168 lines
18 KiB
XML
168 lines
18 KiB
XML
<document id="362B91657667A3C29FE42085F1EEBBAA" ID-DOI="10.11646/zootaxa.4576.3.6" ID-GBIF-Dataset="7abdc62c-c350-49dd-8060-4ed64da07362" ID-ISSN="1175-5326" ID-Zenodo-Dep="3715772" ID-ZooBank="56BC8573-D4A1-4B18-9BF6-7AB5F7984BFD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584603665960" checkinUser="plazi" docAuthor="Muona, Jyrki" docDate="2019" docId="03D387E9FFAE7777FF6FEAC6FA8EF835" docLanguage="en" docName="zootaxa.4576.3.6.pdf" docOrigin="Zootaxa 4576 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Potergus superbus Muona 2019, new species" docType="treatment" docVersion="5" lastPageNumber="530" masterDocId="FFEAFF91FFA7777EFFF8EE40FF81FFA7" masterDocTitle="Throscidae (Coleoptera) relationships, with descriptions of new fossil genera and species" masterLastPageNumber="543" masterPageNumber="521" pageNumber="530" updateTime="1698730058212" updateUser="plazi">
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<mods:titleInfo id="2FAECDCA8390666B009621096252E9C3">
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<mods:title id="3FDBB6131B508155E0CC9154784FC6B2">Throscidae (Coleoptera) relationships, with descriptions of new fossil genera and species</mods:title>
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<mods:name id="1AEEAAC80B56A14561056D0DDD65E75D" type="personal">
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<mods:namePart id="2909CC87B0530DA238B5849B78D296EB">Muona, Jyrki</mods:namePart>
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<mods:title id="474DF349D11C91C1A9D1A38810D6E4B8">Zootaxa</mods:title>
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<mods:part id="B1C9B2B62DFFDD9F9F419C62A822CACC">
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<mods:date id="D700E25EDC395A2C29E155EB7DA8BFEF">2019</mods:date>
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<mods:number id="0CC2325BA5EB0BE69F21FEC58A970018">2019-04-03</mods:number>
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<mods:number id="D5A171D553D06187C95DD927CC239536">4576</mods:number>
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<mods:identifier id="2F832D610CACC7A2AA373884CB591D94" type="DOI">10.11646/zootaxa.4576.3.6</mods:identifier>
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<mods:identifier id="2D8469B5B5B7D4B5C517FF293C849FAF" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A5E92B67B6355A0A266EA1E2848258E4" type="Zenodo-Dep">3715772</mods:identifier>
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<treatment id="03D387E9FFAE7777FF6FEAC6FA8EF835" ID-DOI="http://doi.org/10.5281/zenodo.3716784" ID-GBIF-Taxon="162639250" ID-Zenodo-Dep="3716784" LSID="urn:lsid:plazi:treatment:03D387E9FFAE7777FF6FEAC6FA8EF835" httpUri="http://treatment.plazi.org/id/03D387E9FFAE7777FF6FEAC6FA8EF835" lastPageNumber="530" pageId="9" pageNumber="530">
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<subSubSection id="C3606574FFAE7777FF6FEAC6FD8FFB07" box="[151,526,1158,1184]" pageId="9" pageNumber="530" type="nomenclature">
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<paragraph id="8BC536FFFFAE7777FF6FEAC6FD8FFB07" blockId="9.[151,526,1158,1218]" box="[151,526,1158,1184]" pageId="9" pageNumber="530">
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<heading id="D08D8193FFAE7777FF6FEAC6FD8FFB07" bold="true" box="[151,526,1158,1184]" fontSize="11" level="1" pageId="9" pageNumber="530" reason="1">
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<emphasis id="B90EEAEDFFAE7777FF6FEAC6FD8FFB07" bold="true" box="[151,526,1158,1184]" pageId="9" pageNumber="530">
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<taxonomicName id="4C7A4D7CFFAE7777FF6FEAC6FEF6FB07" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[151,375,1158,1184]" class="Insecta" family="Throscidae" genus="Potergus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="genus" status="sp. nov.">
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<emphasis id="B90EEAEDFFAE7777FF6FEAC6FEF6FB07" bold="true" box="[151,375,1158,1184]" italics="true" pageId="9" pageNumber="530">Potergus superbus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23D5796FFAE7777FE86EAC6FD8FFB07" box="[382,526,1158,1184]" pageId="9" pageNumber="530" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFAE7777FF6FEAE9FF67FB65" box="[151,230,1193,1218]" pageId="9" pageNumber="530" type="description">
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<paragraph id="8BC536FFFFAE7777FF6FEAE9FF67FB65" blockId="9.[151,526,1158,1218]" box="[151,230,1193,1218]" pageId="9" pageNumber="530">
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<figureCitation id="13412A7AFFAE7777FF6FEAE9FF67FB65" box="[151,230,1193,1218]" captionStart="FIGURE 11" captionStartId="12.[151,250,1757,1779]" captionTargetBox="[151,1436,1067,1735]" captionTargetId="figure@12.[151,1436,1067,1735]" captionTargetPageId="12" captionText="FIGURE 11. Potergus superbus n. sp., ventrolateral view. AtG = abdominal tarsal groove; MtG = metathoracic tarsal groove." figureDoi="http://doi.org/10.5281/zenodo.3715796" httpUri="https://zenodo.org/record/3715796/files/figure.png" pageId="9" pageNumber="530">Fig. 11</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFAE7777FF6FEAB1FB8CFAF5" pageId="9" pageNumber="530" type="materials_examined">
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<paragraph id="8BC536FFFFAE7777FF6FEAB1FB8CFAF5" blockId="9.[151,1437,1265,1938]" pageId="9" pageNumber="530">
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<emphasis id="B90EEAEDFFAE7777FF6FEAB1FEC8FAAD" bold="true" box="[151,329,1265,1290]" pageId="9" pageNumber="530">Type material.</emphasis>
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<typeStatus id="54C1885DFFAE7777FEACEAB1FE3DFAAD" box="[340,444,1265,1290]" pageId="9" pageNumber="530" type="holotype">Holotype</typeStatus>
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labeled:
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<taxonomicName id="4C7A4D7CFFAE7777FDD4EAB2FC80FAAD" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[556,769,1265,1290]" class="Insecta" family="Throscidae" genus="Potergus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="genus" status="sp. nov.">
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<emphasis id="B90EEAEDFFAE7777FDD4EAB2FC80FAAD" box="[556,769,1265,1290]" italics="true" pageId="9" pageNumber="530">Potergus superbus</emphasis>
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</taxonomicName>
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<emphasis id="B90EEAEDFFAE7777FCF4EAB1FCCEFAAD" bold="true" box="[780,847,1265,1290]" pageId="9" pageNumber="530">
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<taxonomicNameLabel id="A23D5796FFAE7777FCF4EAB1FCCEFAAD" box="[780,847,1265,1290]" pageId="9" pageNumber="530" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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/
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<materialsCitation id="3B123CA2FFAE7777FCA0EAB2FB88FAF5" ID-GBIF-Occurrence="2578498276" collectingDate="2015-10" collectorName="J. Muona" location="Embedded in Baltic Amber" pageId="9" pageNumber="530" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
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<typeStatus id="54C1885DFFAE7777FCA0EAB2FC6EFAAD" box="[856,1007,1266,1290]" pageId="9" pageNumber="530" type="holotype">HOLOTYPE</typeStatus>
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/
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<collectorName id="268F5329FFAE7777FC0FEAB2FBDEFAAE" box="[1015,1119,1266,1290]" pageId="9" pageNumber="530">J. Muona</collectorName>
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des. 10-2015. Found from
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<collectingRegion id="49BEF81DFFAE7777FF6FEB55FEA0FA89" box="[151,289,1301,1326]" country="Russia" name="Kaliningrad" pageId="9" pageNumber="530">Kaliningrad</collectingRegion>
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.
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<location id="8EA56024FFAE7777FEC8EB55FDF5FA89" LSID="urn:lsid:plazi:treatment:03D387E9FFAE7777FF6FEAC6FA8EF835:8EA56024FFAE7777FEC8EB55FDF5FA89" box="[304,628,1301,1326]" name="Embedded in Baltic Amber" pageId="9" pageNumber="530" stateProvince="Kaliningrad">Embedded in Baltic Amber</location>
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piece,
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<quantity id="4C829B1AFFAE7777FD37EB56FCA5FA8A" box="[719,804,1302,1326]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="530" unit="mm" value="23.0">23 mm</quantity>
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<date id="FFC4103FFFAE7777FCC8EB56FCE7FA89" box="[816,870,1301,1326]" pageId="9" pageNumber="530" value="2015-10">
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<collectingDate id="EF80E9D7FFAE7777FCC8EB56FCE7FA89" box="[816,870,1301,1326]" pageId="9" pageNumber="530" value="2015-10">
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x
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<quantity id="4C829B1AFFAE7777FCB2EB55FCE7FA89" box="[842,870,1301,1326]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="530" unit="mm" value="15.0">15</quantity>
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</collectingDate>
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</date>
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mm,
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<quantity id="4C829B1AFFAE7777FC4AEB56FC78FA8A" box="[946,1017,1302,1325]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="530" unit="mm" value="7.0">7 mm</quantity>
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thick, sex unknown. Age 50 Mya. Numerous minute, hirsute remains of plant matter are present as well (
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<figureCitation id="13412A7AFFAE7777FC54EB79FB81FAF5" box="[940,1024,1337,1362]" captionStart="FIGURE 24" captionStartId="19.[151,250,1925,1947]" captionTargetBox="[228,1352,1135,1910]" captionTargetId="figure@19.[228,1352,1135,1910]" captionTargetPageId="19" captionText="FIGURE 24. Potergus balticus n. sp., holotype." figureDoi="http://doi.org/10.5281/zenodo.3715822" httpUri="https://zenodo.org/record/3715822/files/figure.png" pageId="9" pageNumber="530">Fig. 24</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFAE7777FF3FEB1DFC16F931" pageId="9" pageNumber="530" type="diagnosis">
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<paragraph id="8BC536FFFFAE7777FF3FEB1DFB2BFAD1" blockId="9.[151,1437,1265,1938]" box="[199,1194,1373,1398]" pageId="9" pageNumber="530">
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<emphasis id="B90EEAEDFFAE7777FF3FEB1DFEBBFAD1" bold="true" box="[199,314,1373,1398]" pageId="9" pageNumber="530">Diagnosis</emphasis>
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. Elateroidea synapomorphy: pro-mesothoracic joint with clicking mechanism.
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</paragraph>
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<paragraph id="8BC536FFFFAE7777FF3FEBC1FC16F931" blockId="9.[151,1437,1265,1938]" pageId="9" pageNumber="530">
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<taxonomicName id="4C7A4D7CFFAE7777FF3FEBC1FEC5FA3D" authorityName="Laporte" authorityYear="1840" box="[199,324,1409,1434]" class="Insecta" family="Throscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="family">Throscidae</taxonomicName>
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synapomorphies: a2 as wide as a3–a4, antennal grooves running along notosternal suture and then bending laterally along proleg cavity. Synapomorphy of Potergini +
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<taxonomicName id="4C7A4D7CFFAE7777FC6EEBE5FB87FA19" box="[918,1030,1445,1470]" class="Insecta" family="Throscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="tribe" tribe="Throscini">Throscini</taxonomicName>
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: tibiae flattened, with tarsal grooves apically. Synapomorphy of
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<taxonomicName id="4C7A4D7CFFAE7777FE32EB8AFDAFFA46" authorityName="Bonvouloir. The" authorityYear="1859" box="[458,558,1482,1505]" class="Insecta" family="Throscidae" genus="Potergus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEAEDFFAE7777FE32EB8AFDAFFA46" box="[458,558,1482,1505]" italics="true" pageId="9" pageNumber="530">Potergus</emphasis>
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</taxonomicName>
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: keeled frons. This in combination with abdominal tarsal grooves is diagnostic for the genus. The extant
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<taxonomicName id="4C7A4D7CFFAE7777FE33EBAEFDAEF9A2" authorityName="Bonvouloir. The" authorityYear="1859" box="[459,559,1518,1541]" class="Insecta" family="Throscidae" genus="Potergus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEAEDFFAE7777FE33EBAEFDAEF9A2" box="[459,559,1518,1541]" italics="true" pageId="9" pageNumber="530">Potergus</emphasis>
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</taxonomicName>
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species and the Eocene fossil
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<taxonomicName id="4C7A4D7CFFAE7777FC65EBAEFBCCF9A2" authority="Muona" authorityName="Muona" authorityYear="1993" box="[925,1101,1517,1542]" class="Insecta" family="Throscidae" genus="Potergus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="frohi">
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<emphasis id="B90EEAEDFFAE7777FC65EBAEFC73F9A1" box="[925,1010,1517,1542]" italics="true" pageId="9" pageNumber="530">P. frohi</emphasis>
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Muona
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</taxonomicName>
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are parallel-sided and much narrower than
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<taxonomicName id="4C7A4D7CFFAE7777FEC1E852FE38F98D" authorityName="Muona" authorityYear="2019" box="[313,441,1553,1578]" class="Insecta" family="Throscidae" genus="Potergus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="superbus">
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<emphasis id="B90EEAEDFFAE7777FEC1E852FE38F98D" box="[313,441,1553,1578]" italics="true" pageId="9" pageNumber="530">P. superbus</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="4C7A4D7CFFAE7777FE3FE852FDC9F98D" authorityName="Muona" authorityYear="2019" box="[455,584,1553,1578]" class="Insecta" family="Throscidae" genus="Potergus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="superbus">
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<emphasis id="B90EEAEDFFAE7777FE3FE852FDC9F98D" box="[455,584,1553,1578]" italics="true" pageId="9" pageNumber="530">P. superbus</emphasis>
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</taxonomicName>
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resembles
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<taxonomicName id="4C7A4D7CFFAE7777FD3DE852FCEFF98E" authority="Muona" authorityName="Muona" authorityYear="1993" box="[709,878,1553,1578]" class="Insecta" family="Throscidae" genus="Potergus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="logei">
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<emphasis id="B90EEAEDFFAE7777FD3DE852FC99F98D" box="[709,792,1553,1578]" italics="true" pageId="9" pageNumber="530">P. logei</emphasis>
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Muona
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</taxonomicName>
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in shape, but has longer abdominal tarsal grooves than that species, these extending to
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<date id="FFC4103FFFAE7777FDBDE876FDE9F9E9" box="[581,616,1590,1614]" pageId="9" pageNumber="530">v4.</date>
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In addition, the metathoracic tarsal grooves are more distant from the metanepisterna in
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<taxonomicName id="4C7A4D7CFFAE7777FE9FE81AFE68F9D5" authorityName="Muona" authorityYear="2019" box="[359,489,1625,1650]" class="Insecta" family="Throscidae" genus="Potergus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="superbus">
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<emphasis id="B90EEAEDFFAE7777FE9FE81AFE68F9D5" box="[359,489,1625,1650]" italics="true" pageId="9" pageNumber="530">P. superbus</emphasis>
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</taxonomicName>
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that in
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<taxonomicName id="4C7A4D7CFFAE7777FDB9E81AFD12F9D5" authorityName="Muona" authorityYear="1993" box="[577,659,1625,1650]" class="Insecta" family="Throscidae" genus="Potergus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="logei">
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<emphasis id="B90EEAEDFFAE7777FDB9E81AFD12F9D5" box="[577,659,1625,1650]" italics="true" pageId="9" pageNumber="530">P. logei</emphasis>
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</taxonomicName>
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. The metallic shine of
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<taxonomicName id="4C7A4D7CFFAE7777FC67E81AFBA0F9D5" authorityName="Muona" authorityYear="2019" box="[927,1057,1625,1650]" class="Insecta" family="Throscidae" genus="Potergus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="superbus">
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<emphasis id="B90EEAEDFFAE7777FC67E81AFBA0F9D5" box="[927,1057,1625,1650]" italics="true" pageId="9" pageNumber="530">P. superbus</emphasis>
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</taxonomicName>
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is diagnostic as well. Because of the strong pronotal punctation, the lateral carina is obscure cranially.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFAE7777FF3FE8E1FC87F8C9" pageId="9" pageNumber="530" type="description">
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<paragraph id="8BC536FFFFAE7777FF3FE8E1FC12F882" blockId="9.[151,1437,1265,1938]" pageId="9" pageNumber="530">
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<emphasis id="B90EEAEDFFAE7777FF3FE8E1FED0F91D" bold="true" box="[199,337,1697,1722]" pageId="9" pageNumber="530">Description</emphasis>
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. Length
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<quantity id="4C829B1AFFAE7777FE41E8E2FD92F91E" box="[441,531,1698,1722]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="9" pageNumber="530" unit="mm" value="4.3">4.3 mm</quantity>
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. Venter shiny with conspicuous metallic sheen, blue to slightly purple. Dorsum poorly visible. Pronotum and venter coarsely and densely punctate, head densely and slightly less coarsely punctate than pronotum, medially keeled. Elytral striae hardly indicated, serial punctures orderly placed, large, interstices with minute punctation, epipleura faintly and very densely punctate.
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</paragraph>
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<paragraph id="8BC536FFFFAE7777FF3FE972FC87F8C9" blockId="9.[151,1437,1265,1938]" pageId="9" pageNumber="530">
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Antennae short and robust, a2–a8 about as long as wide and of even width, a9–a11 forming a distinctly wider club. Position and length of tarsal grooves as in
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<figureCitation id="13412A7AFFAE7777FD49E915FC83F8C9" box="[689,770,1877,1902]" captionStart="FIGURE 11" captionStartId="12.[151,250,1757,1779]" captionTargetBox="[151,1436,1067,1735]" captionTargetId="figure@12.[151,1436,1067,1735]" captionTargetPageId="12" captionText="FIGURE 11. Potergus superbus n. sp., ventrolateral view. AtG = abdominal tarsal groove; MtG = metathoracic tarsal groove." figureDoi="http://doi.org/10.5281/zenodo.3715796" httpUri="https://zenodo.org/record/3715796/files/figure.png" pageId="9" pageNumber="530">Fig. 11</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606574FFAE7777FF3FE939FA8EF835" box="[199,1295,1913,1938]" pageId="9" pageNumber="530" type="etymology">
|
||
<paragraph id="8BC536FFFFAE7777FF3FE939FA8EF835" blockId="9.[151,1437,1265,1938]" box="[199,1295,1913,1938]" pageId="9" pageNumber="530">
|
||
<emphasis id="B90EEAEDFFAE7777FF3FE939FEC7F835" bold="true" box="[199,326,1913,1938]" pageId="9" pageNumber="530">Etymology</emphasis>
|
||
. Contrary to other known throscids, the dorsum of this species is faintly metallic blue.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |