486 lines
57 KiB
XML
486 lines
57 KiB
XML
<document id="CE1A0B50477B5D98EF8835A654A3D055" ID-DOI="10.11646/zootaxa.3869.2.7" ID-GBIF-Dataset="31b52d1d-25b5-4745-b515-e93d6649324a" ID-ISSN="1175-5326" ID-Zenodo-Dep="229003" ID-ZooBank="A984C9E6-4785-4E2C-88C6-52324C1554D0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460923774870" checkinUser="plazi" docAuthor="Kehlmaier, Christian, Gharali, Babak & Jahromi, Bahareh Majnon" docDate="2014" docId="039387F4FFBEFF82D9C8FF4BEEDFF838" docLanguage="en" docName="zt03869p179.pdf" docOrigin="Zootaxa 3869 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Corononcodes ziegleri Kehlmaier, Gharali & Majnon Jahromi, spec. nov." docType="treatment" docVersion="7" lastPageNumber="177" masterDocId="FFAAFF8CFFBCFF84D95FFFDCED50FF8B" masterDocTitle="A new Corononcodes Speiser from the Palaearctic Region, with a key to species (Diptera: Acroceridae)" masterLastPageNumber="179" masterPageNumber="171" pageNumber="173" updateTime="1698362211008" updateUser="plazi">
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<mods:mods id="5E399F347067C8A4B908110878CA72C9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1DBE945D4CD3C0FA0DACBF87D3BD811E">
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<mods:title id="C3376CD9D29D88E74711FEC59F8FE141">A new Corononcodes Speiser from the Palaearctic Region, with a key to species (Diptera: Acroceridae)</mods:title>
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</mods:titleInfo>
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<mods:name id="FCECF67F5C78C5394401DAAB7607E3B2" type="personal">
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<mods:role id="CD5F1605F53A7407294EF2CC3D764EC8">
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<mods:roleTerm id="D4244B970682BB315A311CD1639F021F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="77C540160EF9FE06EE9D344CC59BD7D5">Kehlmaier, Christian</mods:namePart>
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</mods:name>
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</mods:role>
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<mods:namePart id="4C3C0C301EC5ED27F824C51ABF8CBB09">Gharali, Babak</mods:namePart>
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</mods:name>
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<mods:name id="F01D4E59F6B8099AA5011EDEBE105178" type="personal">
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<mods:roleTerm id="D3E5E88F6FC6792500C3FC6909E4AA35">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EE117CB76105679B12BAEF4E3E82F252">Jahromi, Bahareh Majnon</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1FEF44A794011E9FFCC6D93A9CAD7DE5">text</mods:typeOfResource>
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<mods:title id="8089759C2FADF5A4AEFED6C53A083B14">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="74B3E0AB40CCDD89E65AD4C57209871F">
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<mods:date id="3CB67285C10A240BA5C5EF70C8CA2F28">2014</mods:date>
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<mods:detail id="9BA641D238038E23D06D30A5A1D01A59" type="volume">
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<mods:number id="19B5E5A5E62C5B4111B3CCB6953CB787">3869</mods:number>
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</mods:detail>
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<mods:classification id="1AD792532E9E0583C83EF0F1BE0BF53D">journal article</mods:classification>
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<mods:identifier id="DB9735CCD1D13D67AC8A3529A8E7DFD5" type="DOI">10.11646/zootaxa.3869.2.7</mods:identifier>
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<treatment id="039387F4FFBEFF82D9C8FF4BEEDFF838" ID-DOI="http://doi.org/10.5281/zenodo.6139137" ID-GBIF-Taxon="119589394" ID-Zenodo-Dep="6139137" LSID="urn:lsid:plazi:treatment:039387F4FFBEFF82D9C8FF4BEEDFF838" httpUri="http://treatment.plazi.org/id/039387F4FFBEFF82D9C8FF4BEEDFF838" lastPageId="6" lastPageNumber="177" pageId="2" pageNumber="173">
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<subSubSection id="C3206569FFBEFF86D9C8FF4BEC9AFF5F" pageId="2" pageNumber="173" type="nomenclature">
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<paragraph id="8B8536E2FFBEFF86D9C8FF4BEEA0FF3B" blockId="2.[151,1008,151,213]" box="[151,1008,151,176]" pageId="2" pageNumber="173">
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<heading id="D0CD818EFFBEFF86D9C8FF4BEEA0FF3B" bold="true" box="[151,1008,151,176]" fontSize="10" level="2" pageId="2" pageNumber="173" reason="2">
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<emphasis id="B94EEAF0FFBEFF86D9C8FF4BEEA0FF3B" bold="true" box="[151,1008,151,176]" pageId="2" pageNumber="173">
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<taxonomicName id="4C3A4D61FFBEFF86D9C8FF4BEE27FF3B" authority="Kehlmaier, Gharali & Majnon Jahromi" authorityName="Kehlmaier, Gharali & Majnon Jahromi" box="[151,887,151,176]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="173" phylum="Arthropoda" rank="species" species="ziegleri" status="sp. nov.">
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<emphasis id="B94EEAF0FFBEFF86D9C8FF4BECC8FF3B" bold="true" box="[151,408,151,176]" italics="true" pageId="2" pageNumber="173">Corononcodes ziegleri</emphasis>
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Kehlmaier, Gharali & Majnon Jahromi
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</taxonomicName>
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<taxonomicNameLabel id="A27D578BFFBEFF86DA21FF4BEEA0FF3B" box="[894,1008,151,176]" pageId="2" pageNumber="173" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8536E2FFBEFF86D9C8FF60EC9AFF5F" blockId="2.[151,1008,151,213]" box="[151,458,188,213]" pageId="2" pageNumber="173">(figs 2A–B, 2E, 3B, 4A–B)</paragraph>
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</subSubSection>
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<subSubSection id="C3206569FFBEFF86D9C8FED8ECDFFE5B" pageId="2" pageNumber="173" type="materials_examined">
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<paragraph id="8B8536E2FFBEFF86D9C8FED8ECDFFE5B" blockId="2.[151,1437,260,2021]" pageId="2" pageNumber="173">
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<emphasis id="B94EEAF0FFBEFF86D9C8FED8EC56FE96" bold="true" box="[151,262,260,285]" pageId="2" pageNumber="173">Material.</emphasis>
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<typeStatus id="54818840FFBEFF86D84DFED8ECFAFE97" box="[274,426,260,284]" pageId="2" pageNumber="173" type="holotype">HOLOTYPE</typeStatus>
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||
: ♂,
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||
<collectingCountry id="F32D7672FFBEFF86D8BDFED9EF42FE97" box="[482,530,261,284]" name="Iran" pageId="2" pageNumber="173">Iran</collectingCountry>
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||
, Ghazvin Province, Ghazvin city, Zarabad village, Plant Medicinal Research Station,
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<geoCoordinate id="EE0E5025FFBEFF86D9ABFEFBEC01FECB" box="[244,337,295,320]" direction="north" orientation="latitude" pageId="2" pageNumber="173" precision="925" value="36.466667">36°28'N</geoCoordinate>
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<geoCoordinate id="EE0E5025FFBEFF86D807FEFBECE4FECB" box="[344,436,295,320]" direction="east" orientation="longitude" pageId="2" pageNumber="173" precision="925" value="50.4">50°24'E</geoCoordinate>
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,
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<quantity id="4CC29B07FFBEFF86D89FFEFBEF48FEB4" box="[448,536,295,320]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="2" pageNumber="173" unit="m" value="1520.0">1520 m</quantity>
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,
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<date id="FF841022FFBEFF86DB7BFEFBEFAFFECB" box="[548,767,295,320]" pageId="2" pageNumber="173" value="2009-06-05" valueMax="2009-08-11" valueMin="2009-06-05">5.VI.–11.VIII.2009</date>
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||
, leg. Babak Gharali, coll.
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<collectionCode id="ED2BAE27FFBEFF86DD79FEF4E927FECB" box="[1062,1143,296,320]" pageId="2" pageNumber="173">SMTD</collectionCode>
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.
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<typeStatus id="54818840FFBEFF86DDDDFEF4E871FECB" box="[1154,1313,296,320]" pageId="2" pageNumber="173" type="paratype">PARATYPES</typeStatus>
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: same data as
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<typeStatus id="54818840FFBEFF86D9E6FE90EC4BFEEE" box="[185,283,332,357]" pageId="2" pageNumber="173" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="ED2BAE27FFBEFF86D803FE91ECE4FEEF" box="[348,436,333,356]" pageId="2" pageNumber="173">MNHU</collectionCode>
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,
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<specimenCount id="9D3CFD6BFFBEFF86D89CFE90ECB6FEEF" box="[451,486,332,356]" pageId="2" pageNumber="173" type="generic">1♂</specimenCount>
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<collectionCode id="ED2BAE27FFBEFF86D8AEFE90EF6FFEEF" box="[497,575,332,356]" pageId="2" pageNumber="173">SMTD</collectionCode>
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(
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<collectionCode id="ED2BAE27FFBEFF86DB0EFE90EFC0FEEF" LSID="urn:lsid:biocol.org:col:15536" box="[593,656,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="2" pageNumber="173">DNA</collectionCode>
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voucher CK611, GenBank voucher
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<accessionNumber id="9469AB01FFBEFF86DD66FE90E9E9FEEF" box="[1081,1209,332,356]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HG800023" pageId="2" pageNumber="173">HG800023</accessionNumber>
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), 1♀
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<collectionCode id="ED2BAE27FFBEFF86DDA1FE90E81CFEEF" box="[1278,1356,332,356]" pageId="2" pageNumber="173">SMTD</collectionCode>
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(
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<collectionCode id="ED2BAE27FFBEFF86DC01FE90E8CDFEEF" LSID="urn:lsid:biocol.org:col:15536" box="[1374,1437,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="2" pageNumber="173">DNA</collectionCode>
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voucher CK612 GenBank voucher
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<accessionNumber id="9469AB01FFBEFF86DB74FEACEFFCFE03" box="[555,684,368,392]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HG800024" pageId="2" pageNumber="173">HG800024</accessionNumber>
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));
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<collectingCountry id="F32D7672FFBEFF86DB93FEACEFACFE0C" box="[716,764,368,391]" name="Iran" pageId="2" pageNumber="173">Iran</collectingCountry>
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,
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||
<collectingCountry id="F32D7672FFBEFF86DA56FEACEE69FE0C" box="[777,825,368,391]" name="Iran" pageId="2" pageNumber="173">Fars</collectingCountry>
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||
Province, Jahrom,
|
||
<geoCoordinate id="EE0E5025FFBEFF86DD45FEACE926FE03" box="[1050,1142,368,392]" direction="north" orientation="latitude" pageId="2" pageNumber="173" precision="925" value="28.5">28°30'N</geoCoordinate>
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||
53˚35'E,
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||
<date id="FF841022FFBEFF86DDB8FEB3E8C8FE03" box="[1255,1432,367,392]" pageId="2" pageNumber="173" value="2013-03-11" valueMax="2013-03-25" valueMin="2013-03-11">11.–25.III.2013</date>
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||
, leg. Bahareh Majnon Jahromi (
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||
<specimenCount id="9D3CFD6BFFBEFF86DB5BFE48EF7EFE27" box="[516,558,404,428]" pageId="2" pageNumber="173" type="generic">2♂</specimenCount>
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||
1♀
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||
<collectionCode id="ED2BAE27FFBEFF86DB0BFE48EFF0FE27" box="[596,672,404,428]" pageId="2" pageNumber="173">PCCK</collectionCode>
|
||
,
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||
<specimenCount id="9D3CFD6BFFBEFF86DBF0FE48EF87FE27" box="[687,727,404,428]" pageId="2" pageNumber="173" type="generic">1♂</specimenCount>
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||
1♀
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||
<collectionCode id="ED2BAE27FFBEFF86DA5FFE48EE1CFE27" LSID="urn:lsid:biocol.org:col:33156" box="[768,844,404,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="2" pageNumber="173">CSCA</collectionCode>
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,
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||
<specimenCount id="9D3CFD6BFFBEFF86DA03FE48EE2FFE27" box="[860,895,404,428]" pageId="2" pageNumber="173" type="generic">3♂</specimenCount>
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<collectionCode id="ED2BAE27FFBEFF86DAD6FE48EE87FE27" box="[905,983,404,428]" pageId="2" pageNumber="173">SMTD</collectionCode>
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(
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<specimenCount id="9D3CFD6BFFBEFF86DAB2FE48E941FE27" box="[1005,1041,404,428]" pageId="2" pageNumber="173" type="generic">1♂</specimenCount>
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,
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||
<collectionCode id="ED2BAE27FFBEFF86DD7EFE48E90EFE27" LSID="urn:lsid:biocol.org:col:15536" box="[1057,1118,404,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="2" pageNumber="173">DNA</collectionCode>
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||
voucher CK698 (GenBank voucher
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||
<accessionNumber id="9469AB01FFBEFF86D9A7FE6BEC29FE5B" box="[248,377,439,464]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HG973062" pageId="2" pageNumber="173">HG973062</accessionNumber>
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||
)).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3206569FFBEFF86D998FE00E98BFD93" pageId="2" pageNumber="173" type="etymology">
|
||
<paragraph id="8B8536E2FFBEFF86D998FE00E98BFD93" blockId="2.[151,1437,260,2021]" pageId="2" pageNumber="173">
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||
<emphasis id="B94EEAF0FFBEFF86D998FE00EC1EFE7E" bold="true" box="[199,334,476,501]" pageId="2" pageNumber="173">Etymology.</emphasis>
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The new species is dedicated to Joachim Ziegler, curator of the
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<taxonomicName id="4C3A4D61FFBEFF86DD7CFE00E927FE7E" box="[1059,1143,476,501]" kingdom="Plantae" order="Diptera" pageId="2" pageNumber="173" rank="order">Diptera</taxonomicName>
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||
collection at the Museum für Naturkunde in Berlin,
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||
<collectingCountry id="F32D7672FFBEFF86D8E5FDDCEF73FD93" box="[442,547,512,536]" name="Germany" pageId="2" pageNumber="173">Germany</collectingCountry>
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||
, in acknowledgment for his substantial support of our studies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3206569FFBEFF81D998FDF8E82AFE96" lastPageId="5" lastPageNumber="176" pageId="2" pageNumber="173" type="description">
|
||
<paragraph id="8B8536E2FFBEFF86D998FDF8EC4BF9DD" blockId="2.[151,1437,260,2021]" pageId="2" pageNumber="173">
|
||
<emphasis id="B94EEAF0FFBEFF86D998FDF8ECE6FDB6" bold="true" box="[199,438,548,573]" pageId="2" pageNumber="173">Description of male.</emphasis>
|
||
Body length (excluding antennae): 2.0–
|
||
<quantity id="4CC29B07FFBEFF86DA28FDF8EE9BFDB7" box="[887,971,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="173" unit="mm" value="2.8">2.8 mm</quantity>
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||
(
|
||
<emphasis id="B94EEAF0FFBEFF86DA85FDF9EEB8FDB7" box="[986,1000,549,572]" italics="true" pageId="2" pageNumber="173">n</emphasis>
|
||
= 9); wing length:
|
||
<quantity id="4CC29B07FFBEFF86DDE2FDF8E813FDB7" box="[1213,1347,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="2" pageNumber="173" unit="mm" value="2.1" valueMax="2.4" valueMin="1.8">1.8–2.4 mm</quantity>
|
||
(
|
||
<emphasis id="B94EEAF0FFBEFF86DC0DFDF9E830FDB7" box="[1362,1376,549,572]" italics="true" pageId="2" pageNumber="173">n</emphasis>
|
||
= 9); wing width:
|
||
<quantity id="4CC29B07FFBEFF86D875FD94ECE4FDD4" box="[298,436,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="2" pageNumber="173" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.8–1.2 mm</quantity>
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||
(
|
||
<emphasis id="B94EEAF0FFBEFF86D899FD95EC84FDEB" box="[454,468,585,608]" italics="true" pageId="2" pageNumber="173">n</emphasis>
|
||
= 9).
|
||
<emphasis id="B94EEAF0FFBEFF86DB7FFD9BEF30FDEB" bold="true" box="[544,608,583,608]" pageId="2" pageNumber="173">Head</emphasis>
|
||
. Eyes dark reddish brown, dichoptic and bare; occiput visible in dorsal view, black, with white pile; ocellar tubercle slightly surpassing height of scape in lateral view, with short and brownish pile, evenly convex and without ridges; with three ocelli, median ocellus about one-sixth to one-fourth size (HT) of lateral ocellus, situated on anterior face of ocellar tubercle and thus only visible anteriorly (fig. 2E), and not in dorsal or lateral view (use high magnification and powerful light source); scape black, with few white dorsal setae; pedicel yellowish brown, about half length of scape, with short white dorsal setae; flagellum (fig. 2A) yellowish brown, with short white dorsal setae at base, in lateral view subangular apically especially at lower corner, somewhat higher in basal quarter and only weakly bent after basal fourth, length
|
||
<quantity id="4CC29B07FFBEFF86DDC7FC98E871FCD7" box="[1176,1313,836,861]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="2" pageNumber="173" unit="mm" value="0.75" valueMax="0.9" valueMin="0.6">0.6–0.9 mm</quantity>
|
||
(
|
||
<emphasis id="B94EEAF0FFBEFF86DC6BFC99E812FCD7" box="[1332,1346,837,860]" italics="true" pageId="2" pageNumber="173">n</emphasis>
|
||
= 9) or about twice height of compound eye; frons light to mid brown, almost parallel sided, minimum width three ommatidial facets; clypeus white (best seen when proboscis is extended), membranous; proboscis small and black; palpi minute, white with darkened centre.
|
||
<emphasis id="B94EEAF0FFBEFF86DB31FC73EF97FC43" bold="true" box="[622,711,943,968]" pageId="2" pageNumber="173">Thorax</emphasis>
|
||
. Dark brown to black, pleura and postalar callus medium to dark brown; pile short (about same length as on occiput) and white, but pleura almost entirely bare.
|
||
<emphasis id="B94EEAF0FFBEFF86DDB9FC08E84EFC66" bold="true" box="[1254,1310,980,1005]" pageId="2" pageNumber="173">Legs</emphasis>
|
||
. Coxa and trochanter light to medium brown, with short white pile (about same length as on occiput), especially on anterior surface of coxa; femur mainly light to medium brown but white in apical fourth to fifth, with short white pile; tibia white with brown ventral spot at end of basal half (absent on hind tibia in HT), with short white pile, no apical spur or row of small sharply pointed projections present (at 144x magnification); tarsomeres white, with short white pile, longest setae dorsally at apex of distitarsus; claws medium to dark brown; pulvilli and empodium white, slightly longer than half length of claws.
|
||
<emphasis id="B94EEAF0FFBEFF86DB39FB70EFF7FB4E" bold="true" box="[614,679,1196,1221]" pageId="2" pageNumber="173">Wing</emphasis>
|
||
.
|
||
<collectingCountry id="F32D7672FFBEFF86DBEAFB70EF84FB4F" box="[693,724,1196,1220]" name="American Samoa" pageId="2" pageNumber="173">As</collectingCountry>
|
||
in fig. 3A; membrane hyaline, but cell sc white opaque, hardly any microtrichia present; wing venation reduced; veins translucent except brownish at wing base; no longitudinal vein reaching posterior wing margin; veins M1+2 and M3+4 incomplete basally; two folds present that substitute basal extension of veins M1+2 and vein A1+CuA2 (best seen in female); two cross-veins present (nomenclature uncertain); upper calypter white opaque (can be less pronounced than in cell sc), surface covered with white microtrichia; lower calypter hyaline, rim translucent, surface covered with horizontal streaks appearing to be undulations; halter with almost translucent stem and bright white knob.
|
||
<emphasis id="B94EEAF0FFBEFF86DBA5FA55EE3EFA29" bold="true" box="[762,878,1417,1442]" pageId="2" pageNumber="173">Abdomen</emphasis>
|
||
. Medium to dark brown (sternites can be partly paler), dorsally widest at tergite 3, in total about 1.5 times longer than wide, with white pile slightly longer than on thorax (
|
||
<quantity id="4CC29B07FFBEFF86D9B3FA0DECC5FA61" box="[236,405,1489,1514]" metricMagnitude="-5" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="173" unit="mm" value="0.045" valueMax="0.05" valueMin="0.04">0.04–0.05 mm</quantity>
|
||
); cerci white, with white pile longer than on rest of abdomen; genitalia as in figs 4A–B, with aedeagus, parameral sheath of aedeagus, epandrium, gonostyli, ejaculatory apodeme, and small parts of gonocoxite medium to dark brown; larger part of the gonocoxite translucent. Gonostyli small and pointed, fused with gonocoxite.
|
||
</paragraph>
|
||
<paragraph id="8B8536E2FFBEFF86D998F9BDEF3AF86E" blockId="2.[151,1437,260,2021]" pageId="2" pageNumber="173">
|
||
Female differing from the male as follows. Body length (excluding antennae):
|
||
<quantity id="4CC29B07FFBEFF86DD39F9BEE9A2F9F2" box="[1126,1266,1634,1658]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="4.2" metricValueMin="2.4" pageId="2" pageNumber="173" unit="mm" value="3.3" valueMax="4.2" valueMin="2.4">2.4–4.2 mm</quantity>
|
||
(
|
||
<emphasis id="B94EEAF0FFBEFF86DC5AF9BEE843F9F2" box="[1285,1299,1634,1657]" italics="true" pageId="2" pageNumber="173">n</emphasis>
|
||
= 3); wing length:
|
||
<quantity id="4CC29B07FFBEFF86D9B1F95AEC28F916" box="[238,376,1670,1694]" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.8" metricValueMin="3.4" pageId="2" pageNumber="173" unit="mm" value="3.5999999999999996" valueMax="3.8" valueMin="3.4">3.4–3.8 mm</quantity>
|
||
(
|
||
<emphasis id="B94EEAF0FFBEFF86D8D4F95AECC9F916" box="[395,409,1670,1693]" italics="true" pageId="2" pageNumber="173">n</emphasis>
|
||
= 2); wing width:
|
||
<quantity id="4CC29B07FFBEFF86DB27F959EE51F916" box="[632,769,1669,1694]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="2" pageNumber="173" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 mm</quantity>
|
||
(
|
||
<emphasis id="B94EEAF0FFBEFF86DA4BF95AEE72F916" box="[788,802,1670,1693]" italics="true" pageId="2" pageNumber="173">n</emphasis>
|
||
= 2).
|
||
<emphasis id="B94EEAF0FFBEFF86DA32F959EEFDF915" bold="true" box="[877,941,1669,1694]" pageId="2" pageNumber="173">Head</emphasis>
|
||
. Pile on occiput darker (brownish); with 3 ocelli; clypeus only visible as a narrow white band that might be due to the retracted state of the proboscis; pedicel and flagellum dark brown; flagellum (fig. 2B) straight with dorsal setae in basal half, laterally at base as broad as pedicel and evenly tapering towards small rounded apex, length
|
||
<quantity id="4CC29B07FFBEFF86DA31F92DEE83F882" box="[878,979,1777,1802]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="2" pageNumber="173" unit="mm" value="0.36">0.36 mm</quantity>
|
||
(
|
||
<emphasis id="B94EEAF0FFBEFF86DABCF92EEEA1F882" box="[995,1009,1778,1801]" italics="true" pageId="2" pageNumber="173">n</emphasis>
|
||
= 1) or slightly shorter than height of compound eye.
|
||
<emphasis id="B94EEAF0FFBEFF86D80AF8C9ECFEF8A5" bold="true" box="[341,430,1813,1838]" pageId="2" pageNumber="173">Thorax</emphasis>
|
||
. Pile brownish.
|
||
<emphasis id="B94EEAF0FFBEFF86DB28F8C9EFFFF8A5" bold="true" box="[631,687,1813,1838]" pageId="2" pageNumber="173">Legs</emphasis>
|
||
. Entirely medium to dark brown, but femoral-tibial joints and tarsomeres lighter.
|
||
<emphasis id="B94EEAF0FFBEFF86D82FF8E5ECE1F8D9" bold="true" box="[368,433,1849,1874]" pageId="2" pageNumber="173">Wing</emphasis>
|
||
. Membrane including upper and lower calypters rather evenly brownish infuscated (fig. 3B), but with obvious hyaline folds (basal extension of vein M1+2 and where vein A1+CuA2 would be expected). Wing venation medium brown; halter with stem and knob mainly brown, but knob bright white at top and bottom.
|
||
<emphasis id="B94EEAF0FFBEFF86D9C8F874EC5CF84A" bold="true" box="[151,268,1960,1985]" pageId="2" pageNumber="173">Abdomen</emphasis>
|
||
. Ovate, only slightly longer than wide, widest at tergite 3 and 4, pile brownish (
|
||
<quantity id="4CC29B07FFBEFF86DDFCF874E85FF84A" box="[1187,1295,1960,1985]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="173" unit="mm" value="0.06">0.06 mm</quantity>
|
||
). Terminalia placed at apex of abdomen, not dissected.
|
||
</paragraph>
|
||
<caption id="DF45666AFFBFFF87D9C8F88BE9E1F86C" httpUri="https://zenodo.org/record/229005/files/figure.png" pageId="3" pageNumber="174" targetBox="[205,1362,193,1818]" targetPageId="3">
|
||
<paragraph id="8B8536E2FFBFFF87D9C8F88BE9E1F86C" blockId="3.[151,1436,1879,2023]" pageId="3" pageNumber="174">
|
||
<emphasis id="B94EEAF0FFBFFF87D9C8F88BEC44F8E7" bold="true" box="[151,276,1879,1901]" pageId="3" pageNumber="174">FIGURE 2.</emphasis>
|
||
A) Male holotype of
|
||
<taxonomicName id="4C3A4D61FFBFFF87D8AAF88BEF8BF8E6" box="[501,731,1879,1901]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="ziegleri" status="sp. nov.">
|
||
<emphasis id="B94EEAF0FFBFFF87D8AAF88BEF8BF8E6" box="[501,731,1879,1901]" italics="true" pageId="3" pageNumber="174">Corononcodes ziegleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94EEAF0FFBFFF87DBBCF88BEE1AF8E7" bold="true" box="[739,842,1879,1900]" pageId="3" pageNumber="174">
|
||
<taxonomicNameLabel id="A27D578BFFBFFF87DBBCF88BEE1AF8E7" box="[739,842,1879,1900]" pageId="3" pageNumber="174" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Scale bar: 0.5 mm. B) Female paratype of
|
||
<taxonomicName id="4C3A4D61FFBFFF87DC56F88BEDB3F800" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="ziegleri" status="sp. nov.">
|
||
<emphasis id="B94EEAF0FFBFFF87DC56F88BEDB3F800" italics="true" pageId="3" pageNumber="174">Corononcodes ziegleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D578BFFBFFF87D9B3F8AAEC04F800" box="[236,340,1910,1931]" pageId="3" pageNumber="174" rank="species">spec. nov.</taxonomicNameLabel>
|
||
from the type location. Scale bar: 0.5 mm. C) Male
|
||
<taxonomicName id="4C3A4D61FFBFFF87DADBF8A9E9FAF800" authority="Bezzi" authorityName="Bezzi" box="[900,1194,1909,1931]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="siculus">
|
||
<emphasis id="B94EEAF0FFBFFF87DADBF8A9E937F800" box="[900,1127,1909,1931]" italics="true" pageId="3" pageNumber="174">Corononcodes siculus</emphasis>
|
||
Bezzi
|
||
</taxonomicName>
|
||
from Tenerife (Canary Islands, Spain). Scale bar: 0.5 mm. D) Male holotype of
|
||
<taxonomicName id="4C3A4D61FFBFFF87DB80F848E97EF821" authority="Bezzi. No" authorityName="Bezzi. No" box="[735,1070,1940,1962]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="siculus">
|
||
<emphasis id="B94EEAF0FFBFFF87DB80F848EE90F821" box="[735,960,1940,1962]" italics="true" pageId="3" pageNumber="174">Corononcodes siculus</emphasis>
|
||
Bezzi. No
|
||
</taxonomicName>
|
||
scale. E) Head of male holotype of
|
||
<taxonomicName id="4C3A4D61FFBFFF87D9C8F86FEC2EF842" box="[151,382,1971,1993]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="ziegleri" status="sp. nov.">
|
||
<emphasis id="B94EEAF0FFBFFF87D9C8F86FEC2EF842" box="[151,382,1971,1993]" italics="true" pageId="3" pageNumber="174">Corononcodes ziegleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D578BFFBFFF87D8D8F868ECBEF842" box="[391,494,1972,1993]" pageId="3" pageNumber="174" rank="species">spec. nov.</taxonomicNameLabel>
|
||
in anterior view, showing mid ocellus and frons. Scale bar: 0.1 mm. F) Scanning electron micrograph of head of male holotype of
|
||
<taxonomicName id="4C3A4D61FFBFFF87DB6EF80DEE00F86C" authority="Bezzi" authorityName="Bezzi" box="[561,848,2001,2023]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="siculus">
|
||
<emphasis id="B94EEAF0FFBFFF87DB6EF80DEE40F86C" box="[561,784,2001,2023]" italics="true" pageId="3" pageNumber="174">Corononcodes siculus</emphasis>
|
||
Bezzi
|
||
</taxonomicName>
|
||
in lateral view. Scale bar: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF45666AFFB8FF80D9C8FAC1E81BFA04" httpUri="https://zenodo.org/record/229006/files/figure.png" pageId="4" pageNumber="175" targetBox="[212,1374,201,1287]" targetPageId="4">
|
||
<paragraph id="8B8536E2FFB8FF80D9C8FAC1E81BFA04" blockId="4.[151,1437,1309,1423]" pageId="4" pageNumber="175">
|
||
<emphasis id="B94EEAF0FFB8FF80D9C8FAC1EC42FAB9" bold="true" box="[151,274,1309,1331]" pageId="4" pageNumber="175">FIGURE 3.</emphasis>
|
||
Wing venation of
|
||
<taxonomicName id="4C3A4D61FFB8FF80D890FAC1EF32FAB8" box="[463,610,1309,1331]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94EEAF0FFB8FF80D890FAC1EF32FAB8" box="[463,610,1309,1331]" italics="true" pageId="4" pageNumber="175">Corononcodes</emphasis>
|
||
</taxonomicName>
|
||
spp. A) Right wing of male
|
||
<taxonomicName id="4C3A4D61FFB8FF80DADCFAC1E840FAB8" authority="Bezzi (Tenerife)" authorityName="Bezzi (Tenerife" box="[899,1296,1309,1331]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="siculus">
|
||
<emphasis id="B94EEAF0FFB8FF80DADCFAC1E932FAB8" box="[899,1122,1309,1331]" italics="true" pageId="4" pageNumber="175">Corononcodes siculus</emphasis>
|
||
Bezzi (Tenerife)
|
||
</taxonomicName>
|
||
, ventral view. Closely spaced dotted lines are folds in the wing. The widely spaced dotted line is the continuation of the margin of the lower calypter behind the wing membrane. The wing venation is identical in male
|
||
<taxonomicName id="4C3A4D61FFB8FF80DAEFFA86E9C7FAFB" box="[944,1175,1370,1392]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="ziegleri" status="sp. nov.">
|
||
<emphasis id="B94EEAF0FFB8FF80DAEFFA86E9C7FAFB" box="[944,1175,1370,1392]" italics="true" pageId="4" pageNumber="175">Corononcodes ziegleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D578BFFB8FF80DDC0FA87E856FAFB" box="[1183,1286,1371,1392]" pageId="4" pageNumber="175" rank="species">spec. nov.</taxonomicNameLabel>
|
||
Scale bar: 0.5 mm. B) Right wing of female
|
||
<taxonomicName id="4C3A4D61FFB8FF80D896FAA5EFFEFA04" box="[457,686,1401,1423]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="ziegleri" status="sp. nov.">
|
||
<emphasis id="B94EEAF0FFB8FF80D896FAA5EFFEFA04" box="[457,686,1401,1423]" italics="true" pageId="4" pageNumber="175">Corononcodes ziegleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D578BFFB8FF80DBEBFAA6EE49FA04" box="[692,793,1402,1423]" pageId="4" pageNumber="175" rank="species">spec. nov.</taxonomicNameLabel>
|
||
from the type location, dorsal view. Scale bar: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8536E2FFB8FF80D998FA67EE92F9CB" blockId="4.[151,1437,1467,2035]" pageId="4" pageNumber="175">
|
||
Material of
|
||
<taxonomicName id="4C3A4D61FFB8FF80D806FA60EC9FFA5F" box="[345,463,1467,1492]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="siculus">
|
||
<emphasis id="B94EEAF0FFB8FF80D806FA60EC9FFA5F" box="[345,463,1467,1492]" italics="true" pageId="4" pageNumber="175">C. siculus</emphasis>
|
||
</taxonomicName>
|
||
studied for comparison (figs 2D, 2F).
|
||
<collectingCountry id="F32D7672FFB8FF80DAF0FA67EEA2FA5F" box="[943,1010,1467,1492]" name="Spain" pageId="4" pageNumber="175">Spain</collectingCountry>
|
||
, Canary Islands, Tenerife, Medano,
|
||
<date id="FF841022FFB8FF80D9C8FA3CEC5CFA73" box="[151,268,1504,1528]" pageId="4" pageNumber="175" value="1930-05-18">18.V.1930</date>
|
||
, leg. A. Cabrera (
|
||
<specimenCount id="9D3CFD6BFFB8FF80D885FA3CECADFA73" box="[474,509,1504,1528]" pageId="4" pageNumber="175" type="generic">1♂</specimenCount>
|
||
MZH);
|
||
<collectingCountry id="F32D7672FFB8FF80DB02FA03EFF0FA73" box="[605,672,1503,1528]" name="Spain" pageId="4" pageNumber="175">Spain</collectingCountry>
|
||
, Canary Islands, Fuerteventura, Los Lagos, ES9675,
|
||
<date id="FF841022FFB8FF80DDA0FA03E836FA73" box="[1279,1382,1503,1528]" pageId="4" pageNumber="175" value="2002-05-06">6.V.2002</date>
|
||
, leg. J.T. Smit (1♀ PCTZ);
|
||
<collectingCountry id="F32D7672FFB8FF80D8C0F9DFECB2F997" box="[415,482,1539,1564]" name="Spain" pageId="4" pageNumber="175">Spain</collectingCountry>
|
||
, Cádiz Province, Urbanización Costa Ballena, Playa Ballena,
|
||
<geoCoordinate id="EE0E5025FFB8FF80DDECF9DFE807F997" box="[1203,1367,1539,1564]" direction="north" orientation="latitude" pageId="4" pageNumber="175" precision="1" value="36.680588">36°40'50.11''N</geoCoordinate>
|
||
<geoCoordinate id="EE0E5025FFB8FF80DC3DF9DFEDAAF9CB" direction="west" orientation="longitude" pageId="4" pageNumber="175" precision="1" value="-6.4152584">6°24' 54.93''W</geoCoordinate>
|
||
,
|
||
<date id="FF841022FFB8FF80D85AF9F4ECD7F9CB" box="[261,391,1576,1600]" pageId="4" pageNumber="175" value="2014-06-22">22.VI.2014</date>
|
||
, leg. José Manuel Ayala
|
||
<collectingCountry id="F32D7672FFB8FF80DBC1F9FBEFB4F9CB" box="[670,740,1575,1600]" name="Netherlands" pageId="4" pageNumber="175">Landa</collectingCountry>
|
||
(1♀ coll. PCJMA).
|
||
</paragraph>
|
||
<paragraph id="8B8536E2FFB8FF80D998F997EFA8F8E8" blockId="4.[151,1437,1467,2035]" pageId="4" pageNumber="175">
|
||
<emphasis id="B94EEAF0FFB8FF80D998F997EC9FF9EF" bold="true" box="[199,463,1611,1636]" pageId="4" pageNumber="175">Differential diagnosis.</emphasis>
|
||
Predominantly dark brown (except white tibiae and tarsomeres in male) and covered with short pale pile; compound eyes bare and dichoptic; three ocelli present but mid ocellus smaller and placed on anterior surface of ocellar tubercle; mouthparts present and functional; no tibial spur or row of small sharply pointed projections discernable; wing with reduced venation, vein M3+4 incomplete basally, membrane hyaline in male and brownish infuscated in female. Female terminalia placed at tip of abdomen. The observed degree of sexual dimorphism in
|
||
<taxonomicName id="4C3A4D61FFB8FF80D8C9F8DFEF5DF890" box="[406,525,1794,1819]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="ziegleri">
|
||
<emphasis id="B94EEAF0FFB8FF80D8C9F8DFEF5DF890" box="[406,525,1794,1819]" italics="true" pageId="4" pageNumber="175">C. ziegleri</emphasis>
|
||
</taxonomicName>
|
||
comprises body size (female much larger), shape of flagellum (larger in male), coloration of tibiae and tarsomeres (white in male, brown in female) and coloration of wing veins and membrane (mainly hyaline in males, mainly brownish in female).
|
||
</paragraph>
|
||
<paragraph id="8B8536E2FFB8FF81D998F8B2E82AFE96" blockId="4.[151,1437,1467,2035]" lastBlockId="5.[151,1436,151,285]" lastPageId="5" lastPageNumber="176" pageId="4" pageNumber="175">
|
||
<taxonomicName id="4C3A4D61FFB8FF80D998F8B2EC94F80C" box="[199,452,1902,1927]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="ziegleri">
|
||
<emphasis id="B94EEAF0FFB8FF80D998F8B2EC94F80C" box="[199,452,1902,1927]" italics="true" pageId="4" pageNumber="175">Corononcodes ziegleri</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all African species by the bare eyes that are dichoptic below the antennae. It is closest to
|
||
<taxonomicName id="4C3A4D61FFB8FF80D883F84FEF1CF820" box="[476,588,1938,1963]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="siculus">
|
||
<emphasis id="B94EEAF0FFB8FF80D883F84FEF1CF820" box="[476,588,1938,1963]" italics="true" pageId="4" pageNumber="175">C. siculus</emphasis>
|
||
</taxonomicName>
|
||
, the males differing in the lateral shape of the flagellum, the colour of the tibiae and tarsomeres, the presence of the mid ocellus, and by the morphology of the genitalia. Females can only be separated by the presence of the mid ocellus in
|
||
<taxonomicName id="4C3A4D61FFB8FF80DB98F807EE6FF878" box="[711,831,2010,2035]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="ziegleri">
|
||
<emphasis id="B94EEAF0FFB8FF80DB98F807EE6FF878" box="[711,831,2010,2035]" italics="true" pageId="4" pageNumber="175">C. ziegleri</emphasis>
|
||
</taxonomicName>
|
||
, as no apparent differences in antennal shape, wing venation, body coloration etc. could be discerned, based on the material at hand. However,
|
||
<bibRefCitation id="EFAB4B13FFB9FF81DDE3FF4BE8CCFF3B" author="Carles-Tolra" box="[1212,1436,151,176]" pageId="5" pageNumber="176" refString="Carles-Tolra, M. (2008) Description of the female of Corononcodes siculus Bezzi (Diptera: Acroceridae). Boletin de la Sociedad Entomologica Aragonesa, 42, 135 - 138." type="journal article" year="2008">Carles-Tolrá (2008)</bibRefCitation>
|
||
documented a restricted degree of infuscation of the wing, limited to the basal portion of the membrane in a female specimen from the Spanish mainland. Due to the lack of material it cannot be excluded that the western Palaearctic specimens (Canary Islands, mainland
|
||
<collectingCountry id="F32D7672FFB9FF81DB1FFED8EFD3FE96" box="[576,643,260,285]" name="Spain" pageId="5" pageNumber="176">Spain</collectingCountry>
|
||
, Sicily) assigned to
|
||
<taxonomicName id="4C3A4D61FFB9FF81DA3BFED9EE82FE96" box="[868,978,260,285]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="siculus">
|
||
<emphasis id="B94EEAF0FFB9FF81DA3BFED9EE82FE96" box="[868,978,260,285]" italics="true" pageId="5" pageNumber="176">C. siculus</emphasis>
|
||
</taxonomicName>
|
||
actually represent a species complex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF45666AFFB9FF81D9C8F8BDEFC7F858" httpUri="https://zenodo.org/record/229007/files/figure.png" pageId="5" pageNumber="176" targetBox="[282,1303,343,1860]" targetPageId="5">
|
||
<paragraph id="8B8536E2FFB9FF81D9C8F8BDEFC7F858" blockId="5.[151,1436,1889,2003]" pageId="5" pageNumber="176">
|
||
<emphasis id="B94EEAF0FFB9FF81D9C8F8BDEC43F8FD" bold="true" box="[151,275,1889,1911]" pageId="5" pageNumber="176">FIGURE 4.</emphasis>
|
||
Male genitalia of
|
||
<taxonomicName id="4C3A4D61FFB9FF81D88EF8BDEF34F8FC" box="[465,612,1889,1911]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94EEAF0FFB9FF81D88EF8BDEF34F8FC" box="[465,612,1889,1911]" italics="true" pageId="5" pageNumber="176">Corononcodes</emphasis>
|
||
</taxonomicName>
|
||
spp. A–B)
|
||
<taxonomicName id="4C3A4D61FFB9FF81DB84F8BDEE91F8FC" box="[731,961,1889,1911]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="ziegleri" status="sp. nov.">
|
||
<emphasis id="B94EEAF0FFB9FF81DB84F8BDEE91F8FC" box="[731,961,1889,1911]" italics="true" pageId="5" pageNumber="176">Corononcodes ziegleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D578BFFB9FF81DA96F8BEE97FF8FC" box="[969,1071,1890,1911]" pageId="5" pageNumber="176" rank="species">spec. nov.</taxonomicNameLabel>
|
||
in ventral (A) and lateral (B) view; DNA voucher CK611. C–D)
|
||
<taxonomicName id="4C3A4D61FFB9FF81D899F85CEFBCF81D" authority="Bezzi" authorityName="Bezzi" box="[454,748,1920,1942]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="siculus">
|
||
<emphasis id="B94EEAF0FFB9FF81D899F85CEFF6F81D" box="[454,678,1920,1942]" italics="true" pageId="5" pageNumber="176">Corononcodes siculus</emphasis>
|
||
Bezzi
|
||
</taxonomicName>
|
||
; specimen from Tenerife in ventral (C) and lateral (D) view. Setae omitted. Abbreviations: a: aedeagus; c: cercus; e: epandrium; ea: ejaculatory apodeme; g: gonocoxite; gs: gonostylus; ps: parameral sheath of aedeagus. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF45666AFFBAFF82D9C8FCFEED8FFCDD" httpUri="https://zenodo.org/record/229008/files/figure.png" pageId="6" pageNumber="177" targetBox="[188,1398,193,780]" targetPageId="6">
|
||
<paragraph id="8B8536E2FFBAFF82D9C8FCFEED8FFCDD" blockId="6.[151,1436,802,854]" pageId="6" pageNumber="177">
|
||
<emphasis id="B94EEAF0FFBAFF82D9C8FCFEEC45FCBC" bold="true" box="[151,277,802,824]" pageId="6" pageNumber="177">FIGURE 5.</emphasis>
|
||
Current distribution of
|
||
<taxonomicName id="4C3A4D61FFBAFF82DB50FCFEEE63FCB3" authority="Bezzi" authorityName="Bezzi" box="[527,819,802,824]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="siculus">
|
||
<emphasis id="B94EEAF0FFBAFF82DB50FCFEEFA0FCB3" box="[527,752,802,824]" italics="true" pageId="6" pageNumber="177">Corononcodes siculus</emphasis>
|
||
Bezzi
|
||
</taxonomicName>
|
||
(black cross) and of
|
||
<taxonomicName id="4C3A4D61FFBAFF82DD4BFCFEE9ABFCB3" box="[1044,1275,802,824]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="ziegleri" status="sp. nov.">
|
||
<emphasis id="B94EEAF0FFBAFF82DD4BFCFEE9ABFCB3" box="[1044,1275,802,824]" italics="true" pageId="6" pageNumber="177">Corononcodes ziegleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D578BFFBAFF82DC5BFCFFE83BFCB3" box="[1284,1387,803,824]" pageId="6" pageNumber="177" rank="species">spec. nov.</taxonomicNameLabel>
|
||
(red circle).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3206569FFBAFF82D998FC5FEEDFF838" pageId="6" pageNumber="177" type="discussion">
|
||
<paragraph id="8B8536E2FFBAFF82D998FC5FE80DFBA7" blockId="6.[151,1436,899,1971]" pageId="6" pageNumber="177">
|
||
<emphasis id="B94EEAF0FFBAFF82D998FC5FEC6BFC17" bold="true" box="[199,315,899,924]" pageId="6" pageNumber="177">Remarks.</emphasis>
|
||
The two populations sampled are about
|
||
<quantity id="4CC29B07FFBAFF82DA7AFC58EE2FFC17" box="[805,895,899,924]" metricMagnitude="5" metricUnit="m" metricValue="9.3" pageId="6" pageNumber="177" unit="km" value="930.0">930 km</quantity>
|
||
apart (fig. 5). Uncorrected p-distance in COI between specimens from Ghazvin province in northern
|
||
<collectingCountry id="F32D7672FFBAFF82DA5CFC74EE60FC34" box="[771,816,936,959]" name="Iran" pageId="6" pageNumber="177">Iran</collectingCountry>
|
||
(CK611 & CK612) and from
|
||
<collectingCountry id="F32D7672FFBAFF82DDDFFC74E9E0FC34" box="[1152,1200,936,959]" name="Iran" pageId="6" pageNumber="177">Fars</collectingCountry>
|
||
province in southern
|
||
<collectingCountry id="F32D7672FFBAFF82D9C8FC10ED94FC68" box="[151,196,972,995]" name="Iran" pageId="6" pageNumber="177">Iran</collectingCountry>
|
||
(CK698) was 5.8%. Nevertheless, the material is regarded as conspecific, as there are no distinct morphological features separating the two populations. Also, similar high intraspecific genetic distances have recently been recorded between populations of
|
||
<taxonomicName id="4C3A4D61FFBAFF82DBF6FBC8E809FBA7" authority="(Fabricius) (Kehlmaier & Almeida 2014)" authorityName="(Fabricius) (Kehlmaier & Almeida" authorityYear="2014" box="[681,1369,1043,1068]" class="Insecta" family="Acroceridae" genus="Acrocera" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="orbiculus">
|
||
<emphasis id="B94EEAF0FFBAFF82DBF6FBC8EED1FBA7" box="[681,897,1043,1068]" italics="true" pageId="6" pageNumber="177">Acrocera orbiculus</emphasis>
|
||
(Fabricius) (
|
||
<bibRefCitation id="EFAB4B13FFBAFF82DD4EFBCFE802FBA7" author="Kehlmaier" box="[1041,1362,1043,1068]" pageId="6" pageNumber="177" refString="Kehlmaier, C. & Almeida, J. M. (2014) New host records for European Acroceridae (Diptera), with discussion of species limits of Acrocera orbiculus (Fabricius) based on DNA-barcoding. Zootaxa, 3780 (1), 135 - 152. http: // dx. doi. org / 10.11646 / zootaxa. 3780.1.5." type="journal article" year="2014">Kehlmaier & Almeida 2014</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8536E2FFBAFF82D998FBEBE923FA1C" blockId="6.[151,1436,899,1971]" pageId="6" pageNumber="177">
|
||
<bibRefCitation id="EFAB4B13FFBAFF82D998FBEBEC03FBDB" author="Sack" box="[199,339,1079,1104]" pageId="6" pageNumber="177" refString="Sack, P. (1936) Cyrtidae (Acroceridae). In: Lindner, E. (Ed.), Die Fliegen der palaarktischen Region, Band IV (I), Teil 21. E. Schweizerbart, Stuttgart, 36 pp." type="book" year="1936">Sack (1936)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFAB4B13FFBAFF82D83FFBEBEF4FFBDB" author="Schlinger" box="[352,543,1079,1104]" pageId="6" pageNumber="177" refString="Schlinger, E. I. (1960) A review of the South African Acroceridae (Diptera). Annals of the Natal Museum, 14, 459 - 504." type="journal article" year="1960">Schlinger (1960)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFAB4B13FFBAFF82DB06FBEBEE67FBDB" author="Barraclough" box="[601,823,1079,1104]" pageId="6" pageNumber="177" refString="Barraclough, D. A. (2001) A review of the South African species of Corononcodes Speiser (Diptera: Acroceridae). African Invertebrates, 42, 95 - 104." type="journal article" year="2001">Barraclough (2001)</bibRefCitation>
|
||
provided diagnoses of
|
||
<emphasis id="B94EEAF0FFBAFF82DD1CFBEBE9BCFBDB" box="[1091,1260,1079,1104]" italics="true" pageId="6" pageNumber="177">
|
||
<taxonomicName id="4C3A4D61FFBAFF82DD1CFBEBE9B8FBDB" box="[1091,1256,1079,1104]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">Corononcodes</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but a thorough revision of the
|
||
<taxonomicName id="4C3A4D61FFBAFF82D81DFB87ECE7FBFF" box="[322,439,1115,1140]" class="Insecta" family="Acroceridae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="subFamily" subFamily="Panopinae">Panopinae</taxonomicName>
|
||
on a generic level is needed to fully define the limits of
|
||
<taxonomicName id="4C3A4D61FFBAFF82DD68FB87E989FBFF" box="[1079,1241,1115,1140]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94EEAF0FFBAFF82DD68FB87E989FBFF" box="[1079,1241,1115,1140]" italics="true" pageId="6" pageNumber="177">Corononcodes</emphasis>
|
||
</taxonomicName>
|
||
. This would also help determine whether the Palaearctic species currently placed in
|
||
<taxonomicName id="4C3A4D61FFBAFF82DA20FBA3E971FB13" box="[895,1057,1151,1176]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94EEAF0FFBAFF82DA20FBA3E971FB13" box="[895,1057,1151,1176]" italics="true" pageId="6" pageNumber="177">Corononcodes</emphasis>
|
||
</taxonomicName>
|
||
should be transferred to a distinct genus as proposed by
|
||
<bibRefCitation id="EFAB4B13FFBAFF82D8FAFB7FEFD9FB37" author="Barraclough" box="[421,649,1187,1212]" pageId="6" pageNumber="177" refString="Barraclough, D. A. (2001) A review of the South African species of Corononcodes Speiser (Diptera: Acroceridae). African Invertebrates, 42, 95 - 104." type="journal article" year="2001">Barraclough (2001)</bibRefCitation>
|
||
, who briefly discussed the morphological differences between the regional faunas. The Palaearctic species discussed here are closest to
|
||
<taxonomicName id="4C3A4D61FFBAFF82DAFBFB14E962FB6B" box="[932,1074,1223,1248]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="dimorpha">
|
||
<emphasis id="B94EEAF0FFBAFF82DAFBFB14E962FB6B" box="[932,1074,1223,1248]" italics="true" pageId="6" pageNumber="177">C. dimorpha</emphasis>
|
||
</taxonomicName>
|
||
, sharing the sexually dimorphic wing colour (brownish infuscated in female, hyaline in male), the basally incomplete vein M3+4, and the reduced tibial spurs. Also,
|
||
<taxonomicName id="4C3A4D61FFBAFF82D83EFACFEC85FAA0" box="[353,469,1298,1323]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="ziegleri">
|
||
<emphasis id="B94EEAF0FFBAFF82D83EFACFEC85FAA0" box="[353,469,1298,1323]" italics="true" pageId="6" pageNumber="177">C. ziegleri</emphasis>
|
||
</taxonomicName>
|
||
has three ocelli like all Afrotropical species. However, unlike the Afrotropical species, the Palaearctic ones have a distinct frons in both sexes, with their compound eyes clearly separated beneath the insertion of antennae. Furthermore, female terminalia are placed at the tip of the abdomen and not ventrally on the abdomen as mentioned in the literature (
|
||
<bibRefCitation id="EFAB4B13FFBAFF82DB05FAA2EF85FA1C" author="Sack" box="[602,725,1406,1431]" pageId="6" pageNumber="177" refString="Sack, P. (1936) Cyrtidae (Acroceridae). In: Lindner, E. (Ed.), Die Fliegen der palaarktischen Region, Band IV (I), Teil 21. E. Schweizerbart, Stuttgart, 36 pp." type="book" year="1936">Sack 1936</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAB4B13FFBAFF82DBBFFAA2EEDDFA1C" author="Schlinger" box="[736,909,1406,1431]" pageId="6" pageNumber="177" refString="Schlinger, E. I. (1960) A review of the South African Acroceridae (Diptera). Annals of the Natal Museum, 14, 459 - 504." type="journal article" year="1960">Schlinger 1960</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAB4B13FFBAFF82DAC6FAA2E936FA1C" author="Barraclough" box="[921,1126,1406,1431]" pageId="6" pageNumber="177" refString="Barraclough, D. A. (2001) A review of the South African species of Corononcodes Speiser (Diptera: Acroceridae). African Invertebrates, 42, 95 - 104." type="journal article" year="2001">Barraclough 2001</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8536E2FFBAFF82D998FA7EE80DF9E4" blockId="6.[151,1436,899,1971]" pageId="6" pageNumber="177">
|
||
<bibRefCitation id="EFAB4B13FFBAFF82D998FA7EEF19FA30" author="Kirk-Spriggs" box="[199,585,1442,1467]" pageId="6" pageNumber="177" refString="Kirk-Spriggs, A. H. & McGregor, G. (2009) Disjunctions in the Diptera (Insecta) fauna of the Mediterranean Province and southern Africa and a discussion of biogeographical considerations. Transactions of the Royal Society of South Africa, 64, 32 - 52. http: // dx. doi. org / 10.1080 / 00359190909519236" type="journal article" year="2009">Kirk-Spriggs & McGregor (2009)</bibRefCitation>
|
||
comment on the southern African—Mediterranean disjunct distribution of
|
||
<taxonomicName id="4C3A4D61FFBAFF82D9C8FA1AEC69FA54" box="[151,313,1478,1503]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94EEAF0FFBAFF82D9C8FA1AEC69FA54" box="[151,313,1478,1503]" italics="true" pageId="6" pageNumber="177">Corononcodes</emphasis>
|
||
</taxonomicName>
|
||
, stating that it might be the result of either isolation following the aridification of the Sahara region during the Pliocene/Pleistocene (
|
||
<emphasis id="B94EEAF0FFBAFF82DB7EFA37EF6DF988" box="[545,573,1515,1539]" italics="true" pageId="6" pageNumber="177">cf.</emphasis>
|
||
“central high Africa corridor”) or that it might have been caused by independent dispersal originating from the Sahel zone via the dust-transporting wind systems of the
|
||
<collectingCountry id="F32D7672FFBAFF82DC1EF9D3EDB7F9C0" name="Western Sahara" pageId="6" pageNumber="177">Western Sahara</collectingCountry>
|
||
. With the discovery of an additional species from the Middle East, the distribution of the genus is considerably enlarged, and its biogeographic history extended by the “eastern high Africa corridor” scenario.
|
||
</paragraph>
|
||
<paragraph id="8B8536E2FFBAFF82D998F9A7EEDFF838" blockId="6.[151,1436,899,1971]" pageId="6" pageNumber="177">
|
||
No host records are available for
|
||
<taxonomicName id="4C3A4D61FFBAFF82DB2AF9A6EE47F918" box="[629,791,1658,1683]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94EEAF0FFBAFF82DB2AF9A6EE47F918" box="[629,791,1658,1683]" italics="true" pageId="6" pageNumber="177">Corononcodes</emphasis>
|
||
</taxonomicName>
|
||
, but so far
|
||
<taxonomicName id="4C3A4D61FFBAFF82DAE3F9A6E961F918" box="[956,1073,1658,1683]" class="Insecta" family="Acroceridae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="subFamily" subFamily="Panopinae">Panopinae</taxonomicName>
|
||
have been reared only from mygalomorph spiders (
|
||
<taxonomicName id="4C3A4D61FFBAFF82D8FCF943EF56F93C" box="[419,518,1695,1719]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="177" phylum="Arthropoda" rank="order">Araneae</taxonomicName>
|
||
: Mygalomorphae). On the Canary Islands, the known distribution of the sole species of mygalomorph spiders (
|
||
<taxonomicName id="4C3A4D61FFBAFF82DB79F91EEE93F950" authority="Wunderlich" authorityName="Wunderlich" box="[550,963,1730,1755]" class="Arachnida" family="Idiopidae" genus="Titanidiops" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="canariensis">
|
||
<emphasis id="B94EEAF0FFBAFF82DB79F91EEE62F951" box="[550,818,1730,1755]" italics="true" pageId="6" pageNumber="177">Titanidiops canariensis</emphasis>
|
||
Wunderlich
|
||
</taxonomicName>
|
||
) is restricted to the easternmost islands Fuerteventura, Lanzarote, and La Graciosa (
|
||
<bibRefCitation id="EFAB4B13FFBAFF82DBDEF93AEE3DF974" author="Izquierdo" box="[641,877,1766,1791]" pageId="6" pageNumber="177" refString="Izquierdo, I., Martin, J. L., Zurita, N. & Arechavaleta, M. (2004) Lista de especies silvestres de Canarian (hongos, plantas y animales terrestres) 2004. Consejeria de Medio Ambiente y Ordenacion Territorial, Gobierno de Canarias, 500 pp." type="book" year="2004">
|
||
Izquierdo
|
||
<emphasis id="B94EEAF0FFBAFF82DBABF93BEE7CF974" box="[756,812,1766,1791]" italics="true" pageId="6" pageNumber="177">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAB4B13FFBAFF82DA28F93BE9DDF974" author="Opatova" box="[887,1165,1766,1791]" pageId="6" pageNumber="177" refString="Opatova, V. & Arnedo, M. A. (2011) Colonization pathways and phylogeography of the Canary Islands endemic trap-door spider Titanidiops canariensis (Araneae, Idiopidae). In: Lubin, Y. et al. (Eds.), 26 th European Congress of Arachnology, Israel 2011, Program and Abstracts Book, 4 - 8 September 2011. Ben-Gurion University of the Negev, Blaustein Institutes for Desert Research, Israel, 30 pp. Available from: http: // www. european-arachnology. org / collo / images / 26 _ abstracts. pdf (accessed 28 August 2014)" type="book chapter" year="2011">Opatova & Arnedo 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAB4B13FFBAFF82DDC7F93AE80DF974" author="Wunderlich" box="[1176,1373,1766,1791]" pageId="6" pageNumber="177" refString="Wunderlich, J. (1992) Die Spinnenfauna der Makaronesischen Inseln: Taxonomie, Okologie, Biogeographie und Evolution. Beitrage zur Araneologie, 1, 1 - 619." type="journal article" year="1992">Wunderlich 1992</bibRefCitation>
|
||
). The first record here of
|
||
<taxonomicName id="4C3A4D61FFBAFF82D8CCF8D7EF5BF8A8" box="[403,523,1802,1827]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="siculus">
|
||
<emphasis id="B94EEAF0FFBAFF82D8CCF8D7EF5BF8A8" box="[403,523,1802,1827]" italics="true" pageId="6" pageNumber="177">C. siculus</emphasis>
|
||
</taxonomicName>
|
||
for Fuerteventura makes a parasitoid-host relationship between these taxa conceivable, assuming that
|
||
<taxonomicName id="4C3A4D61FFBAFF82D882F8F3EF2FF8CD" box="[477,639,1839,1862]" class="Arachnida" family="Idiopidae" genus="Titanidiops" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="canariensis">
|
||
<emphasis id="B94EEAF0FFBAFF82D882F8F3EF2FF8CD" box="[477,639,1839,1862]" italics="true" pageId="6" pageNumber="177">T. canariensis</emphasis>
|
||
</taxonomicName>
|
||
is also present on the Western islands. Fully grown females of
|
||
<taxonomicName id="4C3A4D61FFBAFF82DCD8F8F3EC48F8E1" class="Arachnida" family="Idiopidae" genus="Titanidiops" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="canariensis">
|
||
<emphasis id="B94EEAF0FFBAFF82DCD8F8F3EC48F8E1" italics="true" pageId="6" pageNumber="177">T. canariensis</emphasis>
|
||
</taxonomicName>
|
||
range from
|
||
<quantity id="4CC29B07FFBAFF82D8C0F88FEF48F8E1" box="[415,536,1875,1899]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="6" pageNumber="177" unit="mm" value="15.0" valueMax="18.0" valueMin="12.0">12–18 mm</quantity>
|
||
in total body length (
|
||
<bibRefCitation id="EFAB4B13FFBAFF82DA5BF88EEE9CF8E0" author="Wunderlich" box="[772,972,1874,1899]" pageId="6" pageNumber="177" refString="Wunderlich, J. (1992) Die Spinnenfauna der Makaronesischen Inseln: Taxonomie, Okologie, Biogeographie und Evolution. Beitrage zur Araneologie, 1, 1 - 619." type="journal article" year="1992">Wunderlich 1992</bibRefCitation>
|
||
).
|
||
<collectingCountry id="F32D7672FFBAFF82DA80F88FEEAEF8E0" box="[991,1022,1875,1899]" name="American Samoa" pageId="6" pageNumber="177">As</collectingCountry>
|
||
<taxonomicName id="4C3A4D61FFBAFF82DD5BF88EE9DDF8E0" box="[1028,1165,1874,1899]" class="Insecta" family="Acroceridae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="family">Acroceridae</taxonomicName>
|
||
larvae normally emerge from their host before the spider reaches adulthood, the size of the opistosoma of juvenile
|
||
<taxonomicName id="4C3A4D61FFBAFF82DDF6F8ABE818F805" box="[1193,1352,1911,1934]" class="Arachnida" family="Idiopidae" genus="Titanidiops" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="canariensis">
|
||
<emphasis id="B94EEAF0FFBAFF82DDF6F8ABE818F805" box="[1193,1352,1911,1934]" italics="true" pageId="6" pageNumber="177">T. canariensis</emphasis>
|
||
</taxonomicName>
|
||
should therefore be sufficient to harbour fully grown
|
||
<taxonomicName id="4C3A4D61FFBAFF82DBC5F846EE6CF838" box="[666,828,1946,1971]" class="Insecta" family="Acroceridae" genus="Corononcodes" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94EEAF0FFBAFF82DBC5F846EE6CF838" box="[666,828,1946,1971]" italics="true" pageId="6" pageNumber="177">Corononcodes</emphasis>
|
||
</taxonomicName>
|
||
larvae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |