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<document id="C83C8EB9635FC0355A6BD5620A85F0E3" ID-CLB-Dataset="22834" ID-DOI="10.11646/zootaxa.4657.2.1" ID-GBIF-Dataset="2cd7e95f-232f-44b1-8981-ca0cda569d2d" ID-ISSN="1175-5326" ID-Zenodo-Dep="3763876" ID-ZooBank="73A706D5-41CF-4A2F-965F-70C779E6B9EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587670032081" checkinUser="valdenar" docAuthor="Sassi, Davide" docDate="2019" docId="03C287993C452E6442E5FA28FB09B10D" docLanguage="en" docName="zootaxa.4657.2.1.pdf.imf" docOrigin="Zootaxa 4657 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Metallactus praetorius Sassi 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="236" masterDocId="FFFBFFE13C642E474272FFE5FFE7B316" masterDocTitle="Revision of the Metallactus hamifer species-group (Coleoptera: Chrysomelidae: Cryptocephalinae)" masterLastPageNumber="245" masterPageNumber="201" pageNumber="234" updateTime="1698835701105" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7B0C43D832DA167B4E43AA6F881409BE">Revision of the Metallactus hamifer species-group (Coleoptera: Chrysomelidae: Cryptocephalinae)</mods:title>
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<mods:namePart id="B5580A1165100AC6EF5F9CDADCD86B08">Sassi, Davide</mods:namePart>
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<mods:date id="F4BD6D4011CAB1B594E0E36E1104AAAB">2019</mods:date>
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<treatment id="03C287993C452E6442E5FA28FB09B10D" ID-DOI="http://doi.org/10.5281/zenodo.3803963" ID-GBIF-Taxon="163737089" ID-Zenodo-Dep="3803963" LSID="urn:lsid:plazi:treatment:03C287993C452E6442E5FA28FB09B10D" httpUri="http://treatment.plazi.org/id/03C287993C452E6442E5FA28FB09B10D" lastPageId="35" lastPageNumber="236" pageId="33" pageNumber="234">
<subSubSection id="C37165043C452E6642E5FA28FDEAB6F1" box="[151,525,1485,1512]" pageId="33" pageNumber="234" type="nomenclature">
<paragraph id="8BD4368F3C452E6642E5FA28FDEAB6F1" blockId="33.[151,525,1485,1547]" box="[151,525,1485,1512]" pageId="33" pageNumber="234">
<heading id="D09C81E33C452E6642E5FA28FDEAB6F1" bold="true" box="[151,525,1485,1512]" fontSize="11" level="1" pageId="33" pageNumber="234" reason="1">
<emphasis id="B91FEA9D3C452E6642E5FA28FDEAB6F1" bold="true" box="[151,525,1485,1512]" pageId="33" pageNumber="234">
<taxonomicName id="4C6B4D0C3C452E6642E5FA28FE4FB6F1" authority="Sassi, 2019" authorityName="Sassi" authorityYear="2019" box="[151,424,1485,1512]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="praetorius" status="sp. nov.">
<emphasis id="B91FEA9D3C452E6642E5FA28FE4FB6F1" bold="true" box="[151,424,1485,1512]" italics="true" pageId="33" pageNumber="234">Metallactus praetorius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57E63C452E6643C2FA28FDEAB6F1" box="[432,525,1485,1511]" pageId="33" pageNumber="234" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37165043C452E6642E5FA14FD6BB43D" pageId="33" pageNumber="234" type="description">
<paragraph id="8BD4368F3C452E6642E5FA14FECFB51D" blockId="33.[151,525,1485,1547]" box="[151,296,1521,1547]" pageId="33" pageNumber="234">
(
<figureCitation id="13502A0A3C452E6642EDFA14FF10B51D" box="[159,247,1521,1547]" captionStart="FIGURES 1415" captionStartId="36.[151,264,1865,1890]" captionTargetBox="[242,1341,195,1787]" captionTargetId="figure@36.[199,1388,166,1845]" captionTargetPageId="36" captionText="FIGURES 1415. M. praetorius (14), M. sekerkai (15). 14ab: habitus; 14ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.6 mm]; 14f: sclerites of endophallus [0.4 mm]; 14g: spermatheca [0.5 mm]; 14h: antenna [1.6 mm]. 15ab: habitus; 15ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.9 mm]; 15f: sclerites of endophallus [0.5 mm]; 15g: spermatheca [0.9 mm]; 15h: antenna [1.9 mm]. 14ab: holotype; 15ab: holotype. scIIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.3763893" httpUri="https://zenodo.org/record/3763893/files/figure.png" pageId="33" pageNumber="234">Figs 14</figureCitation>
;
<figureCitation id="13502A0A3C452E664371FA14FEC7B51D" box="[259,288,1521,1547]" captionStart="FIGURES 2734" captionStartId="42.[151,264,1930,1955]" captionTargetBox="[223,1364,169,1836]" captionTargetId="figure@42.[151,1435,166,1911]" captionTargetPageId="42" captionText="FIGURES 2734. Distribution maps for Metallactus hamifer species-group. M. luniger (27); M. madefactus (28); M. octoguttatus (29); M. pollinctor (30); M. praetorius (31); M. sekerkai (32); M. tarsalis (33); M. viator (34). Grey squares: localities reported in a generic way." figureDoi="http://doi.org/10.5281/zenodo.3763899" httpUri="https://zenodo.org/record/3763899/files/figure.png" pageId="33" pageNumber="234">31</figureCitation>
)
</paragraph>
<paragraph id="8BD4368F3C452E6642E5F9DCFD6BB43D" blockId="33.[151,1437,1593,2015]" pageId="33" pageNumber="234">
<emphasis id="B91FEA9D3C452E6642E5F9DCFF07B545" bold="true" box="[151,224,1593,1619]" italics="true" pageId="33" pageNumber="234">Types.</emphasis>
<materialsCitation id="3B033CD23C452E66429BF9DCFC4FB58D" ID-GBIF-Occurrence="2608527009" collectorName="Coll. Machatschke" country="Paraguay" location="Paraguay" pageId="33" pageNumber="234" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54D0882D3C452E66429BF9DCFEB1B547" box="[233,342,1593,1619]" pageId="33" pageNumber="234" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D6DFD063C452E664315F9DCFE99B545" box="[359,382,1593,1619]" pageId="33" pageNumber="234" type="male"></specimenCount>
, glued, aedeagus on the same card bearing the specimen, detached abdomen and sclerites of endophallus on a separate card // “
<collectingCountry id="F37C761F3C452E664068F9B8FD6FB561" box="[538,648,1629,1655]" name="Paraguay" pageId="33" pageNumber="234">Paraguay</collectingCountry>
” [white label, printed] // “Staatssamlung München 1975 Erwerb
<collectorName id="269E53593C452E664717F9B8FEC9B58D" pageId="33" pageNumber="234">Coll. Machatschke</collectorName>
” [white label, printed] // “
<taxonomicName id="4C6B4D0C3C452E66403DF964FCA9B58D" authority="Sassi, 2019" authorityName="Sassi" authorityYear="2019" box="[591,846,1665,1691]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="praetorius" status="sp. nov.">
<emphasis id="B91FEA9D3C452E66403DF964FCA9B58D" box="[591,846,1665,1691]" italics="true" pageId="33" pageNumber="234">Metallactus praetorius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57E63C452E664126F964FC4FB58D" box="[852,936,1665,1691]" pageId="33" pageNumber="234" rank="species">
<emphasis id="B91FEA9D3C452E664126F964FC4FB58D" bold="true" box="[852,936,1665,1691]" pageId="33" pageNumber="234">sp. nov</emphasis>
</taxonomicNameLabel>
</materialsCitation>
<emphasis id="B91FEA9D3C452E6641DAF964FC4AB58D" bold="true" box="[936,941,1665,1691]" pageId="33" pageNumber="234">.</emphasis>
<materialsCitation id="3B033CD23C452E6641C1F96DFA70B411" ID-GBIF-Occurrence="2608526977" collectingDate="1895" collectingDateMax="1896" collectingDateMin="1895" collectionCode="ZSM, MNHUB, MNHN" collectorName="D. Sassi &amp; Leg. C. Fiebrig" country="Paraguay" location="Jatahy Etat de Goyaz Ch. Pujol" pageId="33" pageNumber="234" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
<subScript id="17EF34CA3C452E6641C1F96DFBCFB58F" attach="right" box="[947,1064,1672,1689]" fontSize="7" pageId="33" pageNumber="234">
<typeStatus id="54D0882D3C452E6641C1F96DFBCFB58F" box="[947,1064,1672,1689]" pageId="33" pageNumber="234" type="holotype">HOLOTYPUS</typeStatus>
</subScript>
<collectorName id="269E53593C452E66465FF964FB6EB58D" box="[1069,1161,1665,1691]" pageId="33" pageNumber="234">D. Sassi</collectorName>
des.” [red label, printed] // (
<collectionCode id="ED7AAE4A3C452E6642C8F940FF1EB5A9" box="[186,249,1701,1727]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="33" pageNumber="234" type="Museum">ZSM</collectionCode>
).
<typeStatus id="54D0882D3C452E664362F940FE62B5AB" box="[272,389,1701,1727]" pageId="33" pageNumber="234" type="paratype">
<subScript id="17EF34CA3C452E664362F940FE62B5AB" attach="left" box="[272,389,1701,1727]" fontSize="7" pageId="33" pageNumber="234">PARATYPES</subScript>
</typeStatus>
:
<specimenCount id="9D6DFD063C452E6643EAF940FE5CB5A9" box="[408,443,1701,1727]" pageId="33" pageNumber="234" type="female">1♀</specimenCount>
, abdomen heavily damaged // “fliegl [uncertain spelling, white label, handwritten] // “
<collectingCountry id="F37C761F3C452E6642D3F92CFEEBB5F5" box="[161,268,1737,1763]" name="Paraguay" pageId="33" pageNumber="234">Paraguay</collectingCountry>
<collectorName id="269E53593C452E664366F92CFE21B5F5" box="[276,454,1737,1763]" pageId="33" pageNumber="234">Leg. C. Fiebrig</collectorName>
” [white label, printed] // (
<collectionCode id="ED7AAE4A3C452E664082F92CFCB8B5F5" box="[752,863,1737,1763]" pageId="33" pageNumber="234">MNHUB</collectionCode>
);
<specimenCount id="9D6DFD063C452E664101F92CFC71B5F5" box="[883,918,1737,1763]" pageId="33" pageNumber="234" type="female">1♀</specimenCount>
// “
<location id="8EB460543C452E6641B3F92CFACDB5F5" LSID="urn:lsid:plazi:treatment:03C287993C452E6442E5FA28FB09B10D:8EB460543C452E6641B3F92CFACDB5F5" box="[961,1322,1737,1763]" country="Paraguay" name="Jatahy Etat de Goyaz Ch. Pujol" pageId="33" pageNumber="234">Jatahy Etat de Goyaz Ch. Pujol</location>
<date id="FFD5104F3C452E664740F92DFA75B5F5" box="[1330,1426,1736,1763]" pageId="33" pageNumber="234" value="1895" valueMax="1896">
<collectingDate id="EF91E9A73C452E664740F92DFA75B5F5" box="[1330,1426,1736,1763]" pageId="33" pageNumber="234" value="1895" valueMax="1896">1895-96</collectingDate>
</date>
” [white label, printed] // (
<collectionCode id="ED7AAE4A3C452E6643D7F908FDE6B411" box="[421,513,1773,1799]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="234">MNHN</collectionCode>
). Both
<typeStatus id="54D0882D3C452E66403DF908FD5DB411" box="[591,698,1773,1799]" pageId="33" pageNumber="234" type="paratype">paratypes</typeStatus>
provided with additional label: // “
<taxonomicName id="4C6B4D0C3C452E66464CF908FAD9B411" authority="Sassi, 2019" authorityName="Sassi" authorityYear="2019" box="[1086,1342,1773,1799]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="praetorius" status="sp. nov.">
<emphasis id="B91FEA9D3C452E66464CF908FAD9B411" box="[1086,1342,1773,1799]" italics="true" pageId="33" pageNumber="234">Metallactus praetorius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57E63C452E664736F908FA70B411" box="[1348,1431,1773,1799]" pageId="33" pageNumber="234" rank="species">
<emphasis id="B91FEA9D3C452E664736F908FA70B411" bold="true" box="[1348,1431,1773,1799]" pageId="33" pageNumber="234">sp. nov</emphasis>
</taxonomicNameLabel>
</materialsCitation>
<emphasis id="B91FEA9D3C452E6647E5F908FA7BB411" bold="true" box="[1431,1436,1773,1799]" pageId="33" pageNumber="234">.</emphasis>
<subScript id="17EF34CA3C452E6642E5F8FDFEEEB43F" attach="left" box="[151,265,1816,1833]" fontSize="7" pageId="33" pageNumber="234">
<typeStatus id="54D0882D3C452E6642E5F8FDFEEEB43F" box="[151,265,1816,1833]" pageId="33" pageNumber="234" type="paratype">PARATYPUS</typeStatus>
</subScript>
D. Sassi des.” [red label, printed].
</paragraph>
</subSubSection>
<subSubSection id="C37165043C452E6642B5F8D1FC21B459" box="[199,966,1844,1871]" pageId="33" pageNumber="234" type="etymology">
<paragraph id="8BD4368F3C452E6642B5F8D1FC21B459" blockId="33.[151,1437,1593,2015]" box="[199,966,1844,1871]" pageId="33" pageNumber="234">
<emphasis id="B91FEA9D3C452E6642B5F8D1FEAFB459" bold="true" box="[199,328,1844,1871]" italics="true" pageId="33" pageNumber="234">Etymology.</emphasis>
Latin adjective meaning “belonging to the commander”.
</paragraph>
</subSubSection>
<subSubSection id="C37165043C452E6642B5F8BCFE00B465" box="[199,487,1880,1907]" pageId="33" pageNumber="234" type="materials_examined">
<paragraph id="8BD4368F3C452E6642B5F8BCFE00B465" blockId="33.[151,1437,1593,2015]" box="[199,487,1880,1907]" pageId="33" pageNumber="234">
<materialsCitation id="3B033CD23C452E6642B5F8BCFE00B465" ID-GBIF-Occurrence="2608527044" box="[199,487,1880,1907]" country="Paraguay" location="Paraguay" pageId="33" pageNumber="234" specimenCount="1" typeStatus="holotype">
<emphasis id="B91FEA9D3C452E6642B5F8BCFEB8B465" bold="true" box="[199,351,1880,1907]" italics="true" pageId="33" pageNumber="234">
<typeStatus id="54D0882D3C452E6642B5F8BCFF19B465" box="[199,254,1881,1907]" pageId="33" pageNumber="234">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F37C761F3C452E664302F8BCFE39B465" box="[368,478,1881,1907]" name="Paraguay" pageId="33" pageNumber="234">Paraguay</collectingCountry>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37165043C452E6642B5F899FDC5B481" box="[199,546,1916,1943]" pageId="33" pageNumber="234" type="distribution">
<paragraph id="8BD4368F3C452E6642B5F899FDC5B481" blockId="33.[151,1437,1593,2015]" box="[199,546,1916,1943]" pageId="33" pageNumber="234">
<emphasis id="B91FEA9D3C452E6642B5F899FEB4B481" bold="true" box="[199,339,1916,1943]" italics="true" pageId="33" pageNumber="234">Distribution</emphasis>
.
<collectingCountry id="F37C761F3C452E664313F898FE4FB481" box="[353,424,1917,1943]" name="Brazil" pageId="33" pageNumber="234">Brazil</collectingCountry>
,
<collectingCountry id="F37C761F3C452E6643C6F898FDF9B481" box="[436,542,1917,1943]" name="Paraguay" pageId="33" pageNumber="234">Paraguay</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37165043C452E6542B5F844FDE3B1BD" lastPageId="34" lastPageNumber="235" pageId="33" pageNumber="234" type="diagnosis">
<paragraph id="8BD4368F3C452E6542B5F844FDE3B1BD" blockId="33.[151,1437,1593,2015]" lastBlockId="34.[151,1437,153,2015]" lastPageId="34" lastPageNumber="235" pageId="33" pageNumber="234">
<emphasis id="B91FEA9D3C452E6642B5F844FEDDB4AD" bold="true" box="[199,314,1953,1979]" italics="true" pageId="33" pageNumber="234">Diagnosis</emphasis>
. A
<taxonomicName id="4C6B4D0C3C452E664311F844FE00B4AD" box="[355,487,1953,1979]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEA9D3C452E664311F844FE00B4AD" box="[355,487,1953,1979]" italics="true" pageId="33" pageNumber="234">Metallactus</emphasis>
</taxonomicName>
of medium size. It belongs to a subgroup characterized by a couple of longitudinal dark stripes both on pronotum and (except for
<taxonomicName id="4C6B4D0C3C452E6640E9F823FCF3B4C9" authorityName="Suffrian" authorityYear="1866" box="[667,788,1989,2015]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="B91FEA9D3C452E6640E9F823FCF3B4C9" box="[667,788,1989,2015]" italics="true" pageId="33" pageNumber="234">M. tarsalis</emphasis>
</taxonomicName>
) on each elytron.
<taxonomicName id="4C6B4D0C3C452E6641AEF823FB92B4C9" authorityName="Sassi" authorityYear="2019" box="[988,1141,1989,2015]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="chamorroi">
<emphasis id="B91FEA9D3C452E6641AEF823FB92B4C9" box="[988,1141,1989,2015]" italics="true" pageId="33" pageNumber="234">M. chamorroi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6B4D0C3C452E6646F1F823FB1BB4C9" authorityName="Suffrian" authorityYear="1866" box="[1155,1276,1989,2015]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="B91FEA9D3C452E6646F1F823FB1BB4C9" box="[1155,1276,1989,2015]" italics="true" pageId="33" pageNumber="234">M. tarsalis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6B4D0C3C452E66477BF823FA96B4C9" authorityName="Sassi" authorityYear="2019" box="[1289,1393,1990,2015]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="viator">
<emphasis id="B91FEA9D3C452E66477BF823FA96B4C9" box="[1289,1393,1990,2015]" italics="true" pageId="33" pageNumber="234">M. viator</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6B4D0C3C452E65470CF823FEF2B3A5" authorityName="Jacoby" authorityYear="1907" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" lastPageId="34" lastPageNumber="235" order="Coleoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="bivitticollis">
<emphasis id="B91FEA9D3C452E65470CF823FEF2B3A5" italics="true" lastPageId="34" lastPageNumber="235" pageId="33" pageNumber="234">M. bivitticollis</emphasis>
</taxonomicName>
,
<emphasis id="B91FEA9D3C462E654356FF7FFDABB3A5" box="[292,588,153,179]" italics="true" pageId="34" pageNumber="235">
<taxonomicName id="4C6B4D0C3C462E654356FF7FFE5EB3A5" authorityName="Suffrian" authorityYear="1866" box="[292,441,153,179]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="pollinctor">M. pollinctor</taxonomicName>
,
<taxonomicName id="4C6B4D0C3C462E6543B7FF7FFDABB3A5" authorityName=", Suffrian" authorityYear="1866" box="[453,588,154,179]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="agonista">M. agonista</taxonomicName>
</emphasis>
and a color variation of
<taxonomicName id="4C6B4D0C3C462E654114FF7FFC04B3A5" authorityName=", Suffrian" authorityYear="1866" box="[870,995,153,179]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="hamifer">
<emphasis id="B91FEA9D3C462E654114FF7FFC04B3A5" box="[870,995,153,179]" italics="true" pageId="34" pageNumber="235">M. hamifer</emphasis>
</taxonomicName>
also belong to the same subgroup. Together with
<emphasis id="B91FEA9D3C462E65436EFF5BFDAFB3C1" box="[284,584,189,215]" italics="true" pageId="34" pageNumber="235">
<taxonomicName id="4C6B4D0C3C462E65436EFF5BFE43B3C1" authorityName=", Suffrian" authorityYear="1866" box="[284,420,190,215]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="agonista">M. agonista</taxonomicName>
,
<taxonomicName id="4C6B4D0C3C462E6543DDFF5BFDAFB3C1" authorityName="Sassi" authorityYear="2019" box="[431,584,189,215]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="chamorroi">M. chamorroi</taxonomicName>
</emphasis>
and
<taxonomicName id="4C6B4D0C3C462E65400CFF5BFD10B3C1" authorityName="Suffrian" authorityYear="1866" box="[638,759,189,215]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="B91FEA9D3C462E65400CFF5BFD10B3C1" box="[638,759,189,215]" italics="true" pageId="34" pageNumber="235">M. tarsalis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6B4D0C3C462E654176FF5BFC79B3C1" authorityName="Sassi" authorityYear="2019" box="[772,926,190,215]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="praetorius">
<emphasis id="B91FEA9D3C462E654176FF5BFC79B3C1" box="[772,926,190,215]" italics="true" pageId="34" pageNumber="235">M. praetorius</emphasis>
</taxonomicName>
is distinguishable in the dark pattern which is decidedly partly red and not totally black as in the remaining species cited above. However, this distinction, purely based on a single chromatic character and useful, at present, for diagnostic purpose, could turn out to be inconsistent if more material was available in the future.
<taxonomicName id="4C6B4D0C3C462E6540F7FECFFCEDB255" authorityName=", Suffrian" authorityYear="1866" box="[645,778,298,323]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="agonista">
<emphasis id="B91FEA9D3C462E6540F7FECFFCEDB255" box="[645,778,298,323]" italics="true" pageId="34" pageNumber="235">M. agonista</emphasis>
</taxonomicName>
has a unique pronotum mostly reddish or rusty, with six or four black spots lined up along longitudinal median yellow line.
<taxonomicName id="4C6B4D0C3C462E654113FEABFC1DB271" authorityName="Sassi" authorityYear="2019" box="[865,1018,333,359]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="chamorroi">
<emphasis id="B91FEA9D3C462E654113FEABFC1DB271" box="[865,1018,333,359]" italics="true" pageId="34" pageNumber="235">M. chamorroi</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C6B4D0C3C462E654757FEABFF24B29D" authorityName="Sassi" authorityYear="2019" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="praetorius">
<emphasis id="B91FEA9D3C462E654757FEABFF24B29D" italics="true" pageId="34" pageNumber="235">M. praetorius</emphasis>
</taxonomicName>
by the following traits: higher pronotal /elytral ratio (namely, pronotum is relatively shorter and slightly larger), more lengthened antennae and male rear tarsi, pronotal punctation stronger and also visible in middle of disc, elytral punctation generally closer, interocular distance clearly narrower, ocular line well marked by an irregular rows of punctures clearly perceptible up to canthus, head surface shinier, with more pronounced reddish pattern and stronger and more sparsely arranged punctation. Besides, in the studied specimens of
<taxonomicName id="4C6B4D0C3C462E65418FFDE7FB7FB10D" authorityName="Sassi" authorityYear="2019" box="[1021,1176,513,539]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="chamorroi">
<emphasis id="B91FEA9D3C462E65418FFDE7FB7FB10D" box="[1021,1176,513,539]" italics="true" pageId="34" pageNumber="235">M. chamorroi</emphasis>
</taxonomicName>
the elytral dark stripes are partly reddish (totally black in
<taxonomicName id="4C6B4D0C3C462E654056FDC3FD27B129" authorityName="Sassi" authorityYear="2019" box="[548,704,550,575]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="praetorius">
<emphasis id="B91FEA9D3C462E654056FDC3FD27B129" box="[548,704,550,575]" italics="true" pageId="34" pageNumber="235">M. praetorius</emphasis>
</taxonomicName>
).
<taxonomicName id="4C6B4D0C3C462E6540ABFDC3FCB3B129" authorityName="Suffrian" authorityYear="1866" box="[729,852,549,575]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="B91FEA9D3C462E6540ABFDC3FCB3B129" box="[729,852,549,575]" italics="true" pageId="34" pageNumber="235">M. tarsalis</emphasis>
</taxonomicName>
is actually the closest species, despite the apparent clear difference in elytral color pattern of the available specimens. The most convincing difference in male is given by the interocular distance, much greater in
<taxonomicName id="4C6B4D0C3C462E6540FEFD8BFCCFB191" authorityName="Sassi" authorityYear="2019" box="[652,808,622,647]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="praetorius">
<emphasis id="B91FEA9D3C462E6540FEFD8BFCCFB191" box="[652,808,622,647]" italics="true" pageId="34" pageNumber="235">M. praetorius</emphasis>
</taxonomicName>
. Besides, in the latter species the frontal punctuation is much more irregularly arranged.
</paragraph>
</subSubSection>
<subSubSection id="C37165043C462E6442B5FD50FB09B10D" lastPageId="35" lastPageNumber="236" pageId="34" pageNumber="235" type="description">
<paragraph id="8BD4368F3C462E6542B5FD50FDF2B1E5" blockId="34.[151,1437,153,2015]" pageId="34" pageNumber="235">
<emphasis id="B91FEA9D3C462E6542B5FD50FE55B1D9" bold="true" box="[199,434,692,719]" italics="true" pageId="34" pageNumber="235">Description of male.</emphasis>
Habitus in
<figureCitation id="13502A0A3C462E654044FD50FD63B1D9" box="[566,644,693,719]" captionStart="FIGURES 1415" captionStartId="36.[151,264,1865,1890]" captionTargetBox="[242,1341,195,1787]" captionTargetId="figure@36.[199,1388,166,1845]" captionTargetPageId="36" captionText="FIGURES 1415. M. praetorius (14), M. sekerkai (15). 14ab: habitus; 14ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.6 mm]; 14f: sclerites of endophallus [0.4 mm]; 14g: spermatheca [0.5 mm]; 14h: antenna [1.6 mm]. 15ab: habitus; 15ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.9 mm]; 15f: sclerites of endophallus [0.5 mm]; 15g: spermatheca [0.9 mm]; 15h: antenna [1.9 mm]. 14ab: holotype; 15ab: holotype. scIIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.3763893" httpUri="https://zenodo.org/record/3763893/files/figure.png" pageId="34" pageNumber="235">Fig. 14</figureCitation>
ab (HT). BL = 5.0 mm, BW =
<quantity id="4C939B6A3C462E65419DFD50FBA0B1D9" box="[1007,1095,693,719]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="34" pageNumber="235" unit="mm" value="2.8">2.8 mm</quantity>
, PL =
<quantity id="4C939B6A3C462E6546E0FD50FB0CB1D9" box="[1170,1259,693,719]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="34" pageNumber="235" unit="mm" value="1.8">1.8 mm</quantity>
, PW =
<quantity id="4C939B6A3C462E65474DFD50FA7EB1D9" box="[1343,1433,693,719]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="34" pageNumber="235" unit="mm" value="2.6">2.6 mm</quantity>
. Interocular distance 18.0% of BL.
</paragraph>
<paragraph id="8BD4368F3C462E6542B5FD18FB6EB0B1" blockId="34.[151,1437,153,2015]" pageId="34" pageNumber="235">Head light brown with lighter border along internal rim of eyes. Labrum yellow. Vertex quite dull, bare, with few punctures scattered on yellow spot. Frontoclypeal area matt, bald with dense shallowly impressed fine punctures. Ocular lines visible only along the upper rim of eyes, on frons basically obliterated. Mid-cranial suture shaped as quite large, barely delimited hollow impression, so that surrounding surface is somewhat swollen. Antennae reddish with yellow patches on first five antennomeres, slightly darkened toward apical ones.</paragraph>
<paragraph id="8BD4368F3C462E6542B5FC54FE97B769" blockId="34.[151,1437,153,2015]" pageId="34" pageNumber="235">Pronotum yellow with two large longitudinal stripes extending from anterior margin to posterior one. Those stripes are black with posterior section red towards outside. Pronotal shape tronco-conical, rather lengthened. Lateral margins thin, almost not visible from above, regularly curved with maximum width just behind half length. Posterolateral impressions obliterated so that posterior margin not salient behind them. Surface regularly convex, moderately shiny with fine, shallow, scattered punctures on sides and along posterior margins, almost absent on central part of disc.</paragraph>
<paragraph id="8BD4368F3C462E6542B5FB6CFD63B7D1" blockId="34.[151,1437,153,2015]" pageId="34" pageNumber="235">Scutellum reddish, moderately shiny, bald, distinctly raised, apex truncated in a straight line. Surface almost regularly covered with fine micropunctures.</paragraph>
<paragraph id="8BD4368F3C462E6542B5FB34FECAB689" blockId="34.[151,1437,153,2015]" pageId="34" pageNumber="235">Elytron surface yellow with two longitudinal black stripes extending from anterior margin to apical clivus. Inner stripe in continuity with the pronotal one, fairly tapered posteriorly. Outer stripe slender, running over humerous, tapered towards apex. Suture narrowly black. Elytral outline parallel-sided, rather short, very weakly flattened on disc. Postscutellar area not raised. Humeral callus scarcely prominent, impunctate. Surface moderately shiny with fine, shallow punctures irregularly distributed in anterior inner part of disc, laterally arranged in irregular rows. Intervals flat.</paragraph>
<paragraph id="8BD4368F3C462E6542B5FA4CFDD5B6F1" blockId="34.[151,1437,153,2015]" pageId="34" pageNumber="235">Pygidium yellow with whitish apex, with two shallow impressions on sides, rather lustrous, covered with sparse shallow punctures and whitish setae.</paragraph>
<paragraph id="8BD4368F3C462E6542B5FA14FF02B5A9" blockId="34.[151,1437,153,2015]" pageId="34" pageNumber="235">Inferior parts of thorax light brown, prosternum, mesosternum, metaepisterna and anterior part of metasternum perceptibly darker. Hypomera shallowly punctured, bare. Mesoepimera and mesoepisterna almost bare with scarce shallow punctures. Metaepisterna and metasternum with irregular punctation and wrinkles, bearing sparse setae. Prosternal process coarsely punctured with long setae and raised short triangular apex. Abdominal ventrites completely yellow, shallowly and sparsely punctured, with long, sparse setae. Legs light brownish, with tarsi slightly darker.</paragraph>
<paragraph id="8BD4368F3C462E6442B5F92CFBF9B3C1" blockId="34.[151,1437,153,2015]" lastBlockId="35.[151,1437,153,539]" lastPageId="35" lastPageNumber="236" pageId="34" pageNumber="235">
Median depression on fifth abdominal ventrite well impressed, bald and shiny. Posterior margin of fifth abdominal ventrite weakly notched. Median lobe of aedeagus (
<figureCitation id="13502A0A3C462E654145F908FC61B411" box="[823,902,1773,1799]" captionStart="FIGURES 1415" captionStartId="36.[151,264,1865,1890]" captionTargetBox="[242,1341,195,1787]" captionTargetId="figure@36.[199,1388,166,1845]" captionTargetPageId="36" captionText="FIGURES 1415. M. praetorius (14), M. sekerkai (15). 14ab: habitus; 14ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.6 mm]; 14f: sclerites of endophallus [0.4 mm]; 14g: spermatheca [0.5 mm]; 14h: antenna [1.6 mm]. 15ab: habitus; 15ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.9 mm]; 15f: sclerites of endophallus [0.5 mm]; 15g: spermatheca [0.9 mm]; 15h: antenna [1.9 mm]. 14ab: holotype; 15ab: holotype. scIIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.3763893" httpUri="https://zenodo.org/record/3763893/files/figure.png" pageId="34" pageNumber="235">Fig. 14</figureCitation>
ce) rather small, cylindrical with apex scarcely separated from the rest of aedeagus, blunt. In lateral view apex faintly bent ventrally. Hairy dents almost obliterated, barely visible, bearing scattered, rather long setae. Aedeagal ventral surface not swollen in lateral view, devoid of particular structures. Endophallus (
<figureCitation id="13502A0A3C462E654051F8BCFD98B462" box="[547,639,1881,1908]" captionStart="FIGURES 1415" captionStartId="36.[151,264,1865,1890]" captionTargetBox="[242,1341,195,1787]" captionTargetId="figure@36.[199,1388,166,1845]" captionTargetPageId="36" captionText="FIGURES 1415. M. praetorius (14), M. sekerkai (15). 14ab: habitus; 14ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.6 mm]; 14f: sclerites of endophallus [0.4 mm]; 14g: spermatheca [0.5 mm]; 14h: antenna [1.6 mm]. 15ab: habitus; 15ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.9 mm]; 15f: sclerites of endophallus [0.5 mm]; 15g: spermatheca [0.9 mm]; 15h: antenna [1.9 mm]. 14ab: holotype; 15ab: holotype. scIIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.3763893" httpUri="https://zenodo.org/record/3763893/files/figure.png" pageId="34" pageNumber="235">Fig. 14f</figureCitation>
) with sclerite I very small, almost reduced to only sharp, slender denticle. Dorsal spicule strongly reduced to simple membranous, ribbon-like, fairly pigmented lamina. Sclerite II distinctly bent at base and moderately tapered towards apex. Arch of sclerite III short, scarcely raised. Apex of sclerite III barely separable from arch, arched and pointed, regularly tapered, not expanded on its proximal half, with long, straight, sharpened tip. Branches of sclerite IV longer than sclerite III in the folded-up structure, almost straight, with blunt, almost symmetrical, microdenticulate apex and surface fairly rugose.
</paragraph>
<paragraph id="8BD4368F3C472E6442B5FF04FF06B209" blockId="35.[151,1437,153,539]" pageId="35" pageNumber="236">
Female. BL =
<quantity id="4C939B6A3C472E644318FF05FE25B3ED" box="[362,450,224,251]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="35" pageNumber="236" unit="mm" value="5.8">5.8 mm</quantity>
, BW =
<quantity id="4C939B6A3C472E64406BFF04FD95B3ED" box="[537,626,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="35" pageNumber="236" unit="mm" value="3.3">3.3 mm</quantity>
, PL =
<quantity id="4C939B6A3C472E6440CEFF04FCA1B3ED" box="[700,838,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="35" pageNumber="236" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.82.1 mm</quantity>
, PW =
<quantity id="4C939B6A3C472E6441E8FF04FBC3B3ED" box="[922,1060,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="35" pageNumber="236" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.72.9 mm</quantity>
. Interocular distance 15.517.2% of BL.
</paragraph>
<paragraph id="8BD4368F3C472E6442B5FECCFBB7B255" blockId="35.[151,1437,153,539]" box="[199,1104,297,323]" pageId="35" pageNumber="236">In the two examined specimens pronotal longitudinal stripes completely reddish.</paragraph>
<paragraph id="8BD4368F3C472E6442B5FEA8FB09B10D" blockId="35.[151,1437,153,539]" pageId="35" pageNumber="236">
Fifth abdominal ventrite in females with quite deep, slightly transverse pit. Bottom of pit subluster with few tiny punctures. Vasculum of spermatheca (
<figureCitation id="13502A0A3C472E644032FE94FD78B29D" box="[576,671,369,395]" captionStart="FIGURES 1415" captionStartId="36.[151,264,1865,1890]" captionTargetBox="[242,1341,195,1787]" captionTargetId="figure@36.[199,1388,166,1845]" captionTargetPageId="36" captionText="FIGURES 1415. M. praetorius (14), M. sekerkai (15). 14ab: habitus; 14ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.6 mm]; 14f: sclerites of endophallus [0.4 mm]; 14g: spermatheca [0.5 mm]; 14h: antenna [1.6 mm]. 15ab: habitus; 15ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.9 mm]; 15f: sclerites of endophallus [0.5 mm]; 15g: spermatheca [0.9 mm]; 15h: antenna [1.9 mm]. 14ab: holotype; 15ab: holotype. scIIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.3763893" httpUri="https://zenodo.org/record/3763893/files/figure.png" pageId="35" pageNumber="236">
Fig.
<quantity id="4C939B6A3C472E644006FE94FD78B29D" box="[628,671,369,395]" metricMagnitude="-2" metricUnit="kg" metricValue="1.4" pageId="35" pageNumber="236" unit="g" value="14.0">14g</quantity>
</figureCitation>
) robust, scarcely pigmented with straight proximal branch not swollen at base, short, swollen distal branch and botton-like apex. Ampulla not pigmented, sticking out from basal rim of proximal branch, therefore not shifted on dorsal side of vasculum. Duct insertion and sperm gland insertion barely distinct. Duct uniform in size, slender, coiled with coils thick, rather regularly arranged. Distal not coiled portion of duct rather short, straight. Insertion on bursa copulatrix small, bent, neither swollen nor pigmented.
</paragraph>
</subSubSection>
</treatment>
</document>