refId="ref7341" refString="Evans, J. W. (1943) Upper Permian Homoptera from New South Wales. Records of the Australian Museum, 21, 180 - 198. https: // doi. org / 10.3853 / j. 0067 - 1975.21.1943.534" type="journal article" year="1943">Evans, 1943</bibRefCitation>
|
||
(Paraknightiidae)
|
||
</taxonomicName>
|
||
is similar to the
|
||
<taxonomicName id="6CB32EEBFF85FFFF60A9FA4E2505F9F8" authorityName="Tillyard" authorityYear="1920" box="[972,1095,1523,1549]" class="Insecta" family="Ipsviciidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="family">Ipsviciidae</taxonomicName>
|
||
in the shape of the pronotum with paranotal-like lobes and the presence of a siphonate ovipositor, although its mesoscutellum appears to be somewhat larger (
|
||
<bibRefCitation id="CF222899FF85FFFF6206F98620AEF9A0" author="Evans, J. W." box="[355,492,1595,1621]" pageId="3" pageNumber="506" pagination="180 - 198" refId="ref7341" refString="Evans, J. W. (1943) Upper Permian Homoptera from New South Wales. Records of the Australian Museum, 21, 180 - 198. https: // doi. org / 10.3853 / j. 0067 - 1975.21.1943.534" type="journal article" year="1943">Evans 1943</bibRefCitation>
|
||
, figs 24a, 24b). On the other hand,
|
||
<taxonomicName id="6CB32EEBFF85FFFF60E0F986201BF98F" authority="Becker-Migdisova, 1948" authorityName="Becker-Migdisova" authorityYear="1948" class="Insecta" family="Scytinopteridae" genus="Scytinoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="cubitalis">
|
||
<emphasis id="99C7897AFF85FFFF60E0F986253EF9A0" box="[901,1148,1595,1621]" italics="true" pageId="3" pageNumber="506">Scytinoptera cubitalis</emphasis>
|
||
<bibRefCitation id="CF222899FF85FFFF67E0F98624DEF9A0" author="Becker-Migdisova, E. E." box="[1157,1436,1595,1621]" pageId="3" pageNumber="506" pagination="1 - 42" refId="ref6881" refString="Becker-Migdisova, E. E. (1948) Permian cicadas of the family Scytinopteridae from the Soyana River. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR, 15, 1 - 42. [in Russian]" type="journal article" year="1948">Becker-Migdisova, 1948</bibRefCitation>
|
||
(Scytinopteridae)
|
||
</taxonomicName>
|
||
has a much broader head, pronotum without paranotal lobes, and also a much larger mesoscutellum (
|
||
<bibRefCitation id="CF222899FF85FFFF63B4F93E20A7F968" author="Becker-Migdisova, E. E." box="[209,485,1667,1693]" pageId="3" pageNumber="506" pagination="1 - 42" refId="ref6881" refString="Becker-Migdisova, E. E. (1948) Permian cicadas of the family Scytinopteridae from the Soyana River. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR, 15, 1 - 42. [in Russian]" type="journal article" year="1948">Becker-Migdisova 1948</bibRefCitation>
|
||
, fig. 49).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |